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<mods:title id="AC2EDECA4D2ED0EFF0D1DC0901AD1B24">On the taxonomy of the Zodarion thoni species-group (Araneae, Zodariidae), with description of a new species with an unmodified embolus</mods:title>
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<emphasis id="B914EAE59935FFE9FF44C8A8FEE5FC0C" bold="true" box="[151,734,907,934]" pageId="1" pageNumber="113">
<taxonomicName id="4C604D749935FFE9FF44C8A8FE5DFC0C" ID-CoL="5D6CV" authority="Caporiacco, 1948" authorityName="Caporiacco" authorityYear="1948" box="[151,614,907,934]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59935FFE9FF44C8A8FDBDFC0F" bold="true" box="[151,390,907,934]" italics="true" pageId="1" pageNumber="113">Zodarion rhodiense</emphasis>
<bibRefCitation id="EFF14B069935FFE9FE5EC8A8FE5DFC0C" author="Caporiacco, L." box="[397,614,907,933]" pageId="1" pageNumber="113" pagination="27 - 75" refId="ref3672" refString="Caporiacco, L. (1948) L'arachnofauna di Rodi. Redia 33, 27 - 75. https: // doi. org / 10.1007 / BF 02415559" type="journal article" year="1948">Caporiacco, 1948</bibRefCitation>
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sp. reval.
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<paragraph id="8BDF36F79935FFE9FF44C88CFCC2FC60" blockId="1.[151,734,907,969]" box="[151,249,943,969]" pageId="1" pageNumber="113">
<figureCitation id="135B2A729935FFE9FF44C88CFCC2FC60" box="[151,249,943,969]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 49" captionStartId-0="1.[151,264,1864,1888]" captionStartId-1="3.[151,264,1845,1869]" captionTargetBox-0="[151,1435,1180,1838]" captionTargetBox-1="[158,1421,189,1821]" captionTargetId-0="figure-292@1.[151,1435,1180,1838]" captionTargetId-1="figure-36@3.[151,1435,181,1821]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="FIGURES 13. Zodarion rhodiense sp. reval., male, habitus. 1 Z. rhodiense, syntype (MZUF 619); 2 Z. rhodiense, syntype (MZUF 618); 3 Z. r. nigrifemur syn. n., syntype (MZUF 621). Scale bar = 1 mm." captionText-1="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10781907" figureDoi-1="http://doi.org/10.5281/zenodo.10781909" httpUri-0="https://zenodo.org/record/10781907/files/figure.png" httpUri-1="https://zenodo.org/record/10781909/files/figure.png" pageId="1" pageNumber="113">Figs 19</figureCitation>
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<paragraph id="8BDF36F79935FFE9FF44C8D7F965FBA2" blockId="1.[151,1437,1011,1158]" box="[151,1374,1011,1036]" pageId="1" pageNumber="113">
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<treatmentCitation id="0AC110E69935FFE9FF44C8D7FE90FBA5" author="Caporiacco, L." box="[151,683,1012,1036]" page="47" pageId="1" pageNumber="113" year="1948">
<taxonomicName id="4C604D749935FFE9FF44C8D7FE90FBA5" ID-CoL="5D6CV" authority="Caporiacco, 1948: 47" authorityName="Caporiacco" authorityPageNumber="47" authorityYear="1948" box="[151,683,1012,1036]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="subSpecies" species="rhodiense" subSpecies="rhodiense">
<emphasis id="B914EAE59935FFE9FF44C8D7FDF2FBA5" box="[151,457,1012,1036]" italics="true" pageId="1" pageNumber="113">Zodarion rhodiense rhodiense</emphasis>
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, figs 56 [
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;
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misidentified, belongs to
<taxonomicName id="4C604D749935FFE9FB81C8D7F96FFBA2" ID-CoL="93S8V" authority="Simon, 1885" authorityName="Simon" authorityYear="1885" box="[1106,1364,1011,1036]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="species" species="frenatum">
<emphasis id="B914EAE59935FFE9FB81C8D7F8F3FBA5" box="[1106,1224,1012,1036]" italics="true" pageId="1" pageNumber="113">Z. frenatum</emphasis>
<bibRefCitation id="EFF14B069935FFE9FB1EC8D7F96FFBA2" author="Simon, E." box="[1229,1364,1011,1036]" pageId="1" pageNumber="113" pagination="305 - 356" refId="ref4153" refString="Simon, E. (1885) Etudes arachnologiques. 16 e Memoire. XXIII. Materiaux pour servir a la faune des arachnides de la Grece. Annales de la Societe Entomologique de France, Series 6, 4, 305 - 356." type="journal article" year="1885">Simon, 1885</bibRefCitation>
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].
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<treatmentCitationGroup id="AB7011D99935FFE9FF44CF31F8F2FB83" box="[151,1225,1042,1067]" pageId="1" pageNumber="113">
<treatmentCitation id="0AC110E69935FFE9FF44CF31FE8AFB83" author="Caporiacco, L." box="[151,689,1042,1066]" page="49" pageId="1" pageNumber="113" year="1948">
<taxonomicName id="4C604D749935FFE9FF44CF31FE8AFB83" ID-CoL="5D6C7" authority="Caporiacco, 1948: 49" authorityName="Caporiacco" authorityPageNumber="49" authorityYear="1948" box="[151,689,1042,1066]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="subSpecies" species="rhodiense" subSpecies="nigrifemur">
<emphasis id="B914EAE59935FFE9FF44CF31FDE9FB83" box="[151,466,1042,1066]" italics="true" pageId="1" pageNumber="113">Zodarion rhodiense nigrifemur</emphasis>
<bibRefCitation id="EFF14B069935FFE9FE0BCF31FE8AFB83" author="Caporiacco, L." box="[472,689,1042,1066]" pageId="1" pageNumber="113" pagination="27 - 75" refId="ref3672" refString="Caporiacco, L. (1948) L'arachnofauna di Rodi. Redia 33, 27 - 75. https: // doi. org / 10.1007 / BF 02415559" type="journal article" year="1948">Caporiacco, 1948: 49</bibRefCitation>
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[
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;
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misidentified, belongs to
<taxonomicName id="4C604D749935FFE9FC2FCF30F84AFB83" ID-CoL="93S8V" authorityName="Simon" authorityYear="1885" box="[1020,1137,1042,1066]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="species" species="frenatum">
<emphasis id="B914EAE59935FFE9FC2FCF30F84AFB83" box="[1020,1137,1042,1066]" italics="true" pageId="1" pageNumber="113">Z. frenatum</emphasis>
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]. syn. n.
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<taxonomicName id="4C604D749935FFE9FF44CF12FD57FBE0" ID-CoL="5D6C7" authorityName="Caporiacco" authorityYear="1948" box="[151,364,1073,1097]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="species" species="nigrifemur">
<emphasis id="B914EAE59935FFE9FF44CF12FD57FBE0" box="[151,364,1073,1097]" italics="true" pageId="1" pageNumber="113">Zodarion nigrifemur</emphasis>
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:
<treatmentCitation id="0AC110E69935FFE9FEADCF12FE62FBE0" author="Bosmans, R." box="[382,601,1073,1097]" page="279" pageId="1" pageNumber="113" year="2009">
<bibRefCitation id="EFF14B069935FFE9FEADCF12FE62FBE0" author="Bosmans, R." box="[382,601,1073,1097]" pageId="1" pageNumber="113" pagination="211 - 295" refId="ref3566" refString="Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295." type="journal article" year="2009">Bosmans, 2009: 279</bibRefCitation>
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, figs 172173, 180181 [
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; elevated to species, misidentified].—
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<bibRefCitation id="EFF14B069935FFE9FAF8CF12FDB9FBC1" author="Lecigne, S. &amp; Henrard, A." pageId="1" pageNumber="113" pagination="32 - 40" refId="ref4012" refString="Lecigne, S. &amp; Henrard, A. (2022) Zodarion nigrifemur Caporiacco, 1948 newly recorded for the Turkish fauna (Araneae, Zodariidae). Journal of the Belgian Arachnological Society, 37 (1), 32 - 40." type="journal article" year="2022">Lecigne &amp; Henrard, 2022: 33</bibRefCitation>
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, figs 1AD, 2AG, 3AE, 4AH, 5A [
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; misidentified]., n. syn.
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<treatmentCitationGroup id="AB7011D99935FFE9FF44CF4DF8BFFB2F" box="[151,1156,1134,1158]" pageId="1" pageNumber="113">
<taxonomicName id="4C604D749935FFE9FF44CF4DFD0BFB2F" ID-CoL="5D6DL" authorityName="Nosek" authorityYear="1905" box="[151,304,1134,1158]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="species" species="thoni">
<emphasis id="B914EAE59935FFE9FF44CF4DFD0BFB2F" box="[151,304,1134,1158]" italics="true" pageId="1" pageNumber="113">Zodarion thoni</emphasis>
</taxonomicName>
:
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<bibRefCitation id="EFF14B069935FFE9FEEECF4DFDC3FB2F" author="Lecigne, S." box="[317,504,1134,1158]" pageId="1" pageNumber="113" pagination="11 - 34" refId="ref3957" refString="Lecigne, S. (2021) A new species of Sintula (Linyphiidae), redescription of Brigittea innocens (Dictynidae) and eight spider species newly recorded for Turkey (Araneae). Arachnologische Mitteilungen, 62, 11 - 34. https: // doi. org / 10.30963 / aramit 6204" type="journal article" year="2021">Lecigne, 2021: 34</bibRefCitation>
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, fig. 55ab [
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; misidentified per
<treatmentCitation id="0AC110E69935FFE9FC99CF4DF842FB2F" author="Lecigne, S. &amp; Henrard, A." box="[842,1145,1134,1158]" page="33" pageId="1" pageNumber="113" year="2022">
<bibRefCitation id="EFF14B069935FFE9FC99CF4DF842FB2F" author="Lecigne, S. &amp; Henrard, A." box="[842,1145,1134,1158]" pageId="1" pageNumber="113" pagination="32 - 40" refId="ref4012" refString="Lecigne, S. &amp; Henrard, A. (2022) Zodarion nigrifemur Caporiacco, 1948 newly recorded for the Turkish fauna (Araneae, Zodariidae). Journal of the Belgian Arachnological Society, 37 (1), 32 - 40." type="journal article" year="2022">Lecigne &amp; Henrard, 2022: 33</bibRefCitation>
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].
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<emphasis id="B914EAE59935FFE9FF44CC6BFD07F8C9" bold="true" box="[151,316,1864,1888]" pageId="1" pageNumber="113">FIGURES 13.</emphasis>
<taxonomicName id="4C604D749935FFE9FE97CC6BFE36F8C9" authorityName="Caporiacco" authorityYear="1948" box="[324,525,1864,1888]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59935FFE9FE97CC6BFE36F8C9" box="[324,525,1864,1888]" italics="true" pageId="1" pageNumber="113">Zodarion rhodiense</emphasis>
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<emphasis id="B914EAE59935FFE9FDC6CC6BFE42F8C9" bold="true" box="[533,633,1864,1888]" pageId="1" pageNumber="113">sp. reval.</emphasis>
, male, habitus. 1
<taxonomicName id="4C604D749935FFE9FCE1CC6AFF8AF8C9" authorityName="Caporiacco" authorityYear="1948" box="[818,945,1864,1888]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59935FFE9FCE1CC6AFF8AF8C9" box="[818,945,1864,1888]" italics="true" pageId="1" pageNumber="113">Z. rhodiense</emphasis>
</taxonomicName>
, syntype (MZUF 619); 2
<taxonomicName id="4C604D749935FFE9FB6CCC6AF904F8C9" authorityName="Caporiacco" authorityYear="1948" box="[1215,1343,1864,1888]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59935FFE9FB6CCC6AF904F8C9" box="[1215,1343,1864,1888]" italics="true" pageId="1" pageNumber="113">Z. rhodiense</emphasis>
</taxonomicName>
, syntype (MZUF 618); 3
<taxonomicName id="4C604D749935FFE9FEE8CC4EFDEDF82D" authorityName="Caporiacco" authorityYear="1948" box="[315,470,1900,1924]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="113" phylum="Arthropoda" rank="subSpecies" species="rhodiense" status="syn. nov." subSpecies="nigrifemur">
<emphasis id="B914EAE59935FFE9FEE8CC4EFDEDF82D" box="[315,470,1900,1924]" italics="true" pageId="1" pageNumber="113">Z. r. nigrifemur</emphasis>
</taxonomicName>
<emphasis id="B914EAE59935FFE9FE0FCC4FFE19F82D" bold="true" box="[476,546,1900,1924]" pageId="1" pageNumber="113">
<taxonomicNameLabel id="A227579E9935FFE9FE0FCC4FFE19F82D" box="[476,546,1900,1924]" pageId="1" pageNumber="113">syn. n.</taxonomicNameLabel>
</emphasis>
, syntype (MZUF 621). Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C37A657C9936FFEAFF44CBB4F819FF50" pageId="2" pageNumber="114" type="materials_examined">
<paragraph id="8BDF36F79936FFEAFF44CBB4F819FF50" blockId="2.[151,1437,150,1797]" pageId="2" pageNumber="114">
<emphasis id="B914EAE59936FFEAFF44CBB4FD7CFF19" bold="true" box="[151,327,150,177]" pageId="2" pageNumber="114">Type material.</emphasis>
<materialsCitation id="3B083CAA9936FFEAFE83CBB4F904FF18" box="[336,1343,151,177]" collectionCode="MZUF" country="Greece" location="Rhodes" municipality="Scafe" pageId="2" pageNumber="114" specimenCode="MZUF 618" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="9D66FD7E9936FFEAFE83CBB4FD48FF18" box="[336,371,151,177]" count="1" pageId="2" pageNumber="114" type="male">1♂</specimenCount>
<typeStatus id="54DB88559936FFEAFEAFCBB4FDEDFF18" box="[380,470,151,177]" pageId="2" pageNumber="114" type="syntype">syntype</typeStatus>
,
<collectingCountry id="F37776679936FFEAFE30CBB4FE0EFF18" box="[483,565,151,177]" name="Greece" pageId="2" pageNumber="114">Greece</collectingCountry>
,
<location id="8EBF602C9936FFEAFD90CBB4FEA1FF18" LSID="urn:lsid:plazi:treatment:03C987E19935FFEAFF44C8A8F857F8AC:8EBF602C9936FFEAFD90CBB4FEA1FF18" box="[579,666,151,177]" country="Greece" municipality="Scafe" name="Rhodes" pageId="2" pageNumber="114">Rhodes</location>
,
<collectingMunicipality id="6BBBAC8D9936FFEAFD74CBB4FEDEFF18" box="[679,741,151,177]" pageId="2" pageNumber="114">Scafe</collectingMunicipality>
(near Platania-Campochiaro), 04.1942 (
<specimenCode id="DBC69E8C9936FFEAFB63CBB4F90CFF18" box="[1200,1335,151,177]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="2" pageNumber="114">MZUF 618</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B083CAA9936FFEAFA9ECBB4F8E6FF7C" collectionCode="MZUF" country="Greece" location="Rhodes" municipality="San Demetrio" pageId="2" pageNumber="114" specimenCode="MZUF 619" specimenCount="6" specimenCount-adult="4" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="9D66FD7E9936FFEAFA9ECBB4F9A7FF18" box="[1357,1436,151,177]" count="4" pageId="2" pageNumber="114" type="adult">4 adult</specimenCount>
&amp;
<specimenCount id="9D66FD7E9936FFEAFF62CB98FD26FF7C" box="[177,285,187,213]" count="1" pageId="2" pageNumber="114" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="9D66FD7E9936FFEAFEF6CB98FD74FF7C" box="[293,335,187,213]" count="1" pageId="2" pageNumber="114" type="male">♂♂</specimenCount>
<typeStatus id="54DB88559936FFEAFE84CB98FD87FF7C" box="[343,444,187,213]" pageId="2" pageNumber="114" type="syntype">syntypes</typeStatus>
,
<collectingCountry id="F37776679936FFEAFE1BCB98FE21FF7C" box="[456,538,187,213]" name="Greece" pageId="2" pageNumber="114">Greece</collectingCountry>
,
<location id="8EBF602C9936FFEAFDF5CB98FE46FF7C" LSID="urn:lsid:plazi:treatment:03C987E19935FFEAFF44C8A8F857F8AC:8EBF602C9936FFEAFDF5CB98FE46FF7C" box="[550,637,187,213]" country="Greece" municipality="San Demetrio" name="Rhodes" pageId="2" pageNumber="114">Rhodes</location>
,
<collectingMunicipality id="6BBBAC8D9936FFEAFD5ACB98FF1EFF7C" box="[649,805,187,213]" pageId="2" pageNumber="114">San Demetrio</collectingMunicipality>
(Campochiaro), 05.1942 (
<specimenCode id="DBC69E8C9936FFEAFB9CCB98F8EEFF7C" box="[1103,1237,187,213]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="2" pageNumber="114">MZUF 619</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B083CAA9936FFEAFB39CB98F825FF50" collectionCode="MZUF" country="Greece" location="Rhodes" municipality="Draco" pageId="2" pageNumber="114" specimenCode="MZUF 621" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="9D66FD7E9936FFEAFB39CB98F92EFF7C" box="[1258,1301,187,213]" count="1" pageId="2" pageNumber="114" type="male">1 ♂</specimenCount>
(
<typeStatus id="54DB88559936FFEAFAF7CB98F946FF7C" box="[1316,1405,187,213]" pageId="2" pageNumber="114" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C604D749936FFEAFF44CBFCFD99FF50" authorityName="Caporiacco" authorityYear="1948" box="[151,418,223,249]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="rhodiense" subSpecies="nigrifemur">
<emphasis id="B914EAE59936FFEAFF44CBFCFD99FF50" box="[151,418,223,249]" italics="true" pageId="2" pageNumber="114">Z. rhodiense nigrifemur</emphasis>
</taxonomicName>
),
<collectingCountry id="F37776679936FFEAFE6ACBFCFE30FF50" box="[441,523,223,249]" name="Greece" pageId="2" pageNumber="114">Greece</collectingCountry>
,
<location id="8EBF602C9936FFEAFDC5CBFCFE56FF50" LSID="urn:lsid:plazi:treatment:03C987E19935FFEAFF44C8A8F857F8AC:8EBF602C9936FFEAFDC5CBFCFE56FF50" box="[534,621,223,249]" country="Greece" municipality="Draco" name="Rhodes" pageId="2" pageNumber="114">Rhodes</location>
,
<collectingMunicipality id="6BBBAC8D9936FFEAFDABCBFCFE87FF50" box="[632,700,223,249]" pageId="2" pageNumber="114">Draco</collectingMunicipality>
(Lardo), 04.1941 (
<specimenCode id="DBC69E8C9936FFEAFC43CBFCF82EFF50" box="[912,1045,223,249]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="2" pageNumber="114">MZUF 621</specimenCode>
)
</materialsCitation>
;
</paragraph>
</subSubSection>
<subSubSection id="C37A657C9936FFEAFF14CA20FCC8FE20" pageId="2" pageNumber="114" type="discussion">
<paragraph id="8BDF36F79936FFEAFF14CA20FCC8FE20" blockId="2.[151,1437,150,1797]" pageId="2" pageNumber="114">
<emphasis id="B914EAE59936FFEAFF14CA20FD00FEB4" bold="true" box="[199,315,259,285]" pageId="2" pageNumber="114">Remarks.</emphasis>
The type series is sufficiently small, and the diagnostic characters remain stable (see
<figureCitation id="135B2A729936FFEAFAF4CA20F9B4FEB4" box="[1319,1423,259,285]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 49" captionStartId-0="1.[151,264,1864,1888]" captionStartId-1="3.[151,264,1845,1869]" captionTargetBox-0="[151,1435,1180,1838]" captionTargetBox-1="[158,1421,189,1821]" captionTargetId-0="figure-292@1.[151,1435,1180,1838]" captionTargetId-1="figure-36@3.[151,1435,181,1821]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="FIGURES 13. Zodarion rhodiense sp. reval., male, habitus. 1 Z. rhodiense, syntype (MZUF 619); 2 Z. rhodiense, syntype (MZUF 618); 3 Z. r. nigrifemur syn. n., syntype (MZUF 621). Scale bar = 1 mm." captionText-1="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10781907" figureDoi-1="http://doi.org/10.5281/zenodo.10781909" httpUri-0="https://zenodo.org/record/10781907/files/figure.png" httpUri-1="https://zenodo.org/record/10781909/files/figure.png" pageId="2" pageNumber="114">Figs 19</figureCitation>
). Consequently, the designation of a
<typeStatus id="54DB88559936FFEAFDFECA04FEA6FEE8" box="[557,669,295,321]" pageId="2" pageNumber="114" type="lectotype">Lectotype</typeStatus>
(and
<typeStatus id="54DB88559936FFEAFD31CA04FFBCFEE8" box="[738,903,295,321]" pageId="2" pageNumber="114" type="paralectotype">Paralectotypes</typeStatus>
) seems unnecessary, as it does not confer any discernible nomenclatural advantages (See Article 74.7.3 of the International Code of
<bibRefCitation id="EFF14B069936FFEAFBA9CA68FCD5FE20" author="International Commission on Zoological Nomenclature" pageId="2" pageNumber="114" refId="ref3792" refString="International Commission on Zoological Nomenclature. (1999) International Code of Zoological Nomenclature. Fourth edition. International Trust for Zoological Nomenclature. London, xxix + 306 pp." type="book" year="1999">Zoological Nomenclature (1999))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A657C9936FFEAFF14CAB1FD66FDC8" pageId="2" pageNumber="114" type="diagnosis">
<paragraph id="8BDF36F79936FFEAFF14CAB1FD66FDC8" blockId="2.[151,1437,150,1797]" pageId="2" pageNumber="114">
<emphasis id="B914EAE59936FFEAFF14CAB1FDCCFE05" bold="true" box="[199,503,402,428]" pageId="2" pageNumber="114">Diagnosis (partial, male).</emphasis>
<taxonomicName id="4C604D749936FFEAFDD3CAB0FEE4FE04" authorityName="Caporiacco" authorityYear="1948" box="[512,735,403,429]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59936FFEAFDD3CAB0FEE4FE04" box="[512,735,403,429]" italics="true" pageId="2" pageNumber="114">Zodarion rhodiense</emphasis>
</taxonomicName>
sp. reval. belongs to
<taxonomicName id="4C604D749936FFEAFC04CAB0F80BFE04" authorityName="Nosek" authorityYear="1905" box="[983,1072,403,429]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="thoni">
<emphasis id="B914EAE59936FFEAFC04CAB0F80BFE04" box="[983,1072,403,429]" italics="true" pageId="2" pageNumber="114">Z. thoni</emphasis>
</taxonomicName>
species-group, judging from its large rectangular median apophysis and long, arch-shaped embolus (
<figureCitation id="135B2A729936FFEAFC49CA94F83AFE78" box="[922,1025,439,465]" captionStart="FIGURES 49" captionStartId="3.[151,264,1845,1869]" captionTargetBox="[158,1421,189,1821]" captionTargetId="figure-36@3.[151,1435,181,1821]" captionTargetPageId="3" captionText="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10781909" httpUri="https://zenodo.org/record/10781909/files/figure.png" pageId="2" pageNumber="114">Figs 46</figureCitation>
). It is morphologically most close to
<taxonomicName id="4C604D749936FFEAFF44CAF8FCCBFE5C" authorityName="Nosek" authorityYear="1905" box="[151,240,475,501]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="thoni">
<emphasis id="B914EAE59936FFEAFF44CAF8FCCBFE5C" box="[151,240,475,501]" italics="true" pageId="2" pageNumber="114">Z. thoni</emphasis>
</taxonomicName>
because of its very similar lanceolate tip of the embolus, with a specific knob on the prolateral side. The only difference between the two is that in
<taxonomicName id="4C604D749936FFEAFD57CADCFF28FDB0" authorityName="Caporiacco" authorityYear="1948" box="[644,787,511,537]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59936FFEAFD57CADCFF28FDB0" box="[644,787,511,537]" italics="true" pageId="2" pageNumber="114">Z. rhodiense</emphasis>
</taxonomicName>
sp. reval. the embolic tip is thinner, longer, and twisted backward (
<figureCitation id="135B2A729936FFEAFEC7C900FD47FD94" box="[276,380,547,573]" captionStart="FIGURES 49" captionStartId="3.[151,264,1845,1869]" captionTargetBox="[158,1421,189,1821]" captionTargetId="figure-36@3.[151,1435,181,1821]" captionTargetPageId="3" captionText="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10781909" httpUri="https://zenodo.org/record/10781909/files/figure.png" pageId="2" pageNumber="114">Figs 46</figureCitation>
), while in
<taxonomicName id="4C604D749936FFEAFE25C900FE74FD94" authorityName="Nosek" authorityYear="1905" box="[502,591,547,573]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="thoni">
<emphasis id="B914EAE59936FFEAFE25C900FE74FD94" box="[502,591,547,573]" italics="true" pageId="2" pageNumber="114">Z. thoni</emphasis>
</taxonomicName>
it is wider and shorter (
<bibRefCitation id="EFF14B069936FFEAFCB1C900F834FD94" author="Bosmans, R." box="[866,1039,547,573]" pageId="2" pageNumber="114" pagination="211 - 295" refId="ref3566" refString="Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295." type="journal article" year="2009">Bosmans 2009</bibRefCitation>
: figs 168169;
<bibRefCitation id="EFF14B069936FFEAFB13C900FCE8FDC8" author="Lecigne, S. &amp; Henrard, A." pageId="2" pageNumber="114" pagination="32 - 40" refId="ref4012" refString="Lecigne, S. &amp; Henrard, A. (2022) Zodarion nigrifemur Caporiacco, 1948 newly recorded for the Turkish fauna (Araneae, Zodariidae). Journal of the Belgian Arachnological Society, 37 (1), 32 - 40." type="journal article" year="2022">Lecigne &amp; Henrard 2022</bibRefCitation>
: fig. 5BC).
</paragraph>
</subSubSection>
<subSubSection id="C37A657C9936FFEAFF14C949F951FAFC" pageId="2" pageNumber="114" type="description">
<paragraph id="8BDF36F79936FFEAFF14C949F91DFCBC" blockId="2.[151,1437,150,1797]" pageId="2" pageNumber="114">
<emphasis id="B914EAE59936FFEAFF14C949FDA4FD2C" bold="true" box="[199,415,618,645]" pageId="2" pageNumber="114">Description. Male</emphasis>
(
<figureCitation id="135B2A729936FFEAFE62C948FE23FD2C" box="[433,536,619,645]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 49" captionStartId-0="1.[151,264,1864,1888]" captionStartId-1="3.[151,264,1845,1869]" captionTargetBox-0="[151,1435,1180,1838]" captionTargetBox-1="[158,1421,189,1821]" captionTargetId-0="figure-292@1.[151,1435,1180,1838]" captionTargetId-1="figure-36@3.[151,1435,181,1821]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionText-0="FIGURES 13. Zodarion rhodiense sp. reval., male, habitus. 1 Z. rhodiense, syntype (MZUF 619); 2 Z. rhodiense, syntype (MZUF 618); 3 Z. r. nigrifemur syn. n., syntype (MZUF 621). Scale bar = 1 mm." captionText-1="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10781907" figureDoi-1="http://doi.org/10.5281/zenodo.10781909" httpUri-0="https://zenodo.org/record/10781907/files/figure.png" httpUri-1="https://zenodo.org/record/10781909/files/figure.png" pageId="2" pageNumber="114">Figs 19</figureCitation>
). Measurements. Total length 4.25; carapace length 2.05, width 1.50; sternum length 1.20, width 1.01; chelicerae length 0.66, width 0.39; clypeus height (measured under the AME) 0.39; cymbium length 0.94; eye diameters: AME 0.13, ALE 0.09, PME 0.8, PLE 0.09; abdomen length 2.02, width 1.67; spinnerets length 0.3; leg I—7.88 (0.88, 1.97, 0.72, 1.54, 1.72, 1.05), leg II—7.18 (0.81, 1.72, 0.66, 1.25, 1.84, 0.90), leg III—7.48 (0.76, 1.82, 0.63, 1.42, 1.87, 0.98), leg IV—8.45 (1.13, 2.43, 0.75, 1.30, 1.92, 0.92).
</paragraph>
<paragraph id="8BDF36F79936FFEAFF14C83CFDD1FB08" blockId="2.[151,1437,150,1797]" pageId="2" pageNumber="114">
Colouration (
<figureCitation id="135B2A729936FFEAFE8FC83CFDF9FC90" box="[348,450,799,825]" captionStart="FIGURES 13" captionStartId="1.[151,264,1864,1888]" captionTargetBox="[151,1435,1180,1838]" captionTargetId="figure-292@1.[151,1435,1180,1838]" captionTargetPageId="1" captionText="FIGURES 13. Zodarion rhodiense sp. reval., male, habitus. 1 Z. rhodiense, syntype (MZUF 619); 2 Z. rhodiense, syntype (MZUF 618); 3 Z. r. nigrifemur syn. n., syntype (MZUF 621). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10781907" httpUri="https://zenodo.org/record/10781907/files/figure.png" pageId="2" pageNumber="114">Figs 13</figureCitation>
). Carapace brown with dark brown pattern. The fovea is well-visible, dark brown. Legs with dark brown femora, as dark as the carapace pattern. The other leg segments yellowish. Sternum yellowish with light brown pattern. Chelicerae the same color as the carapace. The palpal femora brown, the other segments yellowish to light brown. Dorsal abdominal scutum black, with a metallic glint, without a pattern. Abdomen almost black, but lighter than the scutum. There is a small white spot above the spinnerets, another (bigger) one ventrally, and a broad diagonal white stripe on each side. The lateral and abdominal white areas connected. Colulus well visible, yellowish, with brown rim. Spinnerets yellowish with light-brown pattern in the base. AME significantly larger than the other eyes. Chelicerae with opistognathic position. Femoral organ with 3 setae on femur I [Since the material is in very bad condition, it was not possible to examined them on the other legs, but see
<bibRefCitation id="EFF14B069936FFEAFB6DCF1CFCC0FBD4" author="Lecigne, S. &amp; Henrard, A." pageId="2" pageNumber="114" pagination="32 - 40" refId="ref4012" refString="Lecigne, S. &amp; Henrard, A. (2022) Zodarion nigrifemur Caporiacco, 1948 newly recorded for the Turkish fauna (Araneae, Zodariidae). Journal of the Belgian Arachnological Society, 37 (1), 32 - 40." type="journal article" year="2022">Lecigne &amp; Henrard, 2022: 36</bibRefCitation>
, fig. 3AD]. Abdomen with 2 spinnerets and narrow dorsal scutum, not covering its whole length. Colulus shaped as a broad, hairy plate.
</paragraph>
<paragraph id="8BDF36F79936FFEAFF14CF88FFF1FA98" blockId="2.[151,1437,150,1797]" pageId="2" pageNumber="114">
Palp (
<figureCitation id="135B2A729936FFEAFED4CF88FD56FB6C" box="[263,365,1195,1221]" captionStart="FIGURES 49" captionStartId="3.[151,264,1845,1869]" captionTargetBox="[158,1421,189,1821]" captionTargetId="figure-36@3.[151,1435,181,1821]" captionTargetPageId="3" captionText="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10781909" httpUri="https://zenodo.org/record/10781909/files/figure.png" pageId="2" pageNumber="114">Figs 49</figureCitation>
). RTA elongated, curved distally, with a pointed tip (
<figureCitation id="135B2A729936FFEAFC60CF88F822FB6C" box="[947,1049,1195,1221]" captionStart="FIGURES 49" captionStartId="3.[151,264,1845,1869]" captionTargetBox="[158,1421,189,1821]" captionTargetId="figure-36@3.[151,1435,181,1821]" captionTargetPageId="3" captionText="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10781909" httpUri="https://zenodo.org/record/10781909/files/figure.png" pageId="2" pageNumber="114">Figs 79</figureCitation>
). Bulbus oval, elongated. Embolus long and thin, arch-shaped, starting approximately at 8 oclock position and ending at approximately 3 oclock. The tip of the embolus wide, lanceolate, with a specific knob on the prolateral side. MA large, shaped as an irregular rectangle, with characteristically twisted prolateral extension (
<figureCitation id="135B2A729936FFEAFC83CE34FF86FA98" box="[848,957,1303,1329]" captionStart="FIGURES 49" captionStartId="3.[151,264,1845,1869]" captionTargetBox="[158,1421,189,1821]" captionTargetId="figure-36@3.[151,1435,181,1821]" captionTargetPageId="3" captionText="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10781909" httpUri="https://zenodo.org/record/10781909/files/figure.png" pageId="2" pageNumber="114">Figs. 46</figureCitation>
).
</paragraph>
<paragraph id="8BDF36F79936FFEAFF14CE19F951FAFC" blockId="2.[151,1437,150,1797]" box="[199,1386,1338,1365]" pageId="2" pageNumber="114">
<emphasis id="B914EAE59936FFEAFF14CE19FE39FAFD" bold="true" box="[199,514,1338,1365]" pageId="2" pageNumber="114">Variations of the body size</emphasis>
(based on the adult
<typeStatus id="54DB88559936FFEAFD37CE18FF7DFAFC" box="[740,838,1339,1365]" pageId="2" pageNumber="114" type="syntype">syntypes</typeStatus>
only). Carapace 1.802.08; abdomen 1.802.02.
</paragraph>
</subSubSection>
<subSubSection id="C37A657C9936FFEAFF14CE7CF857F8AC" pageId="2" pageNumber="114" type="discussion">
<paragraph id="8BDF36F79936FFEAFF14CE7CF857F8AC" blockId="2.[151,1437,150,1797]" pageId="2" pageNumber="114">
<emphasis id="B914EAE59936FFEAFF14CE7CFD00FAD0" bold="true" box="[199,315,1375,1401]" pageId="2" pageNumber="114">Remarks.</emphasis>
When synonymizing
<taxonomicName id="4C604D749936FFEAFDE6CE7CFEF9FAD0" authorityName="Caporiacco" authorityYear="1948" box="[565,706,1375,1401]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59936FFEAFDE6CE7CFEF9FAD0" box="[565,706,1375,1401]" italics="true" pageId="2" pageNumber="114">Z. rhodiense</emphasis>
</taxonomicName>
with
<taxonomicName id="4C604D749936FFEAFCD7CE7CF855FAD0" authority=", Bosmans (2009)" authorityName=", Bosmans" authorityYear="2009" box="[772,1134,1375,1401]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="B914EAE59936FFEAFCD7CE7CFF9DFAD0" box="[772,934,1375,1401]" italics="true" pageId="2" pageNumber="114">Z. granulatum</emphasis>
,
<bibRefCitation id="EFF14B069936FFEAFC66CE7CF855FAD0" author="Bosmans, R." box="[949,1134,1375,1401]" pageId="2" pageNumber="114" pagination="211 - 295" refId="ref3566" refString="Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295." type="journal article" year="2009">Bosmans (2009)</bibRefCitation>
</taxonomicName>
referred to the description and drawings of
<taxonomicName id="4C604D749936FFEAFE9ECEA0FDECFA34" authorityName="Caporiacco" authorityYear="1948" box="[333,471,1411,1437]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59936FFEAFE9ECEA0FDECFA34" box="[333,471,1411,1437]" italics="true" pageId="2" pageNumber="114">Z. rhodiense</emphasis>
</taxonomicName>
provided by Brignoli (1984: figs 5053). They indeed correspond to
<taxonomicName id="4C604D749936FFEAFB1DCEA0F957FA34" authorityName=", Bosmans" authorityYear="2009" box="[1230,1388,1411,1437]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="B914EAE59936FFEAFB1DCEA0F957FA34" box="[1230,1388,1411,1437]" italics="true" pageId="2" pageNumber="114">Z. granulatum</emphasis>
</taxonomicName>
, but this is due to misidentification. Brignoli (1984) has described and depicted
<taxonomicName id="4C604D749936FFEAFC0CCE84F844FA68" authorityName=", Bosmans" authorityYear="2009" box="[991,1151,1447,1473]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="B914EAE59936FFEAFC0CCE84F844FA68" box="[991,1151,1447,1473]" italics="true" pageId="2" pageNumber="114">Z. granulatum</emphasis>
</taxonomicName>
instead of
<taxonomicName id="4C604D749936FFEAFB29CE84F9BEFA68" authorityName="Caporiacco" authorityYear="1948" box="[1274,1413,1447,1473]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59936FFEAFB29CE84F9BEFA68" box="[1274,1413,1447,1473]" italics="true" pageId="2" pageNumber="114">Z. rhodiense</emphasis>
</taxonomicName>
. I was able to find and examine the
<typeStatus id="54DB88559936FFEAFDD7CEE8FE5DFA4C" box="[516,614,1483,1509]" pageId="2" pageNumber="114" type="syntype">syntypes</typeStatus>
of the two subspecies
<taxonomicName id="4C604D749936FFEAFC8ECEE8F860FA4C" authorityName="Caporiacco" authorityYear="1948" box="[861,1115,1483,1509]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="rhodiense" subSpecies="rhodiense">
<emphasis id="B914EAE59936FFEAFC8ECEE8F860FA4C" box="[861,1115,1483,1509]" italics="true" pageId="2" pageNumber="114">Z. rhodiense rhodiense</emphasis>
</taxonomicName>
and
<taxonomicName id="4C604D749936FFEAFB5DCEE8F9AEFA4C" authorityName="Caporiacco" authorityYear="1948" box="[1166,1429,1483,1509]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="rhodiense" subSpecies="nigrifemur">
<emphasis id="B914EAE59936FFEAFB5DCEE8F9AEFA4C" box="[1166,1429,1483,1509]" italics="true" pageId="2" pageNumber="114">Z. rhodiense nigrifemur</emphasis>
</taxonomicName>
. Males have fully identical tips of the embolus (
<figureCitation id="135B2A729936FFEAFD6ECECCFF1FF9A0" box="[701,804,1519,1545]" captionStart="FIGURES 49" captionStartId="3.[151,264,1845,1869]" captionTargetBox="[158,1421,189,1821]" captionTargetId="figure-36@3.[151,1435,181,1821]" captionTargetPageId="3" captionText="FIGURES 49. Zodarion rhodiense sp. reval., male, left palp. 4, 7 Z. rhodiense, syntype (MZUF 619); 5, 8 Z. rhodiense, syntype (MZUF 618); 6, 9 Z. r. nigrifemur syn. n., syntype (MZUF 621); 46 ventral; 79 retrolateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10781909" httpUri="https://zenodo.org/record/10781909/files/figure.png" pageId="2" pageNumber="114">Figs 46</figureCitation>
) and no other somatic or genital differences. Even the main diagnostic feature (the color of the femur) is identical between the two (
<figureCitation id="135B2A729936FFEAFBC7CD30F847F984" box="[1044,1148,1555,1581]" captionStart="FIGURES 13" captionStartId="1.[151,264,1864,1888]" captionTargetBox="[151,1435,1180,1838]" captionTargetId="figure-292@1.[151,1435,1180,1838]" captionTargetPageId="1" captionText="FIGURES 13. Zodarion rhodiense sp. reval., male, habitus. 1 Z. rhodiense, syntype (MZUF 619); 2 Z. rhodiense, syntype (MZUF 618); 3 Z. r. nigrifemur syn. n., syntype (MZUF 621). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10781907" httpUri="https://zenodo.org/record/10781907/files/figure.png" pageId="2" pageNumber="114">Figs 13</figureCitation>
). However, the tip of the embolus clearly differs from those of
<taxonomicName id="4C604D749936FFEAFD81CD14FECEF9F8" authorityName=", Bosmans" authorityYear="2009" box="[594,757,1591,1617]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="B914EAE59936FFEAFD81CD14FECEF9F8" box="[594,757,1591,1617]" italics="true" pageId="2" pageNumber="114">Z. granulatum</emphasis>
</taxonomicName>
(
<figureCitation id="135B2A729936FFEAFCDBCD14FF65F9F8" box="[776,862,1591,1617]" captionStart="FIGURES 1518" captionStartId="5.[151,264,1629,1653]" captionTargetBox="[223,1362,181,1605]" captionTargetId="figure-36@5.[218,1370,181,1605]" captionTargetPageId="5" captionText="FIGURES 1518. 1517 Z. namrun sp. n., male holotype. 18 Z. granulatum, male holotype (ARA025383); 15 habitus dorsal; 16, 18 left palp, ventral; 17 left palp, retrolateral. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10781913" httpUri="https://zenodo.org/record/10781913/files/figure.png" pageId="2" pageNumber="114">Fig. 18</figureCitation>
). The females collected with both
<taxonomicName id="4C604D749936FFEAFB27CD14F9A7F9F8" authorityName="Caporiacco" authorityYear="1948" box="[1268,1436,1591,1617]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="rhodiense" subSpecies="rhodiense">
<emphasis id="B914EAE59936FFEAFB27CD14F9A7F9F8" box="[1268,1436,1591,1617]" italics="true" pageId="2" pageNumber="114">Z. r. rhodiense</emphasis>
</taxonomicName>
and
<taxonomicName id="4C604D749936FFEAFF1BCD78FD4EF9DC" authorityName="Caporiacco" authorityYear="1948" box="[200,373,1627,1653]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="rhodiense" subSpecies="nigrifemur">
<emphasis id="B914EAE59936FFEAFF1BCD78FD4EF9DC" box="[200,373,1627,1653]" italics="true" pageId="2" pageNumber="114">Z. r. nigrifemur</emphasis>
</taxonomicName>
were not conspecific with the males and belong to
<taxonomicName id="4C604D749936FFEAFC6ECD78F8AAF9DC" authorityName="Simon" authorityYear="1885" box="[957,1169,1627,1653]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="frenatum">
<emphasis id="B914EAE59936FFEAFC6ECD78F8AAF9DC" box="[957,1169,1627,1653]" italics="true" pageId="2" pageNumber="114">Zodarion frenatum</emphasis>
</taxonomicName>
(
<figureCitation id="135B2A729936FFEAFB72CD78F91EF9DC" box="[1185,1317,1627,1653]" captionStart="FIGURES 1014" captionStartId="4.[151,264,836,860]" captionTargetBox="[151,1430,181,812]" captionTargetId="figure-15@4.[151,1435,181,812]" captionTargetPageId="4" captionText="FIGURES 1014. Zodarion frenatum. 10 female syntype of Z. rhodiense (MZUF 619); 11, 12 female syntype of Z. rhodiense nigrifemur (MZUF 621); 13, 14 female and male specimens from Bulgaria. 10, 11 epigyne, 12, 13 vulva, 14 male palp, ventral. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10781911" httpUri="https://zenodo.org/record/10781911/files/figure.png" pageId="2" pageNumber="114">Figs 1012</figureCitation>
). I include also photographs of
<taxonomicName id="4C604D749936FFEAFEADCD5CFE3BF930" authorityName="Simon" authorityYear="1885" box="[382,512,1663,1689]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="frenatum">
<emphasis id="B914EAE59936FFEAFEADCD5CFE3BF930" box="[382,512,1663,1689]" italics="true" pageId="2" pageNumber="114">Z. frenatum</emphasis>
</taxonomicName>
from
<collectingCountry id="F37776679936FFEAFD95CD5CFE9DF930" box="[582,678,1663,1689]" name="Bulgaria" pageId="2" pageNumber="114">Bulgaria</collectingCountry>
for comparison (
<figureCitation id="135B2A729936FFEAFCBACD5CFFD6F930" box="[873,1005,1663,1689]" captionStart="FIGURES 1014" captionStartId="4.[151,264,836,860]" captionTargetBox="[151,1430,181,812]" captionTargetId="figure-15@4.[151,1435,181,812]" captionTargetPageId="4" captionText="FIGURES 1014. Zodarion frenatum. 10 female syntype of Z. rhodiense (MZUF 619); 11, 12 female syntype of Z. rhodiense nigrifemur (MZUF 621); 13, 14 female and male specimens from Bulgaria. 10, 11 epigyne, 12, 13 vulva, 14 male palp, ventral. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10781911" httpUri="https://zenodo.org/record/10781911/files/figure.png" pageId="2" pageNumber="114">Figs 1314</figureCitation>
). I do not update the female diagnosis because there were no females belonging to
<taxonomicName id="4C604D749936FFEAFD5ECD80FF21F914" authorityName="Caporiacco" authorityYear="1948" box="[653,794,1699,1725]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59936FFEAFD5ECD80FF21F914" box="[653,794,1699,1725]" italics="true" pageId="2" pageNumber="114">Z. rhodiense</emphasis>
</taxonomicName>
<emphasis id="B914EAE59936FFEAFCF2CD80FFB5F914" bold="true" box="[801,910,1699,1725]" pageId="2" pageNumber="114">sp. reval.</emphasis>
among the
<typeStatus id="54DB88559936FFEAFBC7CD80F841F914" box="[1044,1146,1699,1725]" pageId="2" pageNumber="114" type="syntype">syntypes</typeStatus>
. Since
<bibRefCitation id="EFF14B069936FFEAFB19CD80F9A7F914" author="Caporiacco, L." box="[1226,1436,1699,1725]" pageId="2" pageNumber="114" pagination="27 - 75" refId="ref3672" refString="Caporiacco, L. (1948) L'arachnofauna di Rodi. Redia 33, 27 - 75. https: // doi. org / 10.1007 / BF 02415559" type="journal article" year="1948">Caporiacco (1948)</bibRefCitation>
named the species
<taxonomicName id="4C604D749936FFEAFEA2CDE4FDC5F948" authorityName="Caporiacco" authorityYear="1948" box="[369,510,1735,1761]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="B914EAE59936FFEAFEA2CDE4FDC5F948" box="[369,510,1735,1761]" italics="true" pageId="2" pageNumber="114">Z. rhodiense</emphasis>
</taxonomicName>
in the original description, this name has priority and is revalidated, whereas its subspecies
<taxonomicName id="4C604D749936FFEAFEC5CDC8FDFAF8AC" authorityName="Caporiacco" authorityYear="1948" box="[278,449,1771,1797]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="114" phylum="Arthropoda" rank="subSpecies" species="rhodiense" status="syn. nov." subSpecies="nigrifemur">
<emphasis id="B914EAE59936FFEAFEC5CDC8FDFAF8AC" box="[278,449,1771,1797]" italics="true" pageId="2" pageNumber="114">Z. r. nigrifemur</emphasis>
</taxonomicName>
<emphasis id="B914EAE59936FFEAFE1BCDC8FE2EF8AC" bold="true" box="[456,533,1771,1797]" pageId="2" pageNumber="114">
<taxonomicNameLabel id="A227579E9936FFEAFE1BCDC8FE2EF8AC" box="[456,533,1771,1797]" pageId="2" pageNumber="114">n. syn.</taxonomicNameLabel>
</emphasis>
is assigned as a junior synonym to the nominate one.
</paragraph>
</subSubSection>
</treatment>
</document>