treatments-xml/data/03/D6/1C/03D61C0EFF96F967350461B819076EF0.xml
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<document id="C34F9358CCC3A95B667D9421EC40FE39" ID-DOI="10.5281/zenodo.170561" ID-GBIF-Dataset="0db5b8ca-3eb1-4d70-9715-a0e3c8a9e266" ID-ISSN="1175­5326" ID-Zenodo-Dep="170561" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459713742509" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Osawa, Masayuki" docDate="2005" docId="03D61C0EFF96F967350461B819076EF0" docLanguage="en" docName="zt00801.pdf" docOrigin="Zootaxa 801" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Trichopagurus macrochela Komai &amp; Osawa, 2005, n. sp." docType="treatment" docVersion="7" lastPageNumber="10" masterDocId="FFEF6476FF94F96D340C672C1B346F76" masterDocTitle="Decapoda: Anomura: Paguridae) from the Ryukyu and Yap Islands, and redescription of T. trichophthalmus (Forest)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="3" updateTime="1698207428881" updateUser="plazi">
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<mods:title id="4A3F0221A99FC513988F11BD5DA8F78A">Decapoda: Anomura: Paguridae) from the Ryukyu and Yap Islands, and redescription of T. trichophthalmus (Forest)</mods:title>
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<mods:namePart id="C3A892E8C6201D50E3C2181B119CBB37">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="F481FF23336964C878A3B4FF23795364">Osawa, Masayuki</mods:namePart>
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<mods:date id="BCD9B728E0811D1D0A6078A9831421D0">2005</mods:date>
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<treatment id="03D61C0EFF96F967350461B819076EF0" ID-DOI="http://doi.org/10.5281/zenodo.3509185" ID-GBIF-Taxon="119334063" ID-Zenodo-Dep="3509185" LSID="urn:lsid:plazi:treatment:03D61C0EFF96F967350461B819076EF0" httpUri="http://treatment.plazi.org/id/03D61C0EFF96F967350461B819076EF0" lastPageId="10" lastPageNumber="10" pageId="2" pageNumber="3">
<subSubSection id="C365FE93FF96F96F350461B8185A6983" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BC0AD18FF96F96F350461B8181869D8" blockId="2.[264,812,1684,1710]" box="[264,812,1684,1710]" pageId="2" pageNumber="3">
<heading id="D0881A74FF96F96F350461B8181869D8" bold="true" box="[264,812,1684,1710]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B90B710AFF96F96F350461B8181869D8" bold="true" box="[264,812,1684,1710]" pageId="2" pageNumber="3">
<taxonomicName id="4C7FD69BFF96F96F350461B8196469D8" ID-CoL="892200a4-dedf-440d-bf4d-299f0acd824e" box="[264,592,1684,1710]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="macrochela" status="sp. nov.">
<emphasis id="B90B710AFF96F96F350461B8196469D8" bold="true" box="[264,592,1684,1710]" italics="true" pageId="2" pageNumber="3">Trichopagurus macrochela</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A238CC71FF96F96F365361B8199669D8" box="[607,674,1684,1710]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="1344B19DFF96F96F36BD61B8181669D8" box="[689,802,1684,1710]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[264,367,1646,1670]" captionStartId-1="5.[264,368,1732,1756]" captionStartId-2="7.[264,368,1652,1676]" captionTargetBox-0="[303,1285,316,1616]" captionTargetBox-1="[292,1308,320,1714]" captionTargetBox-2="[293,1318,292,1625]" captionTargetId-0="figure@4.[264,1323,303,1633]" captionTargetId-1="figure@5.[264,1323,307,1723]" captionTargetId-2="figure@7.[264,1323,286,1647]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." captionText-1="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." captionText-2="FIGURE 3. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, chela and carpus of right cheliped, dorsal view (setae omitted); B, right cheliped, mesial view; C, same, lateral view; D, chela and carpus of left cheliped, dorsal view (setae omitted); E, left cheliped, lateral view; F, same, mesial view (setae omitted)." httpUri-0="https://zenodo.org/record/170562/files/figure.png" httpUri-1="https://zenodo.org/record/170563/files/figure.png" httpUri-2="https://zenodo.org/record/170564/files/figure.png" pageId="2" pageNumber="3">Figs. 13</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0AD18FF96F96F350461F3185A6983" blockId="2.[264,878,1758,1782]" box="[264,878,1758,1782]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="AB6F8A36FF96F96F350461F3185A6983" box="[264,878,1758,1782]" pageId="2" pageNumber="3">
<emphasis id="B90B710AFF96F96F350461F319566980" box="[264,610,1759,1782]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C7FD69BFF96F96F350461F3196A6980" ID-CoL="8b12299f-470b-4040-bf2e-d09d0160abc1" box="[264,606,1759,1782]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="trichophthalmus">Trichopagurus trichophthalmus</taxonomicName>
.
</emphasis>
<treatmentCitation id="0ADE8B09FF96F96F368861F218116983" author="Baba" box="[644,805,1758,1781]" page="68" pageId="2" pageNumber="3" year="1982">
<bibRefCitation id="EFEED0E9FF96F96F368861F218346983" author="Baba" box="[644,768,1758,1781]" pageId="2" pageNumber="19" refString="Baba, K. (1982) Galatheids and pagurids of the Palau Islands (Crustacea: Anomura). Proceedings of the Japanese Society of the Systematic Zoology, 23, 56 - 70." type="journal article" year="1982">Baba, 1982</bibRefCitation>
: 68
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C365FE93FF96F96E3504600B18BC6E88" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BC0AD18FF96F96F3504600B1E1E681F" blockId="2.[264,1324,1831,1897]" pageId="2" pageNumber="3">
<emphasis id="B90B710AFF96F96F3504600B1A8D6836" box="[264,441,1831,1856]" italics="true" pageId="2" pageNumber="3">
<typeStatus id="54C413BAFF96F96F3504600B1A0B6836" box="[264,319,1831,1856]" pageId="2" pageNumber="3">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54C413BAFF96F96F35DA600B19426837" box="[470,630,1831,1857]" pageId="2" pageNumber="3" type="holotype">HOLOTYPE</typeStatus>
: male SL
<quantity id="4C8700FDFF96F96F370C600B18516837" box="[768,869,1831,1857]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="3" unit="mm" value="1.8">1.8 mm</quantity>
, Hiyajo, Kume­jima
<collectingCountry id="F368ED88FF96F96F307F600B1F8A6837" box="[1139,1214,1831,1857]" name="Iceland" pageId="2" pageNumber="3">Island</collectingCountry>
, Ryukyu Islands,
<quantity id="4C8700FDFF96F96F357860631A89681F" box="[372,445,1871,1897]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="10.0">10 m</quantity>
,
<date id="FFC18BD8FF96F96F35D8606319BC681F" box="[468,648,1871,1897]" pageId="2" pageNumber="3" value="1995-06-12">12 June 1995</date>
,
<collectionCode id="ED6E35DDFF96F96F369360631835681F" box="[671,769,1871,1897]" pageId="2" pageNumber="3">SCUBA</collectionCode>
diving, coll. K. Nomura, CBM­ZC 7853.
</paragraph>
<paragraph id="8BC0AD18FF97F96E350466301E136ED8" blockId="3.[264,1324,284,1830]" pageId="3" pageNumber="4">
<typeStatus id="54C413BAFF97F96E350466301A9B6E40" box="[264,431,284,310]" pageId="3" pageNumber="4" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D796691FF97F96E35B2663019286E40" box="[446,540,284,310]" pageId="3" pageNumber="4" type="male">2 males</specimenCount>
SL 1.6, 2.0 mm, 1 ovigerous female SL
<quantity id="4C8700FDFF97F96E302F66301FB76E40" box="[1059,1155,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="4" unit="mm" value="2.5">2.5 mm</quantity>
, Ahra Beach, Kume­jima
<collectingCountry id="F368ED88FF97F96E359B66681AD56E28" box="[407,481,324,350]" name="Iceland" pageId="3" pageNumber="4">Island</collectingCountry>
, Ryukyu Islands, subtidal,
<date id="FFC18BD8FF97F96E372A666818FD6E28" box="[806,969,324,350]" pageId="3" pageNumber="4" value="1995-06-15">15 June 1995</date>
, hand, coll. T. Komai, CBM­ ZC 7854;
<specimenCount id="9D796691FF97F96E358D66401AE66EF0" box="[385,466,364,390]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
SL
<quantity id="4C8700FDFF97F96E360A664019506EF0" box="[518,612,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="4" unit="mm" value="1.5">1.5 mm</quantity>
,
<specimenCount id="9D796691FF97F96E367D664019ED6EF0" box="[625,729,364,390]" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
SL
<quantity id="4C8700FDFF97F96E37006640185F6EF0" box="[780,875,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="4" unit="mm" value="1.3">1.3 mm</quantity>
, Urabishi, Kuroshima
<collectingCountry id="F368ED88FF97F96E307B66401FF56EF0" box="[1143,1217,364,390]" name="Iceland" pageId="3" pageNumber="4">Island</collectingCountry>
, Ryukyu Islands,
<quantity id="4C8700FDFF97F96E356666B81A826ED8" box="[362,438,404,430]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="4" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">34 m</quantity>
,
<date id="FFC18BD8FF97F96E35CE66B819686ED8" box="[450,604,404,430]" pageId="3" pageNumber="4" value="1998-07-10">10 July 1998</date>
, skin diving, coll. K. Nomura and T. Komai, CBM­ZC 7855.
</paragraph>
<paragraph id="8BC0AD18FF97F96E3534669018BC6E88" blockId="3.[264,1324,284,1830]" pageId="3" pageNumber="4">
<emphasis id="B90B710AFF97F96E35346690195E6EA3" box="[312,618,444,469]" italics="true" pageId="3" pageNumber="4">Other material examined</emphasis>
. 1 ovigerous female SL
<quantity id="4C8700FDFF97F96E37B866901F3D6EA0" box="[948,1033,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="4" unit="mm" value="1.7">1.7mm</quantity>
, Off Inuf, Yap Islands, lagoon,
<date id="FFC18BD8FF97F96E356A66C819316E88" box="[358,517,484,510]" pageId="3" pageNumber="4" value="1980-06-29">29 June 1980</date>
, coll. K. Baba, NSMT­Cr 14559.
</paragraph>
</subSubSection>
<subSubSection id="C365FE93FF97F9643534652019606B08" lastPageId="9" lastPageNumber="10" pageId="3" pageNumber="4" type="description">
<paragraph id="8BC0AD18FF97F96E353465201A086D38" blockId="3.[264,1324,284,1830]" pageId="3" pageNumber="4">
<emphasis id="B90B710AFF97F96E353465201AF66D53" box="[312,450,524,549]" italics="true" pageId="3" pageNumber="4">Description</emphasis>
. Gill lamellae narrowed distally with concave distolateral margins (
<figureCitation id="1344B19DFF97F96E30F065201A276D38" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
A).
</paragraph>
<paragraph id="8BC0AD18FF97F96E353465701F7D6CA8" blockId="3.[264,1324,284,1830]" pageId="3" pageNumber="4">
Shield (
<figureCitation id="1344B19DFF97F96E359865701AEC6D00" box="[404,472,604,630]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
B, C) approximately as long as broad; anterior margin between rostrum and lateral projections concave; anterolateral margins sloping; posterior margin truncate; dorsal surface weakly convex, with 1 median and 25 dorsolateral tufts of setae. Rostrum triangular or roundly subacute, distinctly overreaching lateral projections, terminating bluntly or subacutely. Lateral projections obsolete, with prominent marginal or submarginal spine. Carapace lateral lobes (
<figureCitation id="1344B19DFF97F96E36BC640819C56C48" box="[688,753,804,830]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
C) narrow, moderately calcified. Posterior carapace (
<figureCitation id="1344B19DFF97F96E355B64601A976C10" box="[343,419,844,870]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
C) generally membranous except for weakly calcified anterior part of posteromedian plate; posteromedian plate with 1 pair of tufts of setae; cardiac sulci obsolescent; sulci cardiobranchialis appearing as opaque lines on either side of posteromedian plate, extending posterolaterally; branchial region with few short setae.
</paragraph>
<paragraph id="8BC0AD18FF97F96E353464C01F066B08" blockId="3.[264,1324,284,1830]" pageId="3" pageNumber="4">
Ocular peduncles (including cornea) (
<figureCitation id="1344B19DFF97F96E36FB64C0180E6B70" box="[759,826,1004,1030]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
B) stout, about 0.7 times of shield length; not inflated basally; cornea slightly dilated; dorsal surface with tuft of long setae near base of cornea. Ocular acicles subtriangular or subovate, separated basally by width of one acicle, with small submarginal spine terminally; dorsal surface concave.
</paragraph>
<paragraph id="8BC0AD18FF97F96E353463A018246A18" blockId="3.[264,1324,284,1830]" pageId="3" pageNumber="4">
Antennular peduncles (
<figureCitation id="1344B19DFF97F96E364763A019BB6BD0" box="[587,655,1164,1190]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
B), when fully extended, overreaching ocular peduncles (including cornea) by 0.50.7 length of ultimate segment. Ultimate segment approximately twice length of penultimate segment, somewhat widened distally in lateral view, with tuft of long setae at dorsodistal lateral angle. Penultimate segment short, without setae. Basal segment with distolateral margin distinctly produced as short process, statocyst lobe weakly inflated, unarmed laterally.
</paragraph>
<paragraph id="8BC0AD18FF97F96E3534625018466850" blockId="3.[264,1324,284,1830]" pageId="3" pageNumber="4">
Antennal peduncles (
<figureCitation id="1344B19DFF97F96E3623625019456AE0" box="[559,625,1404,1430]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
B) overreaching ocular peduncles by 0.20.3 length of fifth segment, with supernumerary segmentation. Fifth and fourth segments with few setae distally. Third segment with spinule at ventromesial distal margin. Second segment with dorsolateral distal angle produced, not reaching midlength of fourth segment, terminating in simple, acute spine; dorsomesial distal angle with small spine; mesial and lateral faces with few setae. First segment with small spine at dorsolateral distal angle; ventromesial distal margin strongly produced anteriorly, with 1 minute spinule laterally. Antennal acicle reaching or slightly overreaching distal margin of cornea, weakly arcuate, distal part abruptly tapering and terminating in small spine; dorsomesial margin with sparse setae. Antennal flagellum (missing in
<typeStatus id="54C413BAFF97F96E368A61C819DA6988" box="[646,750,1764,1790]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
) 4.05.0 length of shield; each article with 35 very short to moderately long setae on distal margin.
</paragraph>
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<paragraph id="8BC0AD18FF90F9693504614219BB6811" blockId="4.[264,1323,1646,1895]" pageId="4" pageNumber="5">
<emphasis id="B90B710AFF90F969350461421AB869F0" bold="true" box="[264,396,1646,1670]" pageId="4" pageNumber="5">FIGURE 1.</emphasis>
<taxonomicName id="4C7FD69BFF90F9693592614319F469F0" box="[414,704,1647,1670]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="macrochela" status="sp. nov.">
<emphasis id="B90B710AFF90F9693592614319F469F0" box="[414,704,1647,1670]" italics="true" pageId="4" pageNumber="5">Trichopagurus macrochela</emphasis>
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<taxonomicNameLabel id="A238CC71FF90F96936C56142183069F3" box="[713,772,1646,1669]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
A, B, E, F, H, J, K, holotype male (SL 1.8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view.
</paragraph>
</caption>
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<paragraph id="8BC0AD18FF91F968350461E81FE26810" blockId="5.[264,1323,1732,1894]" pageId="5" pageNumber="6">
<emphasis id="B90B710AFF91F968350461E81AB869AA" bold="true" box="[264,396,1732,1756]" pageId="5" pageNumber="6">FIGURE 2.</emphasis>
<taxonomicName id="4C7FD69BFF91F968359161E9198B69AA" box="[413,703,1733,1756]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="macrochela" status="sp. nov.">
<emphasis id="B90B710AFF91F968359161E9198B69AA" box="[413,703,1733,1756]" italics="true" pageId="5" pageNumber="6">Trichopagurus macrochela</emphasis>
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<taxonomicNameLabel id="A238CC71FF91F96836CB61E9183669AA" box="[711,770,1733,1756]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view.
</paragraph>
</caption>
<paragraph id="8BC0AD18FF92F96B3534663019916DB0" blockId="6.[264,1324,284,1830]" pageId="6" pageNumber="7">
Mandible without distinctive features. Maxillule (
<figureCitation id="1344B19DFF92F96B3793663018DD6E40" box="[927,1001,284,310]" captionStart="FIGURE 2" captionStartId="5.[264,368,1732,1756]" captionTargetBox="[292,1308,320,1714]" captionTargetId="figure@5.[264,1323,307,1723]" captionTargetPageId="5" captionText="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." httpUri="https://zenodo.org/record/170563/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
A) with subquadrate coxal endite; endopod with outer lobe obsolete, inner lobe slightly produced, with 1 apical seta. Maxilla (
<figureCitation id="1344B19DFF92F96B357F66401A8C6EF0" box="[371,440,364,390]" captionStart="FIGURE 2" captionStartId="5.[264,368,1732,1756]" captionTargetBox="[292,1308,320,1714]" captionTargetId="figure@5.[264,1323,307,1723]" captionTargetPageId="5" captionText="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." httpUri="https://zenodo.org/record/170563/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
B) with broad proximal lobe of coxal endite; endopod overreaching anterior margin of moderately broad scaphognathite. First maxilliped (
<figureCitation id="1344B19DFF92F96B37F666B81F766ED8" box="[1018,1090,404,430]" captionStart="FIGURE 2" captionStartId="5.[264,368,1732,1756]" captionTargetBox="[292,1308,320,1714]" captionTargetId="figure@5.[264,1323,307,1723]" captionTargetPageId="5" captionText="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." httpUri="https://zenodo.org/record/170563/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
C) with moderately broad exopod; endopod distinctly overreaching anterior margin of basial endite. Second maxilliped (
<figureCitation id="1344B19DFF92F96B359766C81AD16E88" box="[411,485,484,510]" captionStart="FIGURE 2" captionStartId="5.[264,368,1732,1756]" captionTargetBox="[292,1308,320,1714]" captionTargetId="figure@5.[264,1323,307,1723]" captionTargetPageId="5" captionText="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." httpUri="https://zenodo.org/record/170563/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
D) with basis­ischium fusion incomplete; dactylus and carpus stout. Third maxilliped (
<figureCitation id="1344B19DFF92F96B35D0652019166D50" box="[476,546,524,550]" captionStart="FIGURE 2" captionStartId="5.[264,368,1732,1756]" captionTargetBox="[292,1308,320,1714]" captionTargetId="figure@5.[264,1323,307,1723]" captionTargetPageId="5" captionText="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." httpUri="https://zenodo.org/record/170563/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
E) moderately stout; carpus unarmed on dorsodistal margin; merus with strong dorsodistal spine and small ventromesial spine; ischium (
<figureCitation id="1344B19DFF92F96B304A65181FBA6D38" box="[1094,1166,564,590]" captionStart="FIGURE 2" captionStartId="5.[264,368,1732,1756]" captionTargetBox="[292,1308,320,1714]" captionTargetId="figure@5.[264,1323,307,1723]" captionTargetPageId="5" captionText="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." httpUri="https://zenodo.org/record/170563/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
F) with welldeveloped crista dentata composed of very small corneous teeth and 1 accessory tooth; basis­ischium fusion incomplete; basis apparently unarmed on ventromesial margin; exopod slightly falling short of carpus.
</paragraph>
<paragraph id="8BC0AD18FF92F96B353465F8194D6850" blockId="6.[264,1324,284,1830]" pageId="6" pageNumber="7">
Chelipeds strongly unequal in both sexes. Right cheliped (
<figureCitation id="1344B19DFF92F96B301F65F81F6F6D98" box="[1043,1115,724,750]" captionStart="FIGURE 3" captionStartId="7.[264,368,1652,1676]" captionTargetBox="[293,1318,292,1625]" captionTargetId="figure@7.[264,1323,286,1647]" captionTargetPageId="7" captionText="FIGURE 3. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, chela and carpus of right cheliped, dorsal view (setae omitted); B, right cheliped, mesial view; C, same, lateral view; D, chela and carpus of left cheliped, dorsal view (setae omitted); E, left cheliped, lateral view; F, same, mesial view (setae omitted)." httpUri="https://zenodo.org/record/170564/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
AC) very large, moderately stout. Chela elongate, subovate in dorsal view, 2.02.1 times as long as broad (greatest width at about midlength of palm). Dactyl about 0.7 length of palm, weakly curved ventrally; dorsal surface minutely granular proximally, dorsomesial margin not delimited; all surfaces with scattered tufts of short to moderately short setae; cutting edge with 2 or 3 large calcareous teeth and very small corneous teeth interspersed, terminating in small calcareous claw. Palm equal in length to carpus; dorsomesial margin not delimited, dorsolateral margin slightly delimited by multiple rows of small granules; dorsal surface weakly convex, nearly smooth medially and granular laterally and mesially, and with few short to moderately short setae; lateral and mesial surfaces minutely granular; ventral surface convex, smooth or microscopically granular, with scattered tufts of long setae extending onto fixed finger. Fixed finger with scattered tufts of short setae on dorsal surface; cutting edge with 1 large tooth proximally and row of small calcareous claw, terminating in small calcareous claw. Carpus flattened dorsoventrally, approximately equal to merus in length; dorsomesial margin clearly delimited by row of moderately large spines and some distal tubercles, dorsodistal margin unarmed, dorsolateral margin weakly delimited by row of low tubercles or protuberances; dorsal surface with some small, low tubercles laterally and scattered tufts of moderately long setae; lateral surface granular, ventrolateral margin with row of small, low protuberances; mesial surface shallowly concave, microscopically granular; distal part of ventral surface concave, bordered by distinct tuberculate ridge bearing tufts of long setae, posterior part convex, with few small tubercles and tufts of long setae. Merus with subdistal transverse ridge extending onto mesial face and few tufts of setae on dorsal surface, dorsodistal margin unarmed; lateral and mesial faces smooth, ventrolateral margin with row of small spines becoming smaller and blunt proximally and tufts of long setae, ventromesial margin with large, blunt tooth proximally and row of long stiff setae; ventral surface with tufts of long setae distally. Ischium with short spiniform setae and moderately short setae on surfaces, ventromesial margin nearly smooth. Coxa unarmed.
</paragraph>
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<paragraph id="8BC0AD18FF93F96A3504615818556985" blockId="7.[264,1323,1652,1779]" pageId="7" pageNumber="8">
<emphasis id="B90B710AFF93F96A350461581ABF69FA" bold="true" box="[264,395,1652,1676]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
<taxonomicName id="4C7FD69BFF93F96A35916159198B69FA" box="[413,703,1653,1676]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="macrochela" status="sp. nov.">
<emphasis id="B90B710AFF93F96A35916159198B69FA" box="[413,703,1653,1676]" italics="true" pageId="7" pageNumber="8">Trichopagurus macrochela</emphasis>
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<taxonomicNameLabel id="A238CC71FF93F96A36CB6158183669FD" box="[711,770,1652,1675]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
Holotype male (SL 1.8 mm), CBM­ZC 7853. A, chela and carpus of right cheliped, dorsal view (setae omitted); B, right cheliped, mesial view; C, same, lateral view; D, chela and carpus of left cheliped, dorsal view (setae omitted); E, left cheliped, lateral view; F, same, mesial view (setae omitted).
</paragraph>
</caption>
<paragraph id="8BC0AD18FF93F9653534600B1E376B70" blockId="7.[264,1323,1831,1897]" lastBlockId="8.[264,1324,284,1870]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Left cheliped (
<figureCitation id="1344B19DFF93F96A35E4600B191A6837" box="[488,558,1831,1857]" captionStart="FIGURE 3" captionStartId="7.[264,368,1652,1676]" captionTargetBox="[293,1318,292,1625]" captionTargetId="figure@7.[264,1323,286,1647]" captionTargetPageId="7" captionText="FIGURE 3. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, chela and carpus of right cheliped, dorsal view (setae omitted); B, right cheliped, mesial view; C, same, lateral view; D, chela and carpus of left cheliped, dorsal view (setae omitted); E, left cheliped, lateral view; F, same, mesial view (setae omitted)." httpUri="https://zenodo.org/record/170564/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
DF) slender. Chela about 3.5 times as long as broad (greatest breadth at about base of fingers). Dactylus slightly longer than palm; dorsomesial margin not delimited, surfaces nearly smooth, with tufts of short to long setae; cutting edge with row of small, slender corneous teeth, terminating in small corneous claw. Palm with dorsal surface weakly elevated in midline, bearing several very small tubercles and scattered short setae; dorsolateral margin of palm and proximal half of fixed finger weakly delimited by row of small, low protuberances, dorsomesial margin not delimited; lateral face with scattered low tubercles; mesial face with low protuberances and long setae; ventral surface convex, with few low protuberances and long setae. Cutting edge of fixed finger with row of small, blunt corneous teeth, terminating in small corneous claw. Carpus equal to or slightly shorter than merus; dorsolateral and dorsomesial margins each with row of small, low protuberances accompanied by spiniform setae; lateral and mesial faces with scattered low protuberances, and short to long setae; ventral surface slightly convex, with low protuberances and scattered long setae. Merus with row of low transverse ridges and few tufts of short setae on dorsal surface, dorsodistal margin unarmed; lateral face with scattered very small, low tubercles and short stiff setae, ventrolateral margin with 2 or 3 small spines distally and row of long stiff setae; mesial face with short, low transverse ridges bearing long stiff setae adjacent to ventromesial margin, ventromesial margin also with row of long stiff setae; ventral surface without distinct spines or tubercles, but with short oblique rows of long stiff setae. Ischium with short to long setae on dorsal and ventral surfaces and cluster of short spiniform setae at ventrolateral angle. Coxa unarmed.
</paragraph>
<paragraph id="8BC0AD18FF9CF965353463381ACC69F0" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
Ambulatory legs (
<figureCitation id="1344B19DFF9CF9653601633819666B58" box="[525,594,1044,1070]" captionStart="FIGURE 2" captionStartId="5.[264,368,1732,1756]" captionTargetBox="[292,1308,320,1714]" captionTargetId="figure@5.[264,1323,307,1723]" captionTargetPageId="5" captionText="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." httpUri="https://zenodo.org/record/170563/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
G, H) elongate, overreaching tip of extended left cheliped, but not reaching tip of right cheliped. Dactyli slender, in dorsal view straight, in lateral view almost straight with slightly curved claw, 1.11.3 times longer than propodi (third pair slightly longer than second); dorsal margins with row of tufts of short to moderately short setae; lateral faces with few short setae; mesial faces (
<figureCitation id="1344B19DFF9CF9653789639818F86BB8" box="[901,972,1204,1230]" captionStart="FIGURE 2" captionStartId="5.[264,368,1732,1756]" captionTargetBox="[292,1308,320,1714]" captionTargetId="figure@5.[264,1323,307,1723]" captionTargetPageId="5" captionText="FIGURE 2. Trichopagurus macrochela n. sp. Holotype male (SL 1.8 mm), CBM­ZC 7853. A, left maxillule, ventral view; B, left maxilla, ventral view; C, left first maxilliped, ventral view; D, left second maxilliped, ventral view; E, left third maxilliped, ventral view; F, same, basis and ischium, dorsal view; G, right second pereopod, lateral view; H, left third pereopod, lateral view; I, dactylus of right second pereopod, mesial view; J, dactylus of left third pereopod, mesial view." httpUri="https://zenodo.org/record/170563/files/figure.png" pageId="8" pageNumber="9">
Fig.
<date id="FFC18BD8FF9CF96537B0639818F86BB8" box="[956,972,1204,1230]" pageId="8" pageNumber="9">2</date>
</figureCitation>
I, J) also with few short setae, unarmed on second, but armed with corneous spinules dorsally on third; ventral margins each with 810 long corneous spines. Propodi 1.752.00 times length of carpi; dorsal surfaces each with widely­spaced tufts of moderately long stiff setae; lateral and mesial faces with few setae; ventral surface with 3 or 4 widely­spaced, long corneous spines, including 2 or 3 spines on ventrodistal margin. Carpi short, 0.500.70 length of meri; dorsal surfaces unarmed but with moderately short to long stiff setae; lateral faces each with row of tufts of setae on midline; ventral surfaces with few short to long, stiff setae. Meri with row of short transverse ridges and stiff setae on dorsal surfaces; lateral and mesial faces mostly naked; ventral surfaces smooth or with row of low protuberances and tufts of moderately long setae, ventrolateral distal margins unarmed. Ischia with short to long setae on dorsal and ventral margins.
</paragraph>
<paragraph id="8BC0AD18FF9CF965353461B8186C6838" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
Fourth pereopods (
<figureCitation id="1344B19DFF9CF965361661B8196A69D8" box="[538,606,1684,1710]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
D) semichelate. Dactyli each with row of minute corneous teeth on distal half of ventral margin, terminating in tiny corneous claw; no preungual process. Propodal rasp composed of single row of small corneous scales on distal 0.7 of ventral margin. Carpi with tufts of long setae on dorsal surface, dorsodistal margin unarmed. Meri with tuft of long setae on dorsodistal margin.
</paragraph>
<paragraph id="8BC0AD18FF9DF964353466301FCC6EF0" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Fifth pereopods (
<figureCitation id="1344B19DFF9DF96436086630197C6E40" box="[516,584,284,310]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
E) chelate.
<collectingCountry id="F368ED88FF9DF96436D16630182D6E40" box="[733,793,284,310]" name="Malta" pageId="9" pageNumber="10">Male</collectingCountry>
coxae (
<figureCitation id="1344B19DFF9DF9643774663018886E40" box="[888,956,284,310]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
H) slightly asymmetrical, both with short stiff setae on anterior and posterior margins. Female coxae (
<figureCitation id="1344B19DFF9DF964304A66681FBC6E28" box="[1094,1160,324,350]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">
Fig.
<date id="FFC18BD8FF9DF964307066681FBC6E28" box="[1148,1160,324,350]" pageId="9" pageNumber="10">1</date>
</figureCitation>
I) asymmetrical, left larger, with oblique transverse row of numerous long setae on ventral surface.
</paragraph>
<paragraph id="8BC0AD18FF9DF964353466B8194E6D50" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Third thoracic sternite with anterior margin slightly produced medially, unarmed. Anterior lobe of sixth thoracic sternite (
<figureCitation id="1344B19DFF9DF96436D1669018146EA0" box="[733,800,444,470]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
F, G) transversely oblong or subrectangular, with row of short setae on anterior margin. Eighth thoracic sternite (
<figureCitation id="1344B19DFF9DF964303D66C81F476E88" box="[1073,1139,484,510]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
H, I) composed of 2 rounded, contiguous lobes.
</paragraph>
<paragraph id="8BC0AD18FF9DF964353465181FC86DE8" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<collectingCountry id="F368ED88FF9DF964353465181A406D38" box="[312,372,564,590]" name="Malta" pageId="9" pageNumber="10">Male</collectingCountry>
with moderately long (34 times of coxal length) right sexual tube directed toward exterior and upward along lateral side of body, distal part transparent and coiled (
<figureCitation id="1344B19DFF9DF964351D65A81A676DE8" box="[273,339,644,670]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
E, H). Right sexual tube (
<figureCitation id="1344B19DFF9DF964368665A819F86DE8" box="[650,716,644,670]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
H) very short. Female with only left gonopore.
</paragraph>
<paragraph id="8BC0AD18FF9DF964353465801A816C48" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Abdomen twisted.
<collectingCountry id="F368ED88FF9DF9643617658019636DB0" box="[539,599,684,710]" name="Malta" pageId="9" pageNumber="10">Male</collectingCountry>
with 3 unpaired, very unequally biramous left pleopods (third to fifth pleopods). Female with 4 unpaired left pleopods, second to fourth subequally biramous; fifth pleopod very small, uniramous. Uropods markedly asymmetrical; protopods unarmed.
</paragraph>
<paragraph id="8BC0AD18FF9DF9643534646019EE6CC0" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Telson (
<figureCitation id="1344B19DFF9DF964359764601AD76C10" box="[411,483,844,870]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
J) with only slight lateral indentations; posterior lobes triangular or rounded, terminal margins oblique, each with row of 47 very small spinules, lateral margins with fringe of long setae (
<figureCitation id="1344B19DFF9DF964366364B019846CC0" box="[623,688,924,950]" captionStart="FIGURE 1" captionStartId="4.[264,367,1646,1670]" captionTargetBox="[303,1285,316,1616]" captionTargetId="figure@4.[264,1323,303,1633]" captionTargetPageId="4" captionText="FIGURE 1. Trichopagurus macrochela n. sp. A, B, E, F, H, J, K, holotype male (SL 1. 8 mm), CBM­ZC 7853; C, paratype male (SL 2.0 mm), CBM­ZC 7854; D, G, I, paratype female (SL 2.5 mm), CBM­ZC 7854. A, gill lamella; B, shield and cephalic appendages, dorsal view; C, carapace, dorsal view; D, left fourth pereopod, lateral view; E, right fifth pereopod, lateral view; F, sixth thoracic sternite, ventral view; G, coxae of third pereopods and sixth thoracic sternite, ventral view; H, I, coxae of fifth pereopods and eighth thoracic sternite, ventral view; J, telson, dorsal view; K, left posterior lobe of telson, lateral view." httpUri="https://zenodo.org/record/170562/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
K).
</paragraph>
<paragraph id="8BC0AD18FF9DF964353464E81A8D6B58" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<emphasis id="B90B710AFF9DF964353464E81AE66CAB" box="[312,466,964,989]" italics="true" pageId="9" pageNumber="10">Color in life.</emphasis>
Carapace, antennae, chelipeds and ambulatory legs generally pale reddish brown, without conspicuous markings; setae at base of cornea accentuated with dark reddish brown.
</paragraph>
<paragraph id="8BC0AD18FF9DF9643534631019606B08" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<emphasis id="B90B710AFF9DF964353463101AA26B23" box="[312,406,1084,1109]" italics="true" pageId="9" pageNumber="10">Ecology</emphasis>
. Found to inhabit small crevices or holes on dead coral. Various gastropod shells were used as housing.
</paragraph>
</subSubSection>
<subSubSection id="C365FE93FF9DF964353463A01E236BD0" box="[312,1303,1164,1190]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BC0AD18FF9DF964353463A01E236BD0" blockId="9.[264,1325,284,1870]" box="[312,1303,1164,1190]" pageId="9" pageNumber="10">
<emphasis id="B90B710AFF9DF964353463A01AF36BD3" box="[312,455,1164,1189]" italics="true" pageId="9" pageNumber="10">Distribution</emphasis>
. Presently known only from Ryukyu and Yap Islands; subtidal to
<specimenCount id="9D796691FF9DF96430D863A01E236BD0" box="[1236,1303,1164,1190]" pageId="9" pageNumber="10" type="generic">
<quantity id="4C8700FDFF9DF96430D863A01E276BD0" box="[1236,1299,1164,1190]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="m" value="10.0">10 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C365FE93FF9DF9643534639819D16B80" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BC0AD18FF9DF9643534639819D16B80" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<emphasis id="B90B710AFF9DF964353463981A876BBB" box="[312,435,1204,1229]" italics="true" pageId="9" pageNumber="10">Etymology</emphasis>
. From the Latin
<emphasis id="B90B710AFF9DF9643699639819EB6BBB" box="[661,735,1204,1229]" italics="true" pageId="9" pageNumber="10">macro</emphasis>
meaning large and
<emphasis id="B90B710AFF9DF96437DA63981F216BBB" box="[982,1045,1204,1229]" italics="true" pageId="9" pageNumber="10">chela</emphasis>
meaning cheliped and referring to the very large right cheliped.
</paragraph>
</subSubSection>
<subSubSection id="C365FE93FF9DF9673534622819076EF0" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BC0AD18FF9DF9643534622818B169A0" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<emphasis id="B90B710AFF9DF964353462281AAA6A6B" box="[312,414,1284,1309]" italics="true" pageId="9" pageNumber="10">Remarks</emphasis>
. This new species is characterized by the very large right cheliped with a dorsoventrally flattened carpus and markedly asymmetrical coxae of the female fifth pereopods, of which the left is the largest and is provided with numerous long setae on the ventral surface. Furthermore, the following notable differences are found between
<taxonomicName id="4C7FD69BFF9DF964311962501AA56ACB" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macrochela">
<emphasis id="B90B710AFF9DF964311962501AA56ACB" italics="true" pageId="9" pageNumber="10">T. macrochela</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7FD69BFF9DF96435DD628819856ACB" box="[465,689,1444,1469]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF9DF96435DD628819856ACB" box="[465,689,1444,1469]" italics="true" pageId="9" pageNumber="10">T. trichophthalmus</emphasis>
</taxonomicName>
. The dorsomesial margin of the carpus of the right cheliped is distinctly delimited by a row of spines in
<taxonomicName id="4C7FD69BFF9DF964378C62E01F136A93" box="[896,1063,1484,1509]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macrochela">
<emphasis id="B90B710AFF9DF964378C62E01F136A93" box="[896,1063,1484,1509]" italics="true" pageId="9" pageNumber="10">T. macrochela</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C7FD69BFF9DF964309362E01A51697B" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF9DF964309362E01A51697B" italics="true" pageId="9" pageNumber="10">T. trichophthalmus</emphasis>
</taxonomicName>
, the carpus is subcylindrical or slightly flattened, without distinct delineation of the dorsomesial margin. The surfaces of the right cheliped are granular or tuberculate in
<taxonomicName id="4C7FD69BFF9DF964350461681A86692B" box="[264,434,1604,1629]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macrochela">
<emphasis id="B90B710AFF9DF964350461681A86692B" box="[264,434,1604,1629]" italics="true" pageId="9" pageNumber="10">A. macrochela</emphasis>
</taxonomicName>
, rather than smooth and polished in
<taxonomicName id="4C7FD69BFF9DF964376D61681F0A692B" box="[865,1086,1604,1629]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF9DF964376D61681F0A692B" box="[865,1086,1604,1629]" italics="true" pageId="9" pageNumber="10">T. trichophthalmus</emphasis>
</taxonomicName>
. The chelipeds and ambulatory legs are distinctly less setose in
<taxonomicName id="4C7FD69BFF9DF964371F6140188C69F3" box="[787,952,1644,1669]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macrochela">
<emphasis id="B90B710AFF9DF964371F6140188C69F3" box="[787,952,1644,1669]" italics="true" pageId="9" pageNumber="10">T. macrochela</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C7FD69BFF9DF964301061401FCE69F3" box="[1052,1274,1644,1669]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF9DF964301061401FCE69F3" box="[1052,1274,1644,1669]" italics="true" pageId="9" pageNumber="10">T. trichophthalmus</emphasis>
</taxonomicName>
. In addition, males of
<taxonomicName id="4C7FD69BFF9DF96435F861B819AB69DB" box="[500,671,1684,1709]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="macrochela">
<emphasis id="B90B710AFF9DF96435F861B819AB69DB" box="[500,671,1684,1709]" italics="true" pageId="9" pageNumber="10">T. macrochela</emphasis>
</taxonomicName>
possess unpaired, unequally biramous pleopods, but those of
<taxonomicName id="4C7FD69BFF9DF96435616190197E69A3" box="[365,586,1724,1749]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF9DF96435616190197E69A3" box="[365,586,1724,1749]" italics="true" pageId="9" pageNumber="10">T. trichophthalmus</emphasis>
</taxonomicName>
have uniramous pleopods.
</paragraph>
<paragraph id="8BC0AD18FF9DF967353461C819076EF0" blockId="9.[264,1325,284,1870]" lastBlockId="10.[264,1323,284,390]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<bibRefCitation id="EFEED0E9FF9DF964353461C81AF96988" author="Baba" box="[312,461,1764,1790]" pageId="9" pageNumber="19" refString="Baba, K. (1982) Galatheids and pagurids of the Palau Islands (Crustacea: Anomura). Proceedings of the Japanese Society of the Systematic Zoology, 23, 56 - 70." type="journal article" year="1982">Baba (1982)</bibRefCitation>
recorded
<taxonomicName id="4C7FD69BFF9DF964364561C8181E698B" box="[585,810,1764,1789]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF9DF964364561C8181E698B" box="[585,810,1764,1789]" italics="true" pageId="9" pageNumber="10">T. trichophthalmus</emphasis>
</taxonomicName>
from the
<collectingCountry id="F368ED88FF9DF96437AB61C818DC6988" box="[935,1000,1764,1790]" name="Palau" pageId="9" pageNumber="10">Palau</collectingCountry>
and Yap Islands based on four specimens. Two of the four specimens are now deposited in the collection of the National Science Museum, Tokyo, although other two were not located. Reexamination of these two specimens revealed that the male (NSMT­Cr 14558) from
<collectingCountry id="F368ED88FF9EF967303066301F496E40" box="[1084,1149,284,310]" name="Palau" pageId="10" pageNumber="11">Palau</collectingCountry>
is referable to true
<taxonomicName id="4C7FD69BFF9EF9673532666819286E2B" box="[318,540,324,349]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="trichophthalmus">
<emphasis id="B90B710AFF9EF9673532666819286E2B" box="[318,540,324,349]" italics="true" pageId="10" pageNumber="11">T. trichophthalmus</emphasis>
</taxonomicName>
, but an ovigerous female (NSMT­Cr 14559) from the Yap Islands represents
<taxonomicName id="4C7FD69BFF9EF967358A664019186EF3" box="[390,556,364,389]" class="Malacostraca" family="Paguridae" genus="Trichopagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="macrochela">
<emphasis id="B90B710AFF9EF967358A664019186EF3" box="[390,556,364,389]" italics="true" pageId="10" pageNumber="11">T. macrochela</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>