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<mods:title id="0D069BA026E398536D2ADAD474B9A6C7">Revised taxonomy and distributions of Costa Rican moss salamanders (Caudata: Plethodontidae: Nototriton), with descriptions of new taxa</mods:title>
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<mods:affiliation id="585B0CA717F37C715F908E990DB787A9">Costa Rican Amphibian Research Center, Guayacán, Provincia de Limón, Costa Rica.</mods:affiliation>
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<mods:namePart id="C30E1CEF9B36F226F05F1FED8DF15E12">Reyes, Aura</mods:namePart>
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<mods:affiliation id="AEAE6FC0DCCF827AADE6AD8CAB885556">Costa Rican Amphibian Research Center, Guayacán, Provincia de Limón, Costa Rica. &amp; auralr 22 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5845 - 0528</mods:affiliation>
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<mods:affiliation id="03B3BADAB84BF84B9FA31B0AE580CCAE">Departamento de Zoología, Instituto de Biología, UNAM, AP 70 - 153 Ciudad Universitaria, CP 04510, México, D. F., México. eapiedra @ gmail. com; https: // orcid. org / 0000 - 0002 - 4449 - 1070 &amp; Escuela de Biología, Universidad de Costa Rica, San Pedro, 11501 - 2060 San José, Costa Rica.</mods:affiliation>
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<mods:title id="1B91197FBC5FDB84CB6BCB6215D79D2B">Zootaxa</mods:title>
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<mods:date id="9D31C0F37A3F50FB123A1D3999E29C52">2022</mods:date>
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<treatment id="03C687EDFFC2FFFFA4C540B02CCFF998" ID-DOI="http://doi.org/10.5281/zenodo.7157723" ID-GBIF-Taxon="202157128" ID-Zenodo-Dep="7157723" LSID="urn:lsid:plazi:treatment:03C687EDFFC2FFFFA4C540B02CCFF998" httpUri="http://treatment.plazi.org/id/03C687EDFFC2FFFFA4C540B02CCFF998" lastPageNumber="489" pageId="38" pageNumber="489">
<subSubSection id="C3756570FFC2FFFFA4C540B02EA1FCD2" pageId="38" pageNumber="489" type="nomenclature">
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<heading id="D0988197FFC2FFFFA4C540B02DFCFD3E" bold="true" box="[151,739,716,743]" fontSize="11" level="1" pageId="38" pageNumber="489" reason="1">
<taxonomicName id="4C6F4D78FFC2FFFFA4C540B02DFCFD3E" ID-CoL="74DDT" authority="(Stejneger, 1911)" baseAuthorityName="Stejneger" baseAuthorityYear="1911" box="[151,739,716,743]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi" subGenus="Nototriton">
<emphasis id="B91BEAE9FFC2FFFFA4C540B02DFCFD3E" bold="true" box="[151,739,716,743]" pageId="38" pageNumber="489">
<emphasis id="B91BEAE9FFC2FFFFA4C540B02E0AFD3E" bold="true" box="[151,277,716,742]" italics="true" pageId="38" pageNumber="489">Nototriton</emphasis>
(
<emphasis id="B91BEAE9FFC2FFFFA57440B02EBBFD3E" bold="true" box="[294,420,716,742]" italics="true" pageId="38" pageNumber="489">Nototriton</emphasis>
)
<emphasis id="B91BEAE9FFC2FFFFA5E640B02D13FD3F" bold="true" box="[436,524,716,743]" italics="true" pageId="38" pageNumber="489">picadoi</emphasis>
(
<bibRefCitation id="EFFE4B0AFFC2FFFFA64940B02DC4FD3E" author="Stejneger, L. H." box="[539,731,716,742]" pageId="38" pageNumber="489" pagination="285 - 288" refId="ref27542" refString="Stejneger, L. H. (1911) Descriptions of three new batrachians from Costa Rica and Panama. Proceedings of the United States National Museum, 41, 285 - 288. https: // doi. org / 10.5479 / si. 00963801.1857.285" type="journal article" year="1911">Stejneger, 1911</bibRefCitation>
)
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<paragraph id="8BD036FBFFC2FFFFA4C5408C2EA1FCD2" blockId="38.[151,739,716,779]" box="[151,446,752,779]" pageId="38" pageNumber="489">Picado’s moss salamander</paragraph>
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<subSubSection id="C3756570FFC2FFFFA4C541492C27FBFC" pageId="38" pageNumber="489" type="reference_group">
<paragraph id="8BD036FBFFC2FFFFA4C541492D2AFC95" blockId="38.[151,824,821,1060]" box="[151,565,821,845]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C541492D2AFC95" box="[151,565,821,845]" pageId="38" pageNumber="489">
<treatmentCitation id="0ACE10EAFFC2FFFFA4C541492D2DFC95" author="Stejneger, L. H." box="[151,562,821,845]" page=")" pageId="38" pageNumber="489" year="1911">
<taxonomicName id="4C6F4D78FFC2FFFFA4C541492D2DFC95" ID-CoL="6Z8RX" authority="Stejneger (1911: 285)" authorityName="Stejneger" authorityPageNumber="285" authorityYear="1911" box="[151,562,821,845]" class="Amphibia" family="Plethodontidae" genus="Spelerpes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi">
<emphasis id="B91BEAE9FFC2FFFFA4C541492E53FC95" box="[151,332,821,845]" italics="true" pageId="38" pageNumber="489">Spelerpes picadoi</emphasis>
<bibRefCitation id="EFFE4B0AFFC2FFFFA50041492D2DFC95" author="Stejneger, L. H." box="[338,562,821,845]" pageId="38" pageNumber="489" pagination="285 - 288" refId="ref27542" refString="Stejneger, L. H. (1911) Descriptions of three new batrachians from Costa Rica and Panama. Proceedings of the United States National Museum, 41, 285 - 288. https: // doi. org / 10.5479 / si. 00963801.1857.285" type="journal article" year="1911">Stejneger (1911: 285)</bibRefCitation>
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.
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<paragraph id="8BD036FBFFC2FFFFA4C541282EE4FCB4" blockId="38.[151,824,821,1060]" box="[151,507,852,876]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C541282EE4FCB4" box="[151,507,852,876]" pageId="38" pageNumber="489">
<treatmentCitation id="0ACE10EAFFC2FFFFA4C541282EE8FCB4" author="Dunn, E. R." box="[151,503,852,876]" page=")" pageId="38" pageNumber="489" year="1924">
<taxonomicName id="4C6F4D78FFC2FFFFA4C541282EE8FCB4" authority=": Dunn (1924: 99)" authorityName="Dunn" authorityPageNumber="99" authorityYear="1924" box="[151,503,852,876]" class="Insecta" family="Plethodontidae" genus="Oedipus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi">
<emphasis id="B91BEAE9FFC2FFFFA4C541282E21FCB4" box="[151,318,852,876]" italics="true" pageId="38" pageNumber="489">Oedipus picadoi</emphasis>
:
<bibRefCitation id="EFFE4B0AFFC2FFFFA51941282EE8FCB4" author="Dunn, E. R." box="[331,503,852,876]" pageId="38" pageNumber="489" pagination="95 - 100" refId="ref26233" refString="Dunn, E. R. (1924) New salamanders of the genus Oedipus with a synoptical key. Field Museum Natural History, Zoological Series, 12 (7), 95 - 100." type="journal article" year="1924">Dunn (1924: 99)</bibRefCitation>
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.
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<paragraph id="8BD036FBFFC2FFFFA4C5410E2D76FC52" blockId="38.[151,824,821,1060]" box="[151,617,882,906]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C5410E2D76FC52" box="[151,617,882,906]" pageId="38" pageNumber="489">
?
<treatmentCitation id="0ACE10EAFFC2FFFFA4FB410E2D7AFC52" author="Taylor, E. H." box="[169,613,882,906]" page=")" pageId="38" pageNumber="489" year="1949">
<taxonomicName id="4C6F4D78FFC2FFFFA4FB410E2D7AFC52" ID-CoL="4NWRV" authority=": Taylor (1949: 279)" authorityName="Taylor" authorityPageNumber="279" authorityYear="1949" box="[169,613,882,906]" class="Amphibia" family="Plethodontidae" genus="Pseudoeurycea" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi">
<emphasis id="B91BEAE9FFC2FFFFA4FB410E2E89FC52" box="[169,406,882,906]" italics="true" pageId="38" pageNumber="489">Pseudoeurycea picadoi</emphasis>
:
<bibRefCitation id="EFFE4B0AFFC2FFFFA5F1410E2D7AFC52" author="Taylor, E. H." box="[419,613,882,906]" pageId="38" pageNumber="489" pagination="279 - 288" refId="ref27676" refString="Taylor, E. H. (1949) New salamanders of Costa Rica. University of Kansas Science Bulletin, 33, 279 - 288. https: // doi. org / 10.5962 / bhl. part. 16130" type="journal article" year="1949">Taylor (1949: 279)</bibRefCitation>
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<paragraph id="8BD036FBFFC2FFFFA4C541ED2DB1FC71" blockId="38.[151,824,821,1060]" box="[151,686,913,937]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C541ED2DB1FC71" box="[151,686,913,937]" pageId="38" pageNumber="489">
<taxonomicName id="4C6F4D78FFC2FFFFA4C541ED2E8EFC71" ID-CoL="5Y6LH" baseAuthorityName="Stejneger" baseAuthorityYear="1911" box="[151,401,913,937]" class="Amphibia" family="Plethodontidae" genus="Chiropterotriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi">
<emphasis id="B91BEAE9FFC2FFFFA4C541ED2E8EFC71" box="[151,401,913,937]" italics="true" pageId="38" pageNumber="489">Chiropterotriton picadoi</emphasis>
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:
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<bibRefCitation id="EFFE4B0AFFC2FFFFA5CF41ED2DB5FC71" author="Wake, D. B. &amp; Lynch, J. F." box="[413,682,913,937]" pageId="38" pageNumber="489" pagination="1 - 65" refId="ref27787" refString="Wake, D. B. &amp; Lynch, J. F. (1976) The distribution, ecology, and evolutionary history of plethodontid salamanders in tropical America. Scientific Bulletin Natural History Museum of Los Angeles Country, 25, 1 - 65." type="journal article" year="1976">
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&amp; Lynch (1976: 59)
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<paragraph id="8BD036FBFFC2FFFFA4C541CC2D7DFC10" blockId="38.[151,824,821,1060]" box="[151,610,944,968]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C541CC2D7DFC10" box="[151,610,944,968]" pageId="38" pageNumber="489">
<taxonomicName id="4C6F4D78FFC2FFFFA4C541CC2E4CFC10" ID-CoL="74DDT" baseAuthorityName="Stejneger" baseAuthorityYear="1911" box="[151,339,944,968]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi">
<emphasis id="B91BEAE9FFC2FFFFA4C541CC2E4CFC10" box="[151,339,944,968]" italics="true" pageId="38" pageNumber="489">Nototriton picadoi</emphasis>
</taxonomicName>
:
<treatmentCitation id="0ACE10EAFFC2FFFFA53241CC2D40FC10" author="Wake, D. B. &amp; Elias, P." box="[352,607,944,968]" page="11" pageId="38" pageNumber="489" year="1983">
<bibRefCitation id="EFFE4B0AFFC2FFFFA53241CC2D40FC10" author="Wake, D. B. &amp; Elias, P." box="[352,607,944,968]" pageId="38" pageNumber="489" pagination="1 - 19" refId="ref27717" refString="Wake, D. B. &amp; Elias, P. (1983) New genera and a new species of Central American salamanders, with a review of the tropical genera (Amphibia, Caudata, Plethodontidae). Contributions in Science. Natural History Museum of Los Angeles County, 345, 1 - 19. https: // doi. org / 10.5962 / p. 208170" type="journal article" year="1983">
<collectingCountry id="F378766BFFC2FFFFA53241CC2E86FC10" box="[352,409,944,968]" name="Wake Island" pageId="38" pageNumber="489">Wake</collectingCountry>
&amp; Elias (1983: 11)
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<paragraph id="8BD036FBFFC2FFFFA4C541B32D40FC3E" blockId="38.[151,824,821,1060]" box="[151,607,974,998]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C541B32D40FC3E" box="[151,607,974,998]" pageId="38" pageNumber="489">
<treatmentCitation id="0ACE10EAFFC2FFFFA4C541B32D44FC3E" author="Good, D. A. &amp; Wake, D. B." box="[151,603,974,998]" page=")" pageId="38" pageNumber="489" year="1993">
<taxonomicName id="4C6F4D78FFC2FFFFA4C541B32D44FC3E" ID-CoL="47Z9X" authority="Good &amp; Wake (1993: 137)" authorityName="Good &amp; Wake" authorityYear="1993" box="[151,603,974,998]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="major">
<emphasis id="B91BEAE9FFC2FFFFA4C541B32E5BFC3E" box="[151,324,975,998]" italics="true" pageId="38" pageNumber="489">Nototriton major</emphasis>
<bibRefCitation id="EFFE4B0AFFC2FFFFA51841B22D44FC3E" author="Good, D. A. &amp; Wake, D. B." box="[330,603,974,998]" pageId="38" pageNumber="489" pagination="131 - 159" refId="ref26443" refString="Good, D. A. &amp; Wake, D. B. (1993) Systematic studies of the Costa Rican moss salamanders, genus Nototriton, with descriptions of three new species. Herpetological Monographs, 7, 131 - 159. https: // doi. org / 10.2307 / 1466956" type="journal article" year="1993">Good &amp; Wake (1993: 137)</bibRefCitation>
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<paragraph id="8BD036FBFFC2FFFFA4C541922D88FBDD" blockId="38.[151,824,821,1060]" box="[151,663,1005,1029]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C541922D88FBDD" box="[151,663,1005,1029]" pageId="38" pageNumber="489">
<treatmentCitation id="0ACE10EAFFC2FFFFA4C541922D8CFBDD" ID-DOI="http://doi.org/10.5281/zenodo.5621478" ID-Zenodo-Dep="5621478" author="Boza-Oviedo, E. &amp; Rovito, S. M. &amp; Chaves, G. &amp; Garcia-Rodriguez, A. &amp; Artavia, L. G. &amp; Bolanos, F. &amp; Wake, D. B." box="[151,659,1005,1029]" httpUri="http://treatment.plazi.org/id/547D87DCFF81FFBFFF1E1545F3DBD4F3" page=")" pageId="38" pageNumber="489" year="2012">
<taxonomicName id="4C6F4D78FFC2FFFFA4C541922D8CFBDD" ID-CoL="47Z9Y" authority="Boza-Oviedo et al. (2012: 48)" authorityName="Boza-Oviedo" authorityPageNumber="48" authorityYear="2012" box="[151,659,1005,1029]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="matama">
<emphasis id="B91BEAE9FFC2FFFFA4C541922E46FBDD" box="[151,345,1006,1029]" italics="true" pageId="38" pageNumber="489">Nototriton matama</emphasis>
<bibRefCitation id="EFFE4B0AFFC2FFFFA50D41912D8CFBDD" author="Boza-Oviedo, E. &amp; Rovito, S. M. &amp; Chaves, G. &amp; Garcia-Rodriguez, A. &amp; Artavia, L. G. &amp; Bolanos, F. &amp; Wake, D. B." box="[351,659,1005,1029]" pageId="38" pageNumber="489" pagination="36 - 61" refId="ref26015" refString="Boza-Oviedo, E., Rovito, S. M., Chaves, G., Garcia-Rodriguez, A., Artavia, L. G., Bolanos, F. &amp; Wake, D. B. (2012) Salamanders from the eastern Cordillera de Talamanca, Costa Rica, with descriptions of five new species (Plethodontidae: Bolitoglossa, Nototriton, and Oedipina) and natural history notes from recent expeditions. Zootaxa, 3309 (1), 36 - 61. https: // doi. org / 10.11646 / zootaxa. 3309.1.2" type="journal article" year="2012">
Boza-Oviedo
<emphasis id="B91BEAE9FFC2FFFFA5BE41922D06FBDD" box="[492,537,1005,1029]" italics="true" pageId="38" pageNumber="489">et al</emphasis>
. (2012: 48)
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD036FBFFC2FFFFA4C546702C27FBFC" blockId="38.[151,824,821,1060]" box="[151,824,1036,1060]" pageId="38" pageNumber="489">
<treatmentCitationGroup id="AB7F11D5FFC2FFFFA4C546702C27FBFC" box="[151,824,1036,1060]" pageId="38" pageNumber="489">
<treatmentCitation id="0ACE10EAFFC2FFFFA4C546702C2BFBFC" author="Dubois, A. &amp; Raffaelli, J." box="[151,820,1036,1060]" page=")" pageId="38" pageNumber="489" year="2012">
<taxonomicName id="4C6F4D78FFC2FFFFA4C546702C2BFBFC" ID-CoL="74DDT" authority=": Dubois and Raffaelli (2012: 141)" authorityName=": Dubois and Raffaelli" authorityPageNumber="141" authorityYear="2012" box="[151,820,1036,1060]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi" subGenus="Nototriton">
<emphasis id="B91BEAE9FFC2FFFFA4C546702E1EFBFB" box="[151,257,1036,1059]" italics="true" pageId="38" pageNumber="489">Nototriton</emphasis>
(
<emphasis id="B91BEAE9FFC2FFFFA54246702E65FBFB" box="[272,378,1036,1059]" italics="true" pageId="38" pageNumber="489">Nototriton</emphasis>
)
<emphasis id="B91BEAE9FFC2FFFFA5DA46702ECBFBFC" box="[392,468,1036,1060]" italics="true" pageId="38" pageNumber="489">picadoi</emphasis>
:
<bibRefCitation id="EFFE4B0AFFC2FFFFA5B346702C2BFBFC" author="Dubois, A. &amp; Raffaelli, J." box="[481,820,1036,1060]" pageId="38" pageNumber="489" pagination="77 - 161" refId="ref26197" refString="Dubois, A. &amp; Raffaelli, J. (2012) A new ergotaxonomy of the order Urodela Dumeril, 1805 (Amphibia, Batrachia). Alytes, 28, 77 - 161." type="journal article" year="2012">Dubois and Raffaëlli (2012: 141)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3756570FFC2FFFFA4C5462E2EB3FB6C" pageId="38" pageNumber="489" type="materials_examined">
<paragraph id="8BD036FBFFC2FFFFA4C5462E2EB3FB6C" blockId="38.[151,1437,1106,1600]" pageId="38" pageNumber="489">
<materialsCitation id="3B073CA6FFC2FFFFA4C5462E2EB3FB6C" ID-GBIF-Occurrence="3921375319" collectionCode="USNM" collectorName="La Estrella" elevation="2000" location="Distrito de San Isidro" municipality="Canton de El Guarco" pageId="38" pageNumber="489" specimenCode="USNM 48280" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
<emphasis id="B91BEAE9FFC2FFFFA4C5462E2E14FBB4" bold="true" box="[151,267,1106,1132]" pageId="38" pageNumber="489">
<typeStatus id="54D48859FFC2FFFFA4C5462E2E19FBB4" box="[151,262,1106,1132]" pageId="38" pageNumber="489" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBC99E80FFC2FFFFA546462E2EA7FBB4" box="[276,440,1106,1132]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="489" type="Museum">USNM 48280</specimenCode>
, by original designation, gender not provided in the description:
<collectingRegion id="49ABF819FFC2FFFFA0F1462E2A86FBB4" box="[1187,1433,1106,1132]" country="Costa Rica" name="Cartago" pageId="38" pageNumber="489">Provincia de Cartago</collectingRegion>
:
<collectingMunicipality id="6BB4AC81FFC2FFFFA4C5460A2E97FB48" box="[151,392,1142,1168]" pageId="38" pageNumber="489">Cantón de El Guarco</collectingMunicipality>
:
<location id="8EB06020FFC2FFFFA5C1460A2D96FB48" LSID="urn:lsid:plazi:treatment:03C687EDFFC2FFFFA4C540B02CCFF998:8EB06020FFC2FFFFA5C1460A2D96FB48" box="[403,649,1142,1168]" municipality="Canton de El Guarco" name="Distrito de San Isidro" pageId="38" pageNumber="489" stateProvince="Cartago">Distrito de San Isidro</location>
:
<collectorName id="269A532DFFC2FFFFA6C6460A2C0EFB48" box="[660,785,1142,1168]" pageId="38" pageNumber="489">La Estrella</collectorName>
, ca.
<elevation id="0042D1C8FFC2FFFFA716460A2CCAFB48" box="[836,981,1142,1168]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="489" unit="m" value="2000.0">
<quantity id="4C979B1EFFC2FFFFA716460A2C86FB48" box="[836,921,1142,1168]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="489" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
(Picado 1913), Collected by Clodomido Picado (
<bibRefCitation id="EFFE4B0AFFC2FFFFA4A046E62E80FB6C" author="Stejneger, L. H." box="[242,415,1178,1204]" pageId="38" pageNumber="489" pagination="285 - 288" refId="ref27542" refString="Stejneger, L. H. (1911) Descriptions of three new batrachians from Costa Rica and Panama. Proceedings of the United States National Museum, 41, 285 - 288. https: // doi. org / 10.5479 / si. 00963801.1857.285" type="journal article" year="1911">Stejneger 1911</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3756570FFC2FFFFA49546C22D36FAF8" pageId="38" pageNumber="489" type="etymology">
<paragraph id="8BD036FBFFC2FFFFA49546C22D36FAF8" blockId="38.[151,1437,1106,1600]" pageId="38" pageNumber="489">
<emphasis id="B91BEAE9FFC2FFFFA49546C22E51FB00" bold="true" box="[199,334,1214,1240]" pageId="38" pageNumber="489">Etymology.</emphasis>
The specific epithet is a patronym, used as a noun in the masculine genitive case, in dedication to Dr. Clodomiro Picado Twight (1887–1944), who was a very distinguished Costa Rican scientist and the person that collected the
<typeStatus id="54D48859FFC2FFFFA57E477A2E91FAF8" box="[300,398,1286,1312]" pageId="38" pageNumber="489" type="holotype">holotype</typeStatus>
of this taxon.
</paragraph>
</subSubSection>
<subSubSection id="C3756570FFC2FFFFA49547552CCFF998" pageId="38" pageNumber="489" type="distribution">
<paragraph id="8BD036FBFFC2FFFFA49547552D4DFA0C" blockId="38.[151,1437,1106,1600]" pageId="38" pageNumber="489">
<emphasis id="B91BEAE9FFC2FFFFA49547552E7EFA9B" bold="true" box="[199,353,1321,1347]" pageId="38" pageNumber="489">Distribution.</emphasis>
<taxonomicName id="4C6F4D78FFC2FFFFA53A47572D26FA9C" authorityName=": Dubois and Raffaelli" authorityYear="2012" box="[360,569,1322,1348]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi">
<emphasis id="B91BEAE9FFC2FFFFA53A47572D26FA9C" box="[360,569,1322,1348]" italics="true" pageId="38" pageNumber="489">Nototriton picadoi</emphasis>
</taxonomicName>
is endemic to
<collectingCountry id="F378766BFFC2FFFFA6B347562C44FA9C" box="[737,859,1322,1348]" name="Costa Rica" pageId="38" pageNumber="489">Costa Rica</collectingCountry>
and is only known to inhabit the Caribbean slopes of extreme eastern portion of the Cordillera Volcánica Central, specifically along the eastern flank of the Turrialba Volcano into the northern and eastern sections of the Cordillera de Talamanca. The reported altitudinal range for this taxon is from
<quantity id="4C979B1EFFC2FFFFA56147EA2EF2FA68" box="[307,493,1430,1456]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="2.2" metricValueMin="1.2" pageId="38" pageNumber="489" unit="m" value="1700.0" valueMax="2200.0" valueMin="1200.0">1200–2200 masl</quantity>
(
<bibRefCitation id="EFFE4B0AFFC2FFFFA5AE47EA2D8EFA68" author="Savage, J. M." box="[508,657,1430,1456]" pageId="38" pageNumber="489" refId="ref27408" refString="Savage, J. M. (2002) The Amphibians and Reptiles of Costa Rica: A Herpetofauna between Two Continents, between Two Seas. The University of Chicago Press, Chicago, Illinois, 934 pp." type="book" year="2002">Savage 2002</bibRefCitation>
), but during this study the lower limit was discovered to extend down to
<quantity id="4C979B1EFFC2FFFFA4E647C62E05FA0C" box="[180,282,1466,1492]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="489" unit="m" value="900.0">900 masl</quantity>
(BK personal observation).
</paragraph>
<paragraph id="8BD036FBFFC2FFFFA49547A22CCFF998" blockId="38.[151,1437,1106,1600]" pageId="38" pageNumber="489">
<emphasis id="B91BEAE9FFC2FFFFA49547A22D9AFA20" bold="true" box="[199,645,1501,1528]" pageId="38" pageNumber="489">Noteworthy observations or remarks.</emphasis>
Due to the very low genetic distances, adjoining distributional ranges, and weakly supported morphological diagnostics, we herein recognized
<taxonomicName id="4C6F4D78FFC2FFFFA784447F2B23F9C4" authorityName="Good &amp; Wake" authorityYear="1993" box="[982,1084,1539,1564]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="major">
<emphasis id="B91BEAE9FFC2FFFFA784447F2B23F9C4" box="[982,1084,1539,1564]" italics="true" pageId="38" pageNumber="489">N. major</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFE4B0AFFC2FFFFA01F447E2A21F9C4" author="Good, D. A. &amp; Wake, D. B." box="[1101,1342,1538,1564]" pageId="38" pageNumber="489" pagination="131 - 159" refId="ref26443" refString="Good, D. A. &amp; Wake, D. B. (1993) Systematic studies of the Costa Rican moss salamanders, genus Nototriton, with descriptions of three new species. Herpetological Monographs, 7, 131 - 159. https: // doi. org / 10.2307 / 1466956" type="journal article" year="1993">
Good &amp;
<collectingCountry id="F378766BFFC2FFFFA0E4447E2BE7F9C4" box="[1206,1272,1538,1564]" name="Wake Island" pageId="38" pageNumber="489">Wake</collectingCountry>
, 1993
</bibRefCitation>
) and
<taxonomicName id="4C6F4D78FFC2FFFFA1D1447F2D39F998" authority="(Boza-Oviedo et al., 2012)" baseAuthorityName="Boza-Oviedo" baseAuthorityYear="2012" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="matama">
<emphasis id="B91BEAE9FFC2FFFFA1D1447F2FEEF998" italics="true" pageId="38" pageNumber="489">N. matama</emphasis>
(
<bibRefCitation id="EFFE4B0AFFC2FFFFA552445A2D01F998" author="Boza-Oviedo, E. &amp; Rovito, S. M. &amp; Chaves, G. &amp; Garcia-Rodriguez, A. &amp; Artavia, L. G. &amp; Bolanos, F. &amp; Wake, D. B." box="[256,542,1574,1600]" pageId="38" pageNumber="489" pagination="36 - 61" refId="ref26015" refString="Boza-Oviedo, E., Rovito, S. M., Chaves, G., Garcia-Rodriguez, A., Artavia, L. G., Bolanos, F. &amp; Wake, D. B. (2012) Salamanders from the eastern Cordillera de Talamanca, Costa Rica, with descriptions of five new species (Plethodontidae: Bolitoglossa, Nototriton, and Oedipina) and natural history notes from recent expeditions. Zootaxa, 3309 (1), 36 - 61. https: // doi. org / 10.11646 / zootaxa. 3309.1.2" type="journal article" year="2012">
Boza-Oviedo
<emphasis id="B91BEAE9FFC2FFFFA5CC445B2ED0F998" box="[414,463,1574,1600]" italics="true" pageId="38" pageNumber="489">et al</emphasis>
., 2012
</bibRefCitation>
)
</taxonomicName>
as junior synonyms under
<taxonomicName id="4C6F4D78FFC2FFFFA706445B2CD6F998" authorityName=": Dubois and Raffaelli" authorityYear="2012" box="[852,969,1574,1600]" class="Amphibia" family="Plethodontidae" genus="Nototriton" kingdom="Animalia" order="Caudata" pageId="38" pageNumber="489" phylum="Chordata" rank="species" species="picadoi">
<emphasis id="B91BEAE9FFC2FFFFA706445B2CD6F998" box="[852,969,1574,1600]" italics="true" pageId="38" pageNumber="489">N. picadoi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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