159 lines
18 KiB
XML
159 lines
18 KiB
XML
<document id="4DC544A19829448F67375E27EC6DCB07" ID-DOI="10.5281/zenodo.276822" ID-GBIF-Dataset="f5489381-8fe8-48a0-9fb6-5addeebaa3f8" ID-ISSN="1175-5326" ID-Zenodo-Dep="276822" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460237113383" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2011" docId="03C687E4FFD0FF9AFF1BEA14156CF85E" docLanguage="en" docName="zt02763p033.pdf" docOrigin="Zootaxa 2763" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hlavaciellus clandestinus Jałoszyński, 2011, sp. n." docType="treatment" docVersion="6" lastPageNumber="16" masterDocId="FFFFFF9CFFDFFF95FF8CEB58101EFFCA" masterDocTitle="Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="16" updateTime="1698239820214" updateUser="plazi">
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<mods:title id="E4C784AADFBC49C16787797FDA58D6CE">Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="AB21EAA92B41F8CEDD39249B3C76C1AB" type="personal">
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<mods:roleTerm id="0B561626938E5A05B96A4406AC18D469">Author</mods:roleTerm>
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<mods:namePart id="F8A712B4B7D17C9794A6E52067882710">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="14114CDECF5214DDEDBD469D5274B5B1">text</mods:typeOfResource>
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<mods:title id="80B9FA213BA5DCAEC8CA40B2EFC8ECE0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5155D5AE493299330C063B0E7F8F16A5">
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<mods:date id="C56DA8E8BFD98BFD91788116E080FF74">2011</mods:date>
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<mods:detail id="E023E44877A30A4C8785FB083AF97D3B" type="volume">
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<mods:number id="F5DF295D6C61B478FA2A41ACA1A24740">2763</mods:number>
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<mods:classification id="3AA3FD5BB1E4798AF2924D95C9278A14">journal article</mods:classification>
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<mods:identifier id="77D951669A4C156A882BC3E2E9132E47" type="DOI">10.5281/zenodo.276822</mods:identifier>
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<mods:identifier id="DEEA3806E0D3B34694FA4E945F5EBE80" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="14D1642CBEF1AE8588A0F3AF8CB81C2A" type="Zenodo-Dep">276822</mods:identifier>
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<treatment id="03C687E4FFD0FF9AFF1BEA14156CF85E" ID-DOI="http://doi.org/10.5281/zenodo.6192613" ID-GBIF-Taxon="119388737" ID-Zenodo-Dep="6192613" LSID="urn:lsid:plazi:treatment:03C687E4FFD0FF9AFF1BEA14156CF85E" httpUri="http://treatment.plazi.org/id/03C687E4FFD0FF9AFF1BEA14156CF85E" lastPageNumber="16" pageId="15" pageNumber="16">
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<subSubSection id="C3756579FFD0FF9AFF1BEA1411B5FE4C" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="8BD036F2FFD0FF9AFF1BEA141208FEAF" blockId="15.[151,534,331,390]" box="[151,534,331,358]" pageId="15" pageNumber="16">
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<heading id="D098819EFFD0FF9AFF1BEA141208FEAF" bold="true" box="[151,534,331,358]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<emphasis id="B91BEAE0FFD0FF9AFF1BEA141208FEAF" bold="true" box="[151,534,331,358]" pageId="15" pageNumber="16">
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<taxonomicName id="4C6F4D71FFD0FF9AFF1BEA1411D2FEAC" ID-CoL="3M5PS" box="[151,460,332,358]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="clandestinus" status="sp. nov.">
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<emphasis id="B91BEAE0FFD0FF9AFF1BEA1411D2FEAC" bold="true" box="[151,460,332,358]" italics="true" pageId="15" pageNumber="16">Hlavaciellus clandestinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A228579BFFD0FF9AFE58EA131208FEAF" box="[468,534,331,357]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD036F2FFD0FF9AFF1BEA3611B5FE4C" blockId="15.[151,534,331,390]" box="[151,427,366,390]" pageId="15" pageNumber="16">
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(
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<figureCitation id="13542A77FFD0FF9AFF13EA3610E3FE4C" box="[159,253,366,390]" captionStart="FIGURES 27 – 28" captionStartId="16.[151,264,1178,1201]" captionTargetBox="[305,1293,212,1160]" captionTargetId="figure@16.[281,1310,193,1169]" captionTargetPageId="16" captionText="FIGURES 27 – 28. Dorsal view of holotypes of Hlavaciellus, males. 27, H. clandestinus sp. n. 28, H. extremalis sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276828/files/figure.png" pageId="15" pageNumber="16">Figs. 27</figureCitation>
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, 37, 38, 57, 58)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD0FF9AFF1BEAE01277FDDD" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="8BD036F2FFD0FF9AFF1BEAE01277FDDD" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">
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<emphasis id="B91BEAE0FFD0FF9AFF1BEAE0126DFE1A" bold="true" box="[151,627,439,464]" pageId="15" pageNumber="16">
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<typeStatus id="54D48850FFD0FF9AFF1BEAE010CCFE1A" box="[151,210,440,464]" pageId="15" pageNumber="16">Type</typeStatus>
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material.
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<typeStatus id="54D48850FFD0FF9AFEDFEAE011DAFE1A" box="[339,452,440,464]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3787662FFD0FF9AFE58EAEF1270FE1A" box="[468,622,439,464]" name="Malaysia" pageId="15" pageNumber="16">MALAYSIA</collectingCountry>
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:
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</emphasis>
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ɗ, "
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<collectionCode id="ED7EAE37FFD0FF9AFD20EAEF130CFE05" box="[684,786,439,463]" pageId="15" pageNumber="16">SABAH</collectionCode>
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: E Mt. Kinabalu \
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<quantity id="4C979B17FFD0FF9AFC7FEAEF1453FE05" box="[1011,1101,439,463]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="15" pageNumber="16" unit="m" value="1150.0">1150 m</quantity>
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, rte Ranau–Kota \ Kinabalu,
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<date id="FFD11032FFD0FF9AFF1BEA841117FE3E" box="[151,265,476,500]" pageId="15" pageNumber="16" value="1987-05-24">24.V.1987</date>
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\ Burckhardt - Löbl" [white, printed], "
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<taxonomicName id="4C6F4D71FFD0FF9AFD4AEA841396FE3E" box="[710,904,476,500]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
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<emphasis id="B91BEAE0FFD0FF9AFD4AEA841396FE3E" box="[710,904,476,500]" italics="true" pageId="15" pageNumber="16">HLAVACIELLUS</emphasis>
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</taxonomicName>
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\
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<taxonomicName id="4C6F4D71FFD0FF9AFC13EA841435FE3E" box="[927,1067,476,500]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="clandestinus">
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<emphasis id="B91BEAE0FFD0FF9AFC13EA841435FE3E" box="[927,1067,476,500]" italics="true" pageId="15" pageNumber="16">clandestinus</emphasis>
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</taxonomicName>
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m. \ det. P. JAŁOSZYŃSKI, '09 \
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<typeStatus id="54D48850FFD0FF9AFF29EAA71151FDDD" box="[165,335,511,535]" pageId="15" pageNumber="16" type="holotype">HOLOTYPUS</typeStatus>
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" [red, printed] (
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<collectionCode id="ED7EAE37FFD0FF9AFD8DEAA71243FDDD" box="[513,605,511,535]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16">MHNG</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD0FF9AFF4BE97B1466FD95" pageId="15" pageNumber="16" type="diagnosis">
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<paragraph id="8BD036F2FFD0FF9AFF4BE97B1466FD95" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">
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<emphasis id="B91BEAE0FFD0FF9AFF4BE97B115FFDF6" bold="true" box="[199,321,547,572]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
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Head of males non-modified; antennomere III subquadrate; antennomere XI stout, less than twice as long as broad; aedeagus in ventral view very slender, feebly narrowing in apical third.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD0FF9AFF4BE93311BCF976" pageId="15" pageNumber="16" type="description">
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<paragraph id="8BD036F2FFD0FF9AFF4BE93310F7FD6D" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">
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<emphasis id="B91BEAE0FFD0FF9AFF4BE9331147FD4E" bold="true" box="[199,345,619,644]" pageId="15" pageNumber="16">Description.</emphasis>
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BL 2.00. Body of male (
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<figureCitation id="13542A77FFD0FF9AFDF8E93412D8FD4E" box="[628,710,620,644]" captionStart="FIGURES 27 – 28" captionStartId="16.[151,264,1178,1201]" captionTargetBox="[305,1293,212,1160]" captionTargetId="figure@16.[281,1310,193,1169]" captionTargetPageId="16" captionText="FIGURES 27 – 28. Dorsal view of holotypes of Hlavaciellus, males. 27, H. clandestinus sp. n. 28, H. extremalis sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276828/files/figure.png" pageId="15" pageNumber="16">Fig. 27</figureCitation>
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) moderately convex, pigmentation reddish-brown, vestiture yellowish.
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</paragraph>
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<paragraph id="8BD036F2FFD0FF9AFF4BE9EC13FDFBDA" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">
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Head large, HL 0.25, HW 0.45; vertex convex, not separated from frons; frons slightly flattened; clypeus convex; supraantennal tubercles small but distinct, clearly but not sharply delimited from median part of frons; eyes large, strongly convex and coarsely faceted. Punctures on dorsal surface of head very fine, separated by spaces 1–
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<date id="FFD11032FFD0FF9AFF1BE84710BBFCFD" box="[151,165,799,823]" pageId="15" pageNumber="16">2</date>
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x as wide as puncture diameters; setae short, relatively dense, suberect. Antennae as long as 0.70 BL, relatively stout and strongly thickened up to antennomere X, uniformly covered with thin, long, dense and suberect setae, AnL 1.43, antennomere I 1.5x as long as broad; II distinctly narrower and shorter than I, subquadrate; III slightly broader and longer than II, 1.1x as long as broad; IV slightly larger than III, 1.2x as long as broad; V slightly broader and much longer than IV, 1.5x as long as broad; VI–VIII slightly and gradually increasing in length and width, each about 1.6x as long as broad; IX slightly larger than VIII, 1.5x as long as broad; X yet larger, 1.3x as long as broad; XI nearly as broad as X, slightly shorter than IX–X together.
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</paragraph>
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<paragraph id="8BD036F2FFD0FF9AFF4BEF45154DFAC6" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">
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Pronotum in dorsal view trapezoidal with weakly convex anterior margin, broadest at base but very weakly narrowing anteriorly, PL 0.63, PW 0.85; sides strongly rounded in anterior third, barely noticeably concave just behind middle and then slightly divergent towards nearly right and blunt hind angles; posterior margin shallowly bi-emarginate; ante-basal pits shallow and diffused, each located slightly closer to posterior than to lateral margin of pronotum; lateral edges not swollen. Pronotal disc glossy, in middle covered with extremely small and shallow punctures separated by spaces 1–
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<date id="FFD11032FFD0FF9AFD9FEF88123CFB22" box="[531,546,1232,1256]" pageId="15" pageNumber="16">2</date>
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x as wide as puncture diameters, punctures close to margins of pronotum are larger and denser, those near posterior angles appear coarse. Setae long, moderately dense, suberect to erect.
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</paragraph>
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<paragraph id="8BD036F2FFD0FF9AFF4BEE4010F1F9C2" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">
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Elytra oval, relatively long, broadest between middle and anterior third, EL 1.10, EW 0.85, EI 1.29; basal pit on each elytron small but distinct, located close to scutellum; subhumeral lines sharply marked, as long as 0.32x EL, moderately strongly divergent, each developed as border between higher humeral region and lower adsutural area; apices of elytra separately rounded. Punctures on basal third of each elytron more distinct than those on pronotum but very shallow, separated by spaces comparable to puncture diameters, gradually reducing in diameter and depth toward lateral margins and apices; setae similar to those on pronotum but slightly thicker.
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<collectingCountry id="F3787662FFD0FF9AFB67EE94153DFA2E" box="[1259,1315,1484,1508]" name="India" pageId="15" pageNumber="16">Hind</collectingCountry>
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wings not studied.
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</paragraph>
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<paragraph id="8BD036F2FFD0FF9AFF4BED4D12C2F9E6" blockId="15.[151,1437,439,1940]" box="[199,732,1556,1581]" pageId="15" pageNumber="16">Legs slender and long, all tibiae nearly straight.</paragraph>
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<paragraph id="8BD036F2FFD0FF9AFF4BED60148DF99A" blockId="15.[151,1437,439,1940]" box="[199,1171,1592,1616]" pageId="15" pageNumber="16">Metaventrite with deep postmesocoxal impressions, occupying about half of its length.</paragraph>
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<paragraph id="8BD036F2FFD0FF9AFF4BED04151CF952" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">Aedeagus (Figs. 37, 38) very slender, AeL 0.41; median lobe with short, trapezoidal apical part; internal armature strongly asymmetrical; parameres in lateral view moderately broad, with strongly curved apices.</paragraph>
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<paragraph id="8BD036F2FFD0FF9AFF4BEDFC11BCF976" blockId="15.[151,1437,439,1940]" box="[199,418,1700,1724]" pageId="15" pageNumber="16">
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<emphasis id="B91BEAE0FFD0FF9AFF4BEDFC113BF976" bold="true" box="[199,293,1700,1724]" pageId="15" pageNumber="16">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD0FF9AFF4BED9F12DEF92A" box="[199,704,1735,1760]" pageId="15" pageNumber="16" type="distribution">
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<paragraph id="8BD036F2FFD0FF9AFF4BED9F12DEF92A" blockId="15.[151,1437,439,1940]" box="[199,704,1735,1760]" pageId="15" pageNumber="16">
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<emphasis id="B91BEAE0FFD0FF9AFF4BED9F117EF92A" bold="true" box="[199,352,1735,1760]" pageId="15" pageNumber="16">Distribution.</emphasis>
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East
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<collectingCountry id="F3787662FFD0FF9AFE12ED901217F92A" box="[414,521,1736,1760]" name="Malaysia" pageId="15" pageNumber="16">Malaysia</collectingCountry>
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: Borneo, Sabah.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD0FF9AFF4BEDB41364F8E2" pageId="15" pageNumber="16" type="etymology">
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<paragraph id="8BD036F2FFD0FF9AFF4BEDB41364F8E2" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">
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<emphasis id="B91BEAE0FFD0FF9AFF4BEDB41150F8CE" bold="true" box="[199,334,1772,1796]" pageId="15" pageNumber="16">Etymology.</emphasis>
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The Latin adjective
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<taxonomicName id="4C6F4D71FFD0FF9AFDB7EDB412D9F8CE" box="[571,711,1772,1796]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="clandestinus">
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<emphasis id="B91BEAE0FFD0FF9AFDB7EDB412D9F8CE" box="[571,711,1772,1796]" italics="true" pageId="15" pageNumber="16">clandestinus</emphasis>
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</taxonomicName>
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("secret, hidden, concealed, clandestine") was chosen to underline unremarkable morphology of this difficult to identify species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD0FF9AFF4BEC6C156CF85E" pageId="15" pageNumber="16" type="discussion">
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<paragraph id="8BD036F2FFD0FF9AFF4BEC6C156CF85E" blockId="15.[151,1437,439,1940]" pageId="15" pageNumber="16">
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<emphasis id="B91BEAE0FFD0FF9AFF4BEC6C1125F886" bold="true" box="[199,315,1844,1868]" pageId="15" pageNumber="16">Remarks.</emphasis>
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<taxonomicName id="4C6F4D71FFD0FF9AFECEEC6C127CF886" box="[322,610,1844,1868]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="clandestinus">
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<emphasis id="B91BEAE0FFD0FF9AFECEEC6C127CF886" box="[322,610,1844,1868]" italics="true" pageId="15" pageNumber="16">Hlavaciellus clandestinus</emphasis>
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</taxonomicName>
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is externally most similar to
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<taxonomicName id="4C6F4D71FFD0FF9AFC25EC6D143EF886" box="[937,1056,1845,1868]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="metrios">
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<emphasis id="B91BEAE0FFD0FF9AFC25EC6D143EF886" box="[937,1056,1845,1868]" italics="true" pageId="15" pageNumber="16">H. metrios</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6F4D71FFD0FF9AFBDAEC6D14A9F886" box="[1110,1207,1845,1868]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="miser">
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<emphasis id="B91BEAE0FFD0FF9AFBDAEC6D14A9F886" box="[1110,1207,1845,1868]" italics="true" pageId="15" pageNumber="16">H. miser</emphasis>
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</taxonomicName>
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; differences are discussed in the remarks for these two species. The aedeagus in ventral view is hardly distinguishable from that of
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<taxonomicName id="4C6F4D71FFD0FF9AFA0DEC0110E2F85E" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="adelphos">
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<emphasis id="B91BEAE0FFD0FF9AFA0DEC0110E2F85E" italics="true" pageId="15" pageNumber="16">H. adelphos</emphasis>
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</taxonomicName>
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, but the external morphology, especially of the head of males of these two species is clearly different.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |