266 lines
31 KiB
XML
266 lines
31 KiB
XML
<document id="8CEDFEEA59539864D3C7D8C2D65AF062" ID-DOI="10.5281/zenodo.197721" ID-GBIF-Dataset="7fbbbf17-731f-4905-a25b-cea2c544bf38" ID-ISSN="1175-5326" ID-Zenodo-Dep="197721" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460276041053" checkinUser="plazi" docAuthor="Symonds, Malcolm Francis & Pacaud, Jean-Michel" docDate="2010" docId="03CFAD7AC0571F69FF1AF9914C5CFB80" docLanguage="en" docName="zt02606p068.pdf" docOrigin="Zootaxa 2606" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tomostoma Deshayes 1824" docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="FFF6D502C0571F68FF8DFFA34871FFB2" masterDocTitle="New species of Neritidae (Neritimorpha) from the Ypresian and Bartonian of the Paris and Basse-Loire Basins, France" masterLastPageNumber="68" masterPageNumber="55" pageNumber="55" updateTime="1698248446951" updateUser="plazi">
|
|
<mods:mods id="01C437E4BF49E21F724174F82EA5A45D" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="F4A72649172BE4B4C144883581EFEE3C">
|
|
<mods:title id="C6B50645364AA2E0B474DEB55CED9EEC">New species of Neritidae (Neritimorpha) from the Ypresian and Bartonian of the Paris and Basse-Loire Basins, France</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="1B19AAF47D23B29AAD2BE9EB598492FD" type="personal">
|
|
<mods:role id="2D36554D43346CECCCA6EDECFFABF7C8">
|
|
<mods:roleTerm id="A392E41E1007FD1AC9DA355D396CA3E0">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="56A16F1B7F256C91D9A92D1E2863D436">Symonds, Malcolm Francis</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="8BC8CF64F7D9532495787D25EA2F8DD4" type="personal">
|
|
<mods:role id="1630CFFFD4EA65909158B03540B35A0D">
|
|
<mods:roleTerm id="A844C12C33C0B65C803DC06E22201184">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="61F7F196658B1F785A0AEAD7EC1FF072">Pacaud, Jean-Michel</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="512420BEE86EF544F44F806B54A23D00">text</mods:typeOfResource>
|
|
<mods:relatedItem id="2AAC562BE1C4D5DCE41A07413A175EAE" type="host">
|
|
<mods:titleInfo id="FC28A92E4DF421C84C2FE2D824286409">
|
|
<mods:title id="F7426D5C7D7A6D78707FE6A69D2A9A35">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="C11DE7FB0210052F84C1AA708BE60809">
|
|
<mods:date id="E15B2B02E2206F32317A340B5C7254C5">2010</mods:date>
|
|
<mods:detail id="0598F27EC3ECB271B7E08E7D162FE0A1" type="volume">
|
|
<mods:number id="CF8B695FC50F09FD725832B674309BB1">2606</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="CD028B18DE4D97EEAC8BBA52B49475EB" unit="page">
|
|
<mods:start id="0A766F8B06D19CA97347310D1AA2194A">55</mods:start>
|
|
<mods:end id="69FC2F24C542B18F09A87DE30EADFD51">68</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="89B6352C806988922642C90C5E08C972">journal article</mods:classification>
|
|
<mods:identifier id="8E17C6ADAC39BC41F851784B5C9B6A16" type="DOI">10.5281/zenodo.197721</mods:identifier>
|
|
<mods:identifier id="95A03C6FE3814E303A0D573290DC324F" type="GBIF-Dataset">7fbbbf17-731f-4905-a25b-cea2c544bf38</mods:identifier>
|
|
<mods:identifier id="F4DFEEEB19918BE142C2C565223AE274" type="ISSN">1175-5326</mods:identifier>
|
|
<mods:identifier id="9AD91B4052993490204550F879437222" type="Zenodo-Dep">197721</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="03CFAD7AC0571F69FF1AF9914C5CFB80" ID-DOI="http://doi.org/10.5281/zenodo.6209649" ID-GBIF-Taxon="119396031" ID-Zenodo-Dep="6209649" LSID="urn:lsid:plazi:treatment:03CFAD7AC0571F69FF1AF9914C5CFB80" httpUri="http://treatment.plazi.org/id/03CFAD7AC0571F69FF1AF9914C5CFB80" lastPageId="1" lastPageNumber="55" pageId="0" pageNumber="55">
|
|
<subSubSection id="C37C4FE7C0571F68FF1AF9914A4CF9FE" box="[151,573,1586,1612]" pageId="0" pageNumber="55" type="nomenclature">
|
|
<paragraph id="8BD91C6CC0571F68FF1AF9914A4CF9FE" blockId="0.[151,573,1586,1612]" box="[151,573,1586,1612]" pageId="0" pageNumber="55">
|
|
<heading id="D091AB00C0571F68FF1AF9914A4CF9FE" bold="true" box="[151,573,1586,1612]" fontSize="11" level="1" pageId="0" pageNumber="55" reason="1">
|
|
<emphasis id="B912C07EC0571F68FF1AF9914A4CF9FE" bold="true" box="[151,573,1586,1612]" pageId="0" pageNumber="55">
|
|
Genus
|
|
<taxonomicName id="4C6667EFC0571F68FF63F9914A4CF9FE" authority="Deshayes, 1824" authorityName="Deshayes" authorityYear="1824" box="[238,573,1586,1612]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="0" pageNumber="55" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0571F68FF63F9914908F9FE" bold="true" box="[238,377,1586,1612]" italics="true" pageId="0" pageNumber="55">Tomostoma</emphasis>
|
|
Deshayes, 1824
|
|
</taxonomicName>
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C37C4FE7C0571F68FF1AF9D94885F908" pageId="0" pageNumber="55" type="materials_examined">
|
|
<paragraph id="8BD91C6CC0571F68FF1AF9D94885F908" blockId="0.[151,1436,1658,2020]" pageId="0" pageNumber="55">
|
|
<emphasis id="B912C07EC0571F68FF1AF9D9494BF926" bold="true" box="[151,314,1658,1684]" pageId="0" pageNumber="55">
|
|
<typeStatus id="54DDA2CEC0571F68FF1AF9D948A5F926" box="[151,212,1658,1684]" pageId="0" pageNumber="55">Type</typeStatus>
|
|
species.
|
|
</emphasis>
|
|
By subsequent designation,
|
|
<bibRefCitation id="EFF7619DC0571F68FD00F9D94B0AF926" author="Fischer" box="[653,891,1658,1684]" pageId="0" pageNumber="55" refString="Fischer, S. D. (1885) Manuel de Conchyliologie et de Paleontologie Conchyliologique ou Histoire Naturelle des Mollusques Vivantes et Fossiles. Savy, Paris, 1369 pp., 23 pls." type="book" year="1885">Fischer (1885: 803)</bibRefCitation>
|
|
:
|
|
<taxonomicName id="4C6667EFC0571F68FC0AF9D84D43F926" authority="Deshayes, 1824" authorityName="Deshayes" authorityYear="1824" box="[903,1330,1658,1684]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="0" pageNumber="55" phylum="Mollusca" rank="species" species="neritoides">
|
|
<emphasis id="B912C07EC0571F68FC0AF9D84C1CF926" box="[903,1133,1659,1684]" italics="true" pageId="0" pageNumber="55">Pileolus neritoides,</emphasis>
|
|
Deshayes, 1824
|
|
</taxonomicName>
|
|
. Eocene, Europe.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C37C4FE7C0571F68FF48F9664B3BF8FD" pageId="0" pageNumber="55" type="diagnosis">
|
|
<paragraph id="8BD91C6CC0571F68FF48F9664AACF8B6" blockId="0.[151,1436,1658,2020]" pageId="0" pageNumber="55">
|
|
<emphasis id="B912C07EC0571F68FF48F9664935F96D" bold="true" box="[197,324,1733,1759]" pageId="0" pageNumber="55">Diagnosis.</emphasis>
|
|
Capuliform, smooth, apex not terminal, aperture trapezoidal, inner lip with sinus,
|
|
<typeStatus id="54DDA2CEC0571F68FA80F9664D31F96D" box="[1293,1344,1733,1759]" pageId="0" pageNumber="55">type</typeStatus>
|
|
species
|
|
<quantity id="4C9EB189C0571F68FF1AF949497EF8B6" box="[151,271,1770,1796]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="0" pageNumber="55" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7 to 9 mm</quantity>
|
|
in length. (Amended after
|
|
<bibRefCitation id="EFF7619DC0571F68FDC1F9494AA1F8B6" author="Keen" box="[588,720,1770,1796]" pageId="0" pageNumber="55" refString="Keen, M. (1960) In: Moore, R. (Ed.), Treatise on Invertebrate Palaeontology. Part 1. Mollusca 1. The Geological Society of America & The University of Kansas, U. S. A., 351 pp., 216 figs." type="book" year="1960">Keen 1960</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BD91C6CC0571F68FF1AF8B34B3BF8FD" blockId="0.[151,1436,1658,2020]" pageId="0" pageNumber="55">
|
|
<taxonomicName id="4C6667EFC0571F68FF1AF8B3496BF89B" box="[151,282,1808,1833]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="0" pageNumber="55" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0571F68FF1AF8B3496BF89B" box="[151,282,1808,1833]" italics="true" pageId="0" pageNumber="55">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
differs from most other genera in
|
|
<taxonomicName id="4C6667EFC0571F68FD26F8B34B69F898" box="[683,792,1808,1834]" class="Gastropoda" family="Neritidae" kingdom="Animalia" order="Cycloneritimorpha" pageId="0" pageNumber="55" phylum="Mollusca" rank="family">Neritidae</taxonomicName>
|
|
in being capuliform. For the differences from the other Tertiary capuliform genus see
|
|
<taxonomicName id="4C6667EFC0571F68FE76F8964AF5F8FC" box="[507,644,1845,1870]" class="Gastropoda" family="Neritidae" genus="Cuisenerita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="0" pageNumber="55" phylum="Mollusca" rank="genus" status="gen. nov.">
|
|
<emphasis id="B912C07EC0571F68FE76F8964AF5F8FC" box="[507,644,1845,1870]" italics="true" pageId="0" pageNumber="55">Cuisenerita</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A2217D05C0571F68FD06F8964A87F8FD" box="[651,758,1845,1871]" pageId="0" pageNumber="55" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
below.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C37C4FE7C0571F69FF4BF8F94C5CFB80" lastPageId="1" lastPageNumber="56" pageId="0" pageNumber="55" type="discussion">
|
|
<paragraph id="8BD91C6CC0571F69FF4BF8F94C5CFB80" blockId="0.[151,1436,1658,2020]" lastBlockId="1.[151,1437,152,1074]" lastPageId="1" lastPageNumber="56" pageId="0" pageNumber="55">
|
|
<emphasis id="B912C07EC0571F68FF4BF8F9494EF8C6" bold="true" box="[198,319,1882,1908]" pageId="0" pageNumber="55">Remarks.</emphasis>
|
|
<bibRefCitation id="EFF7619DC0571F68FEC1F8F94A50F8C6" author="Cossmann" box="[332,545,1882,1908]" pageId="0" pageNumber="55" refString="Cossmann, M. (1925) Essais de Paleoconchologie Comparee. Presses Universitaires de France, Paris, 345 pp., 11 pls." type="book" year="1925">Cossmann (1925)</bibRefCitation>
|
|
and
|
|
<bibRefCitation id="EFF7619DC0571F68FDE8F8F94A8CF8C6" author="Keen" box="[613,765,1882,1908]" pageId="0" pageNumber="55" refString="Keen, M. (1960) In: Moore, R. (Ed.), Treatise on Invertebrate Palaeontology. Part 1. Mollusca 1. The Geological Society of America & The University of Kansas, U. S. A., 351 pp., 216 figs." type="book" year="1960">Keen (1960)</bibRefCitation>
|
|
regarded
|
|
<taxonomicName id="4C6667EFC0571F68FCF3F8F84C70F8C6" box="[894,1025,1883,1908]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="0" pageNumber="55" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0571F68FCF3F8F84C70F8C6" box="[894,1025,1883,1908]" italics="true" pageId="0" pageNumber="55">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
as a subgenus of
|
|
<taxonomicName id="4C6667EFC0571F68FB63F8F8493AF828" authority="G. B. Sowerby, 1823" authorityName="G. B. Sowerby" authorityYear="1823" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="0" pageNumber="55" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0571F68FB63F8F84D3FF8C6" box="[1262,1358,1883,1908]" italics="true" pageId="0" pageNumber="55">Pileolus</emphasis>
|
|
<bibRefCitation id="EFF7619DC0571F68FAD6F8F9493AF828" author="Sowerby" pageId="0" pageNumber="55" refString="Sowerby, G. B. I (1823) The Genera of Recent and Fossil Shells. Sowerby, London, parts 12 - 21." type="book chapter" year="1823">G. B. Sowerby, 1823</bibRefCitation>
|
|
</taxonomicName>
|
|
.
|
|
<bibRefCitation id="EFF7619DC0571F68FEDAF8234A35F828" box="[343,580,1920,1946]" pageId="0" pageNumber="55" refString="Bandel, K., Grundel, J. & Maxwell, P. (2000) Gastropods from the upper Early Jurassic / early Middle Jurassic of Kaiwara Valley, North Canterbury, New Zealand. Freiberger Forschungshefte, C 490, 67 - 132." type="journal article">
|
|
Bandel
|
|
<emphasis id="B912C07EC0571F68FE3FF823499FF82B" box="[434,494,1920,1945]" italics="true" pageId="0" pageNumber="55">et al.</emphasis>
|
|
(2000)
|
|
</bibRefCitation>
|
|
erected a new family
|
|
<taxonomicName id="4C6667EFC0571F68FCC1F8234BB0F828" box="[844,961,1920,1946]" class="Gastropoda" family="Pileolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="0" pageNumber="55" phylum="Mollusca" rank="family">Pileolidae</taxonomicName>
|
|
based on the genus
|
|
<taxonomicName id="4C6667EFC0571F68FB3EF8234D65F82B" box="[1203,1300,1920,1945]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="0" pageNumber="55" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0571F68FB3EF8234D65F82B" box="[1203,1300,1920,1945]" italics="true" pageId="0" pageNumber="55">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
but did not comment on whether they regarded
|
|
<taxonomicName id="4C6667EFC0571F68FDDCF8064AA8F80C" box="[593,729,1957,1982]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="0" pageNumber="55" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0571F68FDDCF8064AA8F80C" box="[593,729,1957,1982]" italics="true" pageId="0" pageNumber="55">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
as a subgenus of
|
|
<taxonomicName id="4C6667EFC0571F68FC35F8064C6DF80C" box="[952,1052,1957,1982]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="0" pageNumber="55" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0571F68FC35F8064C6DF80C" box="[952,1052,1957,1982]" italics="true" pageId="0" pageNumber="55">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
. They placed
|
|
<taxonomicName id="4C6667EFC0571F68FB41F8064D37F80D" box="[1228,1350,1957,1983]" class="Gastropoda" family="Pileolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="0" pageNumber="55" phylum="Mollusca" rank="family">Pileolidae</taxonomicName>
|
|
within Neritoidea
|
|
<bibRefCitation id="EFF7619DC0571F68FE97F8694999F856" author="Rafinesque" box="[282,488,1994,2020]" pageId="0" pageNumber="55" refString="Rafinesque, C. S. (1815) Analyse de la Nature ou Tableau de l'Univers et des Corps Organises. Privately published, Palermo, 224 pp." type="book" year="1815">Rafinesque, 1815</bibRefCitation>
|
|
for which they gave the diagnosis: “In these cycloneritimorphs the internal shell walls are resorbed” (2000: 85). However Sowerby (1823: 443) stated that
|
|
<taxonomicName id="4C6667EFC0561F69FB96FF3B4C0FFF03" box="[1051,1150,152,177]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FB96FF3B4C0FFF03" box="[1051,1150,152,177]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
has an “internal spire”. This is clearly shown by
|
|
<bibRefCitation id="EFF7619DC0561F69FE49FF1E4B71FF65" author="Pana" box="[452,768,189,215]" pageId="1" pageNumber="56" refString="Pana, I. (1998) New taxa of small-sized gastropods from the Lower Cretaceous deposits of South Dobrogea. Revue Roumaine de Geologie, 42, 69 - 90." type="journal article" year="1998">Pană (1998: Pl. 2, Fig. 13)</bibRefCitation>
|
|
in a section through the teleoconch of
|
|
<taxonomicName id="4C6667EFC0561F69FB59FF1E4D44FF64" box="[1236,1333,189,214]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FB59FF1E4D44FF64" box="[1236,1333,189,214]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
. On this basis
|
|
<taxonomicName id="4C6667EFC0561F69FF51FF414927FF4E" box="[220,342,226,252]" class="Gastropoda" family="Pileolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="1" pageNumber="56" phylum="Mollusca" rank="family">Pileolidae</taxonomicName>
|
|
should not be included within Neritoidea.
|
|
<bibRefCitation id="EFF7619DC0561F69FCE5FF414C42FF4E" author="Symonds" box="[872,1075,226,252]" pageId="1" pageNumber="56" refString="Symonds, M. F. (2009) The Neritopsidae and the Neritidae of the Bracklesham Group (Early and Middle Eocene) of the Hampshire Basin. Cainozoic Research, 6, 37 - 51." type="journal article" year="2009">Symonds (2009)</bibRefCitation>
|
|
did not see a close similarity between
|
|
<taxonomicName id="4C6667EFC0561F69FE8DFEAB4911FE93" box="[256,352,264,289]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FE8DFEAB4911FE93" box="[256,352,264,289]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C6667EFC0561F69FE16FEAB4A6FFE93" box="[411,542,264,289]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FE16FEAB4A6FFE93" box="[411,542,264,289]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
as the protoconch of
|
|
<taxonomicName id="4C6667EFC0561F69FCAFFEAB4BF3FE93" box="[802,898,264,289]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FCAFFEAB4BF3FE93" box="[802,898,264,289]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
is spherical and rather large in relation to the rest of the shell while the protoconch of
|
|
<taxonomicName id="4C6667EFC0561F69FD11FE8E4B56FEF4" box="[668,807,301,326]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FD11FE8E4B56FEF4" box="[668,807,301,326]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
is obovate and typical of those genera within the
|
|
<taxonomicName id="4C6667EFC0561F69FF1AFEF14975FEDE" box="[151,260,338,364]" class="Gastropoda" family="Neritidae" kingdom="Animalia" order="Cycloneritimorpha" pageId="1" pageNumber="56" phylum="Mollusca" rank="family">Neritidae</taxonomicName>
|
|
that have a planktotrophic larval stage. The teleoconch of the former is distinctly patelliform unlike that of the latter, which is capuliform. The aperture of
|
|
<taxonomicName id="4C6667EFC0561F69FCB3FEDB4BD2FE23" box="[830,931,376,401]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FCB3FEDB4BD2FE23" box="[830,931,376,401]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
is rather narrow and semilunar whilst in
|
|
<taxonomicName id="4C6667EFC0561F69FF1AFE3E496FFE04" box="[151,286,413,438]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FF1AFE3E496FFE04" box="[151,286,413,438]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
the aperture is much broader.
|
|
<taxonomicName id="4C6667EFC0561F69FD19FE3E4A86FE04" box="[660,759,413,438]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FD19FE3E4A86FE04" box="[660,759,413,438]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
has a straight or slightly convex septum edge, which is smooth or regularly crenulated; in
|
|
<taxonomicName id="4C6667EFC0561F69FDBFFE614AC7FE69" box="[562,694,450,475]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FDBFFE614AC7FE69" box="[562,694,450,475]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
the septum edge is concave and the dentition distinct but very irregular. Additionally in
|
|
<taxonomicName id="4C6667EFC0561F69FE59FE4B4A49FDB3" box="[468,568,488,513]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FE59FE4B4A49FDB3" box="[468,568,488,513]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
the septum is thickened to form a prominent basal plate, a feature not present in
|
|
<taxonomicName id="4C6667EFC0561F69FE94FDAE49D3FD94" box="[281,418,525,550]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FE94FDAE49D3FD94" box="[281,418,525,550]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
. Finally, as mentioned above,
|
|
<taxonomicName id="4C6667EFC0561F69FCADFDAE4BF4FD94" box="[800,901,525,550]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FCADFDAE4BF4FD94" box="[800,901,525,550]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
has a columella while the internal walls of
|
|
<taxonomicName id="4C6667EFC0561F69FF1AFD914956FDF9" box="[151,295,562,587]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FF1AFD914956FDF9" box="[151,295,562,587]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
are entirely reabsorbed.
|
|
<collectingCountry id="F3715CFCC0561F69FDFCFD914AE5FDFE" box="[625,660,562,588]" name="American Samoa" pageId="1" pageNumber="56">As</collectingCountry>
|
|
the morphology of both the protoconch and the teleoconch corresponds to
|
|
<taxonomicName id="4C6667EFC0561F69FEC3FDFB4AE3FDC0" authority="Symonds (2009)" authorityName="Symonds" authorityYear="2009" box="[334,658,600,626]" class="Gastropoda" family="Neritidae" kingdom="Animalia" order="Cycloneritimorpha" pageId="1" pageNumber="56" phylum="Mollusca" rank="family">
|
|
Neritidae,
|
|
<bibRefCitation id="EFF7619DC0561F69FE41FDFB4AE3FDC0" author="Symonds" box="[460,658,600,626]" pageId="1" pageNumber="56" refString="Symonds, M. F. (2009) The Neritopsidae and the Neritidae of the Bracklesham Group (Early and Middle Eocene) of the Hampshire Basin. Cainozoic Research, 6, 37 - 51." type="journal article" year="2009">Symonds (2009)</bibRefCitation>
|
|
</taxonomicName>
|
|
regarded
|
|
<taxonomicName id="4C6667EFC0561F69FC86FDFB4BE2FDC3" box="[779,915,600,625]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FC86FDFB4BE2FDC3" box="[779,915,600,625]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
as being in
|
|
<taxonomicName id="4C6667EFC0561F69FBA8FDFB4CE5FDC0" box="[1061,1172,600,626]" class="Gastropoda" family="Neritidae" kingdom="Animalia" order="Cycloneritimorpha" pageId="1" pageNumber="56" phylum="Mollusca" rank="family">Neritidae</taxonomicName>
|
|
rather than
|
|
<taxonomicName id="4C6667EFC0561F69FAA8FDFB4DEDFDC0" box="[1317,1436,600,626]" class="Gastropoda" family="Pileolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="1" pageNumber="56" phylum="Mollusca" rank="family">Pileolidae</taxonomicName>
|
|
and this is followed here.
|
|
<bibRefCitation id="EFF7619DC0561F69FE5BFDDE4B6EFD25" author="Woods" box="[470,799,637,663]" pageId="1" pageNumber="56" refString="Woods, A. J. C. & Saul, L. R. (1986) New Neritidae from southwestern North America. Journal of Paleontology, 60, 636 - 655." type="journal article" year="1986">Woods & Saul (1986: 650)</bibRefCitation>
|
|
commented that “Deshayes is usually credited with proposing
|
|
<taxonomicName id="4C6667EFC0561F69FE9EFD0149E7FD09" box="[275,406,674,699]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FE9EFD0149E7FD09" box="[275,406,674,699]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
in 1824; but he did not do so.” They considered that it was not until 1864 that he validly proposed
|
|
<taxonomicName id="4C6667EFC0561F69FE80FD6B49E2FD53" box="[269,403,712,737]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FE80FD6B49E2FD53" box="[269,403,712,737]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
and that as a result
|
|
<emphasis id="B912C07EC0561F69FD06FD6B4A92FD53" box="[651,739,712,737]" italics="true" pageId="1" pageNumber="56">Culana</emphasis>
|
|
<bibRefCitation id="EFF7619DC0561F69FD61FD6B4B03FD50" author="Gray" box="[748,882,712,738]" pageId="1" pageNumber="56" refString="Gray, J. E. (1842) Synopsis of the Contents of the British Museum. 44 th edition. British Museum, London, 307 pp." type="book" year="1842">Gray, 1842</bibRefCitation>
|
|
(
|
|
<typeStatus id="54DDA2CEC0561F69FC0EFD6B4BC9FD50" box="[899,952,712,738]" pageId="1" pageNumber="56">type</typeStatus>
|
|
species
|
|
<taxonomicName id="4C6667EFC0561F69FBACFD6B48A2FCB5" authority="Defrance, 1818" authorityName="Defrance" authorityYear="1818" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="species" species="altavillensis">
|
|
<emphasis id="B912C07EC0561F69FBACFD6B4D6FFD53" box="[1057,1310,712,737]" italics="true" pageId="1" pageNumber="56">Pileolus altavillensis</emphasis>
|
|
<bibRefCitation id="EFF7619DC0561F69FAABFD6B48A2FCB5" author="Defrance" pageId="1" pageNumber="56" refString="Defrance, F. (1818) In: Levrault (Ed.), [Palaeontological Descriptions] Dictionnaire des Sciences Naturelles. Levrault, Strasbourg, 11, 615 pp." type="book" year="1818">Defrance, 1818</bibRefCitation>
|
|
</taxonomicName>
|
|
) should have priority.
|
|
<bibRefCitation id="EFF7619DC0561F69FE52FD4E4ADDFCB5" author="Deshayes" box="[479,684,749,775]" pageId="1" pageNumber="56" refString="Deshayes, G. P. (1824 a) Note sur un nouveau genre de la famille des Neritacees. Annales des Sciences Naturelles, 1, 187 - 192." type="journal article" year="1824" yearSuffix="a">Deshayes (1824a</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF7619DC0561F69FD34FD4E4B75FCB5" author="Deshayes" box="[697,772,749,775]" pageId="1" pageNumber="56" refString="Deshayes, G. P. (1824 b) Atlas des tomes 1, 2, 3 comprenant 83 planches. Annales des Sciences Naturelles, 1, pls 1 - 83." type="journal article" year="1824" yearSuffix="b">1824b</bibRefCitation>
|
|
) made it clear that he had intended to propose the name
|
|
<emphasis id="B912C07EC0561F69FF1AFCB14955FC99" box="[151,292,786,811]" italics="true" pageId="1" pageNumber="56">
|
|
<taxonomicName id="4C6667EFC0561F69FF1AFCB14951FC99" box="[151,288,786,811]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">Tomostoma</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
but he did not do so at the time as he considered that it was synonymous with
|
|
<taxonomicName id="4C6667EFC0561F69FB38FCB14D64FC99" box="[1205,1301,786,811]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FB38FCB14D64FC99" box="[1205,1301,786,811]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
, which had just been proposed by Sowerby (Sowerby, 1823). Later,
|
|
<bibRefCitation id="EFF7619DC0561F69FCBEFC9B4C59FCE0" author="Deshayes" box="[819,1064,824,850]" pageId="1" pageNumber="56" refString="Deshayes, G. P. (1864) Description des Animaux sans Vertebres Decouverts dans le Bassin de Paris. J. B. Bailliere et fils, Paris, 3, 1 - 200, pls 63 - 85." type="journal article" year="1864">Deshayes (1864: 25)</bibRefCitation>
|
|
considered that there were two distinct forms of
|
|
<taxonomicName id="4C6667EFC0561F69FEEEFCFE49B5FCC4" box="[355,452,861,886]" class="Gastropoda" family="Pileolidae" genus="Pileolus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FEEEFCFE49B5FCC4" box="[355,452,861,886]" italics="true" pageId="1" pageNumber="56">Pileolus</emphasis>
|
|
</taxonomicName>
|
|
: those from the Mesozoic with a circular base and central apex and those from the Tertiary with an elliptical base and an apex towards the posterior end; to the latter group could be attached the name
|
|
<taxonomicName id="4C6667EFC0561F69FF53FC0B4913FC73" box="[222,354,936,961]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FF53FC0B4913FC73" box="[222,354,936,961]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
“que nous avons proposé autrefois”. Article 11.6.1 of the International Code of Zoological Nomenclature (the Code) provides that if a name first published as a junior synonym is subsequently used as a valid name before 1961, it dates from its first publication as a synonym. Accordingly
|
|
<taxonomicName id="4C6667EFC0561F69FB5BFC514D10FBB9" box="[1238,1377,1010,1035]" class="Gastropoda" family="Neritidae" genus="Tomostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="1" pageNumber="56" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B912C07EC0561F69FB5BFC514D10FBB9" box="[1238,1377,1010,1035]" italics="true" pageId="1" pageNumber="56">Tomostoma</emphasis>
|
|
</taxonomicName>
|
|
was published in 1824 and
|
|
<emphasis id="B912C07EC0561F69FE2FFBBB4989FB83" box="[418,504,1048,1073]" italics="true" pageId="1" pageNumber="56">Culana</emphasis>
|
|
is here considered a subjective junior synonym.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |