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<document id="631733BFE87CE37CA60FB8E9733EA0A2" ID-DOI="10.5281/zenodo.5188356" ID-ISSN="1942-1354" ID-Zenodo-Dep="5188356" ID-ZooBank="AC2597CC-301F-4E91-9711-5C17399C9AA2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628346141153" checkinUser="felipe" docAuthor="King, Jonas G. &amp; Lago, Paul K." docDate="2012" docId="03E15618982A0B2FFF19FEDB4F9E1FD9" docLanguage="en" docName="InsectaMundi.0275.1-53.pdf" docOrigin="Insecta Mundi 2012 (275)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Heterocerus gnatho LeConte 1863" docType="treatment" docVersion="4" lastPageNumber="16" masterDocId="FFD82E60983B0B3EFFD9FFF94D431A43" masterDocTitle="The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Mississippi and Alabama, with discussion and keys to the species occurring in the southeastern United States" masterLastPageNumber="53" masterPageNumber="1" pageNumber="16" updateTime="1699045744038" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="7435F0E7CCE9CDA2999E25F2408487AC">The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Mississippi and Alabama, with discussion and keys to the species occurring in the southeastern United States</mods:title>
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<mods:namePart id="F59C351C1A516B8AE34AACFC31963AC9">King, Jonas G.</mods:namePart>
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<mods:namePart id="12BDEF6CA57A3FF6032C85F5B0127ED6">Lago, Paul K.</mods:namePart>
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<mods:date id="60AC395F990BA592CEA682B7667842B2">2012</mods:date>
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<mods:number id="245BCEC937337285489968CBD954F55C">2012-12-28</mods:number>
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<treatment id="03E15618982A0B2FFF19FEDB4F9E1FD9" ID-DOI="http://doi.org/10.5281/zenodo.5187837" ID-Zenodo-Dep="5187837" LSID="urn:lsid:plazi:treatment:03E15618982A0B2FFF19FEDB4F9E1FD9" httpUri="http://treatment.plazi.org/id/03E15618982A0B2FFF19FEDB4F9E1FD9" lastPageNumber="16" pageId="17" pageNumber="16">
<subSubSection id="C352B485982A0B2FFF19FEDB4F231B79" box="[192,608,290,314]" pageId="17" pageNumber="16" type="nomenclature">
<paragraph id="8BF7E70E982A0B2FFF19FEDB4F231B79" blockId="17.[192,608,290,346]" box="[192,608,290,314]" pageId="17" pageNumber="16">
<heading id="D0BF5062982A0B2FFF19FEDB4F231B79" bold="true" box="[192,608,290,314]" fontSize="10" level="2" pageId="17" pageNumber="16" reason="3">
<emphasis id="B93C3B1C982A0B2FFF19FEDB4F231B79" bold="true" box="[192,608,290,314]" pageId="17" pageNumber="16">
<emphasis id="B93C3B1C982A0B2FFF19FEDB4D9B1B79" bold="true" box="[192,216,290,314]" italics="true" pageId="17" pageNumber="16">2.</emphasis>
<taxonomicName id="4C489C8D982A0B2FFF04FEDA4F231B79" authority="LeConte" authorityName="LeConte" authorityYear="1863" box="[221,608,290,314]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gnatho">
<emphasis id="B93C3B1C982A0B2FFF04FEDA4CA51B79" bold="true" box="[221,486,290,314]" italics="true" pageId="17" pageNumber="16">Heterocerus gnatho</emphasis>
LeConte
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C352B485982A0B2FFF19FEBB4CC11B19" box="[192,386,322,346]" pageId="17" pageNumber="16" type="description">
<paragraph id="8BF7E70E982A0B2FFF19FEBB4CC11B19" blockId="17.[192,608,290,346]" box="[192,386,322,346]" pageId="17" pageNumber="16">
(
<figureCitation id="1373FB8B982A0B2FFF10FEBB4C621B19" box="[201,289,322,346]" captionStart="Figures 11-17" captionStartId="14.[192,290,1646,1668]" captionTargetBox="[192,1440,191,1615]" captionTargetId="figure-878@14.[192,1440,858,954]" captionTargetPageId="14" captionText="Figures 11-17. Habiti of large Heterocerus species lacking post-mesocoxal lines. 11) H. pallidus (male is hypermandibulate). 12) H.intermuralis (male is hypermandibulate). 13) H.fatuus. 14) H. gnatho. 15) H.longilobulus. 16) H. sandersoni. 17) H. angustatus.Scale line = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5188368" httpUri="https://zenodo.org/record/5188368/files/figure.png" pageId="17" pageNumber="16">Fig. 14</figureCitation>
,
<figureCitation id="1373FB8B982A0B2FFEF7FEBB4C0D1B19" box="[302,334,322,346]" captionStart="Figures 32-44" captionStartId="38.[192,286,1682,1704]" captionTargetBox="[192,1357,192,1649]" captionTargetId="figure-419@38.[688,779,856,946]" captionTargetPageId="38" captionText="Figures 32-44. Male genitalia of southeastern Heterocerus species. The ventral view is shown unless otherwise stated. 32-39) Species that lack post-mesocoxal lines. 32) H. fatuus. 33) H. glicki. 34) H. intermuralis. 35) H. angustatus; left - ventral view, right - dorsal view. 36) H. longilobulus. 37) H. pallidus; left - ventral view, right - dorsal view. 38) H. gnatho; left - ventral view, right - dorsal view. 39) H. sandersoni; left - ventral view, center - dorsal view, left - lateral view. 40-44) Species with post-mesocoxal lines. 40) H. sinuosus; left - dorsal view, right - ventral aspect. 41) H. mollinus. 42) H. insolens; left - dorsal view, right - ventral aspect. 43) H. parrotus. 44) H. brunneus." figureDoi="http://doi.org/10.5281/zenodo.5188382" httpUri="https://zenodo.org/record/5188382/files/figure.png" pageId="17" pageNumber="16">38</figureCitation>
,
<figureCitation id="1373FB8B982A0B2FFE82FEBB4C3A1B19" box="[347,377,322,346]" captionStart="Figures 54-57" captionStartId="39.[192,286,1864,1886]" captionTargetBox="[192,1436,1076,1850]" captionTargetId="figure-932@39.[788,832,1441,1510]" captionTargetPageId="39" captionText="Figures 54-57. Distribution by county in the United States. 54) Heterocerus pallidus. 55) Heterocerus gnatho. 56) Heterocerus longilobulus. 57) Heterocerus insolens." figureDoi="http://doi.org/10.5281/zenodo.5188388" httpUri="https://zenodo.org/record/5188388/files/figure.png" pageId="17" pageNumber="16">55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C352B485982A0B2FFF19FE7A4E7D1BF9" pageId="17" pageNumber="16" type="reference_group">
<paragraph id="8BF7E70E982A0B2FFF19FE7A4FCD1BD9" blockId="17.[192,830,386,442]" box="[192,654,386,410]" pageId="17" pageNumber="16">
<treatmentCitationGroup id="AB58C020982A0B2FFF19FE7A4FCD1BD9" box="[192,654,386,410]" pageId="17" pageNumber="16">
<treatmentCitation id="0AE9C11F982A0B2FFF19FE7A4FC81BD9" author="LeConte, J. L." box="[192,651,386,410]" page="74" pageId="17" pageNumber="16" year="1863">
<taxonomicName id="4C489C8D982A0B2FFF19FE7A4FC81BD9" authority="LeConte 1863: 74" authorityName="LeConte" authorityPageNumber="74" authorityYear="1863" box="[192,651,386,410]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gnatho">
<emphasis id="B93C3B1C982A0B2FFF19FE7A4CE81BD9" box="[192,427,386,410]" italics="true" pageId="17" pageNumber="16">Heterocerus gnatho</emphasis>
<bibRefCitation id="EFD99AFF982A0B2FFE68FE7B4FC81BD9" author="LeConte, J. L." box="[433,651,386,410]" pageId="17" pageNumber="16" pagination="1 - 86" refId="ref21411" refString="LeConte, J. L. 1863. New species of North American Coleoptera. Smithsonian Miscellaneous Collection 167: 1 - 86." type="journal article" year="1863">LeConte 1863: 74</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF7E70E982A0B2FFF19FE5B4E7D1BF9" blockId="17.[192,830,386,442]" box="[192,830,418,442]" pageId="17" pageNumber="16">
<treatmentCitationGroup id="AB58C020982A0B2FFF19FE5B4E7D1BF9" box="[192,830,418,442]" pageId="17" pageNumber="16">
<taxonomicName id="4C489C8D982A0B2FFF19FE5B4F191BF9" authority="(LeConte)" baseAuthorityName="LeConte" baseAuthorityYear="1863" box="[192,602,418,442]" class="Insecta" family="Heteroceridae" genus="Neoheterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gnatho">
<emphasis id="B93C3B1C982A0B2FFF19FE5B4C971BF9" box="[192,468,418,442]" italics="true" pageId="17" pageNumber="16">Neoheterocerus gnatho</emphasis>
(LeConte)
</taxonomicName>
:
<treatmentCitation id="0AE9C11F982A0B2FFDBCFE5B4E781BF9" author="Pacheco, F." box="[613,827,418,442]" page="85" pageId="17" pageNumber="16" year="1964">
<bibRefCitation id="EFD99AFF982A0B2FFDBCFE5B4E781BF9" author="Pacheco, F." box="[613,827,418,442]" pageId="17" pageNumber="16" pagination="1 - 155" refId="ref22030" refString="Pacheco, F. 1964. Sistematica, filogenia y distribucion de los Heteroceridos de America (Coleoptera; Heteroceridae). Monografias del Colegio de Post-graduados, No. 1, Escuela Nacional de Agricultura, Chapingo, Mexico, Vol. 1: 1 - 155." type="journal article" year="1964">Pacheco 1964: 85</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C352B485982A0B2FFF19FE1B4F2018F9" pageId="17" pageNumber="16" type="description">
<paragraph id="8BF7E70E982A0B2FFF19FE1B4F2018F9" blockId="17.[192,1441,482,698]" pageId="17" pageNumber="16">
<emphasis id="B93C3B1C982A0B2FFF19FE1B4C2F1BB9" bold="true" box="[192,364,482,506]" pageId="17" pageNumber="16">Description.</emphasis>
Length 3.5 - 7.0 mm. Dark brown to yellowish brown. Elytra dominated by pale markings that are highly variable in shape and definition, vaguely trifasciate (
<figureCitation id="1373FB8B982A0B2FFC34FDFB49001859" box="[1005,1091,514,538]" captionStart="Figures 11-17" captionStartId="14.[192,290,1646,1668]" captionTargetBox="[192,1440,191,1615]" captionTargetId="figure-878@14.[192,1440,858,954]" captionTargetPageId="14" captionText="Figures 11-17. Habiti of large Heterocerus species lacking post-mesocoxal lines. 11) H. pallidus (male is hypermandibulate). 12) H.intermuralis (male is hypermandibulate). 13) H.fatuus. 14) H. gnatho. 15) H.longilobulus. 16) H. sandersoni. 17) H. angustatus.Scale line = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5188368" httpUri="https://zenodo.org/record/5188368/files/figure.png" pageId="17" pageNumber="16">Fig. 14</figureCitation>
); striae absent. Pronotal disc usually darker than elytra, apical angles of the pronotum pale, apical spot sometimes elongated posteriorly along pronotal border, narrowing posteriorly only slightly. Post-metathoracic coxal and post-mesothoracic coxal lines absent. Male genitalia (
<figureCitation id="1373FB8B982A0B2FFD32FD9B4E021839" box="[747,833,610,634]" captionStart="Figures 32-44" captionStartId="38.[192,286,1682,1704]" captionTargetBox="[192,1357,192,1649]" captionTargetId="figure-419@38.[688,779,856,946]" captionTargetPageId="38" captionText="Figures 32-44. Male genitalia of southeastern Heterocerus species. The ventral view is shown unless otherwise stated. 32-39) Species that lack post-mesocoxal lines. 32) H. fatuus. 33) H. glicki. 34) H. intermuralis. 35) H. angustatus; left - ventral view, right - dorsal view. 36) H. longilobulus. 37) H. pallidus; left - ventral view, right - dorsal view. 38) H. gnatho; left - ventral view, right - dorsal view. 39) H. sandersoni; left - ventral view, center - dorsal view, left - lateral view. 40-44) Species with post-mesocoxal lines. 40) H. sinuosus; left - dorsal view, right - ventral aspect. 41) H. mollinus. 42) H. insolens; left - dorsal view, right - ventral aspect. 43) H. parrotus. 44) H. brunneus." figureDoi="http://doi.org/10.5281/zenodo.5188382" httpUri="https://zenodo.org/record/5188382/files/figure.png" pageId="17" pageNumber="16">Fig. 38</figureCitation>
) of the typical
<taxonomicName id="4C489C8D982A0B2FFC2EFD9A49C41839" authorityName="Fabricius" authorityYear="1792" box="[1015,1159,611,634]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B93C3B1C982A0B2FFC2EFD9A49C41839" box="[1015,1159,611,634]" italics="true" pageId="17" pageNumber="16">Heterocerus</emphasis>
</taxonomicName>
<typeStatus id="54F359AC982A0B2FFB54FD9A49861838" box="[1165,1221,611,635]" pageId="17" pageNumber="16">type</typeStatus>
, parameres triangular; dorsal plate of aedeagus depressed on the right anterior edge, and convex on its posterior edge. Hypermandibulate males common.
</paragraph>
</subSubSection>
<subSubSection id="C352B485982A0B2FFF19FD1B48E21E79" pageId="17" pageNumber="16" type="diagnosis">
<paragraph id="8BF7E70E982A0B2FFF19FD1B4FC219D9" blockId="17.[192,1440,738,922]" pageId="17" pageNumber="16">
<emphasis id="B93C3B1C982A0B2FFF19FD1B4C1318B9" bold="true" box="[192,336,738,762]" pageId="17" pageNumber="16">Diagnosis.</emphasis>
<taxonomicName id="4C489C8D982A0B2FFE81FD1A4F0618B9" authorityName="LeConte" authorityYear="1863" box="[344,581,738,762]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gnatho">
<emphasis id="B93C3B1C982A0B2FFE81FD1A4F0618B9" box="[344,581,738,762]" italics="true" pageId="17" pageNumber="16">Heterocerus gnatho</emphasis>
</taxonomicName>
is the largest heterocerid species in North America, although its size varies considerably. While strikingly similar in appearance to
<taxonomicName id="4C489C8D982A0B2FFC72FCFA49701959" authorityName="Say" authorityYear="1823" box="[939,1075,770,794]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="B93C3B1C982A0B2FFC72FCFA49701959" box="[939,1075,770,794]" italics="true" pageId="17" pageNumber="16">H. pallidus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C489C8D982A0B2FFBE6FCFA49F41959" authorityName="LeConte" authorityYear="1863" box="[1087,1207,770,794]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gnatho">
<emphasis id="B93C3B1C982A0B2FFBE6FCFA49F41959" box="[1087,1207,770,794]" italics="true" pageId="17" pageNumber="16">H. gnatho</emphasis>
</taxonomicName>
is generally darker and has less prominent light pronotal borders. Enough variation is present in these characters, however, to warrant inspection of male genitalia. A strong notch on the posterior edge of the dorsal plate (compare dorsal views in
<figureCitation id="1373FB8B982A0B2FFEA1FC9B4F611939" box="[376,546,866,890]" captionStart="Figures 32-44" captionStartId="38.[192,286,1682,1704]" captionTargetBox="[192,1357,192,1649]" captionTargetId="figure-419@38.[688,779,856,946]" captionTargetPageId="38" captionText="Figures 32-44. Male genitalia of southeastern Heterocerus species. The ventral view is shown unless otherwise stated. 32-39) Species that lack post-mesocoxal lines. 32) H. fatuus. 33) H. glicki. 34) H. intermuralis. 35) H. angustatus; left - ventral view, right - dorsal view. 36) H. longilobulus. 37) H. pallidus; left - ventral view, right - dorsal view. 38) H. gnatho; left - ventral view, right - dorsal view. 39) H. sandersoni; left - ventral view, center - dorsal view, left - lateral view. 40-44) Species with post-mesocoxal lines. 40) H. sinuosus; left - dorsal view, right - ventral aspect. 41) H. mollinus. 42) H. insolens; left - dorsal view, right - ventral aspect. 43) H. parrotus. 44) H. brunneus." figureDoi="http://doi.org/10.5281/zenodo.5188382" httpUri="https://zenodo.org/record/5188382/files/figure.png" pageId="17" pageNumber="16">Fig. 37 and 38</figureCitation>
), and having the interior margins of the parameres diverging slightly from one another, distinguish this species.
</paragraph>
<paragraph id="8BF7E70E982A0B2FFF19FC3B49C31999" blockId="17.[192,1441,962,1082]" box="[192,1152,962,986]" pageId="17" pageNumber="16">
<emphasis id="B93C3B1C982A0B2FFF19FC3B4C541999" bold="true" box="[192,279,962,986]" pageId="17" pageNumber="16">Notes.</emphasis>
Southeastern records are mostly from sandy riverine and coastal areas.
</paragraph>
<paragraph id="8BF7E70E982A0B2FFF29FC1A48E21E79" blockId="17.[192,1441,962,1082]" pageId="17" pageNumber="16">
Upon inspection of the male genitalia in previously identified material, it was found that specimens of
<taxonomicName id="4C489C8D982A0B2FFF06FBFA4CDC1E59" authorityName="King &amp; Lago" authorityYear="2012" baseAuthorityName="King &amp; Lago" baseAuthorityYear="2012" box="[223,415,1026,1050]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="intermuralis">
<emphasis id="B93C3B1C982A0B2FFF06FBFA4CDC1E59" box="[223,415,1026,1050]" italics="true" pageId="17" pageNumber="16">H. intermuralis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C489C8D982A0B2FFE02FBFA4F271E59" authorityName="Say" authorityYear="1823" box="[475,612,1026,1050]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="B93C3B1C982A0B2FFE02FBFA4F271E59" box="[475,612,1026,1050]" italics="true" pageId="17" pageNumber="16">H. pallidus</emphasis>
</taxonomicName>
were commonly misidentified as
<taxonomicName id="4C489C8D982A0B2FFC24FBFA49341E59" authorityName="LeConte" authorityYear="1863" box="[1021,1143,1026,1050]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gnatho">
<emphasis id="B93C3B1C982A0B2FFC24FBFA49341E59" box="[1021,1143,1026,1050]" italics="true" pageId="17" pageNumber="16">H. gnatho</emphasis>
</taxonomicName>
. This is an easy error to make if one uses
<bibRefCitation id="EFD99AFF982A0B2FFE50FBDB4F781E79" author="Miller, W. V." box="[393,571,1058,1082]" pageId="17" pageNumber="16" pagination="723 - 726" refId="ref21941" refString="Miller, W. V. 1996. Heteroceridae. p. 723 - 726. In: N. M. Downie and R. H. Arnett, Jr. (eds.). The beetles of northeastern North America. Volume I. Sandhill Crane Press; Gainesville, FL. 880 p." type="book chapter" year="1996">Miller’s (1996)</bibRefCitation>
key. Undoubtedly, some published records for this species are erroneous.
</paragraph>
</subSubSection>
<subSubSection id="C352B485982A0B2FFF19FB9B4F9E1FD9" pageId="17" pageNumber="16" type="distribution">
<paragraph id="8BF7E70E982A0B2FFF19FB9B4C3C1F19" blockId="17.[192,1441,1122,1370]" pageId="17" pageNumber="16">
<emphasis id="B93C3B1C982A0B2FFF19FB9B4C281E39" bold="true" box="[192,363,1122,1146]" pageId="17" pageNumber="16">Distribution</emphasis>
.
<taxonomicName id="4C489C8D982A0B2FFEA1FB9A4F231E39" authorityName="LeConte" authorityYear="1863" box="[376,608,1122,1146]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gnatho">
<emphasis id="B93C3B1C982A0B2FFEA1FB9A4F231E39" box="[376,608,1122,1146]" italics="true" pageId="17" pageNumber="16">Heterocerus gnatho</emphasis>
</taxonomicName>
occurs primarily in the northern
<collectingCountry id="F35FA79E982A0B2FFC36FB9B49D11E39" box="[1007,1170,1122,1146]" name="United States of America" pageId="17" pageNumber="16">United States</collectingCountry>
and southern
<collectingCountry id="F35FA79E982A0B2FFAE4FB9B48DF1E39" box="[1341,1436,1122,1146]" name="Canada" pageId="17" pageNumber="16">Canada</collectingCountry>
, being most common from southern
<collectingRegion id="498C29EC982A0B2FFDB2FB7B4F8F1ED9" box="[619,716,1154,1178]" country="Canada" name="Ontario" pageId="17" pageNumber="16">Ontario</collectingRegion>
to
<collectingRegion id="498C29EC982A0B2FFD36FB7B4E831ED9" box="[751,960,1154,1178]" country="Canada" name="British Columbia" pageId="17" pageNumber="16">British Columbia</collectingRegion>
in
<collectingCountry id="F35FA79E982A0B2FFC3DFB7B49031ED9" box="[996,1088,1154,1178]" name="Canada" pageId="17" pageNumber="16">Canada</collectingCountry>
and west of the Rocky Mountains in the
<collectingCountry id="F35FA79E982A0B2FFE89FB5B4CB41EF9" box="[336,503,1186,1210]" name="United States of America" pageId="17" pageNumber="16">United States</collectingCountry>
(
<figureCitation id="1373FB8B982A0B2FFDDFFB5B4F1D1EF9" box="[518,606,1186,1210]" captionStart="Figures 54-57" captionStartId="39.[192,286,1864,1886]" captionTargetBox="[192,1436,1076,1850]" captionTargetId="figure-932@39.[788,832,1441,1510]" captionTargetPageId="39" captionText="Figures 54-57. Distribution by county in the United States. 54) Heterocerus pallidus. 55) Heterocerus gnatho. 56) Heterocerus longilobulus. 57) Heterocerus insolens." figureDoi="http://doi.org/10.5281/zenodo.5188388" httpUri="https://zenodo.org/record/5188388/files/figure.png" pageId="17" pageNumber="16">Fig. 55</figureCitation>
). Scattered records exist for
<collectingRegion id="498C29EC982A0B2FFC6CFB5A49531EF9" box="[949,1040,1187,1210]" country="United States of America" name="Kansas" pageId="17" pageNumber="16">Kansas</collectingRegion>
,
<collectingRegion id="498C29EC982A0B2FFBC5FB5A49181EF9" box="[1052,1115,1187,1210]" country="United States of America" name="Iowa" pageId="17" pageNumber="16">Iowa</collectingRegion>
,
<collectingRegion id="498C29EC982A0B2FFBBCFB5B49FE1EF9" box="[1125,1213,1186,1210]" country="United States of America" name="Illinois" pageId="17" pageNumber="16">Illinois</collectingRegion>
and
<collectingRegion id="498C29EC982A0B2FFB2FFB5B48E31EF9" box="[1270,1440,1186,1210]" country="United States of America" name="North Dakota" pageId="17" pageNumber="16">North Dakota</collectingRegion>
(
<emphasis id="B93C3B1C982A0B2FFF13FB3B4C831E99" bold="true" box="[202,448,1218,1242]" pageId="17" pageNumber="16">New State Record</emphasis>
- Pembina Co., ND, Goschke Dam, Tongue River Game Management Area,
<date id="FFF6C1CE982A0B2FFA9BFB3B4DBD1EB9" pageId="17" pageNumber="16" value="1974-06-28">28 June 1974</date>
, Paul K. Lago; Richland Co., ND, Walcott Dunes,
<date id="FFF6C1CE982A0B2FFCAEFB1B495F1EB9" box="[887,1052,1250,1275]" pageId="17" pageNumber="16" value="1974-05-28">28 May 1974</date>
, Paul K. Lago). In
<collectingRegion id="498C29EC982A0B2FFAC9FB1B48D81EB9" box="[1296,1435,1250,1274]" country="United States of America" name="Mississippi" pageId="17" pageNumber="16">Mississippi</collectingRegion>
, specimens were collected near freshwater and saline habitats along the
<collectingRegion id="498C29EC982A0B2FFBFCFAFB49EE1F59" box="[1061,1197,1282,1306]" country="United States of America" name="Mississippi" pageId="17" pageNumber="16">Mississippi</collectingRegion>
Gulf Coast, at a UV light in Ocean Springs (Harrison Co.), and along the
<collectingRegion id="498C29EC982A0B2FFCE4FADB4E861F79" box="[829,965,1314,1338]" country="United States of America" name="Mississippi" pageId="17" pageNumber="16">Mississippi</collectingRegion>
River (Great River Road State Park) in Bolivar County.
</paragraph>
<paragraph id="8BF7E70E982A0B2FFF19FA7B4F9E1FD9" blockId="17.[192,733,1410,1434]" box="[192,733,1410,1434]" pageId="17" pageNumber="16">
<emphasis id="B93C3B1C982A0B2FFF19FA7B4CA81FD9" bold="true" box="[192,491,1410,1434]" pageId="17" pageNumber="16">Specimens examined.</emphasis>
119 (See Appendix).
</paragraph>
</subSubSection>
</treatment>
</document>
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