154 lines
19 KiB
XML
154 lines
19 KiB
XML
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<mods:title id="7942EFAC380ED7EA7E695F67A04DF75B">Designation of a neotype for the dendrochirotid holothuroid, Thyone fusus (Müller) (Echinodermata: Holothuroidea: Dendrochirotida: Thyonidae)</mods:title>
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<mods:namePart id="D509D39D2A6C70195A3AD271348FEFAF">Thandar, Ahmed S.</mods:namePart>
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<treatment id="03C687C6FF93FFBF3AAF630725F531F5" ID-DOI="http://doi.org/10.5281/zenodo.10168517" ID-GBIF-Taxon="217741894" ID-Zenodo-Dep="10168517" LSID="urn:lsid:plazi:treatment:03C687C6FF93FFBF3AAF630725F531F5" httpUri="http://treatment.plazi.org/id/03C687C6FF93FFBF3AAF630725F531F5" lastPageNumber="587" pageId="2" pageNumber="587">
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<subSubSection id="C375655BFF93FFBF3AAF630726B1363E" box="[151,499,582,609]" pageId="2" pageNumber="587" type="nomenclature">
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<paragraph id="8BD036D0FF93FFBF3AAF630726B1363E" blockId="2.[151,499,582,645]" box="[151,499,582,609]" pageId="2" pageNumber="587">
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<heading id="D09881BCFF93FFBF3AAF630726B1363E" bold="true" box="[151,499,582,609]" fontSize="11" level="1" pageId="2" pageNumber="587" reason="1">
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<taxonomicName id="4C6F4D53FF93FFBF3AAF630726B1363E" ID-CoL="56RM4" authority="(Muller, 1776)" baseAuthorityName="Muller" baseAuthorityYear="1776" box="[151,499,582,609]" class="Holothuroidea" family="Thyonidae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="587" phylum="Echinodermata" rank="species" species="fusus">
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<emphasis id="B91BEAC2FF93FFBF3AAF630726B1363E" bold="true" box="[151,499,582,609]" pageId="2" pageNumber="587">
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<emphasis id="B91BEAC2FF93FFBF3AAF63072679363E" bold="true" box="[151,315,582,609]" italics="true" pageId="2" pageNumber="587">Thyone fusus</emphasis>
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(Müller, 1776)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C375655BFF93FFBF3AAF632A267B36DB" box="[151,313,619,645]" pageId="2" pageNumber="587" type="description">
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<paragraph id="8BD036D0FF93FFBF3AAF632A267B36DB" blockId="2.[151,499,582,645]" box="[151,313,619,645]" pageId="2" pageNumber="587">
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(
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<figureCitation id="13542A55FF93FFBF3AA7632A267336DB" box="[159,305,619,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,1718,1742]" captionTargetBox="[456,1134,708,1688]" captionTargetId="figure-250@3.[451,1137,705,1695]" captionTargetPageId="3" captionText="FIGURE 1. Thyone fusus (Müller). Neotype (designated herein). A. calcareous ring (ir = interradial plate; r = radial plate, labelled according to Ludwig’s system); B. tables of body wall; C. rosettes of introvert; D. supporting tables of tube feet; E. rods and plates of tentacles; F. tables of introvert, G. tables of body wall of T. flexus (reproduced from holotype slides examined at NHMUK)) (A = Scale 1; B–G = Scale 2)." figureDoi="http://doi.org/10.5281/zenodo.10149944" httpUri="https://zenodo.org/record/10149944/files/figure.png" pageId="2" pageNumber="587">Figure 1A–F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C375655BFF93FFBF3AAF63EE25C1377D" pageId="2" pageNumber="587" type="reference_group">
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<paragraph id="8BD036D0FF93FFBF3AAF63EE259C3699" blockId="2.[151,1436,687,803]" box="[151,734,687,711]" pageId="2" pageNumber="587">
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<taxonomicName id="4C6F4D53FF93FFBF3AAF63EE25723699" authority="O. F. Muller, 1776: 232" authorityName="O. F. Muller" authorityPageNumber="232" authorityYear="1776" box="[151,560,687,711]" class="Holothuroidea" family="Holothuriidae" genus="Holothuria" kingdom="Animalia" order="Holothuriida" pageId="2" pageNumber="587" phylum="Echinodermata" rank="species" species="fusus">
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<emphasis id="B91BEAC2FF93FFBF3AAF63EE26003699" box="[151,322,687,711]" italics="true" pageId="2" pageNumber="587">Holothuria fusus</emphasis>
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O.F. Müller, 1776: 232
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</taxonomicName>
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, pl. 10, figs. 5–6.
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</paragraph>
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<paragraph id="8BD036D0FF93FFBF3AAF638F260A375B" blockId="2.[151,1436,687,803]" pageId="2" pageNumber="587">
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<treatmentCitationGroup id="AB7F11FEFF93FFBF3AAF638F260A375B" pageId="2" pageNumber="587">
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<treatmentCitation id="0ACE10C1FF93FFBF3AAF638F26A236B8" author="Madsen, F. J." box="[151,480,718,742]" page="17" pageId="2" pageNumber="587" year="1941">
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<taxonomicName id="4C6F4D53FF93FFBF3AAF638F26A236B8" ID-CoL="56RM4" authority="Madsen, 1941: 17" authorityName="Madsen" authorityPageNumber="17" authorityYear="1941" box="[151,480,718,742]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="587" phylum="Echinodermata" rank="species" species="fusus">
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<emphasis id="B91BEAC2FF93FFBF3AAF638F265C36B8" box="[151,286,718,742]" italics="true" pageId="2" pageNumber="587">Thyone fusus</emphasis>
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<bibRefCitation id="EFFE4B21FF93FFBF3B1E638F26A236B8" author="Madsen, F. J." box="[294,480,718,742]" pageId="2" pageNumber="587" pagination="1 - 30" refId="ref3569" refString="Madsen, F. J. (1941) On Thyone wahrbergi n. sp., a new holothurian from the Skagerrak, with remarks on T. fusus (O. F. M.) and other related species. Goteborgs Kungl. Vetenskaps och Vitterhets-Samhalles Handlingar, Meddelanded fran Goteborgs Musei Zoologiska Avdelning 85, 6 (1), 1 - 30." type="journal article" year="1941">Madsen, 1941: 17</bibRefCitation>
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</taxonomicName>
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, text-figs. 12–16;
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<treatmentCitation id="0ACE10C1FF93FFBF38A6638F248B36B8" author="Madsen, F. J. & Hansen, B." box="[670,969,718,742]" page="40" pageId="2" pageNumber="587" year="1994">
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<bibRefCitation id="EFFE4B21FF93FFBF38A6638F248B36B8" author="Madsen, F. J. & Hansen, B." box="[670,969,718,742]" pageId="2" pageNumber="587" refId="ref3637" refString="Madsen, F. J. & Hansen, B. (1994) Marine Invertebrates of Scandinavia Number 9. Echinodermata. Holothuroidea. Scandinavian University Press, Oslo, 141 pp." type="book" year="1994">Madsen & Hansen, 1994: 40</bibRefCitation>
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</treatmentCitation>
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, fig. 5, 23–24, map 10;
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<treatmentCitation id="0ACE10C1FF93FFBF3EF8638F22DE36B8" author="McKenzie, J. D." box="[1216,1436,718,742]" page="136" pageId="2" pageNumber="587" year="1991">
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<bibRefCitation id="EFFE4B21FF93FFBF3EF8638F22DE36B8" author="McKenzie, J. D." box="[1216,1436,718,742]" pageId="2" pageNumber="587" pagination="123 - 171" refId="ref3671" refString="McKenzie, J. D. (1991) The taxonomy and natural history of north European dendrochirote holothurians (Echinodermata). Journal of Natural History, 25, 123 - 171. https: // doi. org / 10.1080 / 00222939100770091" type="journal article" year="1991">McKenzie, 1991: 136</bibRefCitation>
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</treatmentCitation>
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(synonymy).
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</paragraph>
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<paragraph id="8BD036D0FF93FFBF3AAF624A25C1377D" blockId="2.[151,1436,687,803]" box="[151,643,779,803]" pageId="2" pageNumber="587">
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<treatmentCitationGroup id="AB7F11FEFF93FFBF3AAF624A25C1377D" box="[151,643,779,803]" pageId="2" pageNumber="587">
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<treatmentCitation id="0ACE10C1FF93FFBF3AAF624A2696377D" author="Hodge, G." box="[151,468,779,803]" page="44" pageId="2" pageNumber="587" year="1867">
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||
<taxonomicName id="4C6F4D53FF93FFBF3AAF624A2696377D" ID-CoL="56RLY" authority="Hodge, 1867: 44" authorityName="Hodge" authorityPageNumber="44" authorityYear="1867" box="[151,468,779,803]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="587" phylum="Echinodermata" rank="species" species="flexus">
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<emphasis id="B91BEAC2FF93FFBF3AAF624A2661377D" box="[151,291,779,803]" italics="true" pageId="2" pageNumber="587">Thyone flexus</emphasis>
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<bibRefCitation id="EFFE4B21FF93FFBF3B11624A2696377D" author="Hodge, G." box="[297,468,779,803]" pageId="2" pageNumber="587" pagination="42 - 45" refId="ref3498" refString="Hodge, G. (1867) Report of the Echinodermata. Natural History Transactions of Northumberland, Durham and Newcastleupon-Tyne, 1, 42 - 45." type="journal article" year="1867">Hodge, 1867: 44</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, pl. 10, fig. 2–11.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C375655BFF93FFBF3AAF6210248430A9" pageId="2" pageNumber="587" type="diagnosis">
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<paragraph id="8BD036D0FF93FFBF3AAF621024B03735" blockId="2.[151,1437,849,1452]" box="[151,1010,849,876]" pageId="2" pageNumber="587">
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<emphasis id="B91BEAC2FF93FFBF3AAF621026483735" bold="true" box="[151,266,849,875]" pageId="2" pageNumber="587">Diagnosis</emphasis>
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(from
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<bibRefCitation id="EFFE4B21FF93FFBF3B60621025283735" author="Madsen, F. J. & Hansen, B." box="[344,618,849,876]" pageId="2" pageNumber="587" refId="ref3637" refString="Madsen, F. J. & Hansen, B. (1994) Marine Invertebrates of Scandinavia Number 9. Echinodermata. Holothuroidea. Scandinavian University Press, Oslo, 141 pp." type="book" year="1994">Madsen & Hansen 1994</bibRefCitation>
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,
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<bibRefCitation id="EFFE4B21FF93FFBF384D621024693735" author="McKenzie, J. D." box="[629,811,849,875]" pageId="2" pageNumber="587" pagination="123 - 171" refId="ref3671" refString="McKenzie, J. D. (1991) The taxonomy and natural history of north European dendrochirote holothurians (Echinodermata). Journal of Natural History, 25, 123 - 171. https: // doi. org / 10.1080 / 00222939100770091" type="journal article" year="1991">McKenzie 1991</bibRefCitation>
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, amended herein)
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</paragraph>
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<paragraph id="8BD036D0FF93FFBF3AE96234248430A9" blockId="2.[151,1437,849,1452]" pageId="2" pageNumber="587">
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||
Medium-sized species, reaching about
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<quantity id="4C979B35FF93FFBF38B6623425A237CE" box="[654,736,885,912]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="587" unit="mm" value="70.0">70 mm</quantity>
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||
in length (an exceptional figure of
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<quantity id="4C979B35FF93FFBF3E446234239937CE" box="[1148,1243,885,912]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="587" unit="mm" value="200.0">200 mm</quantity>
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was reported by
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||
<bibRefCitation id="EFFE4B21FF93FFBF3AAF62DB262637ED" author="Mortensen, T." box="[151,356,921,948]" pageId="2" pageNumber="587" refId="ref3767" refString="Mortensen, T. (1927) Handbook of the Echinoderms of the British Isles. Oxford University Press, Oxford, ix + 471 pp., 269 text-figs. https: // doi. org / 10.5962 / bhl. title. 6841" type="book" year="1927">Mortensen (1927)</bibRefCitation>
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||
but this needs verification). Body fusiform to spindle-shaped, tapering posteriorly. Colour in life greyish-white to light brown to fawn, brown in alcohol. Tube feet well developed, distributed over entire body, numerous ventrally, scattered dorsally, often with adhering sand grains or other debris.Anal teeth present. Calcareous ring long, tubular, both ring and processes sub-divided; radial plates deeply notched anteriorly, dorsal radial plates extended posteriorly before bifurcating beyond posterior border of interradial plates, posterior processes of radial plates long; interradials prolonged, two ventral ones slightly longer. Deposits of body wall 2-pillared tables, disc with four primary holes and often a few additional, smaller, peripheral holes around the primary ones; spire of moderate height, ending in inconspicuous teeth. Supporting tables of tube feet with curved disc and endplate, the latter delicate in smaller individuals. Tentacle deposits perforated rods and plates. Introvert supported by rosettes and tables with multilocular disc and a spire with or without a cross-bar..
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C375655BFF93FFBF3AFF644025F531F5" pageId="2" pageNumber="587" type="materials_examined">
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<paragraph id="8BD036D0FF93FFBF3AFF644025F531F5" blockId="2.[151,1437,849,1452]" pageId="2" pageNumber="587">
|
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<emphasis id="B91BEAC2FF93FFBF3AFF644026F03145" bold="true" box="[199,434,1281,1307]" pageId="2" pageNumber="587">Material examined:</emphasis>
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||
<specimenCount id="9D69FD59FF93FFBF3B80644325293145" box="[440,619,1281,1308]" count="4" pageId="2" pageNumber="587" type="generic">Four specimens</specimenCount>
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||
, identified as
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||
<emphasis id="B91BEAC2FF93FFBF393B6443241C3145" box="[771,862,1281,1307]" italics="true" pageId="2" pageNumber="587">
|
||
<taxonomicName id="4C6F4D53FF93FFBF393B644324193145" authorityName="Madsen" authorityYear="1941" box="[771,859,1281,1307]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="587" phylum="Echinodermata" rank="species" species="fusus">T. fusus</taxonomicName>
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||
,
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||
</emphasis>
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||
originating from the type locality, were received on loan from the Natural History Museum,
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<collectingRegion id="49ABF832FF93FFBF386F646425CD3161" box="[599,655,1317,1343]" country="Norway" name="Oslo" pageId="2" pageNumber="587">Oslo</collectingRegion>
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||
,
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||
<collectingCountry id="F3787640FF93FFBF38A2646725B4311E" box="[666,758,1318,1344]" name="Norway" pageId="2" pageNumber="587">Norway</collectingCountry>
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||
. The largest specimen was selected for dissection and ossicle study and is here designated as the
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<typeStatus id="54D48872FF93FFBF3816640B25C8313A" box="[558,650,1354,1380]" pageId="2" pageNumber="587" type="neotype">neotype</typeStatus>
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, while the other three were only studied for their body wall ossicles. The current material corresponds well with the descriptions of the species presented by especially
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<bibRefCitation id="EFFE4B21FF93FFBF3ED2642C22DA31D9" author="Madsen, F. J." box="[1258,1432,1389,1415]" pageId="2" pageNumber="587" pagination="1 - 30" refId="ref3569" refString="Madsen, F. J. (1941) On Thyone wahrbergi n. sp., a new holothurian from the Skagerrak, with remarks on T. fusus (O. F. M.) and other related species. Goteborgs Kungl. Vetenskaps och Vitterhets-Samhalles Handlingar, Meddelanded fran Goteborgs Musei Zoologiska Avdelning 85, 6 (1), 1 - 30." type="journal article" year="1941">Madsen (1941)</bibRefCitation>
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||
,
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||
<bibRefCitation id="EFFE4B21FF93FFBF3AAF64D0261531F5" author="McKenzie, J. D." box="[151,343,1425,1451]" pageId="2" pageNumber="587" pagination="123 - 171" refId="ref3671" refString="McKenzie, J. D. (1991) The taxonomy and natural history of north European dendrochirote holothurians (Echinodermata). Journal of Natural History, 25, 123 - 171. https: // doi. org / 10.1080 / 00222939100770091" type="journal article" year="1991">Mckenzie (1991)</bibRefCitation>
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and
|
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<bibRefCitation id="EFFE4B21FF93FFBF3BB664D025F131F5" author="Madsen, F. J. & Hansen, B." box="[398,691,1425,1452]" pageId="2" pageNumber="587" refId="ref3637" refString="Madsen, F. J. & Hansen, B. (1994) Marine Invertebrates of Scandinavia Number 9. Echinodermata. Holothuroidea. Scandinavian University Press, Oslo, 141 pp." type="book" year="1994">Madsen & Hansen (1994)</bibRefCitation>
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||
.
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||
</paragraph>
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</subSubSection>
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</treatment>
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</document> |