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<document id="94F03EC463059A00F6F1638CC0B0479E" ID-CLB-Dataset="6535" ID-DOI="10.1080/00222933.2014.902140" ID-GBIF-Dataset="58cf5eca-ad81-4c3d-97a3-c6b17ee9137b" ID-ISSN="1464-5262" ID-Zenodo-Dep="4631429" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616520937039" checkinUser="carolina" docAuthor="Dankittipakul, Pakawin &amp; Singtripop, Tippawan" docDate="2014" docId="03C687E26F17D375FE48F9FED6A4FAA0" docLanguage="en" docName="JNATHIST.48.31-32.1923-1936.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 48 (31 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2014.902140" docStyle="DocumentStyle:CD936EBDDF5274E0936345450CC2DAE4.5:JNatHist.2013-2015.journal_article.1cover" docStyleId="CD936EBDDF5274E0936345450CC2DAE4" docStyleName="JNatHist.2013-2015.journal_article.1cover" docStyleVersion="5" docTitle="Clubiona apiculata Dankittipakul &amp; Singtripop 2014" docType="treatment" docVersion="7" lastPageNumber="1928" masterDocId="FFFFFF9A6F15D373FFD8FF85D751FFC8" masterDocTitle="A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae)" masterLastPageNumber="1936" masterPageNumber="1923" pageNumber="1924" updateTime="1698934598660" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D137CD7441A1008F41C30601B322173C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="267BC0908B6EA3E9ABC7C18EC3A6DB11">
<mods:title id="968D8B92957C2C90CC2A5C8E09515AF1">A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae)</mods:title>
</mods:titleInfo>
<mods:name id="8AD8B996BC1703B4B0D296AD38487BDE" type="personal">
<mods:role id="67DC0ED56BEC375051014275C3E43A1F">
<mods:roleTerm id="984AD4C5EC54E13E15754E8523B1F79E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C60738C036FE6EF91440B6303797B44">Dankittipakul, Pakawin</mods:namePart>
<mods:affiliation id="713551D245E1478601CFF50833DA223A">Endocrinology Research Laboratory, Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand; &amp; Manukau Institute of Technology, Auckland, New Zealand</mods:affiliation>
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<mods:roleTerm id="28B44A62348073BF1F21C839736DD5B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="517CD518C0EABC8E0B5CCC7838C7C73F">Singtripop, Tippawan</mods:namePart>
<mods:affiliation id="DDB201D59EBDBD87BC2A2211141EE15F">Endocrinology Research Laboratory, Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand;</mods:affiliation>
</mods:name>
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<mods:title id="FFF65AAFA9AFD72D4443E2A0D87BCE54">Journal of Natural History</mods:title>
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<mods:part id="7BE83B6F0A8B166A237AE581EA03FDF5">
<mods:date id="8A615723AFCF49FE97431950C3438B87">2014</mods:date>
<mods:detail id="950A5FA4B741EBB2A77EA049150EA8A4" type="series">
<mods:title id="E0528846FA3C74FBBE02C2B8DBD0A0D5">J. Nat. Hist.</mods:title>
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<mods:detail id="5C8A6CAEEECA75F1378BE58042A954EC" type="pubDate">
<mods:number id="EDD68324961DC0059202BD533F26B465">2014-04-09</mods:number>
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<mods:detail id="42C0B7591E521E3CC27CE9C9CF3139DD" type="volume">
<mods:number id="C075A8FFFE93BD0760A5A5A3B7D35BC4">48</mods:number>
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<mods:detail id="22EE6F915DE6D407A06FC3FA00E616B6" type="issue">
<mods:number id="2E9BB330865C5C508649C41E2EC9333B">31 - 32</mods:number>
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<mods:start id="57A53BADAB4C282E394F9E0A4F4EACA2">1923</mods:start>
<mods:end id="CE901DA0EDA656A53D1F0F60448CDDC4">1936</mods:end>
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<mods:url id="E2032E61993F0DB901865097705A760E">http://dx.doi.org/10.1080/00222933.2014.902140</mods:url>
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<mods:classification id="9A0276141CC71162DCF8EA9AEB14DE05">journal article</mods:classification>
<mods:identifier id="25A3085D6F38C758B167BB36B27FEE63" type="CLB-Dataset">6535</mods:identifier>
<mods:identifier id="1D64F0A9F905778AE3186BD75B2EE837" type="DOI">10.1080/00222933.2014.902140</mods:identifier>
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<mods:identifier id="ECD90468ED8456015752EBA15BFCEB0A" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03C687E26F17D375FE48F9FED6A4FAA0" ID-DOI="http://doi.org/10.5281/zenodo.4783718" ID-GBIF-Taxon="182383857" ID-Zenodo-Dep="4783718" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0" httpUri="http://treatment.plazi.org/id/03C687E26F17D375FE48F9FED6A4FAA0" lastPageId="6" lastPageNumber="1928" pageId="2" pageNumber="1924">
<subSubSection id="C375657F6F17D371FE48F9FED41EF95B" box="[400,847,1659,1683]" pageId="2" pageNumber="1924" type="nomenclature">
<paragraph id="8BD036F46F17D371FE48F9FED41EF95B" blockId="2.[354,894,1563,1683]" box="[400,847,1659,1683]" pageId="2" pageNumber="1924">
<heading id="D09881986F17D371FE48F9FED41EF95B" box="[400,847,1659,1683]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="1924" reason="2">
The
<emphasis id="B91BEAE66F17D371FE1DF9FED520F95B" bold="true" box="[453,625,1659,1683]" pageId="2" pageNumber="1924">
<taxonomicName id="4C6F4D776F17D371FE1DF9FED57BF95B" ID-CoL="5ZDBH" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[453,554,1659,1683]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="1924" phylum="Arthropoda" rank="species" species="apiculata">
<emphasis id="B91BEAE66F17D371FE1DF9FED57BF95B" bold="true" box="[453,554,1659,1683]" italics="true" pageId="2" pageNumber="1924">apiculata</emphasis>
</taxonomicName>
-group
</emphasis>
, new species-group
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375657F6F16D375FF0CFF2DD542FD4B" lastPageId="6" lastPageNumber="1928" pageId="3" pageNumber="1925" type="diagnosis">
<paragraph id="8BD036F46F16D370FF0CFF2DD611FF08" blockId="3.[212,1172,168,583]" box="[212,320,168,192]" pageId="3" pageNumber="1925">
<heading id="D09881986F16D370FF0CFF2DD611FF08" box="[212,320,168,192]" fontSize="10" level="3" pageId="3" pageNumber="1925" reason="5">
<emphasis id="B91BEAE66F16D370FF0CFF2DD611FF08" box="[212,320,168,192]" italics="true" pageId="3" pageNumber="1925">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F16D370FF0CFF55D510FD8F" blockId="3.[212,1172,168,583]" pageId="3" pageNumber="1925">
Four characters stand out for all members of the
<taxonomicName id="4C6F4D776F16D370FCA8FF55D486FF20" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[880,983,208,232]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="1925" phylum="Arthropoda" rank="species" species="apiculata">
<emphasis id="B91BEAE66F16D370FCA8FF55D486FF20" box="[880,983,208,232]" italics="true" pageId="3" pageNumber="1925">apiculata</emphasis>
</taxonomicName>
-group: (1) the widely separated PME located at least twice their diameter apart (
<figureCitation id="13542A716F16D370FBDFFF75D3D2FEC0" box="[1031,1155,240,264]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="3" pageNumber="1925">Figure 2G</figureCitation>
); (2) the tibiae and metatarsi of the anterior legs carrying pairs of elongated and slender spines (
<figureCitation id="13542A716F16D370FE17FEB5D536FE80" box="[463,615,304,328]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="3" pageNumber="1925">Figure 2H–I</figureCitation>
); (3) the presence of the tubercle at dorsal cymbium (
<figureCitation id="13542A716F16D370FE8EFED5D685FEA0" box="[342,468,336,360]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="3" pageNumber="1925">Figure 1G</figureCitation>
); and (4) the well-developed retrolateral tibial apophysis (
<figureCitation id="13542A716F16D370FF05FEEAD62BFE4F" box="[221,378,367,392]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="3" pageNumber="1925">Figure 1A–D</figureCitation>
). There are three pairs of spines on the ventral surface of tibia I and II (
<figureCitation id="13542A716F16D370FEE8FE0AD690FE6F" box="[304,449,399,423]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="3" pageNumber="1925">Figure 2H–I</figureCitation>
, TS), whereas metatarsus has only one pair, but these spines are much longer, with their base originating on proximal part and their apices almost reaching distal margin of metatarsus (
<figureCitation id="13542A716F16D370FD20FE4AD4DDFE2F" box="[760,908,463,487]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="3" pageNumber="1925">Figure 2H–I</figureCitation>
, MS). Pairs of spines can also be found on the femora (
<figureCitation id="13542A716F16D370FD41FE6AD460FDCF" box="[665,817,495,519]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="3" pageNumber="1925">Figure 2H–I</figureCitation>
, FS). They are curved and elongate, situated on the dorsal side of all femora, whereas those on other leg segments are relatively shorter.
</paragraph>
<caption id="DF10667C6F16D370FF0CF9CDD47AF950" ID-DOI="http://doi.org/10.5281/zenodo.4631431" ID-Zenodo-Dep="4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="3" pageNumber="1925" startId="3.[212,279,1608,1630]" targetBox="[212,1172,671,1576]" targetPageId="3">
<paragraph id="8BD036F46F16D370FF0CF9CDD47AF950" blockId="3.[212,1172,1608,1688]" pageId="3" pageNumber="1925">
Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G).
<taxonomicName id="4C6F4D776F16D370FBE2F9CDD67EF9B2" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="1925" phylum="Arthropoda" rank="species" species="apiculata" status="sp. nov.">
<emphasis id="B91BEAE66F16D370FBE2F9CDD67EF9B2" italics="true" pageId="3" pageNumber="1925">Clubiona apiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F16D370FEE1F9E0D6DFF9B3" box="[313,398,1637,1659]" pageId="3" pageNumber="1925" rank="species">sp. nov.</taxonomicNameLabel>
(A,F,G).
<taxonomicName id="4C6F4D776F16D370FE24F9E0D5B8F9B2" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[508,745,1637,1658]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="1925" phylum="Arthropoda" rank="species" species="cylindriformis" status="sp. nov.">
<emphasis id="B91BEAE66F16D370FE24F9E0D5B8F9B2" box="[508,745,1637,1658]" italics="true" pageId="3" pageNumber="1925">Clubiona cylindriformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F16D370FD2CF9E0D418F9B3" box="[756,841,1637,1659]" pageId="3" pageNumber="1925" rank="species">sp. nov.</taxonomicNameLabel>
(B).
<taxonomicName id="4C6F4D776F16D370FC5BF9E0D364F9B2" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[899,1077,1637,1658]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="1925" phylum="Arthropoda" rank="species" species="cultrata" status="sp. nov.">
<emphasis id="B91BEAE66F16D370FC5BF9E0D364F9B2" box="[899,1077,1637,1658]" italics="true" pageId="3" pageNumber="1925">Clubiona cultrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F16D370FB98F9E0D3C5F9B3" box="[1088,1172,1637,1659]" pageId="3" pageNumber="1925" rank="species">sp. nov.</taxonomicNameLabel>
(C).
<taxonomicName id="4C6F4D776F16D370FEDBF907D6F5F95F" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[259,420,1666,1687]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="1925" phylum="Arthropoda" rank="species" species="conica" status="sp. nov.">
<emphasis id="B91BEAE66F16D370FEDBF907D6F5F95F" box="[259,420,1666,1687]" italics="true" pageId="3" pageNumber="1925">Clubiona conica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F16D370FE74F907D6ACF950" box="[428,509,1666,1688]" pageId="3" pageNumber="1925" rank="species">sp. nov.</taxonomicNameLabel>
(D).
<taxonomicName id="4C6F4D776F16D370FDEEF907D5ACF95F" authorityName="Berland" authorityYear="1938" box="[566,765,1666,1688]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="1925" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="B91BEAE66F16D370FDEEF907D5C1F95F" box="[566,656,1666,1687]" italics="true" pageId="3" pageNumber="1925">Clubiona</emphasis>
cf.
<emphasis id="B91BEAE66F16D370FD60F907D5ACF95F" box="[696,765,1666,1687]" italics="true" pageId="3" pageNumber="1925">hystrix</emphasis>
</taxonomicName>
(E).
</paragraph>
</caption>
<caption id="DF10667C6F11D377FF48FA8DD515FABE" ID-DOI="http://doi.org/10.5281/zenodo.4631433" ID-Zenodo-Dep="4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="4" pageNumber="1926" startId="4.[144,211,1288,1310]" targetBox="[144,1104,168,1256]" targetPageId="4">
<paragraph id="8BD036F46F11D377FF48FA8DD515FABE" blockId="4.[144,1104,1288,1398]" pageId="4" pageNumber="1926">
Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C)
<taxonomicName id="4C6F4D776F11D377FC76FAC6D31EFA90" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[942,1103,1347,1368]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="1926" phylum="Arthropoda" rank="species" species="conica" status="sp. nov.">
<emphasis id="B91BEAE66F11D377FC76FAC6D31EFA90" box="[942,1103,1347,1368]" italics="true" pageId="4" pageNumber="1926">Clubiona conica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F11D377FF48FAE5D7B2FABE" box="[144,227,1376,1398]" pageId="4" pageNumber="1926" rank="species">sp. nov.</taxonomicNameLabel>
; (D–I)
<taxonomicName id="4C6F4D776F11D377FEF7FAE5D6BAFABD" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[303,491,1376,1397]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="1926" phylum="Arthropoda" rank="species" species="apiculata" status="sp. nov.">
<emphasis id="B91BEAE66F11D377FEF7FAE5D6BAFABD" box="[303,491,1376,1397]" italics="true" pageId="4" pageNumber="1926">Clubiona apiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F11D377FE2BFAE5D515FABE" box="[499,580,1376,1398]" pageId="4" pageNumber="1926" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BD036F46F11D375FF62FA21D6E2FF28" blockId="4.[144,1104,1444,1691]" lastBlockId="6.[144,1104,168,479]" lastPageId="6" lastPageNumber="1928" pageId="4" pageNumber="1926">
Males of the
<taxonomicName id="4C6F4D776F11D377FE86FA21D692FA74" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[350,451,1444,1468]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="1926" phylum="Arthropoda" rank="species" species="apiculata">
<emphasis id="B91BEAE66F11D377FE86FA21D692FA74" box="[350,451,1444,1468]" italics="true" pageId="4" pageNumber="1926">apiculata</emphasis>
</taxonomicName>
-group, the
<taxonomicName id="4C6F4D776F11D377FD8AFA21D5CEFA74" authorityName="Berland" authorityYear="1938" box="[594,671,1444,1468]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="1926" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="B91BEAE66F11D377FD8AFA21D5CEFA74" box="[594,671,1444,1468]" italics="true" pageId="4" pageNumber="1926">hystrix</emphasis>
</taxonomicName>
-group and the
<taxonomicName id="4C6F4D776F11D377FC84FA21D355FA74" authorityName="Barrion and Litsinger" authorityYear="1995" box="[860,1028,1444,1468]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="1926" phylum="Arthropoda" rank="species" species="pahilistapyasea">
<emphasis id="B91BEAE66F11D377FC84FA21D355FA74" box="[860,1028,1444,1468]" italics="true" pageId="4" pageNumber="1926">pahilistapyasea</emphasis>
</taxonomicName>
-group possess a simple, U-shaped sperm duct without additional loop. However, the conformation of the palp is uniform in males of the
<taxonomicName id="4C6F4D776F11D377FD2BFA61D409FA34" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[755,856,1508,1532]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="1926" phylum="Arthropoda" rank="species" species="apiculata">
<emphasis id="B91BEAE66F11D377FD2BFA61D409FA34" box="[755,856,1508,1532]" italics="true" pageId="4" pageNumber="1926">apiculata</emphasis>
</taxonomicName>
-group. They can be differentiated from males of all other species-groups by the embolus emerging from a distinct conical base (compare
<figureCitation id="13542A716F11D377FDDEF9A6D5CEF9F3" box="[518,671,1571,1595]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="4" pageNumber="1926">Figure 1A–D</figureCitation>
with 1E), which occupies most of the apico-prolateral extent of the palpal tegulum – embolus with broad base, gradually emerges from the tegulum, and tapering towards apex in
<taxonomicName id="4C6F4D776F11D377FCF1F9E6D427F9B3" authorityName="Berland" authorityYear="1938" box="[809,886,1635,1659]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="1926" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="B91BEAE66F11D377FCF1F9E6D427F9B3" box="[809,886,1635,1659]" italics="true" pageId="4" pageNumber="1926">hystrix</emphasis>
</taxonomicName>
-group (
<figureCitation id="13542A716F11D377FC0CF9E6D316F9B3" box="[980,1095,1635,1659]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="4" pageNumber="1926">Figure 1E</figureCitation>
) and
<taxonomicName id="4C6F4D776F11D377FF1CF906D63DF953" authorityName="Barrion and Litsinger" authorityYear="1995" box="[196,364,1667,1691]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="1926" phylum="Arthropoda" rank="species" species="pahilistapyasea">
<emphasis id="B91BEAE66F11D377FF1CF906D63DF953" box="[196,364,1667,1691]" italics="true" pageId="4" pageNumber="1926">pahilistapyasea</emphasis>
</taxonomicName>
-group (see also
<bibRefCitation id="EFFE4B056F11D377FDF5F906D4F5F953" author="Barrion AT &amp; Litsinger JA" box="[557,932,1667,1691]" pageId="4" pageNumber="1926" refId="ref5489" refString="Barrion AT, Litsinger JA. 1995. Riceland Spiders of South and Southeast Asia. Wallingford: CAB International." type="book" year="1995">Barrion and Litsinger 1995: 145</bibRefCitation>
, fig. 79d). The apex of the embolus points apically instead of moving clockwise along the margin of the tegulum base (compare
<figureCitation id="13542A716F10D376FDCAFF4DD5FAFF28" box="[530,683,200,224]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="5" pageNumber="1927">Figure 1A–D</figureCitation>
with 1E). The conductor represented by a transparent lamina, and is located next to the embolic base on apico-retrolateral portion of the tegulum (
<figureCitation id="13542A716F10D376FDDDFE8DD52EFEE8" box="[517,639,264,288]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="5" pageNumber="1927">Figure 3E</figureCitation>
) (conductor absent in
<taxonomicName id="4C6F4D776F10D376FC78FE8DD4BCFEE8" authorityName="Berland" authorityYear="1938" box="[928,1005,264,288]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="1927" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="B91BEAE66F10D376FC78FE8DD4BCFEE8" box="[928,1005,264,288]" italics="true" pageId="5" pageNumber="1927">hystrix</emphasis>
</taxonomicName>
-group as per
<bibRefCitation id="EFFE4B056F10D376FF0CFEADD6A2FE88" author="Deeleman-Reinhold CL" box="[212,499,296,320]" pageId="5" pageNumber="1927" refId="ref5538" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanteriidae). Leiden: Brill." type="book" year="2001">Deeleman-Reinhold 2001</bibRefCitation>
). The retrolateral apophysis on the palpal tibia is distinctly enlarged and can vary in shape from broad, heavily sclerotized ridge (
<figureCitation id="13542A716F10D376FC2EFECDD3DAFEA8" box="[1014,1163,328,352]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="5" pageNumber="1927">Figure 1A–C</figureCitation>
) to long, thinnish and point (
<figureCitation id="13542A716F10D376FDC7FEE2D5C6FEB7" box="[543,663,359,383]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="5" pageNumber="1927">Figure 1D</figureCitation>
). The ventral tibial apophysis is well developed; it is variable in shape from being an elevated ridge (
<figureCitation id="13542A716F10D376FCA3FE02D4A3FE57" box="[891,1010,391,415]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="5" pageNumber="1927">Figure 1D</figureCitation>
) to a massive column (
<figureCitation id="13542A716F10D376FEE1FE22D696FE77" box="[313,455,423,447]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="5" pageNumber="1927">Figure 1A,F</figureCitation>
). The legs are not lengthened and devoid of a large spine (legs distinctly elongate and provided with a very large apical spine on tibia II in the males of
<taxonomicName id="4C6F4D776F13D375FF77FF4DD606FF28" authorityName="Barrion and Litsinger" authorityYear="1995" box="[175,343,200,224]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="pahilistapyasea">
<emphasis id="B91BEAE66F13D375FF77FF4DD606FF28" box="[175,343,200,224]" italics="true" pageId="6" pageNumber="1928">pahilistapyasea</emphasis>
</taxonomicName>
-group).
</paragraph>
<caption id="DF10667C6F10D376FF0CF9AFD450F950" ID-DOI="http://doi.org/10.5281/zenodo.4631435" ID-Zenodo-Dep="4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="5" pageNumber="1927" startId="5.[212,279,1578,1600]" targetBox="[212,1172,511,1542]" targetPageId="5">
<paragraph id="8BD036F46F10D376FF0CF9AFD450F950" blockId="5.[212,1172,1578,1688]" pageId="5" pageNumber="1927">
Figure 3.
<taxonomicName id="4C6F4D776F10D376FEE3F9AED6A7F988" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[315,502,1579,1600]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="1927" phylum="Arthropoda" rank="species" species="apiculata" status="sp. nov.">
<emphasis id="B91BEAE66F10D376FEE3F9AED6A7F988" box="[315,502,1579,1600]" italics="true" pageId="5" pageNumber="1927">Clubiona apiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F10D376FE24F9AFD51FF988" box="[508,590,1578,1600]" pageId="5" pageNumber="1927" rank="species">sp. nov.</taxonomicNameLabel>
; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BD036F46F13D375FF62FF6DD69BFE17" blockId="6.[144,1104,168,479]" pageId="6" pageNumber="1928">
The conformation of the female genitalia is uniform and closely resembles that of the
<taxonomicName id="4C6F4D776F13D375FF62FE8DD656FEE8" authorityName="Berland" authorityYear="1938" box="[186,263,264,288]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="B91BEAE66F13D375FF62FE8DD656FEE8" box="[186,263,264,288]" italics="true" pageId="6" pageNumber="1928">hystrix</emphasis>
</taxonomicName>
-group in many aspects. Females of both species groups possess sclerotized spermathecae situated anteriorly to hyaline bursae (
<figureCitation id="13542A716F13D375FD23FEADD4C3FE88" box="[763,914,296,320]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="6" pageNumber="1928">Figure 2A,D</figureCitation>
); their epigynal region is marked with posteriorly situated copulatory orifices. However, the concomitantly copulatory ducts are elongate; these ducts are simple and lack convolution (
<figureCitation id="13542A716F13D375FF40FE02D678FE57" box="[152,297,391,415]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="6" pageNumber="1928">Figure 2A,D</figureCitation>
) in the
<taxonomicName id="4C6F4D776F13D375FE58FE0DD6B4FE68" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[384,485,392,416]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="apiculata">
<emphasis id="B91BEAE66F13D375FE58FE0DD6B4FE68" box="[384,485,392,416]" italics="true" pageId="6" pageNumber="1928">apiculata</emphasis>
</taxonomicName>
-group (insemination ducts usually with convolution in
<taxonomicName id="4C6F4D776F13D375FF48FE2DD78CFE08" authorityName="Berland" authorityYear="1938" box="[144,221,424,448]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="B91BEAE66F13D375FF48FE2DD78CFE08" box="[144,221,424,448]" italics="true" pageId="6" pageNumber="1928">hystrix</emphasis>
</taxonomicName>
-group). All known females also possess an additional, glandular head on each spermatheca (
<figureCitation id="13542A716F13D375FEF6FE42D6EBFE17" box="[302,442,455,479]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="6" pageNumber="1928">Figure 2C,E</figureCitation>
).
</paragraph>
<paragraph id="8BD036F46F13D375FF48FDA6D610FDF3" blockId="6.[144,1104,547,643]" box="[144,321,547,571]" pageId="6" pageNumber="1928">
<heading id="D09881986F13D375FF48FDA6D610FDF3" box="[144,321,547,571]" fontSize="10" level="3" pageId="6" pageNumber="1928" reason="5">
<emphasis id="B91BEAE66F13D375FF48FDA6D610FDF3" box="[144,321,547,571]" italics="true" pageId="6" pageNumber="1928">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F13D375FF48FDCED542FD4B" blockId="6.[144,1104,547,643]" pageId="6" pageNumber="1928">
<taxonomicName id="4C6F4D776F13D375FF48FDCED635FDAB" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[144,356,587,611]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="apiculata" status="sp. nov.">
<emphasis id="B91BEAE66F13D375FF48FDCED635FDAB" box="[144,356,587,611]" italics="true" pageId="6" pageNumber="1928">Clubiona apiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F13D375FEB7FDCED69FFDAB" box="[367,462,587,611]" pageId="6" pageNumber="1928" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C6F4D776F13D375FE06FDCED5B7FDAB" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[478,742,587,611]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="cylindriformis" status="sp. nov.">
<emphasis id="B91BEAE66F13D375FE06FDCED5B7FDAB" box="[478,742,587,611]" italics="true" pageId="6" pageNumber="1928">Clubiona cylindriformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F13D375FD29FDCED401FDAB" box="[753,848,587,611]" pageId="6" pageNumber="1928" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C6F4D776F13D375FCB8FDCED374FDAB" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[864,1061,587,611]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="cultrata" status="sp. nov.">
<emphasis id="B91BEAE66F13D375FCB8FDCED374FDAB" box="[864,1061,587,611]" italics="true" pageId="6" pageNumber="1928">Clubiona cultrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F13D375FBE8FDCED790FD4B" pageId="6" pageNumber="1928" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C6F4D776F13D375FF25FDEED6FEFD4B" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[253,431,619,643]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="conica" status="sp. nov.">
<emphasis id="B91BEAE66F13D375FF25FDEED6FEFD4B" box="[253,431,619,643]" italics="true" pageId="6" pageNumber="1928">Clubiona conica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F13D375FE61FDEED542FD4B" box="[441,531,619,643]" pageId="6" pageNumber="1928" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C375657F6F13D375FF48FD42D690FCCE" box="[144,449,711,774]" pageId="6" pageNumber="1928" type="distribution">
<paragraph id="8BD036F46F13D375FF48FD42D645FD17" blockId="6.[144,449,711,774]" box="[144,276,711,735]" pageId="6" pageNumber="1928">
<heading id="D09881986F13D375FF48FD42D645FD17" box="[144,276,711,735]" fontSize="10" level="3" pageId="6" pageNumber="1928" reason="5">
<emphasis id="B91BEAE66F13D375FF48FD42D645FD17" box="[144,276,711,735]" italics="true" pageId="6" pageNumber="1928">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F13D375FF48FD6BD690FCCE" blockId="6.[144,449,711,774]" box="[144,449,750,774]" pageId="6" pageNumber="1928">Known only from Borneo.</paragraph>
</subSubSection>
<subSubSection id="C375657F6F13D375FF48FCCFD6A4FAA0" pageId="6" pageNumber="1928" type="discussion">
<paragraph id="8BD036F46F13D375FF48FCCFD7A3FCAA" blockId="6.[144,1104,842,1384]" box="[144,242,842,866]" pageId="6" pageNumber="1928">
<heading id="D09881986F13D375FF48FCCFD7A3FCAA" box="[144,242,842,866]" fontSize="10" level="3" pageId="6" pageNumber="1928" reason="5">
<emphasis id="B91BEAE66F13D375FF48FCCFD7A3FCAA" box="[144,242,842,866]" italics="true" pageId="6" pageNumber="1928">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F13D375FF48FCF7D536FB81" blockId="6.[144,1104,842,1384]" pageId="6" pageNumber="1928">
<materialsCitation id="3B073CA96F13D375FF48FCF7D533FB81" ID-GBIF-Occurrence="3117074306" collectingDate="2000-12-30" collectingDateMax="2001-01-01" collectingDateMin="2000-12-30" collectorName="A. Riedel" country="Guinea-Bissau" county="Gunnung Meja" elevation="200" location="No" municipality="Manokwari" pageId="6" pageNumber="1928" specimenCount="2">
<location id="8EB0602F6F13D375FF48FCF7D7E5FC42" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0:8EB0602F6F13D375FF48FCF7D7E5FC42" box="[144,180,882,906]" country="Guinea-Bissau" county="Gunnung Meja" municipality="Manokwari" name="No" pageId="6" pageNumber="1928">No</location>
<typeStatus id="54D488566F13D375FF66FCF7D7BFFC42" box="[190,238,882,906]" pageId="6" pageNumber="1928">type</typeStatus>
specimens of
<taxonomicName id="4C6F4D776F13D375FE4DFCF7D5A8FC42" authority="Berland, 1938" authorityName="Berland" authorityYear="1938" box="[405,761,882,906]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="B91BEAE66F13D375FE4DFCF7D501FC42" box="[405,592,882,906]" italics="true" pageId="6" pageNumber="1928">Clubiona hystrix</emphasis>
Berland, 1938
</taxonomicName>
are known to exist.
<location id="8EB0602F6F13D375FC35FCF7D348FC42" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0:8EB0602F6F13D375FC35FCF7D348FC42" box="[1005,1049,882,906]" country="Guinea-Bissau" county="Gunnung Meja" municipality="Manokwari" name="The" pageId="6" pageNumber="1928">The</location>
first author could not locate them in the collections of RMHN.
<location id="8EB0602F6F13D375FCE6FC17D43BFC62" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0:8EB0602F6F13D375FCE6FC17D43BFC62" box="[830,874,914,938]" country="Guinea-Bissau" county="Gunnung Meja" municipality="Manokwari" name="The" pageId="6" pageNumber="1928">The</location>
original description and illustrations of this species correspond well with those provided by
<location id="8EB0602F6F13D375FC0DFC37D7A9FC22" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0:8EB0602F6F13D375FC0DFC37D7A9FC22" country="Guinea-Bissau" county="Gunnung Meja" municipality="Manokwari" name="Reinhold" pageId="6" pageNumber="1928">Deeleman-Reinhold</location>
(2001: 104, fig. 14–16, male from
<location id="8EB0602F6F13D375FD57FC57D5ECFC22" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0:8EB0602F6F13D375FD57FC57D5ECFC22" box="[655,701,978,1002]" country="Guinea-Bissau" county="Gunnung Meja" municipality="Manokwari" name="Bali" pageId="6" pageNumber="1928">Bali</location>
) and the specimen examined here, which was collected near the
<typeStatus id="54D488566F13D375FE2DFC74D574FBC1" box="[501,549,1009,1033]" pageId="6" pageNumber="1928">type</typeStatus>
locality in
<location id="8EB0602F6F13D375FD6EFC74D5BAFBC1" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0:8EB0602F6F13D375FD6EFC74D5BAFBC1" box="[694,747,1009,1033]" country="Guinea-Bissau" county="Gunnung Meja" municipality="Manokwari" name="New" pageId="6" pageNumber="1928">New</location>
<collectingCountry id="F37876646F13D375FD20FC74D41AFBC1" box="[760,843,1009,1033]" name="Guinea-Bissau" pageId="6" pageNumber="1928">Guinea</collectingCountry>
(MHNG,
<location id="8EB0602F6F13D375FC09FC74D31AFBC1" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0:8EB0602F6F13D375FC09FC74D31AFBC1" box="[977,1099,1009,1033]" country="Guinea-Bissau" county="Gunnung Meja" municipality="Manokwari" name="Irian Jaya" pageId="6" pageNumber="1928">Irian Jaya</location>
,
<location id="8EB0602F6F13D375FF48FB94D6D6FBE1" LSID="urn:lsid:plazi:treatment:03C687E26F17D375FE48F9FED6A4FAA0:8EB0602F6F13D375FF48FB94D6D6FBE1" box="[144,391,1041,1065]" country="Guinea-Bissau" county="Gunnung Meja" municipality="Manokwari" name="Manokwari Province" pageId="6" pageNumber="1928">Manokwari Province</location>
,
<collectingMunicipality id="6BB4AC8E6F13D375FE40FB94D54EFBE1" box="[408,543,1041,1065]" pageId="6" pageNumber="1928">Manokwari</collectingMunicipality>
,
<collectingCounty id="62B14E786F13D375FDE9FB94D5B6FBE1" box="[561,743,1041,1065]" pageId="6" pageNumber="1928">Gunnung Meja</collectingCounty>
,
<quantity id="4C979B116F13D375FD21FB94D416FBE1" box="[761,839,1041,1065]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="1928" unit="m" value="200.0">
<elevation id="0042D1C76F13D375FD21FB94D416FBE1" box="[761,839,1041,1065]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="1928" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="FFD110346F13D375FC80FB94D364FBE1" box="[856,1077,1041,1065]" pageId="6" pageNumber="1928" value="2000-12-30">
<collectingDate id="EF95E9DC6F13D375FC80FB94D364FBE1" box="[856,1077,1041,1065]" pageId="6" pageNumber="1928" value="2000-12-30">30 December 2000</collectingDate>
</date>
–
<date id="FFD110346F13D375FB9AFB94D67AFB81" pageId="6" pageNumber="1928" value="2001-01-01">
<collectingDate id="EF95E9DC6F13D375FB9AFB94D67AFB81" pageId="6" pageNumber="1928" value="2001-01-01">1 January 2001</collectingDate>
</date>
,
<collectorName id="269A53226F13D375FEE0FBB4D6F4FB81" box="[312,421,1073,1097]" pageId="6" pageNumber="1928">A. Riedel</collectorName>
leg.,
<figureCitation id="13542A716F13D375FE3DFBB4D506FB81" box="[485,599,1073,1097]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="6" pageNumber="1928">Figure 1E</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD036F46F13D375FF62FBD4D6A4FAA0" blockId="6.[144,1104,842,1384]" pageId="6" pageNumber="1928">
The
<taxonomicName id="4C6F4D776F13D375FF23FBD4D619FBA1" authorityName="Berland" authorityYear="1938" box="[251,328,1105,1129]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="hystrix">
<emphasis id="B91BEAE66F13D375FF23FBD4D619FBA1" box="[251,328,1105,1129]" italics="true" pageId="6" pageNumber="1928">hystrix</emphasis>
</taxonomicName>
-group is one of several species-groups in the spider genus
<taxonomicName id="4C6F4D776F13D375FF48FBF4D7A5FB41" authorityName="Latreille" authorityYear="1804" box="[144,244,1137,1161]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAE66F13D375FF48FBF4D7A5FB41" box="[144,244,1137,1161]" italics="true" pageId="6" pageNumber="1928">Clubiona</emphasis>
</taxonomicName>
– for a full listing of species and synonymies see
<bibRefCitation id="EFFE4B056F13D375FCB5FBF4D655FB61" author="Deeleman-Reinhold CL" pageId="6" pageNumber="1928" refId="ref5538" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanteriidae). Leiden: Brill." type="book" year="2001">Deeleman-Reinhold (2001: 92)</bibRefCitation>
. The males of this group have the following characteristics: (1) the palpal tegulum is marked with U-shaped sperm duct without forming additional loop; (2) the conductor is absent (but clearly present in a specimen treated here:
<figureCitation id="13542A716F13D375FC72FB54D34BFB21" box="[938,1050,1233,1257]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="6" pageNumber="1928">Figure 1E</figureCitation>
); (3) the embolus is situated on the distal margin of tegulum, with its apex pointed retrolaterally. The females can be recognized by: (1) the copulatory orifices located on posterior half of the epigynal region, hidden by fold or hood; (2) the spermathecae and hyaline bursae are distinct.
</paragraph>
</subSubSection>
</treatment>
</document>
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