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<document id="1968632AE25BFA8D6096A69E659BB5D2" ID-DOI="10.5252/zoosystema2024v46a5" ID-ISSN="1638-9387" ID-Zenodo-Dep="10803267" ID-ZooBank="1B0779F3-FAE4-4105-8130-A26DE519721E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710145347001" checkinUser="plazi" docAuthor="Cardoso, Giovanna Monticelli, M, Diego de, Bento, edeiros &amp; Ferreira, Rodrigo Lopes" docDate="2024" docId="03C687BFC61FFFDA13A2F40E68C7FA77" docLanguage="en" docName="zoosystema.2024.46.5.pdf" docOrigin="Zoosystema 46 (5)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a5.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Ctenorillo potiguar Cardoso &amp; Ferreira 2024, n. sp." docType="treatment" docUuid="53D58AC8-70BA-48E8-A228-4F455E72B475" docUuidSource="ZooBank" docVersion="2" lastPageNumber="121" masterDocId="FFFFFFC7C606FFC6136FF25C6C6BFFBA" masterDocTitle="Unveiling a hidden diversity: descriptions of nine new species of Ctenorillo Verhoeff, 1942 (Isopoda, Armadillidae) discovered in Brazilian caves and their importance for conservation" masterLastPageNumber="132" masterPageNumber="95" pageNumber="118" updateTime="1710145921992" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="58F654AEDFF1322B8DFE2A8E51178CB2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8B5926BBABBCB81E9DE3C4532F7F1A59">
<mods:title id="AF570305D4F5E49C5049D929E4A2C40E">Unveiling a hidden diversity: descriptions of nine new species of Ctenorillo Verhoeff, 1942 (Isopoda, Armadillidae) discovered in Brazilian caves and their importance for conservation</mods:title>
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<mods:name id="4285522AFC410E949A7F7EC7289521E0" type="personal">
<mods:role id="0EA3180C153CA576A1CE36982C1E9289">
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<mods:namePart id="2EACC8C7740CC756FD5739ABA31831CC">Cardoso, Giovanna Monticelli</mods:namePart>
<mods:affiliation id="353786DC95E413339215330F21B12775">Universidade Federal de Lavras (UFLA), Centro de Estudos em Biologia Subterrânea, Departamento de Biologia, programa de Pós-graduação em Ecologia Aplicada, Campus Universitário, CP 3037, 37200 - 000 Lavras, Minas Gerais (Brasil) gmcardoso. bio @ gmail. com (corresponding author)</mods:affiliation>
<mods:nameIdentifier id="C0C84278A19A52FAD34414B440177864" type="email">bio@gmail.com</mods:nameIdentifier>
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<mods:role id="4F1407483CED8BCDE2C003D1F45C19A8">
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<mods:namePart id="ABD3023D6C8AEDE8815073C1DAA30268">M, Diego de</mods:namePart>
</mods:name>
<mods:name id="E7516C29472CC20E30848BE32D2506DC" type="personal">
<mods:role id="41FBCF8D36E2EA8231D2CE51DDF0673E">
<mods:roleTerm id="D93BD85A7E8ABAD5C7F7F52F59AE24D3">Author</mods:roleTerm>
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<mods:namePart id="D7DDF63E1386E7D82BD67959E155599E">Bento, edeiros</mods:namePart>
</mods:name>
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<mods:role id="94F810206E1D498CB461D554B1C5F893">
<mods:roleTerm id="60A63547F5CC68D1D28B65AAEC3833D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BEC1D15141E866E691148864AE59238">Ferreira, Rodrigo Lopes</mods:namePart>
<mods:affiliation id="F4E721BBFB1A67A84207545B2D981F84">Universidade Federal de Lavras (UFLA), Centro de Estudos em Biologia Subterrânea, Departamento de Biologia, programa de Pós-graduação em Ecologia Aplicada, Campus Universitário, CP 3037, 37200 - 000 Lavras, Minas Gerais (Brasil) drops @ ufla. br</mods:affiliation>
<mods:nameIdentifier id="C9907EB1C64D90C4DEA9A3B637E94ADD" type="email">drops@ufla.br</mods:nameIdentifier>
</mods:name>
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<mods:title id="F5F956A97E2B5AF27A2FE139993C544C">Zoosystema</mods:title>
</mods:titleInfo>
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<mods:date id="A55B09D40C1AB9541398AE645572B106">2024</mods:date>
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<mods:number id="EED76CC9E41003F269A7C5C38B200AD9">2024-03-05</mods:number>
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<mods:number id="3EB93961079456289B80C0F18A5E061A">46</mods:number>
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<mods:start id="5D9CE0474FF91EFCC51C653E437021E7">95</mods:start>
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<subSubSection id="C3756522C61FFFDF13A2F40E6ED5F9D7" box="[205,702,1618,1645]" pageId="25" pageNumber="118" type="nomenclature">
<paragraph id="8BD036A9C61FFFDF13A2F40E6ED5F9D7" blockId="25.[205,702,1618,1676]" box="[205,702,1618,1645]" pageId="25" pageNumber="118">
<heading id="D09881C5C61FFFDF13A2F40E6ED5F9D7" box="[205,702,1618,1645]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="118" reason="2">
<taxonomicName id="4C6F4D2AC61FFFDF13A2F40E6E12F9D6" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[205,633,1618,1645]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="118" phylum="Arthropoda" rank="species" species="potiguar" status="sp. nov.">
<emphasis id="B91BEABBC61FFFDF13A2F40E6DF6F9D6" bold="true" box="[205,413,1618,1645]" italics="true" pageId="25" pageNumber="118">Ctenorillo potiguar</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61FFFDF11EBF40F6ED5F9D7" box="[644,702,1619,1645]" pageId="25" pageNumber="118" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3756522C61FFFDF1226F42E6E80F907" pageId="25" pageNumber="118" type="description">
<paragraph id="8BD036A9C61FFFDF1226F42E6E29F936" blockId="25.[205,702,1618,1676]" box="[329,578,1650,1676]" pageId="25" pageNumber="118">
(
<figureCitation id="13542A2CC61FFFDF123FF42E6DACF936" box="[336,455,1650,1676]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="26.[133,144,1571,1589]" captionStartId-1="27.[132,143,1340,1358]" captionStartId-2="28.[133,144,1106,1124]" captionTargetBox-0="[151,1437,213,1527]" captionTargetBox-1="[137,1428,218,1293]" captionTargetBox-2="[132,1455,215,1063]" captionTargetId-0="figure-218@26.[151,1437,213,1527]" captionTargetId-1="figure-339@27.[132,1455,210,1301]" captionTargetId-2="figure-483@28.[132,1455,215,1063]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionText-0="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." captionText-1="FIG. 19. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp., paratype (male, 3 mm, ISLA96798): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." captionText-2="FIG. 20. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: A, limestone outcrops at the Furna Feia National Park, with the entrance of the Furna Feia cave (yellow star); B, detail of the Furna Feia cave entrance; C, inner portion of the Furna Nova cave, where specimens are found; D, habitus in natural condition." figureDoi-0="http://doi.org/10.5281/zenodo.10803305" figureDoi-1="http://doi.org/10.5281/zenodo.10803307" figureDoi-2="http://doi.org/10.5281/zenodo.10803309" httpUri-0="https://zenodo.org/record/10803305/files/figure.png" httpUri-1="https://zenodo.org/record/10803307/files/figure.png" httpUri-2="https://zenodo.org/record/10803309/files/figure.png" pageId="25" pageNumber="118">Figs 18-20</figureCitation>
;
<figureCitation id="13542A2CC61FFFDF12BCF42E6E63F936" box="[467,520,1650,1676]" captionStart="FIG" captionStartId="35.[132,143,1225,1243]" captionTargetBox="[132,1455,216,1183]" captionTargetId="figure-480@35.[699,983,655,744]" captionTargetPageId="35" captionText="FIG. 27. — Habitus of the species of the genus Ctenorillo Verhoeff, 1942 discussed in the present paper:A, Ctenorillo ferrarai Campos-Filho, Araujo &amp; Taiti,2014; B, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.; C, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.; D, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.; E, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.; F, Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.; G, Ctenorillo quiteriensis Cardoso &amp; Ferreira,n. sp.; H, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.; I, Ctenorillo iuiuensis n. sp.; J, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10803325" httpUri="https://zenodo.org/record/10803325/files/figure.png" pageId="25" pageNumber="118">27H</figureCitation>
;
<figureCitation id="13542A2CC61FFFDF117CF42E6E52F936" box="[531,569,1650,1676]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="25" pageNumber="118">28I</figureCitation>
)
</paragraph>
<paragraph id="8BD036A9C61FFFDF13CEF4F76E80F907" blockId="25.[161,747,1707,1725]" box="[161,747,1707,1725]" pageId="25" pageNumber="118">
<uri id="FFFE3AABC61FFFDF13CEF4F76E80F907" box="[161,747,1707,1725]" pageId="25" pageNumber="118">
urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
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<paragraph id="8BD036A9C61FFFDF13EBF4806EA1F8FA" blockId="25.[131,776,1756,2028]" pageId="25" pageNumber="118">
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<collectionCode id="ED7EAE6CC61FFFDF13EBF4806CDEF948" box="[132,181,1756,1779]" pageId="25" pageNumber="118">TYPE</collectionCode>
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. —
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<typeStatus id="54D4880BC61FFFDF1229F4806DCEF94E" box="[326,421,1756,1780]" pageId="25" pageNumber="118" type="holotype">Holotype</typeStatus>
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.
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<collectingCountry id="F3787639C61FFFDF12DEF4816D80F94F" box="[433,491,1757,1781]" name="Brazil" pageId="25" pageNumber="118">Brazil</collectingCountry>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B073CF4C61FFFDF116DF4806F6FF89C" collectingDate="2008-06-12" collectionCode="VI" collectorName="D. M. Bento" country="Iceland" latitude="-5.034167" location="Barauna" longLatPrecision="21" longitude="-37.57111" municipality="Furna Nova cave" pageId="25" pageNumber="118" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Norte" typeStatus="holotype">
<specimenCount id="9D69FD20C61FFFDF116DF4806E4DF949" box="[514,550,1756,1780]" count="1" pageId="25" pageNumber="118" type="male">1 ♂</specimenCount>
(
<quantity id="4C979B4CC61FFFDF115AF4806E19F949" box="[565,626,1756,1780]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="118" unit="mm" value="3.0">3 mm</quantity>
, parts in slide);
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;
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municipality;
<collectingMunicipality id="6BB4ACD3C61FFFDF110DF4AA6F68F8B7" box="[610,771,1782,1805]" pageId="25" pageNumber="118">Furna Nova cave</collectingMunicipality>
;
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,
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;
<date id="FFD11069C61FFFDF120DF5536DBCF89D" box="[354,471,1807,1831]" pageId="25" pageNumber="118" value="2008-06-12">
<collectingDate id="EF95E981C61FFFDF120DF5536DBCF89D" box="[354,471,1807,1831]" pageId="25" pageNumber="118" value="2008-06-12">
12.
<collectionCode id="ED7EAE6CC61FFFDF1210F5536DF7F89D" box="[383,412,1807,1831]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="25" pageNumber="118">VI</collectionCode>
.2008
</collectingDate>
</date>
;
<collectorName id="269A537FC61FFFDF128EF5536E31F89C" box="[481,602,1807,1830]" pageId="25" pageNumber="118">D.M. Bento</collectorName>
leg.;
<collectingCountry id="F3787639C61FFFDF11E1F5526EADF89C" box="[654,710,1806,1830]" name="Iceland" pageId="25" pageNumber="118">ISLA</collectingCountry>
96798
</materialsCitation>
.
<materialsCitation id="3B073CF4C61FFFDF13EBF5746D40F8FB" box="[132,299,1832,1857]" country="Brazil" location="Brazil" pageId="25" pageNumber="118" specimenCount="1" typeStatus="paratype">
<emphasis id="B91BEABBC61FFFDF13EBF5746D40F8FB" bold="true" box="[132,299,1832,1857]" pageId="25" pageNumber="118">
<typeStatus id="54D4880BC61FFFDF13EBF5746C8DF8FA" box="[132,230,1832,1856]" pageId="25" pageNumber="118" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F3787639C61FFFDF139EF5756D40F8FB" box="[241,299,1833,1857]" name="Brazil" pageId="25" pageNumber="118">Brazil</collectingCountry>
</emphasis>
</materialsCitation>
•
<materialsCitation id="3B073CF4C61FFFDF122EF5746EADF8FA" box="[321,710,1832,1856]" collectingDate="2008-06-12" collectionCode="VI" collectorName="D. M. Bento" country="Iceland" latitude="-5.034167" location="Barauna" longLatPrecision="21" longitude="-37.57111" municipality="Furna Nova cave" pageId="25" pageNumber="118" specimenCount="2" specimenCount-female="2" stateProvince="Rio Grande do Norte" typeStatus="paratype">
<specimenCount id="9D69FD20C61FFFDF122EF5746D09F885" box="[321,354,1832,1856]" count="2" pageId="25" pageNumber="118" type="female">2 ♀</specimenCount>
; same data as holotype;
<collectingCountry id="F3787639C61FFFDF113EF5746EE3F8FA" box="[593,648,1832,1856]" name="Iceland" pageId="25" pageNumber="118">ISLA</collectingCountry>
96799
</materialsCitation>
.
</paragraph>
<paragraph id="8BD036A9C61FFFDF13EBF5096EF1F851" blockId="25.[131,776,1756,2028]" pageId="25" pageNumber="118">
<materialsCitation id="3B073CF4C61FFFDF13EBF5096C88F83C" collectionCode="OTHER, MATERIAL" collectorName="Barauna" country="Brazil" location="Rio Grande do Norte" pageId="25" pageNumber="118" specimenCount="1" stateProvince="Rio Grande do Norte">
<smallCapsWord id="8D36A075C61FFFDF13EBF5096CA0F8D1" baselines="1895,1895" box="[132,203,1877,1901]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Other" pageId="25" pageNumber="118">
<collectionCode id="ED7EAE6CC61FFFDF13EBF5096CA0F8D1" box="[132,203,1877,1901]" pageId="25" pageNumber="118">OTHER</collectionCode>
</smallCapsWord>
<smallCapsWord id="8D36A075C61FFFDF13BCF5066D5AF8D1" baselines="1895" box="[211,305,1882,1899]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="25" pageNumber="118">
<collectionCode id="ED7EAE6CC61FFFDF13BCF5066D5AF8D1" box="[211,305,1882,1899]" pageId="25" pageNumber="118">MATERIAL</collectionCode>
</smallCapsWord>
. —
<emphasis id="B91BEABBC61FFFDF1231F50A6DF4F8D4" bold="true" box="[350,415,1878,1902]" pageId="25" pageNumber="118">
<collectingCountry id="F3787639C61FFFDF1231F50A6DF0F8D4" box="[350,411,1878,1902]" name="Brazil" pageId="25" pageNumber="118">Brazil</collectingCountry>
.
</emphasis>
<collectingRegion id="49ABF84BC61FFFDF12CAF5096E10F8D6" box="[421,635,1877,1901]" country="Brazil" name="Rio Grande do Norte" pageId="25" pageNumber="118">Rio Grande do Norte</collectingRegion>
;
<collectorName id="269A537FC61FFFDF11E8F5096EBCF8D7" box="[647,727,1877,1901]" pageId="25" pageNumber="118">Baraúna</collectorName>
municipality
</materialsCitation>
<emphasis id="B91BEABBC61FFFDF1383F5326C9DF83C" bold="true" box="[236,246,1902,1926]" pageId="25" pageNumber="118">•</emphasis>
<materialsCitation id="3B073CF4C61FFFDF1390F5326E42F825" collectingDate="2010-07-31" collectorName="D. M. Bento" country="Brazil" latitude="-5.036667" location="Furna Feia cave" longLatPrecision="21" longitude="-37.559998" pageId="25" pageNumber="118" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="9D69FD20C61FFFDF1390F5326D4FF83F" box="[255,292,1902,1926]" count="1" pageId="25" pageNumber="118" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D69FD20C61FFFDF125BF5326D3CF83F" box="[308,343,1902,1926]" count="2" pageId="25" pageNumber="118" type="female">2 ♀</specimenCount>
;
<location id="8EB06072C61FFFDF1209F5336E61F83C" LSID="urn:lsid:plazi:treatment:03C687BFC61FFFDA13A2F40E68C7FA77:8EB06072C61FFFDF1209F5336E61F83C" box="[358,522,1902,1926]" country="Brazil" latitude="-5.036667" longLatPrecision="21" longitude="-37.559998" name="Furna Feia cave" pageId="25" pageNumber="118">Furna Feia cave</location>
;
<geoCoordinate id="EE5B506EC61FFFDF1178F5326E18F83C" box="[535,627,1902,1926]" degrees="5" direction="south" minutes="2" orientation="latitude" pageId="25" pageNumber="118" precision="15" seconds="12" value="-5.036667">5°2’12”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC61FFFDF11EFF5326F6BF83C" box="[640,768,1902,1926]" degrees="37" direction="west" minutes="33" orientation="longitude" pageId="25" pageNumber="118" precision="15" seconds="36" value="-37.559998">37°33’36”W</geoCoordinate>
;
<date id="FFD11069C61FFFDF13EBF5D46D6AF81A" box="[132,257,1928,1952]" pageId="25" pageNumber="118" value="2010-07-31">
<collectingDate id="EF95E981C61FFFDF13EBF5D46D6AF81A" box="[132,257,1928,1952]" pageId="25" pageNumber="118" value="2010-07-31">31.VII.2010</collectingDate>
</date>
;
<collectorName id="269A537FC61FFFDF1264F5D46DEFF825" box="[267,388,1928,1951]" pageId="25" pageNumber="118">D.M. Bento</collectorName>
leg.; ISLA59267
</materialsCitation>
•
<materialsCitation id="3B073CF4C61FFFDF1150F5D46F6CF803" collectingDate="2010-06-12" collectionCode="VI" collectorName="D. M. Bento" country="Brazil" latitude="-5.037778" location="Esquecida cave" longLatPrecision="21" longitude="-37.56111" pageId="25" pageNumber="118" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D69FD20C61FFFDF1150F5D46E08F825" box="[575,611,1928,1952]" count="2" pageId="25" pageNumber="118" type="male">2 ♂</specimenCount>
;
<location id="8EB06072C61FFFDF1100F5DB6F68F825" LSID="urn:lsid:plazi:treatment:03C687BFC61FFFDA13A2F40E68C7FA77:8EB06072C61FFFDF1100F5DB6F68F825" box="[623,771,1927,1951]" country="Brazil" latitude="-5.037778" longLatPrecision="21" longitude="-37.56111" name="Esquecida cave" pageId="25" pageNumber="118">Esquecida cave</location>
;
<geoCoordinate id="EE5B506EC61FFFDF13EBF5FD6CB5F803" box="[132,222,1953,1977]" degrees="5" direction="south" minutes="2" orientation="latitude" pageId="25" pageNumber="118" precision="15" seconds="16" value="-5.037778">5°2’16”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC61FFFDF1388F5FD6D09F803" box="[231,354,1953,1977]" degrees="37" direction="west" minutes="33" orientation="longitude" pageId="25" pageNumber="118" precision="15" seconds="40" value="-37.56111">37°33’40”W</geoCoordinate>
;
<date id="FFD11069C61FFFDF1202F5FD6D8AF803" box="[365,481,1953,1977]" pageId="25" pageNumber="118" value="2010-06-12">
<collectingDate id="EF95E981C61FFFDF1202F5FD6D8AF803" box="[365,481,1953,1977]" pageId="25" pageNumber="118" value="2010-06-12">
12.
<collectionCode id="ED7EAE6CC61FFFDF12E6F5FD6DCDF803" box="[393,422,1953,1977]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="25" pageNumber="118">VI</collectionCode>
.2010
</collectingDate>
</date>
;
<collectorName id="269A537FC61FFFDF1284F5FD6E09F802" box="[491,610,1953,1976]" pageId="25" pageNumber="118">D.M. Bento</collectorName>
leg.; ISLA59283
</materialsCitation>
•
<materialsCitation id="3B073CF4C61FFFDF13FCF5E66EF4F868" box="[147,671,1978,2002]" collectingDate="2010-06-12" collectionCode="VI" collectorName="D. M. Bento" country="Brazil" latitude="-5.037778" location="Esquecida cave" longLatPrecision="21" longitude="-37.56111" pageId="25" pageNumber="118" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D69FD20C61FFFDF13FCF5E66CDFF86B" box="[147,180,1978,2002]" count="1" pageId="25" pageNumber="118" type="female">1 ♀</specimenCount>
; same collection data as for preceding; ISLA59333
</materialsCitation>
•
<materialsCitation id="3B073CF4C61FFFDF11DBF5E66EFDF851" collectingDate="2010-06-11" collectionCode="VI" collectorName="D. M. Bento" country="Brazil" location="Britador cave" pageId="25" pageNumber="118" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D69FD20C61FFFDF11DBF5E66EBEF86B" box="[692,725,1978,2002]" count="1" pageId="25" pageNumber="118" type="female">1 ♀</specimenCount>
;
<location id="8EB06072C61FFFDF118EF5E66C81F851" LSID="urn:lsid:plazi:treatment:03C687BFC61FFFDA13A2F40E68C7FA77:8EB06072C61FFFDF118EF5E66C81F851" country="Brazil" name="Britador cave" pageId="25" pageNumber="118">Britador cave</location>
;
<date id="FFD11069C61FFFDF139BF5886D03F856" box="[244,360,2004,2028]" pageId="25" pageNumber="118" value="2010-06-11">
<collectingDate id="EF95E981C61FFFDF139BF5886D03F856" box="[244,360,2004,2028]" pageId="25" pageNumber="118" value="2010-06-11">
11.
<collectionCode id="ED7EAE6CC61FFFDF1261F5886D47F856" box="[270,300,2004,2028]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="25" pageNumber="118">VI</collectionCode>
.2010
</collectingDate>
</date>
;
<collectorName id="269A537FC61FFFDF121CF5886D87F851" box="[371,492,2004,2027]" pageId="25" pageNumber="118">D.M. Bento</collectorName>
leg.; ISLA59263
</materialsCitation>
.
</paragraph>
<paragraph id="8BD036A9C61FFFDF1042F4096873F96F" blockId="25.[811,1455,1620,1750]" pageId="25" pageNumber="118">
<materialsCitation id="3B073CF4C61FFFDF1042F409692CF9D6" box="[813,1351,1620,1645]" collectorName="Felipe Guerra" country="Brazil" location="Rio Grande do Norte" pageId="25" pageNumber="118" specimenCount="1" stateProvince="Rio Grande do Norte">
<emphasis id="B91BEABBC61FFFDF1042F4096F06F9D7" bold="true" box="[813,877,1621,1645]" pageId="25" pageNumber="118">
<collectingCountry id="F3787639C61FFFDF1042F4096F02F9D7" box="[813,873,1621,1645]" name="Brazil" pageId="25" pageNumber="118">Brazil</collectingCountry>
.
</emphasis>
<collectingRegion id="49ABF84BC61FFFDF101EF4086854F9D1" box="[881,1087,1620,1644]" country="Brazil" name="Rio Grande do Norte" pageId="25" pageNumber="118">Rio Grande do Norte</collectingRegion>
;
<collectorName id="269A537FC61FFFDF1729F40868A2F9D6" box="[1094,1225,1620,1644]" pageId="25" pageNumber="118">Felipe Guerra</collectorName>
municipality
</materialsCitation>
•
<materialsCitation id="3B073CF4C61FFFDF1635F4086FF7F91A" collectingDate="2010-06-04" collectionCode="VI" collectorName="D. M. Bento" country="Brazil" latitude="-5.560278" location="Crotes cave" longLatPrecision="21" longitude="-37.658337" pageId="25" pageNumber="118" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D69FD20C61FFFDF1635F4086916F9D1" box="[1370,1405,1620,1644]" count="1" pageId="25" pageNumber="118" type="male">1♂</specimenCount>
,
<specimenCount id="9D69FD20C61FFFDF16E7F40869C2F9D1" box="[1416,1449,1620,1644]" count="1" pageId="25" pageNumber="118" type="female">1 ♀</specimenCount>
;
<location id="8EB06072C61FFFDF1042F4326FF6F93C" LSID="urn:lsid:plazi:treatment:03C687BFC61FFFDA13A2F40E68C7FA77:8EB06072C61FFFDF1042F4326FF6F93C" box="[813,925,1646,1670]" country="Brazil" latitude="-5.560278" longLatPrecision="21" longitude="-37.658337" name="Crotes cave" pageId="25" pageNumber="118">Crotes cave</location>
;
<geoCoordinate id="EE5B506EC61FFFDF10C9F4326860F93C" box="[934,1035,1646,1670]" degrees="5" direction="south" minutes="33" orientation="latitude" pageId="25" pageNumber="118" precision="15" seconds="37" value="-5.560278">5°33’37”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC61FFFDF177AF43268FBF93C" box="[1045,1168,1646,1670]" degrees="37" direction="west" minutes="39" orientation="longitude" pageId="25" pageNumber="118" precision="15" seconds="30" value="-37.658337">37°39’30”W</geoCoordinate>
;
<date id="FFD11069C61FFFDF17F5F4326968F93C" box="[1178,1283,1646,1670]" pageId="25" pageNumber="118" value="2010-06-04">
<collectingDate id="EF95E981C61FFFDF17F5F4326968F93C" box="[1178,1283,1646,1670]" pageId="25" pageNumber="118" value="2010-06-04">
4.
<collectionCode id="ED7EAE6CC61FFFDF17C4F43268A3F93C" box="[1195,1224,1646,1670]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="25" pageNumber="118">VI</collectionCode>
.2010
</collectingDate>
</date>
;
<collectorName id="269A537FC61FFFDF1664F43369E9F93C" box="[1291,1410,1647,1670]" pageId="25" pageNumber="118">D.M. Bento</collectorName>
leg.; ISLA59330
</materialsCitation>
•
<materialsCitation id="3B073CF4C61FFFDF10C0F4D569C4F91A" box="[943,1455,1672,1697]" collectingDate="2010-06-04" collectionCode="VI" collectorName="D. M. Bento" country="Brazil" latitude="-5.560278" location="Crotes cave" longLatPrecision="21" longitude="-37.658337" pageId="25" pageNumber="118" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D69FD20C61FFFDF10C0F4D56FBAF91A" box="[943,977,1673,1697]" count="2" pageId="25" pageNumber="118" type="female">2 ♀</specimenCount>
; same collection data as for preceding; ISLA59282
</materialsCitation>
•
<materialsCitation id="3B073CF4C61FFFDF1052F4FF687FF96F" collectingDate="2010-08-04" collectorName="D. M. Bento" country="Brazil" latitude="-5.5625" location="Trapia cave" longLatPrecision="21" longitude="-37.62083" pageId="25" pageNumber="118" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D69FD20C61FFFDF1052F4FF6F08F900" box="[829,867,1699,1723]" count="1" pageId="25" pageNumber="118" type="male">1 ♂</specimenCount>
;
<location id="8EB06072C61FFFDF101EF4FF6F8FF901" LSID="urn:lsid:plazi:treatment:03C687BFC61FFFDA13A2F40E68C7FA77:8EB06072C61FFFDF101EF4FF6F8FF901" box="[881,996,1699,1723]" country="Brazil" latitude="-5.5625" longLatPrecision="21" longitude="-37.62083" name="Trapia cave" pageId="25" pageNumber="118">Trapiá cave</location>
;
<geoCoordinate id="EE5B506EC61FFFDF109FF4FF6833F901" box="[1008,1112,1699,1723]" degrees="5" direction="south" minutes="33" orientation="latitude" pageId="25" pageNumber="118" precision="15" seconds="45" value="-5.5625">5°33’45”S</geoCoordinate>
,
<geoCoordinate id="EE5B506EC61FFFDF170BF4FF688AF901" box="[1124,1249,1699,1723]" degrees="37" direction="west" minutes="37" orientation="longitude" pageId="25" pageNumber="118" precision="15" seconds="15" value="-37.62083">37°37’15”W</geoCoordinate>
;
<date id="FFD11069C61FFFDF1780F4FF6900F901" box="[1263,1387,1699,1723]" pageId="25" pageNumber="118" value="2010-08-04">
<collectingDate id="EF95E981C61FFFDF1780F4FF6900F901" box="[1263,1387,1699,1723]" pageId="25" pageNumber="118" value="2010-08-04">4.VIII.2010</collectingDate>
</date>
;
<collectorName id="269A537FC61FFFDF1618F4F86F02F96F" pageId="25" pageNumber="118">D.M. Bento</collectorName>
leg.; ISLA59270
</materialsCitation>
.
</paragraph>
<paragraph id="8BD036A9C61FFFDF1042F4A569C4F891" blockId="25.[813,1455,1784,1835]" pageId="25" pageNumber="118">
<smallCapsWord id="8D36A075C61FFFDF1042F4A56FCDF8B5" baselines="1802,1803" box="[813,934,1785,1808]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="25" pageNumber="118">ETYMOLOGY</smallCapsWord>
. — The new species name is a noun in reference to people who are born in the State of
<collectingRegion id="49ABF84BC61FFFDF17E1F54F690EF890" box="[1166,1381,1811,1835]" country="Brazil" name="Rio Grande do Norte" pageId="25" pageNumber="118">Rio Grande do Norte</collectingRegion>
,
<collectingCountry id="F3787639C61FFFDF1600F54F69C0F891" box="[1391,1451,1811,1835]" name="Brazil" pageId="25" pageNumber="118">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BD036A9C61FFFDF1042F51269C3F850" blockId="25.[811,1456,1870,2026]" pageId="25" pageNumber="118">
<smallCapsWord id="8D36A075C61FFFDF1042F5126FFCF8DE" baselines="1887,1888" box="[813,919,1870,1893]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="25" pageNumber="118">DIAGNOSIS</smallCapsWord>
. — Dorsum covered with round tubercles and ribs with three rows on cephalon (4, 4, 6); four rows on pereonite 1 (4, 5, 8, 9); two rows on pereonites 2-7 (8+ 7); one row of four tubercles on pleonites 3-5 with; two paramedian tubercles on telson. Pleopod 1 exopod with round distal portion, outer margin straight; endopod with distal portion bent outward, four times longer than exopod.
</paragraph>
<caption id="DF106621C61CFFDC13EAF47F6F8FF9D9" ID-DOI="http://doi.org/10.5281/zenodo.10803305" ID-Zenodo-Dep="10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="26" pageNumber="119" startId="26.[133,144,1571,1589]" targetBox="[151,1437,213,1527]" targetPageId="26" targetType="figure">
<paragraph id="8BD036A9C61CFFDC13EAF47F6F8FF9D9" blockId="26.[133,1457,1571,1635]" pageId="26" pageNumber="119">
<smallCapsWord id="8D36A075C61CFFDC13EAF47F6CF6F98E" baselines="1584,1585" box="[133,157,1571,1589]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="26" pageNumber="119">FIG</smallCapsWord>
. 18. —
<taxonomicName id="4C6F4D2AC61CFFDC13B3F47F6E7EF98F" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[220,533,1571,1589]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="26" pageNumber="119" phylum="Arthropoda" rank="species" species="potiguar" status="sp. nov.">
<emphasis id="B91BEABBC61CFFDC13B3F47F6D1EF98F" box="[220,373,1571,1589]" italics="true" pageId="26" pageNumber="119">Ctenorillo potiguar</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61CFFDC1174F47F6E27F98F" box="[539,588,1571,1589]" pageId="26" pageNumber="119" rank="species">n. sp.</taxonomicNameLabel>
,paratype (male,3 mm, ISLA96798):
<emphasis id="B91BEABBC61CFFDC101AF47F6FE9F98F" bold="true" box="[885,898,1571,1589]" pageId="26" pageNumber="119">A</emphasis>
, habitus,lateral view;
<emphasis id="B91BEABBC61CFFDC1759F47F6828F98F" bold="true" box="[1078,1091,1571,1589]" pageId="26" pageNumber="119">B</emphasis>
, disposition of dorsal tubercles;
<emphasis id="B91BEABBC61CFFDC1623F47F6931F98F" bold="true" box="[1356,1370,1571,1589]" pageId="26" pageNumber="119">C</emphasis>
, cephalon and epimeron 1, dorsal view;
<emphasis id="B91BEABBC61CFFDC1218F4666DEEF9F6" bold="true" box="[375,389,1594,1612]" pageId="26" pageNumber="119">D</emphasis>
, cephalon, frontal view;
<emphasis id="B91BEABBC61CFFDC1121F4666E31F9F6" bold="true" box="[590,602,1594,1612]" pageId="26" pageNumber="119">E</emphasis>
, epimera 1-3, ventral view;
<emphasis id="B91BEABBC61CFFDC1052F4666F23F9F6" bold="true" box="[829,840,1594,1612]" pageId="26" pageNumber="119">F</emphasis>
, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;
<emphasis id="B91BEABBC61CFFDC162FF4666925F9F6" bold="true" box="[1344,1358,1594,1612]" pageId="26" pageNumber="119">G</emphasis>
, antennula;
<emphasis id="B91BEABBC61CFFDC13EAF40D6CF8F9D9" bold="true" box="[133,147,1617,1635]" pageId="26" pageNumber="119">H</emphasis>
, antenna;
<emphasis id="B91BEABBC61CFFDC1385F40D6C9BF9D9" bold="true" box="[234,240,1617,1635]" pageId="26" pageNumber="119">I</emphasis>
, right mandible;
<emphasis id="B91BEABBC61CFFDC1213F40D6DEDF9D9" bold="true" box="[380,390,1617,1635]" pageId="26" pageNumber="119">J</emphasis>
, left mandible;
<emphasis id="B91BEABBC61CFFDC1168F40D6E7FF9D9" bold="true" box="[519,532,1617,1635]" pageId="26" pageNumber="119">K</emphasis>
, maxillula;
<emphasis id="B91BEABBC61CFFDC111EF40D6E17F9D9" bold="true" box="[625,636,1617,1635]" pageId="26" pageNumber="119">L</emphasis>
, maxilla;
<emphasis id="B91BEABBC61CFFDC11A4F40D6EB7F9D9" bold="true" box="[715,732,1617,1635]" pageId="26" pageNumber="119">M</emphasis>
, maxilliped. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD036A9C61CFFDC13EBF4CD6DBBF979" blockId="26.[132,775,1681,1731]" pageId="26" pageNumber="119">
<smallCapsWord id="8D36A075C61CFFDC13EBF4CD6D79F91D" baselines="1698,1699" box="[132,274,1681,1704]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="26" pageNumber="119">DISTRIBUTION</smallCapsWord>
. — Baraúna and Felipe Guerra municipalities, west of the
<collectingRegion id="49ABF84BC61CFFDC13ACF4F76DA0F979" box="[195,459,1707,1731]" country="Brazil" name="Rio Grande do Norte" pageId="26" pageNumber="119">Rio Grande do Norte state</collectingRegion>
.
</paragraph>
<paragraph id="8BD036A9C61CFFDC13EBF4AE6D7EF8B0" blockId="26.[130,777,1778,2028]" box="[132,277,1778,1804]" pageId="26" pageNumber="119">
<smallCapsWord id="8D36A075C61CFFDC13EBF4AE6D7EF8B0" baselines="1797,1797" box="[132,277,1778,1804]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="26" pageNumber="119">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BD036A9C61CFFDD13EBF54E6DD3F956" blockId="26.[130,777,1778,2028]" lastBlockId="27.[130,777,1459,1772]" lastPageId="27" lastPageNumber="120" pageId="26" pageNumber="119">
Maximum size:
<specimenCount id="9D69FD20C61CFFDC1252F5496D3FF891" box="[317,340,1813,1835]" count="1" pageId="26" pageNumber="119" type="male">♂</specimenCount>
,
<quantity id="4C979B4CC61CFFDC120AF54E6DC5F896" box="[357,430,1810,1836]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="119" unit="mm" value="3.0">3 mm</quantity>
,
<specimenCount id="9D69FD20C61CFFDC12D4F5496DA4F891" box="[443,463,1813,1835]" count="1" pageId="26" pageNumber="119" type="female">♀</specimenCount>
,
<quantity id="4C979B4CC61CFFDC128FF54E6E42F896" box="[480,553,1810,1836]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="119" unit="mm" value="4.0">4 mm</quantity>
. Color brown with lateral margins depigmented in nature (
<figureCitation id="13542A2CC61CFFDC1126F56D6EDDF8F6" box="[585,694,1841,1868]" captionStart="FIG" captionStartId="28.[133,144,1106,1124]" captionTargetBox="[132,1455,215,1063]" captionTargetId="figure-483@28.[132,1455,215,1063]" captionTargetPageId="28" captionText="FIG. 20. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: A, limestone outcrops at the Furna Feia National Park, with the entrance of the Furna Feia cave (yellow star); B, detail of the Furna Feia cave entrance; C, inner portion of the Furna Nova cave, where specimens are found; D, habitus in natural condition." figureDoi="http://doi.org/10.5281/zenodo.10803309" httpUri="https://zenodo.org/record/10803309/files/figure.png" pageId="26" pageNumber="119">Figs 20D</figureCitation>
;
<figureCitation id="13542A2CC61CFFDC11ABF56E6E97F8F6" box="[708,764,1842,1868]" captionStart="FIG" captionStartId="35.[132,143,1225,1243]" captionTargetBox="[132,1455,216,1183]" captionTargetId="figure-480@35.[699,983,655,744]" captionTargetPageId="35" captionText="FIG. 27. — Habitus of the species of the genus Ctenorillo Verhoeff, 1942 discussed in the present paper:A, Ctenorillo ferrarai Campos-Filho, Araujo &amp; Taiti,2014; B, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.; C, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.; D, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.; E, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.; F, Ctenorillo cearensis Cardoso &amp; Ferreira,n. sp.; G, Ctenorillo quiteriensis Cardoso &amp; Ferreira,n. sp.; H, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.; I, Ctenorillo iuiuensis n. sp.; J, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10803325" httpUri="https://zenodo.org/record/10803325/files/figure.png" pageId="26" pageNumber="119">27H</figureCitation>
). Dorsum covered with round tubercles and ribs (
<figureCitation id="13542A2CC61CFFDC11CFF50D6CF1F831" captionStart="FIG" captionStartId="26.[133,144,1571,1589]" captionTargetBox="[151,1437,213,1527]" captionTargetId="figure-218@26.[151,1437,213,1527]" captionTargetPageId="26" captionText="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="26" pageNumber="119">Fig. 18A, B</figureCitation>
): cephalon with three rows (4, 4, and 6 from front to back of vertex); pereonite 1 with
<quantity id="4C979B4CC61CFFDC12DDF5CD6D80F811" box="[434,491,1937,1963]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="26" pageNumber="119" unit="in" value="26.0">26 in</quantity>
four rows (4, 5, 8 and 9); pereonites 2-7 with 15 (8 +7); pleonites 3-5 with one row of four tubercles; telson with two paramedian tubercles. Dorsal cuticle with short triangular scale setae; pereonites 1-7 bearing one line of
<emphasis id="B91BEABBC61CFFDC10B7F4EE681CF976" box="[984,1143,1714,1740]" italics="true" pageId="26" pageNumber="119">noduli lateralis</emphasis>
per side on outer surface of posterior tubercle of second tubercles (
<figureCitation id="13542A2CC61CFFDC17BBF48E6944F956" box="[1236,1327,1746,1772]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="26" pageNumber="119">Fig. 28I</figureCitation>
). Cephalon with frontal shield protruding above vertex; eye consisting of five ommatidia (
<figureCitation id="13542A2CC61CFFDC1763F54E68CBF896" box="[1036,1184,1810,1836]" captionStart="FIG" captionStartId="26.[133,144,1571,1589]" captionTargetBox="[151,1437,213,1527]" captionTargetId="figure-218@26.[151,1437,213,1527]" captionTargetPageId="26" captionText="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="26" pageNumber="119">Figs 18C, D</figureCitation>
;
<figureCitation id="13542A2CC61CFFDC17C0F54E68B0F896" box="[1199,1243,1810,1836]" captionStart="FIG" captionStartId="36.[133,144,1473,1491]" captionTargetBox="[132,1455,216,1425]" captionTargetId="figure-264@36.[699,984,774,961]" captionTargetPageId="36" captionText="FIG. 28. — SEM images of Ctenorillo Verhoeff, 1942 new species: A-D, Ctenorillo pelado Cardoso &amp; Ferreira, n. sp.: cephalon, dorsal view (A); epimera 1-3, lateral view (B); pleopod 3 exopod (C); epimera 1 and 2, ventral view (D); E, Ctenorillo araguaia Cardoso &amp; Ferreira, n. sp.:, cephalon and epimera 1-3, lateral view; F, G, Ctenorillo intertidalis Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (F); epimera 1-3, lateral view (G); H, Ctenorillo ubajarensis Cardoso &amp; Ferreira,n. sp.: cephalon and epimera 1-3, lateral view;I, Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: cephalon and epimera 1 and 2, lateral view; J, K, Ctenorillo jequitinhonha Cardoso &amp; Ferreira, n. sp.: cephalon, frontal view (J); cephalon and epimera 1 and 2, lateral view (K). Scale bars: A, B, D-K, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10803327" httpUri="https://zenodo.org/record/10803327/files/figure.png" pageId="26" pageNumber="119">28I</figureCitation>
). Pereonite 1 with posterior margin sinuous at sides; inner lobe of schisma rounded, extending beyond posterior margin of outer lobe; pereonite 2 with triangular ventral tooth reaching posterior margin of epimera (
<figureCitation id="13542A2CC61CFFDC1093F5CD6837F811" box="[1020,1116,1937,1963]" captionStart="FIG" captionStartId="26.[133,144,1571,1589]" captionTargetBox="[151,1437,213,1527]" captionTargetId="figure-218@26.[151,1437,213,1527]" captionTargetPageId="26" captionText="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="26" pageNumber="119">Fig. 18E</figureCitation>
). Pereonites 2-7 (
<figureCitation id="13542A2CC61CFFDC167DF5CD691BF811" box="[1298,1392,1937,1963]" captionStart="FIG" captionStartId="26.[133,144,1571,1589]" captionTargetBox="[151,1437,213,1527]" captionTargetId="figure-218@26.[151,1437,213,1527]" captionTargetPageId="26" captionText="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="26" pageNumber="119">Fig. 18A</figureCitation>
) with wide quadrangular epimera slightly bent outwards. Telson (
<figureCitation id="13542A2CC61CFFDC105AF58D6FFAF851" box="[821,913,2001,2027]" captionStart="FIG" captionStartId="26.[133,144,1571,1589]" captionTargetBox="[151,1437,213,1527]" captionTargetId="figure-218@26.[151,1437,213,1527]" captionTargetPageId="26" captionText="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="26" pageNumber="119">Fig. 18F</figureCitation>
) hourglass-shaped, proximal portion broader than distal portion. Antennula (
<figureCitation id="13542A2CC61DFFDD12CFF7EF6E6AFA77" box="[416,513,1459,1485]" captionStart="FIG" captionStartId="26.[133,144,1571,1589]" captionTargetBox="[151,1437,213,1527]" captionTargetId="figure-218@26.[151,1437,213,1527]" captionTargetPageId="26" captionText="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="27" pageNumber="120">Fig. 18G</figureCitation>
) of three articles, second article shorter than first and third, third article with seven apical aesthetascs. Antenna (
<figureCitation id="13542A2CC61DFFDD12BFF7AF6E50F9B7" box="[464,571,1523,1549]" captionStart="FIG" captionStartId="26.[133,144,1571,1589]" captionTargetBox="[151,1437,213,1527]" captionTargetId="figure-218@26.[151,1437,213,1527]" captionTargetPageId="26" captionText="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="27" pageNumber="120">Fig. 18H</figureCitation>
) short and stout, flagellum shorter than fifth article of peduncle; second article of flagellum about three times as long as first. Buccal pieces as
<taxonomicName id="4C6F4D2AC61DFFDD1388F40F6E43F9D6" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[231,552,1618,1645]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="27" pageNumber="120" phylum="Arthropoda" rank="species" species="pelado" status="n. sp.">
<emphasis id="B91BEABBC61DFFDD1388F40F6D20F9D6" box="[231,331,1618,1645]" italics="true" pageId="27" pageNumber="120">C. pelado</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61DFFDD115CF40F6E05F9D7" box="[563,622,1619,1645]" pageId="27" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13542A2CC61DFFDD1113F40E6EA2F9D6" box="[636,713,1618,1644]" captionStart="FIG" captionStartId="26.[133,144,1571,1589]" captionTargetBox="[151,1437,213,1527]" captionTargetId="figure-218@26.[151,1437,213,1527]" captionTargetPageId="26" captionText="FIG. 18. — Ctenorillo potiguar Cardoso &amp; Ferreira,n. sp.,paratype (male,3 mm, ISLA96798):A, habitus,lateral view;B, disposition of dorsal tubercles;C, cephalon and epimeron 1, dorsal view; D, cephalon, frontal view; E, epimera 1-3, ventral view; F, epimeron 7, pleonites 3-5, uropods and telson, dorsal view;G, antennula; H, antenna; I, right mandible; J, left mandible; K, maxillula; L, maxilla; M, maxilliped. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10803305" httpUri="https://zenodo.org/record/10803305/files/figure.png" pageId="27" pageNumber="120">Fig. 18</figureCitation>
I-M). All pleopod exopods with monospiracular covered lungs. Uropod (
<figureCitation id="13542A2CC61DFFDD1381F4CE6D39F916" box="[238,338,1682,1708]" captionStart="FIG" captionStartId="27.[132,143,1340,1358]" captionTargetBox="[137,1428,218,1293]" captionTargetId="figure-339@27.[132,1455,210,1301]" captionTargetPageId="27" captionText="FIG. 19. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp., paratype (male, 3 mm, ISLA96798): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803307" httpUri="https://zenodo.org/record/10803307/files/figure.png" pageId="27" pageNumber="120">Fig. 19A</figureCitation>
) protopod flattened, enlarged on basal portion; exopod outward, short, inserted dorsally close to medial margin of protopod.
</paragraph>
<caption id="DF106621C61DFFDD13EBF7606F0CFADF" ID-DOI="http://doi.org/10.5281/zenodo.10803307" ID-Zenodo-Dep="10803307" httpUri="https://zenodo.org/record/10803307/files/figure.png" pageId="27" pageNumber="120" startId="27.[132,143,1340,1358]" targetBox="[137,1428,218,1293]" targetPageId="27" targetType="figure">
<paragraph id="8BD036A9C61DFFDD13EBF7606F0CFADF" blockId="27.[132,1455,1340,1381]" pageId="27" pageNumber="120">
<smallCapsWord id="8D36A075C61DFFDD13EBF7606CF6FAF7" baselines="1353,1354" box="[132,157,1340,1358]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="27" pageNumber="120">FIG</smallCapsWord>
. 19. —
<taxonomicName id="4C6F4D2AC61DFFDD13B1F7606E70FAF4" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[222,539,1340,1358]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="27" pageNumber="120" phylum="Arthropoda" rank="species" species="potiguar" status="sp. nov.">
<emphasis id="B91BEABBC61DFFDD13B1F7606D13FAF4" box="[222,376,1340,1358]" italics="true" pageId="27" pageNumber="120">Ctenorillo potiguar</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61DFFDD114DF7606E3FFAF4" box="[546,596,1340,1358]" pageId="27" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
, paratype (male, 3 mm, ISLA96798):
<emphasis id="B91BEABBC61DFFDD10ECF7606FFBFAF4" bold="true" box="[899,912,1340,1358]" pageId="27" pageNumber="120">A</emphasis>
, uropod,
<emphasis id="B91BEABBC61DFFDD108FF7606F86FAF4" bold="true" box="[992,1005,1340,1358]" pageId="27" pageNumber="120">B</emphasis>
, pereopod 1;
<emphasis id="B91BEABBC61DFFDD170EF7606804FAF4" bold="true" box="[1121,1135,1340,1358]" pageId="27" pageNumber="120">C</emphasis>
, pereopod 7;
<emphasis id="B91BEABBC61DFFDD178CF760689AFAF4" bold="true" box="[1251,1265,1340,1358]" pageId="27" pageNumber="120">D</emphasis>
, pleopod 1 and genital papilla;
<emphasis id="B91BEABBC61DFFDD13AAF70F6CBAFADF" bold="true" box="[197,209,1363,1381]" pageId="27" pageNumber="120">E</emphasis>
, pleopod 2;
<emphasis id="B91BEABBC61DFFDD1255F70F6D2EFADF" bold="true" box="[314,325,1363,1381]" pageId="27" pageNumber="120">F</emphasis>
, pleopod 3 exopod;
<emphasis id="B91BEABBC61DFFDD129CF70E6E6AFADE" bold="true" box="[499,513,1362,1380]" pageId="27" pageNumber="120">G</emphasis>
, pleopod 4 exopod;
<emphasis id="B91BEABBC61DFFDD11C0F70F6ED6FADF" bold="true" box="[687,701,1363,1381]" pageId="27" pageNumber="120">H</emphasis>
, pleopod 5 exopod.
</paragraph>
</caption>
<paragraph id="8BD036A9C61DFFDD13EBF54E6CD2F896" blockId="27.[132,776,1810,2028]" box="[132,185,1810,1836]" pageId="27" pageNumber="120">
<emphasis id="B91BEABBC61DFFDD13EBF54E6CD2F896" box="[132,185,1810,1836]" italics="true" pageId="27" pageNumber="120">Male</emphasis>
</paragraph>
<paragraph id="8BD036A9C61DFFDD13EBF56D6D31F851" blockId="27.[132,776,1810,2028]" pageId="27" pageNumber="120">
Pereopods without particular modifications (
<figureCitation id="13542A2CC61DFFDD111DF56D6E92F8F6" box="[626,761,1841,1868]" captionStart="FIG" captionStartId="27.[132,143,1340,1358]" captionTargetBox="[137,1428,218,1293]" captionTargetId="figure-339@27.[132,1455,210,1301]" captionTargetPageId="27" captionText="FIG. 19. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp., paratype (male, 3 mm, ISLA96798): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803307" httpUri="https://zenodo.org/record/10803307/files/figure.png" pageId="27" pageNumber="120">Fig. 19B, C</figureCitation>
). Pleopod 1 exopod (
<figureCitation id="13542A2CC61DFFDD1236F50D6DD7F8D6" box="[345,444,1873,1900]" captionStart="FIG" captionStartId="27.[132,143,1340,1358]" captionTargetBox="[137,1428,218,1293]" captionTargetId="figure-339@27.[132,1455,210,1301]" captionTargetPageId="27" captionText="FIG. 19. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp., paratype (male, 3 mm, ISLA96798): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803307" httpUri="https://zenodo.org/record/10803307/files/figure.png" pageId="27" pageNumber="120">Fig. 19D</figureCitation>
) wider than long, round distal portion, outer margin straight; endopod with distal portion bent outward, four times longer than exopod. Pleopod 2 endopod longer than exopod (
<figureCitation id="13542A2CC61DFFDD12C3F5ED6E67F871" box="[428,524,1969,1995]" captionStart="FIG" captionStartId="27.[132,143,1340,1358]" captionTargetBox="[137,1428,218,1293]" captionTargetId="figure-339@27.[132,1455,210,1301]" captionTargetPageId="27" captionText="FIG. 19. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp., paratype (male, 3 mm, ISLA96798): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803307" httpUri="https://zenodo.org/record/10803307/files/figure.png" pageId="27" pageNumber="120">Fig. 19E</figureCitation>
). Pleopods 3-5 exopods as in
<figureCitation id="13542A2CC61DFFDD13D2F58D6D48F851" box="[189,291,2001,2027]" captionStart="FIG" captionStartId="27.[132,143,1340,1358]" captionTargetBox="[137,1428,218,1293]" captionTargetId="figure-339@27.[132,1455,210,1301]" captionTargetPageId="27" captionText="FIG. 19. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp., paratype (male, 3 mm, ISLA96798): A, uropod, B, pereopod 1; C, pereopod 7; D, pleopod 1 and genital papilla; E, pleopod 2; F, pleopod 3 exopod; G, pleopod 4 exopod; H, pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.10803307" httpUri="https://zenodo.org/record/10803307/files/figure.png" pageId="27" pageNumber="120">Figure 19</figureCitation>
F-H.
</paragraph>
<paragraph id="8BD036A9C61DFFDD1042F7EF6FE6FA71" blockId="27.[810,1458,1459,2028]" box="[813,909,1459,1485]" pageId="27" pageNumber="120">
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</paragraph>
<paragraph id="8BD036A9C61DFFDD1042F78F688DF816" blockId="27.[810,1458,1459,2028]" pageId="27" pageNumber="120">
Specimens of
<taxonomicName id="4C6F4D2AC61DFFDD10ABF78F6903FA57" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[964,1384,1491,1517]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="27" pageNumber="120" phylum="Arthropoda" rank="species" species="potiguar" status="sp. nov.">
<emphasis id="B91BEABBC61DFFDD10ABF78F68E2FA57" box="[964,1161,1491,1517]" italics="true" pageId="27" pageNumber="120">Ctenorillo potiguar</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61DFFDD161BF78F69C4FA57" box="[1396,1455,1491,1517]" pageId="27" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
were found in six caves located in the west of the
<collectingRegion id="49ABF84BC61DFFDD165DF7AF6FA4F997" country="Brazil" name="Rio Grande do Norte" pageId="27" pageNumber="120">Rio Grande do Norte state</collectingRegion>
. This region has more than 1000 currently known caves (
<collectionCode id="ED7EAE6CC61DFFDD10D0F46F687AF9F7" box="[959,1041,1587,1613]" pageId="27" pageNumber="120">CANIE</collectionCode>
/
<collectionCode id="ED7EAE6CC61DFFDD1776F46F6818F9F7" box="[1049,1139,1587,1613]" pageId="27" pageNumber="120">CECAV</collectionCode>
2022), but only about 48 have been inventoried (
<bibRefCitation id="EFFE4B58C61DFFDD1084F40E68D9F9D6" author="FERREIRA R. L. &amp; PROUS X. &amp; BERNARDI L. F. O. &amp; SOUZA-SILVA M." box="[1003,1202,1618,1644]" pageId="27" pageNumber="120" pagination="25 - 51" refId="ref32576" refString="FERREIRA R. L., PROUS X., BERNARDI L. F. O. &amp; SOUZA-SILVA M. 2010. - Fauna subterranea do Estado do Rio Grande do Norte: Caracterizacao e impactos. Revista Brasileira de Espeleologia 1: 25 - 51." type="journal article" year="2010">
Ferreira
<emphasis id="B91BEABBC61DFFDD172BF40F681DF9D6" box="[1092,1142,1618,1644]" italics="true" pageId="27" pageNumber="120">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EFFE4B58C61DFFDD17D1F40F6918F9D6" author="BENTO D. &amp; DE M. &amp; SOUZA-SILVA M. &amp; VASCONCELLOS A. &amp; BELLINI B. C. &amp; PROUS X. &amp; FERREIRA R. L." box="[1214,1395,1618,1645]" pageId="27" pageNumber="120" pagination="3837 - 3857" refId="ref31253" refString="BENTO D. DE M., SOUZA-SILVA M., VASCONCELLOS A., BELLINI B. C., PROUS X. &amp; FERREIRA R. L. 2021. - Subterranean &quot; oasis &quot; in the Brazilian semiarid region: neglected sources of biodiversity. Biodiversity and Conservation 30: 3837 - 3857. https: // doi. org / 10.1007 / s 10531 - 021 - 02277 - 6." type="journal article" year="2021">
Bento
<emphasis id="B91BEABBC61DFFDD166BF40F695FF9D6" box="[1284,1332,1618,1644]" italics="true" pageId="27" pageNumber="120">et al.</emphasis>
2021
</bibRefCitation>
). The caves are inserted in the limestones of the Janaíra Formation, deposited during the Upper Cretaceous, which corresponds to a carbonate ramp that emerged along the emersed portion of the Potiguar Basin (
<bibRefCitation id="EFFE4B58C61DFFDD174DF48E6885F956" author="BEZERRA F. H. R. &amp; TAKEYA M. K. &amp; SOUSA M. O. L. &amp; NASCIMENTO A. F." box="[1058,1262,1746,1772]" pageId="27" pageNumber="120" pagination="27 - 39" refId="ref31328" refString="BEZERRA F. H. R., TAKEYA M. K., SOUSA M. O. L. &amp; NASCIMENTO A. F. 2007. - Coseismic reactivation of the Samambaia fault, Brazil. Tectonophysics 430: 27 - 39. https: // doi. org / 10.1016 / j. tecto. 2006.10.007" type="journal article" year="2007">
Bezerra
<emphasis id="B91BEABBC61DFFDD1716F48F68C5F956" box="[1145,1198,1746,1772]" italics="true" pageId="27" pageNumber="120">et al.</emphasis>
2007
</bibRefCitation>
). The region is in the Caatinga Biome, with a hot and semiarid climate (BSh on the Köppen climate classification) (
<bibRefCitation id="EFFE4B58C61DFFDD17BAF54E69CBF896" author="ALVARES C. A. &amp; STAPE J. L. &amp; SENTELHAS P. C. &amp; MORAES G. &amp; DE &amp; LEONARDO J. &amp; SPAROVEK G." box="[1237,1440,1810,1836]" pageId="27" pageNumber="120" pagination="711 - 728" refId="ref31163" refString="ALVARES C. A., STAPE J. L., SENTELHAS P. C., MORAES G. DE, LEONARDO J. &amp; SPAROVEK G. 2013. - Koppen's climate classification map for Brazil. Meteorologische Zeitschrift 22 (6): 711 - 728. https: // doi. org / 10.1127 / 0941 - 2948 / 2013 / 0507" type="journal article" year="2013">
Alvares
<emphasis id="B91BEABBC61DFFDD1645F54E6934F896" box="[1322,1375,1810,1836]" italics="true" pageId="27" pageNumber="120">et al.</emphasis>
2013
</bibRefCitation>
). The average temperature is constant over the year, ranging from 25° to 30°C, with average annual precipitation around
<quantity id="4C979B4CC61DFFDD1042F52D6FE2F836" box="[813,905,1905,1932]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="120" unit="mm" value="800.0">800 mm</quantity>
(concentrated from February to June, but 70% can fall in a single month) (
<bibRefCitation id="EFFE4B58C61DFFDD1744F5CD68BCF811" author="SILVA J. M. C. &amp; LEAL I. &amp; TABARELLI M." box="[1067,1239,1937,1963]" pageId="27" pageNumber="120" refId="ref34056" refString="SILVA J. M. C., LEAL I. &amp; TABARELLI M. 2017. - Caatinga: the largest tropical dry forest region in South America, first ed. Springer, New York, 482 p. https: // doi. org / 10.1007 / 978 - 3 - 319 - 68339 - 3" type="book" year="2017">
Silva
<emphasis id="B91BEABBC61DFFDD170BF5CE68FCF811" box="[1124,1175,1937,1963]" italics="true" pageId="27" pageNumber="120">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD036A9C61DFFDA102CF5ED6D9BF936" blockId="27.[810,1458,1459,2028]" lastBlockId="28.[130,777,1236,2028]" lastPageId="28" lastPageNumber="121" pageId="27" pageNumber="120">
Three caves, Furna Feia (
<figureCitation id="13542A2CC61DFFDD1753F5ED68D8F871" box="[1084,1203,1969,1995]" captionStart="FIG" captionStartId="28.[133,144,1106,1124]" captionTargetBox="[132,1455,215,1063]" captionTargetId="figure-483@28.[132,1455,215,1063]" captionTargetPageId="28" captionText="FIG. 20. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: A, limestone outcrops at the Furna Feia National Park, with the entrance of the Furna Feia cave (yellow star); B, detail of the Furna Feia cave entrance; C, inner portion of the Furna Nova cave, where specimens are found; D, habitus in natural condition." figureDoi="http://doi.org/10.5281/zenodo.10803309" httpUri="https://zenodo.org/record/10803309/files/figure.png" pageId="27" pageNumber="120">Fig. 20A, B</figureCitation>
), Furna Nova (
<figureCitation id="13542A2CC61DFFDD1624F5ED69C3F871" box="[1355,1448,1969,1995]" captionStart="FIG" captionStartId="28.[133,144,1106,1124]" captionTargetBox="[132,1455,215,1063]" captionTargetId="figure-483@28.[132,1455,215,1063]" captionTargetPageId="28" captionText="FIG. 20. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: A, limestone outcrops at the Furna Feia National Park, with the entrance of the Furna Feia cave (yellow star); B, detail of the Furna Feia cave entrance; C, inner portion of the Furna Nova cave, where specimens are found; D, habitus in natural condition." figureDoi="http://doi.org/10.5281/zenodo.10803309" httpUri="https://zenodo.org/record/10803309/files/figure.png" pageId="27" pageNumber="120">Fig.20C</figureCitation>
) and Macacos/Esquecida, are located in the Furna Feia National Park (
<collectionCode id="ED7EAE6CC61AFFDA1263F6886D38FB54" box="[268,339,1236,1262]" pageId="28" pageNumber="121">FFNP</collectionCode>
) in the municipality of Baraúna. Specimens were also found in the Britador cave, near
<collectionCode id="ED7EAE6CC61AFFDA11CDF6A86E8FFAB4" box="[674,740,1268,1294]" pageId="28" pageNumber="121">FFNP</collectionCode>
, in Baraúna. Other specimens were found in Trapiá and Crotes caves, in the municipality of Felipe Guerra (
<collectionCode id="ED7EAE6CC61AFFDA1116F76F6ECDFAF7" box="[633,678,1331,1357]" pageId="28" pageNumber="121">RN</collectionCode>
),
<quantity id="4C979B4CC61AFFDA11D2F76F6F6DFAF7" box="[701,774,1331,1357]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="121" unit="km" value="60.0">60 km</quantity>
from the first ones. These caves exhibit varied morphology, dimensions, and environmental conditions, ranging from small photic caves spanning a few tens of meters (e.g. Britador cave) to large aphotic caves covering hundreds of meters and characterized by more stable climatic conditions (e.g. Furna Feia, Furna Nova, and Trapiá caves). Despite this distribution, individuals (
<figureCitation id="13542A2CC61AFFDA12F2F44E6E6FF997" box="[413,516,1554,1581]" captionStart="FIG" captionStartId="28.[133,144,1106,1124]" captionTargetBox="[132,1455,215,1063]" captionTargetId="figure-483@28.[132,1455,215,1063]" captionTargetPageId="28" captionText="FIG. 20. — Ctenorillo potiguar Cardoso &amp; Ferreira, n. sp.: A, limestone outcrops at the Furna Feia National Park, with the entrance of the Furna Feia cave (yellow star); B, detail of the Furna Feia cave entrance; C, inner portion of the Furna Nova cave, where specimens are found; D, habitus in natural condition." figureDoi="http://doi.org/10.5281/zenodo.10803309" httpUri="https://zenodo.org/record/10803309/files/figure.png" pageId="28" pageNumber="121">Fig. 20D</figureCitation>
) were mainly found in aphotic and humid areas within the caves, often sheltered under rocks. This observation suggests that the species may have a specific habitat preference.
</paragraph>
<caption id="DF106621C61AFFDA13EAF60E6933FBC1" ID-DOI="http://doi.org/10.5281/zenodo.10803309" ID-Zenodo-Dep="10803309" httpUri="https://zenodo.org/record/10803309/files/figure.png" pageId="28" pageNumber="121" startId="28.[133,144,1106,1124]" targetBox="[132,1455,215,1063]" targetPageId="28" targetType="figure">
<paragraph id="8BD036A9C61AFFDA13EAF60E6933FBC1" blockId="28.[133,1455,1106,1147]" pageId="28" pageNumber="121">
<smallCapsWord id="8D36A075C61AFFDA13EAF60E6CF6FBD9" baselines="1119,1120" box="[133,157,1106,1124]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="28" pageNumber="121">FIG</smallCapsWord>
. 20. —
<taxonomicName id="4C6F4D2AC61AFFDA13B1F60E6E70FBDE" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[222,539,1106,1124]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="28" pageNumber="121" phylum="Arthropoda" rank="species" species="potiguar" status="sp. nov.">
<emphasis id="B91BEABBC61AFFDA13B1F60E6D13FBDE" box="[222,376,1106,1124]" italics="true" pageId="28" pageNumber="121">Ctenorillo potiguar</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61AFFDA114CF60E6E3EFBDE" box="[547,597,1106,1124]" pageId="28" pageNumber="121" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B91BEABBC61AFFDA1134F60E6E03FBDE" bold="true" box="[603,616,1106,1124]" pageId="28" pageNumber="121">A</emphasis>
, limestone outcrops at the Furna Feia National Park, with the entrance of the Furna Feia cave (
<emphasis id="B91BEABBC61AFFDA1619F60E6CC3FBC1" bold="true" pageId="28" pageNumber="121">yellow star</emphasis>
);
<emphasis id="B91BEABBC61AFFDA13D9F6356CA8FBC1" bold="true" box="[182,195,1129,1147]" pageId="28" pageNumber="121">B</emphasis>
, detail of the Furna Feia cave entrance;
<emphasis id="B91BEABBC61AFFDA117CF6356E4AFBC1" bold="true" box="[531,545,1129,1147]" pageId="28" pageNumber="121">C</emphasis>
, inner portion of the Furna Nova cave, where specimens are found;
<emphasis id="B91BEABBC61AFFDA1738F635680EFBC1" bold="true" box="[1111,1125,1129,1147]" pageId="28" pageNumber="121">D</emphasis>
, habitus in natural condition.
</paragraph>
</caption>
<paragraph id="8BD036A9C61AFFDA13F4F4CF6E96F856" blockId="28.[130,777,1236,2028]" pageId="28" pageNumber="121">No samplings were conducted outside caves in the area, and it is likely that the species also occurs in surface habitats (especially considering the absence of any obvious troglomorphic traits). Therefore, further studies including external samplings are needed to better understand the species distribution. Although only part of the distribution of the new species occurs within a conservation unit, the other caves are found in relatively conserved areas, without direct anthropic impacts. Three caves (Furna Feia, Furna Nova, and Crotes Caves) are visited by tourists in an orderly manner. Thus, the conservation status of the species can be considered of less concern.</paragraph>
<paragraph id="8BD036A9C61AFFDA1042F6886FECFB56" blockId="28.[811,1455,1236,1485]" box="[813,903,1236,1262]" pageId="28" pageNumber="121">
<smallCapsWord id="8D36A075C61AFFDA1042F6886FECFB56" baselines="1255,1255" box="[813,903,1236,1262]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remark" pageId="28" pageNumber="121">REMARK</smallCapsWord>
</paragraph>
<paragraph id="8BD036A9C61AFFDA1042F6AF68C7FA77" blockId="28.[811,1455,1236,1485]" pageId="28" pageNumber="121">
<taxonomicName id="4C6F4D2AC61AFFDA1042F6AF6880FAB7" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[813,1259,1267,1294]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="28" pageNumber="121" phylum="Arthropoda" rank="species" species="potiguar" status="sp. nov.">
<emphasis id="B91BEABBC61AFFDA1042F6AF6F94FAB7" box="[813,1023,1267,1293]" italics="true" pageId="28" pageNumber="121">Ctenorillo potiguar</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61AFFDA1795F6A86952FAB4" box="[1274,1337,1268,1294]" pageId="28" pageNumber="121" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="4C6F4D2AC61AFFDA1040F74868FBFA97" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[815,1168,1299,1326]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="28" pageNumber="121" phylum="Arthropoda" rank="species" species="ubajarensis" status="n. sp.">
<emphasis id="B91BEABBC61AFFDA1040F7486FD5FA97" box="[815,958,1299,1326]" italics="true" pageId="28" pageNumber="121">C. ubajarensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61AFFDA17F7F74868BAFA94" box="[1176,1233,1300,1326]" pageId="28" pageNumber="121" rank="species">n. sp.</taxonomicNameLabel>
by the less developed tubercles, however they are more prominent on
<taxonomicName id="4C6F4D2AC61AFFDA162BF76F6835FAD7" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="28" pageNumber="121" phylum="Arthropoda" rank="species" species="potiguar" status="sp. nov.">
<emphasis id="B91BEABBC61AFFDA162BF76F6FE8FAD7" italics="true" pageId="28" pageNumber="121">Ctenorillo potiguar</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61AFFDA1707F70868C9FAD4" box="[1128,1186,1364,1390]" pageId="28" pageNumber="121" rank="species">n. sp.</taxonomicNameLabel>
By the number of tubercles on pereonite 1,
<taxonomicName id="4C6F4D2AC61AFFDA1767F72F69C7FA37" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[1032,1452,1395,1421]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="28" pageNumber="121" phylum="Arthropoda" rank="species" species="potiguar" status="sp. nov.">
<emphasis id="B91BEABBC61AFFDA1767F72F68A6FA37" box="[1032,1229,1395,1421]" italics="true" pageId="28" pageNumber="121">Ctenorillo potiguar</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61AFFDA1042F7C86F0DFA14" box="[813,870,1428,1454]" pageId="28" pageNumber="121" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="4C6F4D2AC61AFFDA10B9F7CF6951FA17" authority="Cardoso &amp; Ferreira" authorityName="Cardoso &amp; Ferreira" authorityYear="2024" box="[982,1338,1427,1453]" class="Malacostraca" family="Armadillidae" genus="Ctenorillo" kingdom="Animalia" order="Isopoda" pageId="28" pageNumber="121" phylum="Arthropoda" rank="species" species="quiteriensis" status="n. sp.">
<emphasis id="B91BEABBC61AFFDA10B9F7CF680DFA17" box="[982,1126,1427,1453]" italics="true" pageId="28" pageNumber="121">C. quiteriensis</emphasis>
Cardoso &amp; Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A22857C0C61AFFDA162DF7C86911FA14" box="[1346,1402,1428,1454]" pageId="28" pageNumber="121" rank="species">n. sp.</taxonomicNameLabel>
with 26 tubercles, differing by the shape.
</paragraph>
</subSubSection>
</treatment>
</document>
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