157 lines
20 KiB
XML
157 lines
20 KiB
XML
<document id="72C13AFF17C903DBF17055745AF0C719" ID-CLB-Dataset="93239" ID-DOI="10.11646/zootaxa.5162.4.1" ID-GBIF-Dataset="7d343017-483e-43be-9439-d06e748fc54e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6810290" ID-ZooBank="DD2279FA-E8F1-4951-A5CA-91082E875580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657270124299" checkinUser="plazi" docAuthor="Fortey, Richard A., Wernette, Shelly J. & Hughes, Nigel C." docDate="2022" docId="03CB87846E721D20FF3AC30049D9F7C8" docLanguage="en" docName="zootaxa.5162.4.1.pdf" docOrigin="Zootaxa 5162 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hadromeros submitis Lane 1971" docType="treatment" docVersion="7" lastPageNumber="341" masterDocId="FFF2FFFC6E551D08FFADC4414A1BF54C" masterDocTitle="Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China" masterLastPageNumber="356" masterPageNumber="301" pageNumber="340" updateTime="1699356965331" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="680DC2AD730970234D6EA2B2D20CE87B">Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<mods:name id="8F17D7A7293ABE5B6538376EAA2F2AEF" type="personal">
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<mods:namePart id="A0477540B7CA2F8859E9D32FA39598CA">Fortey, Richard A.</mods:namePart>
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<mods:affiliation id="87047115622F3B6D778F1B1174823441">Department of Earth Sciences, Museum of Natural History, Cromwell Road, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="581271361EE2B10DEDA5BE5B87F8958B">Wernette, Shelly J.</mods:namePart>
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<mods:affiliation id="3F8565F09D139F2820650FC6B34D293B">Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. & Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA</mods:affiliation>
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<mods:namePart id="EBD5DAAB03C9E045C85751FDFFB05DA1">Hughes, Nigel C.</mods:namePart>
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<mods:title id="1AA22D406FFEF00F17200331B5136C11">Zootaxa</mods:title>
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<mods:date id="05A53F592400BD41A0D08F26CAD768B3">2022</mods:date>
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<mods:number id="47749C72B2C210DFE60E038B165D2854">2022-07-08</mods:number>
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<mods:number id="469357D8643D84462BE84033839E0A5C">5162</mods:number>
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<treatment id="03CB87846E721D20FF3AC30049D9F7C8" ID-DOI="http://doi.org/10.5281/zenodo.6816311" ID-GBIF-Taxon="197214812" ID-Zenodo-Dep="6816311" LSID="urn:lsid:plazi:treatment:03CB87846E721D20FF3AC30049D9F7C8" httpUri="http://treatment.plazi.org/id/03CB87846E721D20FF3AC30049D9F7C8" lastPageId="40" lastPageNumber="341" pageId="39" pageNumber="340">
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<subSubSection id="C37865196E721D2FFF3AC3004856F217" box="[151,589,1857,1884]" pageId="39" pageNumber="340" type="nomenclature">
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<paragraph id="8BDD36926E721D2FFF3AC3004856F217" blockId="39.[151,589,1857,1920]" box="[151,589,1857,1884]" pageId="39" pageNumber="340">
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<heading id="D09581FE6E721D2FFF3AC3004856F217" bold="true" box="[151,589,1857,1884]" fontSize="11" level="1" pageId="39" pageNumber="340" reason="1">
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<taxonomicName id="4C624D116E721D2FFF3AC3004856F217" ID-CoL="635ZG" authority="(Reed, 1915)" authorityName="Lane" authorityYear="1971" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[151,589,1857,1884]" class="Trilobita" family="Cheiruridae" genus="Hadromeros" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="340" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E721D2FFF3AC3004856F217" bold="true" box="[151,589,1857,1884]" pageId="39" pageNumber="340">
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<emphasis id="B916EA806E721D2FFF3AC3004B34F210" bold="true" box="[151,303,1857,1884]" italics="true" pageId="39" pageNumber="340">Hadromeros</emphasis>
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?
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<emphasis id="B916EA806E721D2FFEE8C3004BB1F210" bold="true" box="[325,426,1857,1884]" italics="true" pageId="39" pageNumber="340">submitis</emphasis>
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(
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<bibRefCitation id="EFF34B636E721D2FFE14C300485EF217" author="Reed, F. R. C." box="[441,581,1857,1883]" pageId="39" pageNumber="340" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37865196E721D2FFF3AC32748FEF28E" pageId="39" pageNumber="340" type="description">
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<paragraph id="8BDD36926E721D2FFF3AC3274AE7F2CC" blockId="39.[151,589,1857,1920]" box="[151,252,1894,1920]" pageId="39" pageNumber="340">
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<figureCitation id="13592A176E721D2FFF3AC3274AE7F2CC" box="[151,252,1894,1920]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="39" pageNumber="340">Fig. 13.1</figureCitation>
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</paragraph>
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<paragraph id="8BDD36926E721D2FFF3AC3EB48FEF28E" blockId="39.[151,741,1962,1986]" box="[151,741,1962,1986]" pageId="39" pageNumber="340">
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1915
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<taxonomicName id="4C624D116E721D2FFF5AC3EB4BEDF28E" authority="Reed" authorityName="Reed" box="[247,502,1962,1986]" class="Trilobita" family="Cheiruridae" genus="Cheirurus" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="340" phylum="Arthropoda" rank="species" species="submitis">
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<emphasis id="B916EA806E721D2FFF5AC3EB4BA3F28E" box="[247,440,1962,1986]" italics="true" pageId="39" pageNumber="340">Cheirurus submitis</emphasis>
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Reed
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</taxonomicName>
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, p. 49, pl. 8, figs 13,14.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37865196E7D1D20FF3AC4D749BAF599" pageId="40" pageNumber="341" type="materials_examined">
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<paragraph id="8BDD36926E7D1D20FF3AC4D749BAF599" blockId="40.[151,1436,150,645]" pageId="40" pageNumber="341">
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<emphasis id="B916EA806E7D1D20FF3AC4D74AE4F5FC" bold="true" box="[151,255,150,176]" pageId="40" pageNumber="341">Material</emphasis>
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.
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<materialsCitation id="3B0A3CCF6E7D1D20FEBCC4D649BAF599" ID-GBIF-Occurrence="3861306307" accessionNumber="GSI 11560" collectionCode="GSI" country="Myanmar" location="Upper Naungkangyi Beds" municipality="Lilu" pageId="40" pageNumber="341" specimenCount="1" stateProvince="Shan" typeStatus="lectotype">
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<typeStatus id="54D988306E7D1D20FEBCC4D64B9AF5FD" box="[273,385,151,177]" pageId="40" pageNumber="341" type="lectotype">Lectotype</typeStatus>
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(selected herein): cranidium from
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||
<location id="8EBD60496E7D1D20FCB5C4D64E5AF5FC" LSID="urn:lsid:plazi:treatment:03CB87846E721D20FF3AC30049D9F7C8:8EBD60496E7D1D20FCB5C4D64E5AF5FC" box="[792,1089,150,177]" country="Myanmar" municipality="Lilu" name="Upper Naungkangyi Beds" pageId="40" pageNumber="341" stateProvince="Shan">Upper Naungkangyi Beds</location>
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(Katian),
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<collectingMunicipality id="6BB9ACE86E7D1D20FB16C4D74EF7F5FC" box="[1211,1260,150,176]" pageId="40" pageNumber="341">Lilu</collectingMunicipality>
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, northern
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<collectingRegion id="49A6F8706E7D1D20FAC9C4D74AC9F598" country="Myanmar" name="Shan" pageId="40" pageNumber="341">Shan State</collectingRegion>
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,
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<collectingCountry id="F37576026E7D1D20FF70C4FA4B57F599" box="[221,332,187,213]" name="Myanmar" pageId="40" pageNumber="341">Myanmar</collectingCountry>
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,
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<figureCitation id="13592A176E7D1D20FEFAC4FB4BA0F598" box="[343,443,186,212]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="40" pageNumber="341">Fig. 13.1</figureCitation>
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(
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<bibRefCitation id="EFF34B636E7D1D20FE67C4FB4855F599" author="Reed, F. R. C." box="[458,590,186,213]" pageId="40" pageNumber="341" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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, pl. 8, figs 13,14),
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<collectionCode id="ED73AE576E7D1D20FC8DC4FB4956F598" box="[800,845,186,212]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="40" pageNumber="341">GSI</collectionCode>
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11560.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37865196E7D1D20FF6AC49F49D9F7C8" pageId="40" pageNumber="341" type="discussion">
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<paragraph id="8BDD36926E7D1D20FF6AC49F49D9F7C8" blockId="40.[151,1436,150,645]" pageId="40" pageNumber="341">
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<emphasis id="B916EA806E7D1D20FF6AC49F4B57F5B4" bold="true" box="[199,332,222,248]" pageId="40" pageNumber="341">Discussion.</emphasis>
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We illustrate here a cast taken from the
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<typeStatus id="54D988306E7D1D20FCB5C49F4999F5B4" box="[792,898,222,248]" pageId="40" pageNumber="341" type="lectotype">lectotype</typeStatus>
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, original of
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<bibRefCitation id="EFF34B636E7D1D20FBA6C49F4E8BF5B5" author="Reed, F. R. C." box="[1035,1168,222,249]" pageId="40" pageNumber="341" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed (1915</bibRefCitation>
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, pl 8. fig. 14). Only the cephalon is known of this species, which is only generally cheirurine, and invites comparison with those of many other species of similar age. It does, however, have very large, prominent but scattered tubercles on the cheeks and frontal lobe of the glabella. Many somewhat similar later Ordovician Chinese cheirurines have been assigned to the genus
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<taxonomicName id="4C624D116E7D1D20FF4CC52E4B94F4C4" authorityName="M.S.Zhou" authorityYear="1977" box="[225,399,367,392]" class="Trilobita" family="Cheiruridae" genus="Parisoceraurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E7D1D20FF4CC52E4B94F4C4" box="[225,399,367,392]" italics="true" pageId="40" pageNumber="341">Parisoceraurus</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="EFF34B636E7D1D20FE16C52F48B2F4C4" author="Zhou, Z. - Y. & Zhen, Y. - Y." box="[443,681,366,392]" pageId="40" pageNumber="341" refId="ref49762" refString="Zhou, Z. - Y. & Zhen, Y. - Y. (Eds.) (2008) Trilobite Record of China. Science Press, Beijing, 20 pp." type="book" year="2008">Zhou & Zhen (2008)</bibRefCitation>
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but these do not have such prominent tubercles. One species from the Pagoda Formation of Sichuan attributed (under open nomenclature) to
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<taxonomicName id="4C624D116E7D1D20FC79C5D34E44F4E0" authorityName="Lane" authorityYear="1971" box="[980,1119,402,428]" class="Trilobita" family="Cheiruridae" genus="Hadromeros" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E7D1D20FC79C5D34E44F4E0" box="[980,1119,402,428]" italics="true" pageId="40" pageNumber="341">Hadromeros</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="EFF34B636E7D1D20FB25C5D34F50F4E0" author="Zhou, Z. - G. & Zhou, Z. - Y. & Xiang, L. - W." box="[1160,1355,402,428]" pageId="40" pageNumber="341" refId="ref49347" refString="Zhou, Z. - G., Zhou, Z. - Y. & Xiang, L. - W. (2016) Trilobite fauna from the Ordovician Pagoda Limestone Formation of Central and Western Yangtze block, China. Geological Publishing House, Beijing, 422 pp. [in Chinese]" type="book" year="2016">
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Zhou
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<emphasis id="B916EA806E7D1D20FB67C5D24F1AF4E0" box="[1226,1281,402,428]" italics="true" pageId="40" pageNumber="341">et al.</emphasis>
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(2016
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</bibRefCitation>
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, pl. 14, fig. 14) does invite comparison with
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<emphasis id="B916EA806E7D1D20FD99C5F64853F49C" box="[564,584,439,464]" italics="true" pageId="40" pageNumber="341">H</emphasis>
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.?
|
||
<taxonomicName id="4C624D116E7D1D20FDCFC5F748A4F49C" box="[610,703,438,464]" class="Trilobita" family="Cheiruridae" genus="Cheirurus" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="species" species="submitis">
|
||
<emphasis id="B916EA806E7D1D20FDCFC5F748A4F49C" box="[610,703,438,464]" italics="true" pageId="40" pageNumber="341">submitis</emphasis>
|
||
</taxonomicName>
|
||
in showing very coarse, scattered tubercles over the fixed cheeks and forward part of the glabella, and may support a tentative assignment to
|
||
<taxonomicName id="4C624D116E7D1D20FBA8C59B4E8BF4B8" authorityName="Lane" authorityYear="1971" box="[1029,1168,474,500]" class="Trilobita" family="Cheiruridae" genus="Hadromeros" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916EA806E7D1D20FBA8C59B4E8BF4B8" box="[1029,1168,474,500]" italics="true" pageId="40" pageNumber="341">Hadromeros</emphasis>
|
||
</taxonomicName>
|
||
. However, the Sichuan specimen and the type species of
|
||
<taxonomicName id="4C624D116E7D1D20FDBFC5BF4886F754" authorityName="Lane" authorityYear="1971" box="[530,669,510,536]" class="Trilobita" family="Cheiruridae" genus="Hadromeros" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B916EA806E7D1D20FDBFC5BF4886F754" box="[530,669,510,536]" italics="true" pageId="40" pageNumber="341">Hadromeros</emphasis>
|
||
</taxonomicName>
|
||
both show relatively long (tr.) glabellar furrows compared with H.?
|
||
<taxonomicName id="4C624D116E7D1D20FF3AC6634AEFF770" box="[151,244,546,572]" class="Trilobita" family="Cheiruridae" genus="Cheirurus" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="species" species="submitis">
|
||
<emphasis id="B916EA806E7D1D20FF3AC6634AEFF770" box="[151,244,546,572]" italics="true" pageId="40" pageNumber="341">submitis</emphasis>
|
||
</taxonomicName>
|
||
, while the frontal glabellar lobe of the latter is relatively short (sag.) and wide. This structure seems to be unusual in cheirurids, and the distinctive glabellar lobation of the
|
||
<collectingCountry id="F37576026E7D1D20FCC7C60649CCF72D" box="[874,983,583,609]" name="Myanmar" pageId="40" pageNumber="341">Myanmar</collectingCountry>
|
||
species may indicate that it could belong to an undescribed genus, but without a pygidium this cannot be verified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |