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<document id="E7C1BF7C1D294DAAAEC5474D22A1E4DA" ID-DOI="10.5852/ejt.2016.208" ID-GBIF-Dataset="2c5cb62e-9bc7-4fba-a4cb-e5a2fda8c719" ID-ISSN="2118-9773" ID-Zenodo-Dep="269306" ID-ZooBank="urn:lsid:zoobank.org:pub:F807AC79-2615-463B-8272-82523B46210A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468057990865" checkinUser="plazi" docAuthor="Diana Delicado, ladimir Pešić &amp; Peter Glöer" docDate="2016" docId="03C987BFFFA8FFBAFE7A7197FC24FDF3" docLanguage="en" docName="331-1720-1-PB.pdf" docOrigin="European Journal of Taxonomy 208" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Intermaria kermanshahensis Gloer &amp; Pesic 2009, comb. nov." docType="treatment" docVersion="14" lastPageNumber="16" masterDocId="FFF0FFC7FFA3FFB5FFDB776BFFAAFF9C" masterDocTitle="Unraveling a new lineage of Hydrobiidae genera (Caenogastropoda: Truncatelloidea) from the Ponto-Caspian region" masterLastPageNumber="29" masterPageNumber="1" pageNumber="12" updateTime="1698364436418" updateUser="ExternalLinkService">
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<mods:title id="A84F80515B4F72C112F67B293E1377C5">Unraveling a new lineage of Hydrobiidae genera (Caenogastropoda: Truncatelloidea) from the Ponto-Caspian region</mods:title>
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<mods:namePart id="122C3B29D514AEBA705A6AFDD189DDD8">Diana Delicado</mods:namePart>
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<mods:namePart id="779CB0FCECFC7EC37A8EC6C98CBADF94">ladimir Pešić</mods:namePart>
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<mods:namePart id="3858E81B08D5FB28C356BD62355B1CD3">Peter Glöer</mods:namePart>
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<mods:title id="FBADA14030574E07E8F29EDA13349530">European Journal of Taxonomy</mods:title>
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<mods:date id="1EF4739493A6642FC74E52578E81D22A">2016</mods:date>
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<mods:number id="4AF5B97F2D3053961E3A2B8AC6CDCA65">208</mods:number>
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<treatment id="03C987BFFFA8FFBAFE7A7197FC24FDF3" ID-DOI="http://doi.org/10.5281/zenodo.3510526" ID-GBIF-Taxon="123190709" ID-Zenodo-Dep="3510526" LSID="urn:lsid:plazi:treatment:03C987BFFFA8FFBAFE7A7197FC24FDF3" httpUri="http://treatment.plazi.org/id/03C987BFFFA8FFBAFE7A7197FC24FDF3" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12">
<subSubSection id="C37A6522FFA8FFBEFE7A7197FB39F88A" box="[417,1171,1787,1814]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BDF36A9FFA8FFBEFE7A7197FB39F88A" blockId="11.[417,1171,1787,1850]" box="[417,1171,1787,1814]" pageId="11" pageNumber="12">
<heading id="D09781C5FFA8FFBEFE7A7197FB39F88A" box="[417,1171,1787,1814]" centered="true" fontSize="11" level="3" pageId="11" pageNumber="12" reason="4">
<taxonomicName id="4C604D2AFFA8FFBEFE7A7197FBA0F88A" ID-CoL="6N7CD" authority="Gloer &amp; Pesic, 2009" authorityName="Gloer &amp; Pesic" authorityYear="2009" box="[417,1034,1787,1814]" class="Gastropoda" family="Hydrobiidae" genus="Intermaria" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="kermanshahensis" status="comb. nov.">
<emphasis id="B914EABBFFA8FFBEFE7A7197FCAAF88A" bold="true" box="[417,768,1787,1814]" italics="true" pageId="11" pageNumber="12">Intermaria kermanshahensis</emphasis>
(
<bibRefCitation id="EFF14B58FFA8FFBEFCD47197FBA8F88A" author="Gloer P. &amp; Pesic V." box="[783,1026,1788,1814]" journalOrPublisher="Mollusca" pageId="11" pageNumber="12" pagination="33 - 39" refString="Gloer P. &amp; Pesic V. 2009. New freshwater gastropod species of the Iran (Gastropoda: Stenothyridae, Bithyniidae, Hydrobiidae). Mollusca 27: 33 - 39." title="New freshwater gastropod species of the Iran (Gastropoda: Stenothyridae, Bithyniidae, Hydrobiidae)" type="book chapter" year="2009">Glöer &amp; Pešić, 2009</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22757C0FFA8FFBEFBC97197FB39F88A" box="[1042,1171,1788,1814]" pageId="11" pageNumber="12">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A6522FFA8FFBEFD7B7074FC3EF8A5" box="[672,916,1823,1850]" pageId="11" pageNumber="12" type="description">
<paragraph id="8BDF36A9FFA8FFBEFD7B7074FC3EF8A5" blockId="11.[417,1171,1787,1850]" box="[672,916,1823,1850]" pageId="11" pageNumber="12">
<figureCitation id="135B2A2CFFA8FFBEFD7B7074FCA3F8A6" box="[672,777,1823,1850]" captionStart-0="Fig. 4" captionStart-1="Fig. 5" captionStartId-0="12.[189,232,1894,1920]" captionStartId-1="14.[189,232,1860,1886]" captionTargetBox-0="[215,1373,334,1849]" captionTargetBox-1="[370,1225,265,1828]" captionTargetId-0="figure@12.[215,1373,334,1849]" captionTargetId-1="figure@14.[370,1225,265,1829]" captionTargetPageId-0="12" captionTargetPageId-1="14" captionText-0="Fig. 4. Shell, operculum and radula of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A – B. Shell in front view. C. Shell in lateral view. D. Protoconch and microsculpture. E – F. Internal and external side of the operculum. G. Rows of teeth of the radula. H. Central teeth. I. Detail of outer marginal teeth." captionText-1="Fig. 5. Anatomy of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A. Ctenidium and osphradium. B. Partial nervous system. C. Female genitalia. D. Bursa copulatrix and seminal receptacle. E – F. Head of a male and penis. G. Prostate gland. Anatomical abbreviations given in the Material and methods section, except DL (distal lobe)." httpUri-0="https://zenodo.org/record/269310/files/figure.png" httpUri-1="https://zenodo.org/record/269311/files/figure.png" pageId="11" pageNumber="12">Figs 4–5</figureCitation>
,
<tableCitation id="C6E20312FFA8FFBEFCCF7074FC3EF8A5" box="[788,916,1823,1849]" captionStart-0="Table 2" captionStart-1="Table 3" captionStartId-0="9.[189,258,268,294]" captionStartId-1="10.[189,258,268,294]" captionTargetBox-0="[211,1384,459,1599]" captionTargetBox-1="[241,1381,463,2008]" captionText-0="Table 2. Shell measurements (in mm) of the species: 1. Intermaria zagrosensis (Glöer &amp; Pešić, 2009). 2. Intermaria kermanshahensis (Glöer &amp; Pešić, 2009). 3. Persipyrgula saboori (Glöer &amp; Pešić, 2009). 4. Shadinia terpoghassiani (Shadin, 1952). ANOVA results are shown through effect size F, degrees of freedom (df), residuals (r) and the resulting significance (p – value: *** when 0; ** when 0.001; * when 0.01; NS = no significant)." captionText-1="Table 3. Ctenidium, osphradium. Female and male genitalia and nervous system measurements (in mm) of the species: 1. Intermaria zagrosensis (Glöer &amp; Pešić, 2009). 2. Intermaria kermanshahensis (Glöer &amp; Pešić, 2009). 3. Persipyrgula saboori (Glöer &amp; Pešić, 2009). 4. Shadinia terpoghassiani (Shadin, 1952). For ANOVA results effect size F, degrees of freedom (df), residuals (r) and the resulting significance (p – value: *** when 0; ** when 0.001; * when 0.01; NS = not significant) are shown." httpUri-0="http://table.plazi.org/id/DF1F6621FFAAFFBCFF667667FDF8FE3B" httpUri-1="http://table.plazi.org/id/DF1F6621FFA9FFBFFF667667FB0EFE3A" pageId="11" pageNumber="12" tableUuid-0="DF1F6621FFAAFFBCFF667667FDF8FE3B" tableUuid-1="DF1F6621FFA9FFBFFF667667FB0EFE3A">Tables 2–3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37A6522FFA8FFBEFF66700BFBBFF820" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8BDF36A9FFA8FFBEFF66700BFB86F8E6" blockId="11.[189,1068,1888,1914]" box="[189,1068,1888,1914]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB701187FFA8FFBEFF66700BFB86F8E6" box="[189,1068,1888,1914]" pageId="11" pageNumber="12">
<taxonomicName id="4C604D2AFFA8FFBEFF66700BFCC0F8E6" ID-CoL="4NJWR" authority="Gloer &amp; Pesic, 2009: 38" authorityName="Gloer &amp; Pesic" authorityPageNumber="38" authorityYear="2009" box="[189,874,1888,1914]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="kermanshahensis">
<emphasis id="B914EABBFFA8FFBEFF66700BFDE9F8E6" box="[189,579,1888,1914]" italics="true" pageId="11" pageNumber="12">Pseudamnicola kermanshahensis</emphasis>
<treatmentCitation id="0AC110B8FFA8FFBEFD90700BFCC0F8E6" author="Gloer P. &amp; Pesic V." box="[587,874,1888,1914]" page="38" pageId="11" pageNumber="12" year="2009">
<bibRefCitation id="EFF14B58FFA8FFBEFD90700BFC94F8E6" author="Gloer P. &amp; Pesic V." box="[587,830,1888,1914]" journalOrPublisher="Mollusca" pageId="11" pageNumber="12" pagination="33 - 39" refString="Gloer P. &amp; Pesic V. 2009. New freshwater gastropod species of the Iran (Gastropoda: Stenothyridae, Bithyniidae, Hydrobiidae). Mollusca 27: 33 - 39." title="New freshwater gastropod species of the Iran (Gastropoda: Stenothyridae, Bithyniidae, Hydrobiidae)" type="book chapter" year="2009">Glöer &amp; Pešić, 2009</bibRefCitation>
: 38
</treatmentCitation>
</taxonomicName>
, pl. 6, figs 7–10.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDF36A9FFA8FFBEFF6670C9FBBFF820" blockId="11.[189,1045,1954,1980]" box="[189,1045,1954,1980]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB701187FFA8FFBEFF6670C9FBBFF820" box="[189,1045,1954,1980]" pageId="11" pageNumber="12">
<taxonomicName id="4C604D2AFFA8FFBEFF6670C9FE40F820" ID-CoL="6XNZ8" box="[189,490,1954,1980]" class="Gastropoda" family="Hydrobiidae" genus="Sarkhia" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="kermanshahensis">
<emphasis id="B914EABBFFA8FFBEFF6670C9FE40F820" box="[189,490,1954,1980]" italics="true" pageId="11" pageNumber="12">Sarkhia kermanshahensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AC110B8FFA8FFBEFDDC70C9FC8CF820" ID-DOI="http://doi.org/10.5281/zenodo.3509691" ID-Zenodo-Dep="3509691" author="Gloer P. &amp; Pesic V." box="[519,806,1954,1980]" httpUri="http://treatment.plazi.org/id/CA8B2EF976BE53DCAEBCD80D614B3086" page="33" pageId="11" pageNumber="12" year="2012">
<bibRefCitation id="EFF14B58FFA8FFBEFDDC70C9FD51F820" author="Gloer P. &amp; Pesic V." box="[519,763,1954,1980]" journalOrPublisher="Gloer P., Yildirim M. Z. &amp; Kebapci U. 2015. Description of two new species of Pseudamnicola from southern Turkey (Mollusca: Gastropoda: Hydrobiidae). Zoology in the Middle East" pageId="11" pageNumber="12" pagination="139 - 143" publicationUrl="http://dx.doi.org/10.3897/zookeys.219.3406" refString="Gloer P. &amp; Pesic V. 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61. http: // dx. doi. org / 10.3897 / zookeys. 219.3406 Gloer P., Yildirim M. Z. &amp; Kebapci U. 2015. Description of two new species of Pseudamnicola from southern Turkey (Mollusca: Gastropoda: Hydrobiidae). Zoology in the Middle East 61: 139 - 143. http: // dx. doi. org / 10.1080 / 09397140.2015.1008189" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61" type="book chapter" year="2012">
Glöer &amp; Pešić,
<date id="FFDE1069FFA8FFBEFD6770C9FD51F820" box="[700,763,1954,1980]" pageId="11" pageNumber="12" value="2012">2012</date>
</bibRefCitation>
: 33
</treatmentCitation>
, fig. 12h comb. nov.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF1F6621FFAFFFB9FF66700DFAA9F876" httpUri="https://zenodo.org/record/269310/files/figure.png" pageId="12" pageNumber="13" targetBox="[215,1373,334,1849]" targetPageId="12">
<paragraph id="8BDF36A9FFAFFFB9FF66700DFAA9F876" blockId="12.[189,1399,1894,2027]" pageId="12" pageNumber="13">
<emphasis id="B914EABBFFAFFFB9FF66700DFEA5F81C" bold="true" box="[189,271,1894,1920]" pageId="12" pageNumber="13">Fig. 4.</emphasis>
Shell, operculum and radula of
<taxonomicName id="4C604D2AFFAFFFB9FD7E700CFABDF81D" authority="Gloer &amp; Pesic, 2009" authorityName="Gloer &amp; Pesic" authorityYear="2009" box="[677,1303,1894,1921]" class="Gastropoda" family="Hydrobiidae" genus="Intermaria" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="kermanshahensis">
<emphasis id="B914EABBFFAFFFB9FD7E700CFC8EF81C" box="[677,804,1895,1920]" italics="true" pageId="12" pageNumber="13">Intermaria</emphasis>
<taxonomicName id="4C604D2AFFAFFFB9FCEB700CFC50F81D" box="[816,1018,1895,1921]" pageId="12" pageNumber="13" rank="species" species="kermanshahensis">
<emphasis id="B914EABBFFAFFFB9FCEB700CFC50F81D" box="[816,1018,1895,1921]" italics="true" pageId="12" pageNumber="13">kermanshahensis</emphasis>
</taxonomicName>
(Glöer &amp; Pešić, 2009)
</taxonomicName>
gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran.
<emphasis id="B914EABBFFAFFFB9FB5470E1FB6DF838" bold="true" box="[1167,1223,1930,1956]" pageId="12" pageNumber="13">A–B</emphasis>
. Shell in front view.
<emphasis id="B914EABBFFAFFFB9FED870C6FEB2F85B" bold="true" box="[259,280,1965,1991]" pageId="12" pageNumber="13">C</emphasis>
. Shell in lateral view.
<emphasis id="B914EABBFFAFFFB9FDC470C6FD9EF85B" bold="true" box="[543,564,1965,1991]" pageId="12" pageNumber="13">D</emphasis>
. Protoconch and microsculpture.
<emphasis id="B914EABBFFAFFFB9FC1B70C6FC59F85B" bold="true" box="[960,1011,1965,1991]" pageId="12" pageNumber="13">E–F</emphasis>
. Internal and external side of the operculum.
<emphasis id="B914EABBFFAFFFB9FE9270BBFECAF876" bold="true" box="[329,352,2000,2026]" pageId="12" pageNumber="13">G</emphasis>
. Rows of teeth of the radula.
<emphasis id="B914EABBFFAFFFB9FD6770BBFD79F876" bold="true" box="[700,723,2000,2026]" pageId="12" pageNumber="13">H</emphasis>
. Central teeth.
<emphasis id="B914EABBFFAFFFB9FC5C70BBFC38F876" bold="true" box="[903,914,2000,2026]" pageId="12" pageNumber="13">I</emphasis>
. Detail of outer marginal teeth.
</paragraph>
</caption>
<subSubSection id="C37A6522FFAEFFB8FF667660FB8DFE21" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BDF36A9FFAEFFB8FF667660FEDAFEB9" blockId="13.[189,1399,267,446]" box="[189,368,267,293]" pageId="13" pageNumber="14">
<heading id="D09781C5FFAEFFB8FF667660FEDAFEB9" bold="true" box="[189,368,267,293]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<emphasis id="B914EABBFFAEFFB8FF667660FEDAFEB9" bold="true" box="[189,368,267,293]" pageId="13" pageNumber="14">New diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF667652FB8DFE21" blockId="13.[189,1399,267,446]" pageId="13" pageNumber="14">Shell ovate-conic, yellowish, with pyriform aperture; protoconch microsculpture slightly wrinkled; central radular tooth formula 4–C–4/2–2; pyriform bursa copulatrix; one seminal receptacle elongate with short duct; penis gradually tapering with small distal lobe on inner edge, end tapered; nervous system elongated (mean RPG ratio = 0.50) with slight black pigmentation.</paragraph>
</subSubSection>
<subSubSection id="C37A6522FFAEFFB8FF66768FFADDFCBA" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BDF36A9FFAEFFB8FF66768FFE00FE62" blockId="13.[189,426,484,510]" box="[189,426,484,510]" pageId="13" pageNumber="14">
<heading id="D09781C5FFAEFFB8FF66768FFE00FE62" bold="true" box="[189,426,484,510]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B914EABBFFAEFFB8FF66768FFE00FE62" bold="true" box="[189,426,484,510]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF317578FEF6FDB1" blockId="13.[189,772,531,593]" box="[234,348,531,557]" pageId="13" pageNumber="14">
<heading id="D09781C5FFAEFFB8FF317578FEF6FDB1" bold="true" box="[234,348,531,557]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<emphasis id="B914EABBFFAEFFB8FF317578FEF6FDB1" bold="true" box="[234,348,531,557]" pageId="13" pageNumber="14">Holotype</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF66755DFCAEFDCC" blockId="13.[189,772,531,593]" box="[189,772,566,593]" pageId="13" pageNumber="14">IRAN: ZMH 51404: 4 mm height, 2.7 mm width.</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF317513FECDFD0E" blockId="13.[189,908,632,694]" box="[234,359,632,658]" pageId="13" pageNumber="14">
<heading id="D09781C5FFAEFFB8FF317513FECDFD0E" bold="true" box="[234,359,632,658]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<emphasis id="B914EABBFFAEFFB8FF317513FECDFD0E" bold="true" box="[234,359,632,658]" pageId="13" pageNumber="14">Paratypes</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF6675F0FC26FD29" blockId="13.[189,908,632,694]" box="[189,908,667,694]" pageId="13" pageNumber="14">IRAN: ZMH 51405 (5 ex.) and P. Glöeris (66 ex.) collection.</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF6675B7FEF7FD6A" blockId="13.[189,1399,732,806]" box="[189,349,732,758]" pageId="13" pageNumber="14">
<heading id="D09781C5FFAEFFB8FF6675B7FEF7FD6A" bold="true" box="[189,349,732,758]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<materialsCitation id="3B083CF4FFAEFFB8FF6675B7FEF7FD6A" ID-GBIF-Occurrence="1291639911" box="[189,349,732,758]" collectingDate="2005-06-27" country="Iran" latitude="34.45" location="spring between Sarab and Sahneh cities" longLatPrecision="1199" longitude="47.733334" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Kermanshah Province" typeStatus="type">
<emphasis id="B914EABBFFAEFFB8FF6675B7FEF7FD6A" bold="true" box="[189,349,732,758]" pageId="13" pageNumber="14">
<typeStatus id="54DB880BFFAEFFB8FF6675B7FF50FD6A" box="[189,250,732,758]" pageId="13" pageNumber="14">Type</typeStatus>
locality
</emphasis>
</materialsCitation>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF667460FADDFCBA" blockId="13.[189,1399,732,806]" box="[189,1399,779,806]" pageId="13" pageNumber="14">
<collectingCountry id="F3777639FFAEFFB8FF667460FEA3FCB9" box="[189,265,779,805]" name="Iran" pageId="13" pageNumber="14">IRAN</collectingCountry>
:
<location id="8EBF6072FFAEFFB8FEC87460FD4EFCB9" LSID="urn:lsid:plazi:treatment:03C987BFFFA8FFBAFE7A7197FC24FDF3:8EBF6072FFAEFFB8FEC87460FD4EFCB9" box="[275,740,779,805]" country="Iran" latitude="34.45" longLatPrecision="1199" longitude="47.733334" name="spring between Sarab and Sahneh cities" pageId="13" pageNumber="14" stateProvince="Kermanshah Province">spring between Sarab and Sahneh cities</location>
,
<collectingRegion id="49A4F84BFFAEFFB8FD357460FC5EFCB9" box="[750,1012,779,805]" pageId="13" pageNumber="14">Kermanshah Province</collectingRegion>
,
<geoCoordinate id="EE54506EFFAEFFB8FC247460FBCEFCB9" box="[1023,1124,779,805]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="34.45">34º27' N</geoCoordinate>
<geoCoordinate id="EE54506EFFAEFFB8FBB07460FB64FCB9" box="[1131,1230,779,805]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="47.733334">47º44' E</geoCoordinate>
,
<date id="FFDE1069FFAEFFB8FB017460FAD8FCBA" box="[1242,1394,779,806]" pageId="13" pageNumber="14" value="2005-06-27">
<collectingDate id="EF9AE981FFAEFFB8FB017460FAD8FCBA" box="[1242,1394,779,806]" pageId="13" pageNumber="14" value="2005-06-27">27 Jun. 2005</collectingDate>
</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A6522FFAEFFB8FF667427FDBBF8E3" pageId="13" pageNumber="14" type="description">
<paragraph id="8BDF36A9FFAEFFB8FF667427FEE4FCFA" blockId="13.[189,1399,844,1058]" box="[189,334,844,870]" pageId="13" pageNumber="14">
<heading id="D09781C5FFAEFFB8FF667427FEE4FCFA" bold="true" box="[189,334,844,870]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<emphasis id="B914EABBFFAEFFB8FF667427FEE4FCFA" bold="true" box="[189,334,844,870]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF667410FE25FBBE" blockId="13.[189,1399,844,1058]" pageId="13" pageNumber="14">
Shell ovate-conic with 4.25–4.5 whorls, height 3–4 mm (
<figureCitation id="135B2A2CFFAEFFB8FC857410FC0FFC09" box="[862,933,891,917]" captionStart="Fig. 4" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[215,1373,334,1849]" captionTargetId="figure@12.[215,1373,334,1849]" captionTargetPageId="12" captionText="Fig. 4. Shell, operculum and radula of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A – B. Shell in front view. C. Shell in lateral view. D. Protoconch and microsculpture. E – F. Internal and external side of the operculum. G. Rows of teeth of the radula. H. Central teeth. I. Detail of outer marginal teeth." httpUri="https://zenodo.org/record/269310/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
A–C,
<tableCitation id="C6E20312FFAEFFB8FC317410FBEEFC09" box="[1002,1092,891,917]" captionStart="Table 2" captionStartId="9.[189,258,268,294]" captionTargetBox="[211,1384,459,1599]" captionText="Table 2. Shell measurements (in mm) of the species: 1. Intermaria zagrosensis (Glöer &amp; Pešić, 2009). 2. Intermaria kermanshahensis (Glöer &amp; Pešić, 2009). 3. Persipyrgula saboori (Glöer &amp; Pešić, 2009). 4. Shadinia terpoghassiani (Shadin, 1952). ANOVA results are shown through effect size F, degrees of freedom (df), residuals (r) and the resulting significance (p – value: *** when 0; ** when 0.001; * when 0.01; NS = no significant)." httpUri="http://table.plazi.org/id/DF1F6621FFAAFFBCFF667667FDF8FE3B" pageId="13" pageNumber="14" tableUuid="DF1F6621FFAAFFBCFF667667FDF8FE3B">Table 2</tableCitation>
); periostracum yellowish; protoconch approximately 500 µm wide with 1.4 whorls and nucleus around 135 µm long; protoconch microsculpture slightly wrinkled (
<figureCitation id="135B2A2CFFAEFFB8FD8D74AAFD0AFC47" box="[598,672,961,987]" captionStart="Fig. 4" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[215,1373,334,1849]" captionTargetId="figure@12.[215,1373,334,1849]" captionTargetPageId="12" captionText="Fig. 4. Shell, operculum and radula of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A – B. Shell in front view. C. Shell in lateral view. D. Protoconch and microsculpture. E – F. Internal and external side of the operculum. G. Rows of teeth of the radula. H. Central teeth. I. Detail of outer marginal teeth." httpUri="https://zenodo.org/record/269310/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
D); body whorl about ⅔ total length; rest of whorls slightly convex with deep sutures; aperture complete, pyriform, inner lip thicker than outer lip; peristome margin straight (
<figureCitation id="135B2A2CFFAEFFB8FEFE7363FEC7FBBE" box="[293,365,1032,1058]" captionStart="Fig. 4" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[215,1373,334,1849]" captionTargetId="figure@12.[215,1373,334,1849]" captionTargetPageId="12" captionText="Fig. 4. Shell, operculum and radula of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A – B. Shell in front view. C. Shell in lateral view. D. Protoconch and microsculpture. E – F. Internal and external side of the operculum. G. Rows of teeth of the radula. H. Central teeth. I. Detail of outer marginal teeth." httpUri="https://zenodo.org/record/269310/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
C).
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF667322FEE0FB1A" blockId="13.[189,1398,1097,1158]" pageId="13" pageNumber="14">
Operculum with 2 whorls approximately (
<figureCitation id="135B2A2CFFAEFFB8FD717322FD44FBFF" box="[682,750,1097,1123]" captionStart="Fig. 4" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[215,1373,334,1849]" captionTargetId="figure@12.[215,1373,334,1849]" captionTargetPageId="12" captionText="Fig. 4. Shell, operculum and radula of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A – B. Shell in front view. C. Shell in lateral view. D. Protoconch and microsculpture. E – F. Internal and external side of the operculum. G. Rows of teeth of the radula. H. Central teeth. I. Detail of outer marginal teeth." httpUri="https://zenodo.org/record/269310/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
E–F) and muscle attachment area oval and located near the nucleus.
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF6673C5FCFAFA92" blockId="13.[189,1399,1197,1295]" pageId="13" pageNumber="14">
Radula intermediate length (25% total shell length) bearing about 55 rows of teeth; central tooth formula 4–C–4/2–2 (
<figureCitation id="135B2A2CFFAEFFB8FE8A73BAFE33FB77" box="[337,409,1233,1259]" captionStart="Fig. 4" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[215,1373,334,1849]" captionTargetId="figure@12.[215,1373,334,1849]" captionTargetPageId="12" captionText="Fig. 4. Shell, operculum and radula of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A – B. Shell in front view. C. Shell in lateral view. D. Protoconch and microsculpture. E – F. Internal and external side of the operculum. G. Rows of teeth of the radula. H. Central teeth. I. Detail of outer marginal teeth." httpUri="https://zenodo.org/record/269310/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
G, H); lateral teeth formula 3–C–3; inner marginal teeth having 15–18 sharp cusps; outer marginal teeth having 12–14 sharp cusps (
<figureCitation id="135B2A2CFFAEFFB8FD2B739FFC92FA92" box="[752,824,1268,1294]" captionStart="Fig. 4" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[215,1373,334,1849]" captionTargetId="figure@12.[215,1373,334,1849]" captionTargetPageId="12" captionText="Fig. 4. Shell, operculum and radula of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A – B. Shell in front view. C. Shell in lateral view. D. Protoconch and microsculpture. E – F. Internal and external side of the operculum. G. Rows of teeth of the radula. H. Central teeth. I. Detail of outer marginal teeth." httpUri="https://zenodo.org/record/269310/files/figure.png" pageId="13" pageNumber="14">
Fig.
<date id="FFDE1069FFAEFFB8FCFC739FFC92FA92" box="[807,824,1268,1294]" pageId="13" pageNumber="14">4</date>
</figureCitation>
I).
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF66725EFDBAFACC" blockId="13.[189,1399,1333,1512]" box="[189,528,1333,1360]" pageId="13" pageNumber="14">
<heading id="D09781C5FFAEFFB8FF66725EFDBAFACC" bold="true" box="[189,528,1333,1360]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<emphasis id="B914EABBFFAEFFB8FF66725EFDBAFACC" bold="true" box="[189,528,1333,1360]" pageId="13" pageNumber="14">Pigmentation and anatomy</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF66720FFE3DFA74" blockId="13.[189,1399,1333,1512]" pageId="13" pageNumber="14">
Head light brown pigmented from snout to neck; tentacles also brown pigmented except on ocular lobes; snout as long as wide, with medial lobation. Ctenidium extended across most of pallial cavity with 18–21 narrow gill filaments; osphradium three times longer than wide and opposite middle of ctenidium (
<figureCitation id="135B2A2CFFAEFFB8FF1D72A5FEA1FA74" box="[198,267,1485,1512]" captionStart="Fig. 5" captionStartId="14.[189,232,1860,1886]" captionTargetBox="[370,1225,265,1828]" captionTargetId="figure@14.[370,1225,265,1829]" captionTargetPageId="14" captionText="Fig. 5. Anatomy of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A. Ctenidium and osphradium. B. Partial nervous system. C. Female genitalia. D. Bursa copulatrix and seminal receptacle. E – F. Head of a male and penis. G. Prostate gland. Anatomical abbreviations given in the Material and methods section, except DL (distal lobe)." httpUri="https://zenodo.org/record/269311/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
A,
<tableCitation id="C6E20312FFAEFFB8FEF472A5FE2DFA74" box="[303,391,1486,1512]" captionStart="Table 3" captionStartId="10.[189,258,268,294]" captionTargetBox="[241,1381,463,2008]" captionText="Table 3. Ctenidium, osphradium. Female and male genitalia and nervous system measurements (in mm) of the species: 1. Intermaria zagrosensis (Glöer &amp; Pešić, 2009). 2. Intermaria kermanshahensis (Glöer &amp; Pešić, 2009). 3. Persipyrgula saboori (Glöer &amp; Pešić, 2009). 4. Shadinia terpoghassiani (Shadin, 1952). For ANOVA results effect size F, degrees of freedom (df), residuals (r) and the resulting significance (p – value: *** when 0; ** when 0.001; * when 0.01; NS = not significant) are shown." httpUri="http://table.plazi.org/id/DF1F6621FFA9FFBFFF667667FB0EFE3A" pageId="13" pageNumber="14" tableUuid="DF1F6621FFA9FFBFFF667667FB0EFE3A">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF66717BFE60F9D0" blockId="13.[189,1399,1551,1613]" pageId="13" pageNumber="14">
Nervous system with black pigmentation and elongate (mean RPG ratio 0.50); cerebral ganglia equal in size (
<figureCitation id="135B2A2CFFAEFFB8FF207159FEEAF9D1" box="[251,320,1586,1613]" captionStart="Fig. 5" captionStartId="14.[189,232,1860,1886]" captionTargetBox="[370,1225,265,1828]" captionTargetId="figure@14.[370,1225,265,1829]" captionTargetPageId="14" captionText="Fig. 5. Anatomy of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A. Ctenidium and osphradium. B. Partial nervous system. C. Female genitalia. D. Bursa copulatrix and seminal receptacle. E – F. Head of a male and penis. G. Prostate gland. Anatomical abbreviations given in the Material and methods section, except DL (distal lobe)." httpUri="https://zenodo.org/record/269311/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
B,
<tableCitation id="C6E20312FFAEFFB8FEB97159FE10F9D0" box="[354,442,1586,1612]" captionStart="Table 3" captionStartId="10.[189,258,268,294]" captionTargetBox="[241,1381,463,2008]" captionText="Table 3. Ctenidium, osphradium. Female and male genitalia and nervous system measurements (in mm) of the species: 1. Intermaria zagrosensis (Glöer &amp; Pešić, 2009). 2. Intermaria kermanshahensis (Glöer &amp; Pešić, 2009). 3. Persipyrgula saboori (Glöer &amp; Pešić, 2009). 4. Shadinia terpoghassiani (Shadin, 1952). For ANOVA results effect size F, degrees of freedom (df), residuals (r) and the resulting significance (p – value: *** when 0; ** when 0.001; * when 0.01; NS = not significant) are shown." httpUri="http://table.plazi.org/id/DF1F6621FFA9FFBFFF667667FB0EFE3A" pageId="13" pageNumber="14" tableUuid="DF1F6621FFA9FFBFFF667667FB0EFE3A">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF66711FFADCF964" blockId="13.[189,1399,1652,1784]" pageId="13" pageNumber="14">
Female pallial oviduct with a capsule gland slightly shorter than albumen gland (
<figureCitation id="135B2A2CFFAEFFB8FB47711FFB4FF913" box="[1180,1253,1652,1679]" captionStart="Fig. 5" captionStartId="14.[189,232,1860,1886]" captionTargetBox="[370,1225,265,1828]" captionTargetId="figure@14.[370,1225,265,1829]" captionTargetPageId="14" captionText="Fig. 5. Anatomy of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A. Ctenidium and osphradium. B. Partial nervous system. C. Female genitalia. D. Bursa copulatrix and seminal receptacle. E – F. Head of a male and penis. G. Prostate gland. Anatomical abbreviations given in the Material and methods section, except DL (distal lobe)." httpUri="https://zenodo.org/record/269311/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
C,
<tableCitation id="C6E20312FFAEFFB8FAD1711FFACCF912" box="[1290,1382,1652,1678]" captionStart="Table 3" captionStartId="10.[189,258,268,294]" captionTargetBox="[241,1381,463,2008]" captionText="Table 3. Ctenidium, osphradium. Female and male genitalia and nervous system measurements (in mm) of the species: 1. Intermaria zagrosensis (Glöer &amp; Pešić, 2009). 2. Intermaria kermanshahensis (Glöer &amp; Pešić, 2009). 3. Persipyrgula saboori (Glöer &amp; Pešić, 2009). 4. Shadinia terpoghassiani (Shadin, 1952). For ANOVA results effect size F, degrees of freedom (df), residuals (r) and the resulting significance (p – value: *** when 0; ** when 0.001; * when 0.01; NS = not significant) are shown." httpUri="http://table.plazi.org/id/DF1F6621FFA9FFBFFF667667FB0EFE3A" pageId="13" pageNumber="14" tableUuid="DF1F6621FFA9FFBFFF667667FB0EFE3A">Table 3</tableCitation>
); pyriform bursa copulatrix with a duct as long as bursa length, and lying against the middle section of the albumen gland; renal oviduct white from the insertion point of bursal duct to the seminal receptacle and hereafter black making one or two loops; one elongate seminal receptacle with short duct (
<figureCitation id="135B2A2CFFAEFFB8FAD071B6FAFAF964" box="[1291,1360,1757,1784]" captionStart="Fig. 5" captionStartId="14.[189,232,1860,1886]" captionTargetBox="[370,1225,265,1828]" captionTargetId="figure@14.[370,1225,265,1829]" captionTargetPageId="14" captionText="Fig. 5. Anatomy of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A. Ctenidium and osphradium. B. Partial nervous system. C. Female genitalia. D. Bursa copulatrix and seminal receptacle. E – F. Head of a male and penis. G. Prostate gland. Anatomical abbreviations given in the Material and methods section, except DL (distal lobe)." httpUri="https://zenodo.org/record/269311/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
D).
</paragraph>
<paragraph id="8BDF36A9FFAEFFB8FF667074FDBBF8E3" blockId="13.[189,1399,1823,1920]" pageId="13" pageNumber="14">
Male genitalia with penis gradually tapering bearing a small distal lobe in the inner edge; slightly grayish pigmented on the distal section in some specimens (
<figureCitation id="135B2A2CFFAEFFB8FCFD7029FCC1F8C1" box="[806,875,1858,1885]" captionStart="Fig. 5" captionStartId="14.[189,232,1860,1886]" captionTargetBox="[370,1225,265,1828]" captionTargetId="figure@14.[370,1225,265,1829]" captionTargetPageId="14" captionText="Fig. 5. Anatomy of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A. Ctenidium and osphradium. B. Partial nervous system. C. Female genitalia. D. Bursa copulatrix and seminal receptacle. E – F. Head of a male and penis. G. Prostate gland. Anatomical abbreviations given in the Material and methods section, except DL (distal lobe)." httpUri="https://zenodo.org/record/269311/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
E, F); prostate gland about two times longer than wide (
<figureCitation id="135B2A2CFFAEFFB8FE9B700EFE2FF81C" box="[320,389,1893,1920]" captionStart="Fig. 5" captionStartId="14.[189,232,1860,1886]" captionTargetBox="[370,1225,265,1828]" captionTargetId="figure@14.[370,1225,265,1829]" captionTargetPageId="14" captionText="Fig. 5. Anatomy of Intermaria kermanshahensis (Glöer &amp; Pešić, 2009) gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran. A. Ctenidium and osphradium. B. Partial nervous system. C. Female genitalia. D. Bursa copulatrix and seminal receptacle. E – F. Head of a male and penis. G. Prostate gland. Anatomical abbreviations given in the Material and methods section, except DL (distal lobe)." httpUri="https://zenodo.org/record/269311/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
G,
<tableCitation id="C6E20312FFAEFFB8FE72700EFDABF8E3" box="[425,513,1893,1919]" captionStart="Table 3" captionStartId="10.[189,258,268,294]" captionTargetBox="[241,1381,463,2008]" captionText="Table 3. Ctenidium, osphradium. Female and male genitalia and nervous system measurements (in mm) of the species: 1. Intermaria zagrosensis (Glöer &amp; Pešić, 2009). 2. Intermaria kermanshahensis (Glöer &amp; Pešić, 2009). 3. Persipyrgula saboori (Glöer &amp; Pešić, 2009). 4. Shadinia terpoghassiani (Shadin, 1952). For ANOVA results effect size F, degrees of freedom (df), residuals (r) and the resulting significance (p – value: *** when 0; ** when 0.001; * when 0.01; NS = not significant) are shown." httpUri="http://table.plazi.org/id/DF1F6621FFA9FFBFFF667667FB0EFE3A" pageId="13" pageNumber="14" tableUuid="DF1F6621FFA9FFBFFF667667FB0EFE3A">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1F6621FFADFFBBFF66702FFED5F870" httpUri="https://zenodo.org/record/269311/files/figure.png" pageId="14" pageNumber="15" targetBox="[370,1225,265,1828]" targetPageId="14">
<paragraph id="8BDF36A9FFADFFBBFF66702FFED5F870" blockId="14.[189,1399,1860,2028]" pageId="14" pageNumber="15">
<emphasis id="B914EABBFFADFFBBFF66702FFEA0F8C3" bold="true" box="[189,266,1860,1887]" pageId="14" pageNumber="15">Fig. 5.</emphasis>
Anatomy of
<taxonomicName id="4C604D2AFFADFFBBFE7A702EFC52F8C3" authority="Gloer &amp; Pesic, 2009" authorityName="Gloer &amp; Pesic" authorityYear="2009" box="[417,1016,1860,1887]" class="Gastropoda" family="Hydrobiidae" genus="Intermaria" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="kermanshahensis">
<emphasis id="B914EABBFFADFFBBFE7A702EFD8AF8C2" box="[417,544,1861,1886]" italics="true" pageId="14" pageNumber="15">Intermaria</emphasis>
<taxonomicName id="4C604D2AFFADFFBBFDFC702FFD5BF8C2" box="[551,753,1860,1886]" pageId="14" pageNumber="15" rank="species" species="kermanshahensis">
<emphasis id="B914EABBFFADFFBBFDFC702FFD5BF8C2" box="[551,753,1860,1886]" italics="true" pageId="14" pageNumber="15">kermanshahensis</emphasis>
</taxonomicName>
(Glöer &amp; Pešić, 2009)
</taxonomicName>
gen. et comb. nov. from a spring near Sarabe - Sahneh city, Kermanshah Province, Iran.
<emphasis id="B914EABBFFADFFBBFCE1700CFCE5F81D" bold="true" box="[826,847,1895,1921]" pageId="14" pageNumber="15">A</emphasis>
. Ctenidium and osphradium.
<emphasis id="B914EABBFFADFFBBFB7D7003FB10F81E" bold="true" box="[1190,1210,1896,1922]" pageId="14" pageNumber="15">B</emphasis>
. Partial nervous system.
<emphasis id="B914EABBFFADFFBBFEFA70E0FE9CF839" bold="true" box="[289,310,1931,1957]" pageId="14" pageNumber="15">C</emphasis>
. Female genitalia.
<emphasis id="B914EABBFFADFFBBFDFA70E0FD9CF839" bold="true" box="[545,566,1931,1957]" pageId="14" pageNumber="15">D</emphasis>
. Bursa copulatrix and seminal receptacle.
<emphasis id="B914EABBFFADFFBBFBE570E0FBDBF839" bold="true" box="[1086,1137,1931,1957]" pageId="14" pageNumber="15">E–F</emphasis>
. Head of a male and penis.
<emphasis id="B914EABBFFADFFBBFED070C5FE88F854" bold="true" box="[267,290,1966,1992]" pageId="14" pageNumber="15">G</emphasis>
. Prostate gland. Anatomical abbreviations given in the Material and methods section, except DL (distal lobe).
</paragraph>
</caption>
<subSubSection id="C37A6522FFACFFBAFF667661FA87FD9E" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BDF36A9FFACFFBAFF667661FE85FEB8" blockId="15.[189,1399,266,515]" box="[189,303,266,292]" pageId="15" pageNumber="16">
<heading id="D09781C5FFACFFBAFF667661FE85FEB8" bold="true" box="[189,303,266,292]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="6">
<emphasis id="B914EABBFFACFFBAFF667661FE85FEB8" bold="true" box="[189,303,266,292]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFACFFBAFF667652FA87FD9E" blockId="15.[189,1399,266,515]" pageId="15" pageNumber="16">
Apart from differences in body dimensions and in certain anatomical features (explained above), the penial distal lobe is more prominent in this species than in
<taxonomicName id="4C604D2AFFACFFBAFCB17637FBA0FEE9" box="[874,1034,348,373]" class="Gastropoda" family="Hydrobiidae" genus="Intermaria" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="zagrosensis">
<emphasis id="B914EABBFFACFFBAFCB17637FBA0FEE9" box="[874,1034,348,373]" italics="true" pageId="15" pageNumber="16">I. zagrosensis</emphasis>
</taxonomicName>
. Uncorrected genetic distances are, on the contrary, low between these two species (0.2% for COI fragment,
<tableCitation id="C6E20312FFACFFBAFBE77614FB3EFE05" box="[1084,1172,383,409]" captionStart="Table 4" captionStartId="23.[227,296,192,218]" captionTargetBox="[234,1957,254,1230]" captionTargetPageId="23" captionText="Table 4. Uncorrected COI p-distances between species of the subfamilies Pyrgulinae and Pseudamnicolinae." httpUri="http://table.plazi.org/id/DF1F6621FFB4FFA2FF3877ABFA77FF46" pageId="15" pageNumber="16" tableUuid="DF1F6621FFB4FFA2FF3877ABFA77FF46">Table 4</tableCitation>
), though sister taxa in the subfamily
<taxonomicName id="4C604D2AFFACFFBAFE5776C9FDA1FE20" box="[396,523,418,444]" class="Gastropoda" family="Hydrobiidae" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="subFamily" subFamily="Pyrgulinae">Pyrgulinae</taxonomicName>
are often genetically very close (e.g., COI p-distances of 0.3% between species of the genus
<taxonomicName id="4C604D2AFFACFFBAFE1876AEFCE0FE43" authority="Radoman, 1973" authorityName="Radoman" authorityYear="1973" box="[451,842,453,479]" class="Gastropoda" family="Hydrobiidae" genus="Macedopyrgula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
<emphasis id="B914EABBFFACFFBAFE1876AEFDD6FE43" box="[451,636,453,479]" italics="true" pageId="15" pageNumber="16">Macedopyrgula</emphasis>
Radoman, 1973
</taxonomicName>
). Given this minor genetic variation between congeners, these observed anatomical differences could potentially reflect intraspecific variation.
</paragraph>
</subSubSection>
<subSubSection id="C37A6522FFACFFBAFF66754DFC24FDF3" pageId="15" pageNumber="16" type="biology_ecology">
<paragraph id="8BDF36A9FFACFFBAFF66754DFE58FDDC" blockId="15.[189,910,550,623]" box="[189,498,550,576]" pageId="15" pageNumber="16">
<heading id="D09781C5FFACFFBAFF66754DFE58FDDC" bold="true" box="[189,498,550,576]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="6">
<emphasis id="B914EABBFFACFFBAFF66754DFE58FDDC" bold="true" box="[189,498,550,576]" pageId="15" pageNumber="16">Ecology and distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36A9FFACFFBAFF66753EFC24FDF3" blockId="15.[189,910,550,623]" box="[189,910,596,623]" pageId="15" pageNumber="16">Known only from a spring in the Kermanshah Province, Iran.</paragraph>
</subSubSection>
</treatment>
</document>
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