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<mods:title id="2F7C1B333973F38213BB16F869D71EE2">Taxonomy, systematics and biology of the Australian halotolerant wolf spider genus Tetralycosa (Araneae: Lycosidae: Artoriinae)</mods:title>
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<mods:affiliation id="37C9F4A88C7DAAF945EA6B788BB5D01E">South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia. &amp; Email: Peter. Hudson @ samuseum. sa. gov. au</mods:affiliation>
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<paragraph id="8BD036DEFFEA183295B99831B4E6C2ED" blockId="14.[640,948,1401,1428]" box="[640,948,1401,1428]" pageId="14" pageNumber="15">
<heading id="D09881B2FFEA183295B99831B4E6C2ED" box="[640,948,1401,1428]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="4C6F4D5DFFEA183295B99831B4E6C2ED" ID-CoL="63V4M" ID-ENA="332816" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[640,948,1401,1428]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEA183295B99831B45BC2ED" bold="true" box="[640,777,1401,1428]" italics="true" pageId="14" pageNumber="15">Tetralycosa</emphasis>
<bibRefCitation id="EFFE4B2FFFEA183294299832B4E6C2ED" author="Roewer" box="[784,948,1402,1428]" firstAuthor="Roewer" pageId="14" pageNumber="15" pagination="519 - 1040" refId="ref39690" refString="Roewer C. F. 1960. Araneae Lycosaeformia II. (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba - Mission G. F. De Witte 55: 519 - 1040." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
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<treatmentCitation id="0ACE10CFFFEA1832978498F3B565C2AF" author="Roewer C. F." box="[189,567,1467,1494]" page="949" pageId="14" pageNumber="15" year="1960">
<taxonomicName id="4C6F4D5DFFEA1832978498F3B565C2AF" ID-CoL="63V4M" authority="Roewer, 1960: 949" authorityName="Roewer" authorityPageNumber="949" authorityYear="1960" box="[189,567,1467,1494]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEA1832978498F3B617C2AC" box="[189,325,1467,1493]" italics="true" pageId="14" pageNumber="15">Tetralycosa</emphasis>
<bibRefCitation id="EFFE4B2FFFEA1832967698F4B565C2AF" author="Roewer" box="[335,567,1467,1494]" firstAuthor="Roewer" pageId="14" pageNumber="15" pagination="519 - 1040" refId="ref39690" refString="Roewer C. F. 1960. Araneae Lycosaeformia II. (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba - Mission G. F. De Witte 55: 519 - 1040." type="journal article" year="1960">Roewer, 1960: 949</bibRefCitation>
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,
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<typeStatus id="54D4887CFFEA1832957F98F4B52BC2AF" box="[582,633,1468,1494]" pageId="14" pageNumber="15">type</typeStatus>
species
<taxonomicName id="4C6F4D5DFFEA183295DB98F4B301C2AF" authority="Simon, 1909" authorityName="Simon" authorityYear="1909" box="[738,1107,1467,1494]" class="Arachnida" family="Lycosidae" genus="Lycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="meracula">
<emphasis id="B91BEACCFFEA183295DB98F4B4FFC2AC" box="[738,941,1467,1493]" italics="true" pageId="14" pageNumber="15">Lycosa meracula</emphasis>
<bibRefCitation id="EFFE4B2FFFEA1832948E98F3B301C2AF" author="Simon" box="[951,1107,1467,1494]" firstAuthor="Simon" pageId="14" pageNumber="15" pagination="155 - 212" refId="ref39922" refString="Simon E. 1909. Araneae, 2 me partie. In: Michaelsen W. &amp; Hartmeyer R. (eds) Die Fauna Sudwest- Australiens. Ergebnisse der Hamburger sudwest-australischen Forschungsreise 1905: 155 - 212. Gustav Fischer, Jena." type="journal article" year="1909">Simon, 1909</bibRefCitation>
</taxonomicName>
, by original designation (considered a junior synonym of
<taxonomicName id="4C6F4D5DFFEA183295699897B4A8C283" ID-CoL="55P4J" authority="(L. Koch, 1876)" baseAuthorityName="L. Koch" baseAuthorityYear="1876" box="[592,1018,1503,1530]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B91BEACCFFEA183295699897B47FC280" box="[592,813,1503,1529]" italics="true" pageId="14" pageNumber="15">Tetralycosa oraria</emphasis>
(L.
<bibRefCitation id="EFFE4B2FFFEA1832945C9897B4A0C283" author="Koch" box="[869,1010,1503,1530]" firstAuthor="Koch" pageId="14" pageNumber="15" refId="ref38160" refString="Koch L. 1876. Die Arachniden Australiens nach der Natur beschrieben und abgebildet. 1. Theil. 6. Lieferung. Verlag von Bauer und Raspe, Nurnberg." type="book" year="1876">Koch, 1876</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="EFFE4B2FFFEA1832931598A8B27DC283" author="Framenau" box="[1068,1327,1503,1530]" etAl="et al." firstAuthor="Framenau" pageId="14" pageNumber="15" pagination="185 - 202" refId="ref37149" refString="Framenau V. W. &amp; Yoo J. - S. 2006. Systematics of the new Australian wolf spider genus Tuberculosa (Araneae, Lycosidae). Invertebrate Systematics 20: 185 - 202. https: // doi. org / 10.1071 / is 05036" type="journal article" year="2006">
Framenau
<emphasis id="B91BEACCFFEA1832939598A8B3BBC280" box="[1196,1257,1503,1529]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
<date id="FFD1101EFFEA183293CA9897B27DC283" box="[1267,1327,1503,1530]" pageId="14" pageNumber="15" value="2006">2006</date>
</bibRefCitation>
). The gender is feminine.
</paragraph>
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<subSubSection id="C3756555FFEA183397849B0CB4BBC69A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BD036DEFFEA183297849B0CB664C127" blockId="14.[189,310,1604,1630]" box="[189,310,1604,1630]" pageId="14" pageNumber="15">
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<emphasis id="B91BEACCFFEA183297849B0CB664C127" bold="true" box="[189,310,1604,1630]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
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Males of
<taxonomicName id="4C6F4D5DFFEA183296169B3BB6E5C1F4" authorityName="Roewer" authorityYear="1960" box="[303,439,1651,1677]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEA183296169B3BB6E5C1F4" box="[303,439,1651,1677]" italics="true" pageId="14" pageNumber="15">Tetralycosa</emphasis>
</taxonomicName>
differ from other lycosid genera within the
<taxonomicName id="4C6F4D5DFFEA183294FB9B3BB369C1F4" authorityName="Framenau" authorityYear="2007" box="[962,1083,1651,1677]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subFamily" subFamily="Artoriinae">Artoriinae</taxonomicName>
by the combination of the following characters: tegular apophysis originating apically on tegulum and basally curved, opposing an apico-medially directed pointy (lobed in
<taxonomicName id="4C6F4D5DFFEA183295FC9BF3B401C1AD" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[709,851,1722,1748]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="halophila" status="sp. nov.">
<emphasis id="B91BEACCFFEA183295FC9BF3B401C1AD" box="[709,851,1722,1748]" italics="true" pageId="14" pageNumber="15">T. halophila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFEA183294629BF3B4EAC1AC" box="[859,952,1723,1749]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
) protrusion on the retrolateral section of the tegulum; basal lobe of tegular apophysis present; terminal apophysis forms a shaft for the resting embolus; female epigyne variable: with a wide median septum that displays an inverted T-shaped dark central pattern, or a round to oval opening with anterior notch.
</paragraph>
<paragraph id="8BD036DEFFEA183397849A2EB4BBC69A" blockId="14.[189,1399,1894,2027]" lastBlockId="15.[189,1399,267,483]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Putative sister groups of
<taxonomicName id="4C6F4D5DFFEA183296CB9A2EB528C0F9" authorityName="Roewer" authorityYear="1960" box="[498,634,1894,1920]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEA183296CB9A2EB528C0F9" box="[498,634,1894,1920]" italics="true" pageId="14" pageNumber="15">Tetralycosa</emphasis>
</taxonomicName>
are
<taxonomicName id="4C6F4D5DFFEA1832958C9A2FB357C0F9" authority="Framenau, 2010" authorityName="Framenau" authorityYear="2010" box="[693,1029,1894,1921]" class="Arachnida" family="Lycosidae" genus="Kangarosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEA1832958C9A2FB464C0F9" box="[693,822,1895,1920]" italics="true" pageId="14" pageNumber="15">Kangarosa</emphasis>
<bibRefCitation id="EFFE4B2FFFEA1832947B9A2FB357C0F9" author="Framenau" box="[834,1029,1894,1921]" firstAuthor="Framenau" pageId="14" pageNumber="15" pagination="113 - 142" refId="ref36964" refString="Framenau V. W. 2010. Revision of the new Australian wolf spider genus Kangarosa (Araneae: Lycosidae: Artoriinae). Arthropod Systematics and Phylogeny 68: 113 - 142." type="journal article" year="2010">Framenau, 2010</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6F4D5DFFEA1832937F9A2EB220C0F9" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[1094,1394,1894,1921]" class="Arachnida" family="Lycosidae" genus="Diahogna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEA1832937F9A2EB3E9C0F9" box="[1094,1211,1894,1920]" italics="true" pageId="14" pageNumber="15">Diahogna</emphasis>
<bibRefCitation id="EFFE4B2FFFEA183293FE9A2FB220C0F9" author="Roewer" box="[1223,1394,1894,1921]" firstAuthor="Roewer" pageId="14" pageNumber="15" pagination="519 - 1040" refId="ref39690" refString="Roewer C. F. 1960. Araneae Lycosaeformia II. (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba - Mission G. F. De Witte 55: 519 - 1040." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
</taxonomicName>
. The genital morphology within these two genera is very similar to
<taxonomicName id="4C6F4D5DFFEA183294D19AC2B322C0DD" authorityName="Roewer" authorityYear="1960" box="[1000,1136,1930,1956]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEA183294D19AC2B322C0DD" box="[1000,1136,1930,1956]" italics="true" pageId="14" pageNumber="15">Tetralycosa</emphasis>
</taxonomicName>
; however, the tegular apophysis of the male pedipalp is not curved, but is more or less straight. The female epigyne in both genera forms a small open atrium in comparison with
<taxonomicName id="4C6F4D5DFFEA183294759A99B486C092" authorityName="Roewer" authorityYear="1960" box="[844,980,2001,2027]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEA183294759A99B486C092" box="[844,980,2001,2027]" italics="true" pageId="14" pageNumber="15">Tetralycosa</emphasis>
</taxonomicName>
with its distinct median septum or large round to oval opening. Somatic characters shared by the
<taxonomicName id="4C6F4D5DFFEB183394BE9C44B35AC65C" authorityName="Framenau" authorityYear="2010" box="[903,1032,268,293]" class="Arachnida" family="Lycosidae" genus="Kangarosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183394BE9C44B35AC65C" box="[903,1032,268,293]" italics="true" pageId="15" pageNumber="16">Kangarosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6F4D5DFFEB1833932C9C43B3D8C65C" authorityName="Roewer" authorityYear="1960" box="[1045,1162,267,293]" class="Arachnida" family="Lycosidae" genus="Diahogna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB1833932C9C43B3D8C65C" box="[1045,1162,267,293]" italics="true" pageId="15" pageNumber="16">Diahogna</emphasis>
</taxonomicName>
and some species of
<taxonomicName id="4C6F4D5DFFEB183397849C79B617C632" authorityName="Roewer" authorityYear="1960" box="[189,325,305,331]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183397849C79B617C632" box="[189,325,305,331]" italics="true" pageId="15" pageNumber="16">Tetralycosa</emphasis>
</taxonomicName>
include the arrangement of the eyes, in particular the length of the anterior row of eyes that is as wide or wider than the posterior median row of eyes. Our phylogeny (
<figureCitation id="13542A5BFFEB183393139C1FB3D4C608" box="[1066,1158,343,369]" captionStart="Fig" captionStartId="12.[189,232,1880,1906]" captionTargetBox="[187,1402,417,1819]" captionTargetId="figure@12.[160,1415,402,1838]" captionTargetPageId="12" captionText="Fig. 4. Phylogenetic hypothesis of Tetralycosa Roewer, 1960 based on 31 morphological and one ecological characters. A. Basal branches including outgroups. B. Terminal salt lake clade (L = 71, Ci = 61, Ri = 80). Bremer support values&gt; 1 are indicated by numbers in squares below the respective node." figureDoi="http://doi.org/10.5281/zenodo.3832431" httpUri="https://zenodo.org/record/3832431/files/figure.png" pageId="15" pageNumber="16">Fig. 4A</figureCitation>
) suggest
<taxonomicName id="4C6F4D5DFFEB183393CC9C10B224C608" authorityName="Framenau" authorityYear="2010" box="[1269,1398,344,369]" class="Arachnida" family="Lycosidae" genus="Kangarosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183393CC9C10B224C608" box="[1269,1398,344,369]" italics="true" pageId="15" pageNumber="16">Kangarosa</emphasis>
</taxonomicName>
to be the sistergroup of
<taxonomicName id="4C6F4D5DFFEB183396F59C35B506C6EE" authorityName="Roewer" authorityYear="1960" box="[460,596,381,407]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183396F59C35B506C6EE" box="[460,596,381,407]" italics="true" pageId="15" pageNumber="16">Tetralycosa</emphasis>
</taxonomicName>
based on the absence of a median band on the carapace (character 4). However, only a single species of
<taxonomicName id="4C6F4D5DFFEB183395509CECB5B8C6C4" authorityName="Framenau" authorityYear="2010" box="[617,746,420,445]" class="Arachnida" family="Lycosidae" genus="Kangarosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183395509CECB5B8C6C4" box="[617,746,420,445]" italics="true" pageId="15" pageNumber="16">Kangarosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6F4D5DFFEB183394159CEBB4F3C6C4" authorityName="Roewer" authorityYear="1960" box="[812,929,419,445]" class="Arachnida" family="Lycosidae" genus="Diahogna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183394159CEBB4F3C6C4" box="[812,929,419,445]" italics="true" pageId="15" pageNumber="16">Diahogna</emphasis>
</taxonomicName>
were included in our analyses, which focused on establishing intrageneric relationships within
<taxonomicName id="4C6F4D5DFFEB183394629C81B4B1C69A" authorityName="Roewer" authorityYear="1960" box="[859,995,457,483]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183394629C81B4B1C69A" box="[859,995,457,483]" italics="true" pageId="15" pageNumber="16">Tetralycosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756555FFEB183397849F44B420C2A4" pageId="15" pageNumber="16" type="description">
<paragraph id="8BD036DEFFEB183397849F44B61CC55F" blockId="15.[189,334,524,550]" box="[189,334,524,550]" pageId="15" pageNumber="16">
<heading id="D09881B2FFEB183397849F44B61CC55F" bold="true" box="[189,334,524,550]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B91BEACCFFEB183397849F44B61CC55F" bold="true" box="[189,334,524,550]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFEB183397849F76B233C3E9" blockId="15.[189,1399,574,1169]" pageId="15" pageNumber="16">
Small to large wolf spiders (TL
<quantity id="4C979B3BFFEB1833950A9F76B586C521" box="[563,724,574,601]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="2.25" metricValueMin="0.45" pageId="15" pageNumber="16" unit="mm" value="13.5" valueMax="22.5" valueMin="4.5">4.522.5 mm</quantity>
). Males smaller than females. Carapace longer than wide, dorsal profile straight in lateral view (
<taxonomicName id="4C6F4D5DFFEB183395449F2DB59AC507" baseAuthorityName="L. Koch" baseAuthorityYear="1876" box="[637,712,613,638]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B91BEACCFFEB183395449F2DB59AC507" box="[637,712,613,638]" italics="true" pageId="15" pageNumber="16">oraria</emphasis>
</taxonomicName>
-group) (
<figureCitation id="13542A5BFFEB183394099F2CB4DDC506" box="[816,911,612,639]" captionStart="Fig" captionStartId="21.[189,232,1893,1919]" captionTargetBox="[189,1386,412,1815]" captionTargetId="figure@21.[189,1398,394,1852]" captionTargetPageId="21" captionText="Fig. 6. Tetralycosa oraria (L. Koch, 1876), ♂ (WAM 71/360) and female (WAM 71/361). AB. Male carapace, lateral and frontal view. CD. Left male pedipalp, ventral and retrolateral view. E. Left male pedipalp, palea section of bulbus, ventral view. FG. Female epigyne, dorsal and ventral view. Scale bar: A = 4.42 mm; B = 1.88 mm; CD = 1.28 mm; E = 0.56 mm; FG = 0.89 mm." figureDoi="http://doi.org/10.5281/zenodo.3832435" httpUri="https://zenodo.org/record/3832435/files/figure.png" pageId="15" pageNumber="16">Fig. 6A</figureCitation>
), or with an elevated cephalic region and downward slope towards posterior end (
<taxonomicName id="4C6F4D5DFFEB183395A59FC2B5ABC5DD" baseAuthorityName="McKay" baseAuthorityYear="1976" box="[668,761,650,676]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alteripa">
<emphasis id="B91BEACCFFEB183395A59FC2B5ABC5DD" box="[668,761,650,676]" italics="true" pageId="15" pageNumber="16">alteripa</emphasis>
</taxonomicName>
- and
<taxonomicName id="4C6F4D5DFFEB183394069FC2B42BC5DA" baseAuthorityName="Hickman" baseAuthorityYear="1944" box="[831,889,650,675]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="eyrei">
<emphasis id="B91BEACCFFEB183394069FC2B42BC5DA" box="[831,889,650,675]" italics="true" pageId="15" pageNumber="16">eyrei</emphasis>
</taxonomicName>
-groups) (
<figureCitation id="13542A5BFFEB183394D79FC1B30FC5DD" box="[1006,1117,649,676]" captionStart="Fig" captionStartId="37.[189,232,1892,1918]" captionTargetBox="[218,1398,604,1850]" captionTargetId="figure@37.[189,1398,563,1850]" captionTargetPageId="37" captionText="Fig. 15. Tetralycosa alteripa (McKay, 1976), ♂ (WAM T47354) and ♀, paratype (WAM T47356). A. Male carapace, lateral view. BC. Left male pedipalp, ventral and retrolateral view. D. Left male pedipalp, palea section of bulbus, ventral view. EF. Female epigyne, ventral and dorsal view. Scale bar: A = 8.83 mm; BC = 2.56 mm; D = 1.47 mm; EF = 1.91 mm." figureDoi="http://doi.org/10.5281/zenodo.3832455" httpUri="https://zenodo.org/record/3832455/files/figure.png" pageId="15" pageNumber="16">Fig. 15A</figureCitation>
). Head flanks in frontal view a gentle slope (
<taxonomicName id="4C6F4D5DFFEB183396969FF8B6A8C5B0" baseAuthorityName="L. Koch" baseAuthorityYear="1876" box="[431,506,688,713]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B91BEACCFFEB183396969FF8B6A8C5B0" box="[431,506,688,713]" italics="true" pageId="15" pageNumber="16">oraria</emphasis>
</taxonomicName>
-group) or nearly vertical (
<taxonomicName id="4C6F4D5DFFEB1833940B9FE7B4DDC5B0" baseAuthorityName="McKay" baseAuthorityYear="1976" box="[818,911,687,713]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alteripa">
<emphasis id="B91BEACCFFEB1833940B9FE7B4DDC5B0" box="[818,911,687,713]" italics="true" pageId="15" pageNumber="16">alteripa</emphasis>
</taxonomicName>
- and
<taxonomicName id="4C6F4D5DFFEB183394E89FF8B359C5B0" baseAuthorityName="Hickman" baseAuthorityYear="1944" box="[977,1035,688,713]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="eyrei">
<emphasis id="B91BEACCFFEB183394E89FF8B359C5B0" box="[977,1035,688,713]" italics="true" pageId="15" pageNumber="16">eyrei</emphasis>
</taxonomicName>
-groups). Carapace colouration variable from uniform light yellowish-brown (
<taxonomicName id="4C6F4D5DFFEB183395E69F9EB43EC596" authorityName="Framenau, Gotch &amp; Austin" authorityYear="2006" box="[735,876,725,751]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="arabanae">
<emphasis id="B91BEACCFFEB183395E69F9EB43EC596" box="[735,876,725,751]" italics="true" pageId="15" pageNumber="16">T. arabanae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6F4D5DFFEB183394429F9EB4B7C596" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[891,997,725,751]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFEB183394429F9EB4B7C596" box="[891,997,725,751]" italics="true" pageId="15" pageNumber="16">T caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFEB183394D49F9EB319C589" box="[1005,1099,726,752]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
), brown with median and marginal bands (e.g.,
<taxonomicName id="4C6F4D5DFFEB183396879FB4B502C46C" authorityName="Framenau &amp; Hudson" authorityYear="2017" baseAuthorityName="McKay" baseAuthorityYear="1979" box="[446,592,763,789]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="wundurra" status="comb. nov.">
<emphasis id="B91BEACCFFEB183396879FB4B502C46C" box="[446,592,763,789]" italics="true" pageId="15" pageNumber="16">T. wundurra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFEB183395619FB3B58FC46F" box="[600,733,763,790]" pageId="15" pageNumber="16" rank="species">comb. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C6F4D5DFFEB183395D39FB3B415C46C" baseAuthorityName="McKay" baseAuthorityYear="1976" box="[746,839,763,789]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alteripa">
<emphasis id="B91BEACCFFEB183395D39FB3B415C46C" box="[746,839,763,789]" italics="true" pageId="15" pageNumber="16">alteripa</emphasis>
</taxonomicName>
-group) to uniformly dark brown (
<taxonomicName id="4C6F4D5DFFEB183393E79FB4B24AC46C" baseAuthorityName="Hickman" baseAuthorityYear="1944" box="[1246,1304,764,789]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="eyrei">
<emphasis id="B91BEACCFFEB183393E79FB4B24AC46C" box="[1246,1304,764,789]" italics="true" pageId="15" pageNumber="16">eyrei</emphasis>
</taxonomicName>
-group). Anterior median eyes larger than anterior lateral eyes (except in
<taxonomicName id="4C6F4D5DFFEB1833949B9E6AB348C442" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[930,1050,801,827]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="rebecca" status="sp. nov.">
<emphasis id="B91BEACCFFEB1833949B9E6AB348C442" box="[930,1050,801,827]" italics="true" pageId="15" pageNumber="16">T. rebecca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFEB183393269E6AB32EC445" box="[1055,1148,802,828]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
), row of anterior eyes wider or as wide (
<taxonomicName id="4C6F4D5DFFEB183396A09E00B6B6C418" baseAuthorityName="L. Koch" baseAuthorityYear="1876" box="[409,484,840,865]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B91BEACCFFEB183396A09E00B6B6C418" box="[409,484,840,865]" italics="true" pageId="15" pageNumber="16">oraria</emphasis>
</taxonomicName>
-group) or smaller (
<taxonomicName id="4C6F4D5DFFEB183395E99E0FB47FC418" baseAuthorityName="McKay" baseAuthorityYear="1976" box="[720,813,839,865]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alteripa">
<emphasis id="B91BEACCFFEB183395E99E0FB47FC418" box="[720,813,839,865]" italics="true" pageId="15" pageNumber="16">alteripa</emphasis>
</taxonomicName>
- and
<taxonomicName id="4C6F4D5DFFEB1833944D9E00B4FCC418" baseAuthorityName="Hickman" baseAuthorityYear="1944" box="[884,942,840,865]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="eyrei">
<emphasis id="B91BEACCFFEB1833944D9E00B4FCC418" box="[884,942,840,865]" italics="true" pageId="15" pageNumber="16">eyrei</emphasis>
</taxonomicName>
-groups) than row of posterior median eyes; row of anterior eyes straight, procurved or strongly procurved. Chelicerae generally with three promarginal and three retromarginal teeth, but two to four teeth on individual chelicerae possible on both margins. Labium generally as wide as or wider than long (except in
<taxonomicName id="4C6F4D5DFFEB183393749EF1B393C4AB" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[1101,1217,952,978]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFEB183393749EF1B393C4AB" box="[1101,1217,952,978]" italics="true" pageId="15" pageNumber="16">T. caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFEB183393F59EF1B27FC4AA" box="[1228,1325,953,979]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, male
<taxonomicName id="4C6F4D5DFFEB183397849E97B662C481" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[189,304,990,1016]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="adarca" status="sp. nov.">
<emphasis id="B91BEACCFFEB183397849E97B662C481" box="[189,304,990,1016]" italics="true" pageId="15" pageNumber="16">T. adarca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFEB183396029E97B6C8C480" box="[315,410,991,1017]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
and male and female
<taxonomicName id="4C6F4D5DFFEB183395899E97B413C481" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[688,833,990,1016]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="halophila" status="sp. nov.">
<emphasis id="B91BEACCFFEB183395899E97B413C481" box="[688,833,990,1016]" italics="true" pageId="15" pageNumber="16">T. halophila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFEB183394749E97B4E2C480" box="[845,944,991,1017]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
). Abdomen generally with olive-grey heart mark, auxiliary colouration variable. Leg formula variable between species and sexes of the same species, leg I or leg III shortest (except in
<taxonomicName id="4C6F4D5DFFEB1833958F9963B475C33D" authority="Framenau &amp; Hudson, 2017" authorityName="Framenau &amp; Hudson" authorityYear="2017" box="[694,807,1066,1092]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="caudex" status="sp. nov.">
<emphasis id="B91BEACCFFEB1833958F9963B475C33D" box="[694,807,1066,1092]" italics="true" pageId="15" pageNumber="16">T. caudex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857B7FFEB183394169962B4DCC33D" box="[815,910,1066,1092]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, leg II shortest). Femora I generally with three dorsal spines (rarely two or four), patella in males generally with prolateral and retrolateral spine (reduced in most females), tibiae of males generally with two dorsal spines (rarely none or only one).
</paragraph>
<paragraph id="8BD036DEFFEB1833978499F2B420C2A4" blockId="15.[189,1399,1210,1501]" pageId="15" pageNumber="16">
Tegulum of male pedipalp deeply divided. Tegular apophysis located apically at tegulum, basally curved and opposing an apico-medially directed pointy protrusion on the retrolateral section of the tegulum. Tegular apophysis with a basal lobe. Embolus originating prolaterally on and curving ventrally around palea, long and slim. Basoembolic apophysis an unsclerotised lobe. Terminal apophysis well developed and forming a sclerotised shaft for the resting embolus.
<taxonomicName id="4C6F4D5DFFEB183394719819B4B1C212" baseAuthorityName="Framenau" baseAuthorityYear="2007" box="[840,995,1361,1387]" class="Insecta" family="Volutidae" genus="Cymbium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tip">Cymbium tip</taxonomicName>
without or only a few macrosetae. Female epigyne variable with a wide median septum sometimes only partially visible behind the sclerotised margins of the epigyne, which only leave a round or oval opening. Small round or oval spermathecal heads. Spermathecal stalks short and twisted.
</paragraph>
</subSubSection>
<subSubSection id="C3756555FFEB182C97849B4FB39CC579" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BD036DEFFEB183397849B4FB67DC158" blockId="15.[189,303,1543,1569]" box="[189,303,1543,1569]" pageId="15" pageNumber="16">
<heading id="D09881B2FFEB183397849B4FB67DC158" bold="true" box="[189,303,1543,1569]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B91BEACCFFEB183397849B4FB67DC158" bold="true" box="[189,303,1543,1569]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036DEFFEB183397849B70B5A8C1BD" blockId="15.[189,1399,1592,1732]" pageId="15" pageNumber="16">
The
<typeStatus id="54D4887CFFEB183397C89B70B676C12B" box="[241,292,1592,1618]" pageId="15" pageNumber="16">type</typeStatus>
species of
<taxonomicName id="4C6F4D5DFFEB1833969A9B70B579C12B" authorityName="Roewer" authorityYear="1960" box="[419,555,1592,1618]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB1833969A9B70B579C12B" box="[419,555,1592,1618]" italics="true" pageId="15" pageNumber="16">Tetralycosa</emphasis>
</taxonomicName>
is
<taxonomicName id="4C6F4D5DFFEB183395729B71B4E2C12B" authority="Simon, 1909" authorityName="Simon" authorityYear="1909" box="[587,944,1592,1618]" class="Arachnida" family="Lycosidae" genus="Lycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="meracula">
<emphasis id="B91BEACCFFEB183395729B71B5CFC12B" box="[587,669,1593,1618]" italics="true" pageId="15" pageNumber="16">Lycosa</emphasis>
<emphasis id="B91BEACCFFEB1833959A9B70B440C12B" box="[675,786,1592,1618]" italics="true" pageId="15" pageNumber="16">meracula</emphasis>
<bibRefCitation id="EFFE4B2FFFEB183394219B70B4E2C12B" author="Simon" box="[792,944,1592,1618]" firstAuthor="Simon" pageId="15" pageNumber="16" pagination="155 - 212" refId="ref39922" refString="Simon E. 1909. Araneae, 2 me partie. In: Michaelsen W. &amp; Hartmeyer R. (eds) Die Fauna Sudwest- Australiens. Ergebnisse der Hamburger sudwest-australischen Forschungsreise 1905: 155 - 212. Gustav Fischer, Jena." type="journal article" year="1909">Simon, 1909</bibRefCitation>
</taxonomicName>
, which was recently synonymised with
<taxonomicName id="4C6F4D5DFFEB183397849B17B560C100" authority="(Framenau et al. 2006)" baseAuthorityName="Framenau" baseAuthorityYear="2006" box="[189,562,1630,1657]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B91BEACCFFEB183397849B17B781C101" box="[189,211,1631,1656]" italics="true" pageId="15" pageNumber="16">T.</emphasis>
<emphasis id="B91BEACCFFEB183397E19B17B671C101" box="[216,291,1631,1656]" italics="true" pageId="15" pageNumber="16">oraria</emphasis>
(
<bibRefCitation id="EFFE4B2FFFEB183396089B16B578C100" author="Framenau" box="[305,554,1630,1657]" etAl="et al." firstAuthor="Framenau" pageId="15" pageNumber="16" pagination="185 - 202" refId="ref37149" refString="Framenau V. W. &amp; Yoo J. - S. 2006. Systematics of the new Australian wolf spider genus Tuberculosa (Araneae, Lycosidae). Invertebrate Systematics 20: 185 - 202. https: // doi. org / 10.1071 / is 05036" type="journal article" year="2006">
Framenau
<emphasis id="B91BEACCFFEB183396969B17B6BAC101" box="[431,488,1630,1656]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2006
</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C6F4D5DFFEB183395799B17B455C101" authorityName="Simon" authorityYear="1909" box="[576,775,1630,1656]" class="Arachnida" family="Lycosidae" genus="Lycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="meracula">
<emphasis id="B91BEACCFFEB183395799B17B455C101" box="[576,775,1630,1656]" italics="true" pageId="15" pageNumber="16">Lycosa meracula</emphasis>
</taxonomicName>
as illustrated by (
<bibRefCitation id="EFFE4B2FFFEB183394EB9B16B329C101" author="McKay" box="[978,1147,1630,1656]" firstAuthor="McKay" pageId="15" pageNumber="16" pagination="241 - 275" refId="ref38615" refString="McKay R. J. 1979 a. The wolf spiders of Australia (Araneae: Lycosidae): 12. Descriptions of some Western Australian species. Memoirs of the Queensland Museum 19: 241 - 275." type="journal article" year="1979" yearSuffix="a">
McKay
<date id="FFD1101EFFEB1833930B9B16B33CC101" box="[1074,1134,1630,1656]" pageId="15" pageNumber="16" value="1979">1979</date>
a
</bibRefCitation>
) represented material of two different but closely related species of
<taxonomicName id="4C6F4D5DFFEB183395DE9BCCB3F0C1E7" authority="Framenau &amp; Leung, 2013" authorityName="Framenau &amp; Leung" authorityYear="2013" box="[743,1186,1668,1694]" class="Arachnida" family="Lycosidae" genus="Costacosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183395DE9BCCB433C1E7" box="[743,865,1668,1694]" italics="true" pageId="15" pageNumber="16">Costacosa</emphasis>
<bibRefCitation id="EFFE4B2FFFEB183394529BCCB3F0C1E7" author="Framenau &amp; Leung" box="[875,1186,1668,1694]" firstAuthor="Framenau" pageId="15" pageNumber="16" pagination="173 - 184" refId="ref37103" refString="Framenau V. W. &amp; Leung A. E. 2013. Costacosa, a new genus of wolf spider (Araneae, Lycosidae) from coastal north-west Western Australia. Records of the Western Australian Museum, Supplement 83: 173 - 184." type="journal article" year="2013">Framenau &amp; Leung, 2013</bibRefCitation>
</taxonomicName>
, a member of the subfamily
<taxonomicName id="4C6F4D5DFFEB183396039BE1B5A1C1BD" authority="(Framenau &amp; Leung 2013)" baseAuthorityName="Framenau &amp; Leung" baseAuthorityYear="2013" box="[314,755,1705,1732]" family="Null" pageId="15" pageNumber="16" rank="subFamily" subFamily="Lycosinae">
Lycosinae (
<bibRefCitation id="EFFE4B2FFFEB183396F99BE2B5B9C1BD" author="Framenau &amp; Leung" box="[448,747,1706,1732]" firstAuthor="Framenau" pageId="15" pageNumber="16" pagination="173 - 184" refId="ref37103" refString="Framenau V. W. &amp; Leung A. E. 2013. Costacosa, a new genus of wolf spider (Araneae, Lycosidae) from coastal north-west Western Australia. Records of the Western Australian Museum, Supplement 83: 173 - 184." type="journal article" year="2013">Framenau &amp; Leung 2013</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD036DEFFEB183397849BA6B57AC092" blockId="15.[189,1399,1773,2027]" pageId="15" pageNumber="16">
<taxonomicName id="4C6F4D5DFFEB183397849BA6B617C071" authorityName="Roewer" authorityYear="1960" box="[189,325,1774,1800]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183397849BA6B617C071" box="[189,325,1774,1800]" italics="true" pageId="15" pageNumber="16">Tetralycosa</emphasis>
</taxonomicName>
belongs to the subfamily
<taxonomicName id="4C6F4D5DFFEB183395549BA5B4F8C071" authority="Framenau, 2007" authorityName="Framenau" authorityYear="2007" box="[621,938,1773,1800]" class="Arachnida" family="Lycosidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subFamily" subFamily="Artoriinae">
Artoriinae
<bibRefCitation id="EFFE4B2FFFEB183395D59BA6B4F8C071" author="Framenau" box="[748,938,1774,1800]" firstAuthor="Framenau" pageId="15" pageNumber="16" pagination="1 - 34" refId="ref36928" refString="Framenau V. W. 2007. Revision of the new Australian genus Artoriopsis in a new subfamily of wolf spiders, Artoriinae (Araneae: Lycosidae). Zootaxa 1391: 1 - 34." type="journal article" year="2007">Framenau, 2007</bibRefCitation>
</taxonomicName>
that also includes
<taxonomicName id="4C6F4D5DFFEB183393469BA6B7A8C057" authority="L. Koch, 1878" authorityName="L. Koch" authorityYear="1878" class="Arachnida" family="Lycosidae" genus="Anoteropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183393469BA6B258C071" box="[1151,1290,1774,1800]" italics="true" pageId="15" pageNumber="16">Anoteropsis</emphasis>
L. Koch, 1878
</taxonomicName>
,
<taxonomicName id="4C6F4D5DFFEB1833963E9A5CB55AC057" authority="Thorell, 1877" authorityName="Thorell" authorityYear="1877" box="[263,520,1811,1838]" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB1833963E9A5CB60FC057" box="[263,349,1812,1838]" italics="true" pageId="15" pageNumber="16">Artoria</emphasis>
Thorell, 1877
</taxonomicName>
,
<taxonomicName id="4C6F4D5DFFEB1833952C9A5CB436C057" authority="Framenau, 2007" authorityName="Framenau" authorityYear="2007" box="[533,868,1812,1838]" class="Arachnida" family="Lycosidae" genus="Artoriopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB1833952C9A5CB5CBC057" box="[533,665,1812,1838]" italics="true" pageId="15" pageNumber="16">Artoriopsis</emphasis>
<bibRefCitation id="EFFE4B2FFFEB183395989A5CB436C057" author="Framenau" box="[673,868,1812,1838]" firstAuthor="Framenau" pageId="15" pageNumber="16" pagination="1 - 34" refId="ref36928" refString="Framenau V. W. 2007. Revision of the new Australian genus Artoriopsis in a new subfamily of wolf spiders, Artoriinae (Araneae: Lycosidae). Zootaxa 1391: 1 - 34." type="journal article" year="2007">Framenau, 2007</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6F4D5DFFEB183394499A5BB3C6C054" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[880,1172,1811,1838]" class="Arachnida" family="Lycosidae" genus="Diahogna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183394499A5BB4B7C054" box="[880,997,1811,1837]" italics="true" pageId="15" pageNumber="16">Diahogna</emphasis>
<bibRefCitation id="EFFE4B2FFFEB183394D49A5CB3C6C054" author="Roewer" box="[1005,1172,1811,1838]" firstAuthor="Roewer" pageId="15" pageNumber="16" pagination="519 - 1040" refId="ref39690" refString="Roewer C. F. 1960. Araneae Lycosaeformia II. (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba - Mission G. F. De Witte 55: 519 - 1040." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6F4D5DFFEB183393989A5BB7A8C02A" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" class="Arachnida" family="Lycosidae" genus="Lycosella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183393989A5BB243C054" box="[1185,1297,1811,1837]" italics="true" pageId="15" pageNumber="16">Lycosella</emphasis>
Thorell, 1890
</taxonomicName>
,
<taxonomicName id="4C6F4D5DFFEB183396319A72B557C02A" authority="Vink, 2002" authorityName="Vink" authorityYear="2002" box="[264,517,1849,1875]" class="Arachnida" family="Lycosidae" genus="Notocosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183396319A72B625C02A" box="[264,375,1850,1875]" italics="true" pageId="15" pageNumber="16">Notocosa</emphasis>
<bibRefCitation id="EFFE4B2FFFEB183396469A71B557C02A" author="Vink" box="[383,517,1849,1875]" firstAuthor="Vink" pageId="15" pageNumber="16" pagination="576 - 587" refId="ref40140" refString="Vink C. J. 2002. Lycosidae (Arachnida: Araneae). Fauna of New Zealand 44: 1 - 94. Vink C. J. &amp; Paterson A. M. 2003. Combined molecular and morphological phylogenetic analyses of the New Zealand wolf spider genus Anoteropsis (Araneae: Lycosidae). Molecular Phylogenetics and Evolution 28: 576 - 587. https: // doi. org / 10.1016 / s 1055 - 7903 (03) 00219 - 7" type="journal article" year="2002">Vink, 2002</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C6F4D5DFFEB1833957C9A71B404C02A" authority="Simon, 1900" authorityName="Simon" authorityYear="1900" box="[581,854,1849,1875]" class="Arachnida" family="Lycosidae" genus="Syroloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB1833957C9A71B5E6C02A" box="[581,692,1849,1875]" italics="true" pageId="15" pageNumber="16">Syroloma</emphasis>
Simon, 1900
</taxonomicName>
, an undescribed genus listed in
<bibRefCitation id="EFFE4B2FFFEB183393EA9A71B659C003" author="Murphy" etAl="et al." firstAuthor="Murphy" pageId="15" pageNumber="16" pagination="583 - 602" refId="ref38844" refString="Murphy N. P., Framenau V. W., Donnellan S. C., Harvey M. S., Park Y. - C. &amp; Austin A. D. 2006. Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 S rRNA, and NADH 1 genes: implications for classification, biogeography, and the evolution of web building behaviour. Molecular Phylogenetics and Evolution 38: 583 - 602. https: // doi. org / 10.1016 / j. ympev. 2005.09.004" type="journal article" year="2006">
Murphy
<emphasis id="B91BEACCFFEB183392039A72B225C02A" box="[1338,1399,1849,1875]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
(
<date id="FFD1101EFFEB183397FC9A17B651C003" box="[197,259,1887,1914]" pageId="15" pageNumber="16" value="2006">2006</date>
)
</bibRefCitation>
as New Genus 1 (
<bibRefCitation id="EFFE4B2FFFEB183395379A17B59CC000" author="Framenau" box="[526,718,1887,1913]" firstAuthor="Framenau" pageId="15" pageNumber="16" pagination="1 - 34" refId="ref36928" refString="Framenau V. W. 2007. Revision of the new Australian genus Artoriopsis in a new subfamily of wolf spiders, Artoriinae (Araneae: Lycosidae). Zootaxa 1391: 1 - 34." type="journal article" year="2007">
Framenau
<date id="FFD1101EFFEB183395AB9A17B59CC000" box="[658,718,1887,1913]" pageId="15" pageNumber="16" value="2007">2007</date>
</bibRefCitation>
), and
<taxonomicName id="4C6F4D5DFFEB183394189A28B202C000" authority="Framenau" authorityName="Framenau" authorityYear="2010" box="[801,1360,1887,1914]" class="Arachnida" family="Lycosidae" genus="Kangarosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183394189A28B4F0C000" box="[801,930,1888,1913]" italics="true" pageId="15" pageNumber="16">Kangarosa</emphasis>
<bibRefCitation id="EFFE4B2FFFEB183394969A28B326C000" author="Framenau" box="[943,1140,1887,1914]" firstAuthor="Framenau" pageId="15" pageNumber="16" pagination="113 - 142" refId="ref36964" refString="Framenau V. W. 2010. Revision of the new Australian wolf spider genus Kangarosa (Araneae: Lycosidae: Artoriinae). Arthropod Systematics and Phylogeny 68: 113 - 142." type="journal article" year="2010">Framenau, 2010</bibRefCitation>
(
<bibRefCitation id="EFFE4B2FFFEB183393B19A17B21AC000" author="Framenau" box="[1160,1352,1887,1913]" firstAuthor="Framenau" pageId="15" pageNumber="16" pagination="113 - 142" refId="ref36964" refString="Framenau V. W. 2010. Revision of the new Australian wolf spider genus Kangarosa (Araneae: Lycosidae: Artoriinae). Arthropod Systematics and Phylogeny 68: 113 - 142." type="journal article" year="2010">Framenau 2010</bibRefCitation>
)
</taxonomicName>
. It may also include
<collectingCountry id="F378764EFFEB183396B69ACDB562C0E6" box="[399,560,1925,1951]" name="Argentina" pageId="15" pageNumber="16">the Argentine</collectingCountry>
genera
<taxonomicName id="4C6F4D5DFFEB183395B69ACDB31EC0E6" authority="Piacentini" authorityName="Piacentini" authorityYear="2009" box="[655,1100,1925,1951]" class="Arachnida" family="Lycosidae" genus="Lobizon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183395B69ACDB5BCC0E6" box="[655,750,1925,1951]" italics="true" pageId="15" pageNumber="16">Lobizon</emphasis>
<bibRefCitation id="EFFE4B2FFFEB183395CF9ACDB31EC0E6" author="Piacentini &amp; Grismado" box="[758,1100,1925,1951]" firstAuthor="Piacentini" pageId="15" pageNumber="16" pagination="1 - 33" refId="ref39085" refString="Piacentini L. N. &amp; Grismado C. J. 2009. Lobizon and Navira, two new genera of wolf spiders from Argentina (Araneae: Lycosidae). Zootaxa 2195: 1 - 33." type="journal article" year="2009">Piacentini &amp; Grismado 2009</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6F4D5DFFEB183393BE9ACEB6D3C0BC" authority="Piacentini" authorityName="Piacentini" authorityYear="2009" class="Arachnida" family="Lycosidae" genus="Navira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEACCFFEB183393BE9ACEB38AC0E6" box="[1159,1240,1926,1951]" italics="true" pageId="15" pageNumber="16">Navira</emphasis>
<bibRefCitation id="EFFE4B2FFFEB183393D89ACDB6D3C0BC" author="Piacentini &amp; Grismado" firstAuthor="Piacentini" pageId="15" pageNumber="16" pagination="1 - 33" refId="ref39085" refString="Piacentini L. N. &amp; Grismado C. J. 2009. Lobizon and Navira, two new genera of wolf spiders from Argentina (Araneae: Lycosidae). Zootaxa 2195: 1 - 33." type="journal article" year="2009">Piacentini &amp; Grismado, 2009</bibRefCitation>
</taxonomicName>
; however, the placement of these genera within the subfamily remains contentious (
<bibRefCitation id="EFFE4B2FFFEB183397FC9A99B54BC092" author="Piacentini &amp; Grismado" box="[197,537,2001,2027]" firstAuthor="Piacentini" pageId="15" pageNumber="16" pagination="1 - 33" refId="ref39085" refString="Piacentini L. N. &amp; Grismado C. J. 2009. Lobizon and Navira, two new genera of wolf spiders from Argentina (Araneae: Lycosidae). Zootaxa 2195: 1 - 33." type="journal article" year="2009">
Piacentini &amp; Grismado
<date id="FFD1101EFFEB183396E49A99B54BC092" box="[477,537,2001,2027]" pageId="15" pageNumber="16" value="2009">2009</date>
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD036DEFFF4182C97849C44B5F7C608" blockId="16.[189,1398,267,370]" pageId="16" pageNumber="17">
A putative synapomorphy of this subfamily is the presence of the basoembolic apophysis on the male pedipalp. The monophyly of this subfamily is not only supported by the similar genital morphology, but also molecular data (
<bibRefCitation id="EFFE4B2FFFF4182C96899C1FB5C4C60B" author="Murphy" box="[432,662,343,370]" etAl="et al." firstAuthor="Murphy" pageId="16" pageNumber="17" pagination="583 - 602" refId="ref38844" refString="Murphy N. P., Framenau V. W., Donnellan S. C., Harvey M. S., Park Y. - C. &amp; Austin A. D. 2006. Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 S rRNA, and NADH 1 genes: implications for classification, biogeography, and the evolution of web building behaviour. Molecular Phylogenetics and Evolution 38: 583 - 602. https: // doi. org / 10.1016 / j. ympev. 2005.09.004" type="journal article" year="2006">
Murphy
<emphasis id="B91BEACCFFF4182C95219C10B501C608" box="[536,595,343,369]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
<date id="FFD1101EFFF4182C95639C1FB5C4C60B" box="[602,662,343,370]" pageId="16" pageNumber="17" value="2006">2006</date>
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD036DEFFF4182C97849CD3B39CC579" blockId="16.[189,1399,411,513]" pageId="16" pageNumber="17">
The following taxonomic treatment groups the species into (in this order) the
<taxonomicName id="4C6F4D5DFFF4182C930F9CD4B3D3C6CC" box="[1078,1153,412,437]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B91BEACCFFF4182C930F9CD4B3D3C6CC" box="[1078,1153,412,437]" italics="true" pageId="16" pageNumber="17">oraria</emphasis>
</taxonomicName>
-,
<taxonomicName id="4C6F4D5DFFF4182C93AE9CD3B3A6C6CC" baseAuthorityName="McKay" baseAuthorityYear="1976" box="[1175,1268,411,437]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alteripa">
<emphasis id="B91BEACCFFF4182C93AE9CD3B3A6C6CC" box="[1175,1268,411,437]" italics="true" pageId="16" pageNumber="17">alteripa</emphasis>
</taxonomicName>
- and
<taxonomicName id="4C6F4D5DFFF4182C920A9CD4B23FC6CC" baseAuthorityName="Hickman" baseAuthorityYear="1944" box="[1331,1389,412,437]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="eyrei">
<emphasis id="B91BEACCFFF4182C920A9CD4B23FC6CC" box="[1331,1389,412,437]" italics="true" pageId="16" pageNumber="17">eyrei</emphasis>
</taxonomicName>
- groups to facilitate comparison of similar species. Within each group, the species are in alphabetical order with the exception of
<taxonomicName id="4C6F4D5DFFF4182C953B9CAFB538C579" box="[514,618,487,512]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B91BEACCFFF4182C953B9CAFB538C579" box="[514,618,487,512]" italics="true" pageId="16" pageNumber="17">T. oraria</emphasis>
</taxonomicName>
, the
<typeStatus id="54D4887CFFF4182C959D9CAFB585C578" box="[676,727,487,513]" pageId="16" pageNumber="17">type</typeStatus>
species of the genus, which is treated first.
</paragraph>
</subSubSection>
</treatment>
</document>