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<document id="40CA64F1F88991B53AF16F032F206B2F" ID-DOI="10.5281/zenodo.184909" ID-GBIF-Dataset="c08e7d43-53e3-49f0-9567-62b426246a40" ID-ISSN="1175-5326" ID-Zenodo-Dep="184909" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460029948373" checkinUser="plazi" docAuthor="Brown, Brian V. &amp; Oliver, Hugh" docDate="2008" docId="03C6879CD570FF9FFF6345CE2CA76011" docLanguage="en" docName="zt01933p011.pdf" docOrigin="Zootaxa 1933" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Wharia Brown &amp; Oliver, 2008, n. gen." docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FFFFFFE4D570FF9BFFF443232E4C6747" masterDocTitle="Two new genera of Phoridae (Diptera) from New Zealand" masterLastPageNumber="11" masterPageNumber="1" pageNumber="1" updateTime="1698234100721" updateUser="plazi">
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<mods:title id="68B166A04E439CCE782848C417133E50">Two new genera of Phoridae (Diptera) from New Zealand</mods:title>
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<mods:namePart id="4BB87D2226D482274A21233E01BF6333">Brown, Brian V.</mods:namePart>
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<mods:namePart id="778E53D15EC84B2EA302C7F34CA99A6E">Oliver, Hugh</mods:namePart>
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<mods:date id="AA31C1B8315BC0D89AF9989C71730057">2008</mods:date>
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<mods:number id="FEAFDEC8516ED4F6ABEEA86963F7B13F">1933</mods:number>
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<mods:identifier id="544577861DFBF8AAB2994FFB0A5960A9" type="DOI">10.5281/zenodo.184909</mods:identifier>
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<mods:identifier id="0D325872E6F70722A54B8C3FE739405E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C6879CD570FF9FFF6345CE2CA76011" ID-DOI="http://doi.org/10.5281/zenodo.6231753" ID-GBIF-Taxon="119382354" ID-Zenodo-Dep="6231753" LSID="urn:lsid:plazi:treatment:03C6879CD570FF9FFF6345CE2CA76011" httpUri="http://treatment.plazi.org/id/03C6879CD570FF9FFF6345CE2CA76011" lastPageId="4" lastPageNumber="5" pageId="0" pageNumber="1">
<subSubSection id="C3756501D570FF9BFF6345CE2F6A606E" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="8BD0368AD570FF9BFF6345CE2F076041" blockId="0.[151,331,1772,1833]" box="[151,331,1772,1799]" pageId="0" pageNumber="1">
<heading id="D09881E6D570FF9BFF6345CE2F076041" bold="true" box="[151,331,1772,1799]" fontSize="11" level="1" pageId="0" pageNumber="1" reason="1">
<emphasis id="B91BEA98D570FF9BFF6345CE2F076041" bold="true" box="[151,331,1772,1799]" pageId="0" pageNumber="1">
<taxonomicName id="4C6F4D09D570FF9BFF6345CE2EBE6040" ID-CoL="9HP4Z" box="[151,242,1773,1799]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91BEA98D570FF9BFF6345CE2EBE6040" bold="true" box="[151,242,1773,1799]" italics="true" pageId="0" pageNumber="1">Wharia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857E3D570FF9BFF0D45CF2F076041" box="[249,331,1772,1798]" pageId="0" pageNumber="1" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0368AD570FF9BFF63442C2F6A606E" blockId="0.[151,331,1772,1833]" box="[151,294,1807,1833]" pageId="0" pageNumber="1">
(
<figureCitation id="13542A0FD570FF9BFF6B442C2F51606E" box="[159,285,1807,1833]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 12" captionStart-2="FIGURES 13 – 16" captionStartId-0="1.[151,269,1856,1880]" captionStartId-1="2.[151,270,1679,1703]" captionStartId-2="3.[151,269,1926,1950]" captionTargetBox-0="[151,1436,357,1822]" captionTargetBox-1="[151,1436,194,1651]" captionTargetBox-2="[210,1367,187,1886]" captionTargetId-0="figure@1.[151,1436,357,1830]" captionTargetId-1="figure@2.[151,1436,194,1655]" captionTargetId-2="figure@3.[195,1382,159,1904]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 1 – 6. Wharia willcocksorum new species, specimen from Rotopounamu [LACM ENT 223564]. 1, habitus, left lateral. 2, head and thorax, left lateral. 3, head, anterior. 4, thorax, dorsal. 5, hind femur, posterior. 6, hind femur, posterior, basal area." captionText-1="FIGURES 7 – 12. Wharia willcocksorum new species, hind femur, posterior, and hind femur, posterior, basal area. 7 – 8, specimen from Tree Trunk Gorge [LACM ENT 221624]. 9 – 10, specimen from Raglan [LACM ENT 223165]. 11 – 12, specimen from Desert Road at Oturere River [LACM ENT 221483]." captionText-2="FIGURES 13 – 16. Wharia willcocksorum new species, male. 13, head, specimen from Rotopounamu [LACM ENT 223564]. 14, wing, specimen from Tree Trunk Gorge [LACM ENT 221624]. 15 – 16, male terminalia, right lateral and left lateral, specimen (molecular voucher) from Sandy Bay [LACM ENT 240024]." httpUri-0="https://zenodo.org/record/184910/files/figure.png" httpUri-1="https://zenodo.org/record/184912/files/figure.png" httpUri-2="https://zenodo.org/record/184913/files/figure.png" pageId="0" pageNumber="1">Figs. 1–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3756501D570FF9FFF63447C2CA76011" lastPageId="4" lastPageNumber="5" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="8BD0368AD570FF9AFF63447C2DB5666D" blockId="0.[151,1436,1887,2037]" lastBlockId="1.[151,1437,152,298]" lastPageId="1" lastPageNumber="2" pageId="0" pageNumber="1">
<emphasis id="B91BEA98D570FF9BFF63447C2F5B603E" bold="true" box="[151,279,1887,1913]" pageId="0" pageNumber="1">Diagnosis.</emphasis>
Frons lacking medial furrow except for small pit between supra-antennal setae (
<figureCitation id="13542A0FD570FF9BFB39447C2B69603E" box="[1229,1317,1887,1913]" captionStart="FIGURES 13 – 16" captionStartId="3.[151,269,1926,1950]" captionTargetBox="[210,1367,187,1886]" captionTargetId="figure@3.[195,1382,159,1904]" captionTargetPageId="3" captionText="FIGURES 13 – 16. Wharia willcocksorum new species, male. 13, head, specimen from Rotopounamu [LACM ENT 223564]. 14, wing, specimen from Tree Trunk Gorge [LACM ENT 221624]. 15 – 16, male terminalia, right lateral and left lateral, specimen (molecular voucher) from Sandy Bay [LACM ENT 240024]." httpUri="https://zenodo.org/record/184913/files/figure.png" pageId="0" pageNumber="1">Fig. 13</figureCitation>
); one pair of reclinate supra-antennal setae present; ventral fronto-orbital setae absent (
<figureCitation id="13542A0FD570FF9BFBEE44A42A2060E6" box="[1050,1132,1927,1953]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1856,1880]" captionTargetBox="[151,1436,357,1822]" captionTargetId="figure@1.[151,1436,357,1830]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Wharia willcocksorum new species, specimen from Rotopounamu [LACM ENT 223564]. 1, habitus, left lateral. 2, head and thorax, left lateral. 3, head, anterior. 4, thorax, dorsal. 5, hind femur, posterior. 6, hind femur, posterior, basal area." httpUri="https://zenodo.org/record/184910/files/figure.png" pageId="0" pageNumber="1">Figs. 3</figureCitation>
,
<figureCitation id="13542A0FD570FF9BFB8E44A42AD660E6" box="[1146,1178,1927,1953]" captionStart="FIGURES 13 – 16" captionStartId="3.[151,269,1926,1950]" captionTargetBox="[210,1367,187,1886]" captionTargetId="figure@3.[195,1382,159,1904]" captionTargetPageId="3" captionText="FIGURES 13 – 16. Wharia willcocksorum new species, male. 13, head, specimen from Rotopounamu [LACM ENT 223564]. 14, wing, specimen from Tree Trunk Gorge [LACM ENT 221624]. 15 – 16, male terminalia, right lateral and left lateral, specimen (molecular voucher) from Sandy Bay [LACM ENT 240024]." httpUri="https://zenodo.org/record/184913/files/figure.png" pageId="0" pageNumber="1">13</figureCitation>
). Anepisternum without furrows (= undivided), bare (
<figureCitation id="13542A0FD570FF9BFDDA448C2CEF608E" box="[558,675,1967,1993]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1856,1880]" captionTargetBox="[151,1436,357,1822]" captionTargetId="figure@1.[151,1436,357,1830]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Wharia willcocksorum new species, specimen from Rotopounamu [LACM ENT 223564]. 1, habitus, left lateral. 2, head and thorax, left lateral. 3, head, anterior. 4, thorax, dorsal. 5, hind femur, posterior. 6, hind femur, posterior, basal area." httpUri="https://zenodo.org/record/184910/files/figure.png" pageId="0" pageNumber="1">Figs. 1–2</figureCitation>
). Scutellum with one large and one small seta (
<figureCitation id="13542A0FD570FF9BFB01448C2B0E608E" box="[1269,1346,1967,1993]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,269,1856,1880]" captionTargetBox="[151,1436,357,1822]" captionTargetId="figure@1.[151,1436,357,1830]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Wharia willcocksorum new species, specimen from Rotopounamu [LACM ENT 223564]. 1, habitus, left lateral. 2, head and thorax, left lateral. 3, head, anterior. 4, thorax, dorsal. 5, hind femur, posterior. 6, hind femur, posterior, basal area." httpUri="https://zenodo.org/record/184910/files/figure.png" pageId="0" pageNumber="1">Fig. 4</figureCitation>
). Wing present in both sexes, vein R2+3 absent (
<figureCitation id="13542A0FD570FF9BFD9944F42C8B60B6" box="[621,711,2007,2033]" captionStart="FIGURES 13 – 16" captionStartId="3.[151,269,1926,1950]" captionTargetBox="[210,1367,187,1886]" captionTargetId="figure@3.[195,1382,159,1904]" captionTargetPageId="3" captionText="FIGURES 13 – 16. Wharia willcocksorum new species, male. 13, head, specimen from Rotopounamu [LACM ENT 223564]. 14, wing, specimen from Tree Trunk Gorge [LACM ENT 221624]. 15 – 16, male terminalia, right lateral and left lateral, specimen (molecular voucher) from Sandy Bay [LACM ENT 240024]." httpUri="https://zenodo.org/record/184913/files/figure.png" pageId="0" pageNumber="1">Fig. 14</figureCitation>
). Wing vein Rs with one seta near base only. One alular seta present. Midtibia with one anterodorsal and one dorsal seta near base, with one short anterodorsal seta near apex. Posterior face of hind femur often with rounded basal process covered by thin membrane (
<figureCitation id="13542A0FD571FF9AFAF943E32BC2679D" box="[1293,1422,192,218]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 12" captionStartId-0="1.[151,269,1856,1880]" captionStartId-1="2.[151,270,1679,1703]" captionTargetBox-0="[151,1436,357,1822]" captionTargetBox-1="[151,1436,194,1651]" captionTargetId-0="figure@1.[151,1436,357,1830]" captionTargetId-1="figure@2.[151,1436,194,1655]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 6. Wharia willcocksorum new species, specimen from Rotopounamu [LACM ENT 223564]. 1, habitus, left lateral. 2, head and thorax, left lateral. 3, head, anterior. 4, thorax, dorsal. 5, hind femur, posterior. 6, hind femur, posterior, basal area." captionText-1="FIGURES 7 – 12. Wharia willcocksorum new species, hind femur, posterior, and hind femur, posterior, basal area. 7 – 8, specimen from Tree Trunk Gorge [LACM ENT 221624]. 9 – 10, specimen from Raglan [LACM ENT 223165]. 11 – 12, specimen from Desert Road at Oturere River [LACM ENT 221483]." httpUri-0="https://zenodo.org/record/184910/files/figure.png" httpUri-1="https://zenodo.org/record/184912/files/figure.png" pageId="1" pageNumber="2">Figs. 5–10</figureCitation>
), although size varies greatly and absent in some specimens (
<figureCitation id="13542A0FD571FF9AFCA543CB2D936645" box="[849,991,232,258]" captionStart="FIGURES 7 – 12" captionStartId="2.[151,270,1679,1703]" captionTargetBox="[151,1436,194,1651]" captionTargetId="figure@2.[151,1436,194,1655]" captionTargetPageId="2" captionText="FIGURES 7 – 12. Wharia willcocksorum new species, hind femur, posterior, and hind femur, posterior, basal area. 7 – 8, specimen from Tree Trunk Gorge [LACM ENT 221624]. 9 – 10, specimen from Raglan [LACM ENT 223165]. 11 – 12, specimen from Desert Road at Oturere River [LACM ENT 221483]." httpUri="https://zenodo.org/record/184912/files/figure.png" pageId="1" pageNumber="2">Figs. 11–12</figureCitation>
).
<collectingCountry id="F378761AD571FF9AFC0043CB2A636645" box="[1012,1071,232,258]" name="India" pageId="1" pageNumber="2">Hind</collectingCountry>
tibia without dorsal, longitudinal setal palisade; with one anterodorsal seta near base and one near apex.
</paragraph>
<caption id="DF106602D571FF9AFF6344632F0260DB" httpUri="https://zenodo.org/record/184910/files/figure.png" pageId="1" pageNumber="2" targetBox="[151,1436,357,1822]" targetPageId="1">
<paragraph id="8BD0368AD571FF9AFF6344632F0260DB" blockId="1.[151,1436,1856,1948]" pageId="1" pageNumber="2">
<emphasis id="B91BEA98D571FF9AFF6344632F08601F" bold="true" box="[151,324,1856,1880]" pageId="1" pageNumber="2">FIGURES 1–6.</emphasis>
<taxonomicName id="4C6F4D09D571FF9AFEBA44622C0E601F" box="[334,578,1857,1880]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="willcocksorum" status="sp. nov.">
<emphasis id="B91BEA98D571FF9AFEBA44622C0E601F" box="[334,578,1857,1880]" italics="true" pageId="1" pageNumber="2">Wharia willcocksorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857E3D571FF9AFDBF44632C9C6010" box="[587,720,1856,1879]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
, specimen from Rotopounamu [LACM ENT 223564]. 1, habitus, left lateral. 2, head and thorax, left lateral. 3, head, anterior. 4, thorax, dorsal. 5, hind femur, posterior. 6, hind femur, posterior, basal area.
</paragraph>
</caption>
<caption id="DF106602D572FF99FF6345AC2D3F61AB" httpUri="https://zenodo.org/record/184912/files/figure.png" pageId="2" pageNumber="3" targetBox="[151,1436,194,1651]" targetPageId="2">
<paragraph id="8BD0368AD572FF99FF6345AC2D3F61AB" blockId="2.[151,1436,1679,1772]" pageId="2" pageNumber="3">
<emphasis id="B91BEA98D572FF99FF6345AC2F1D61E0" bold="true" box="[151,337,1679,1703]" pageId="2" pageNumber="3">FIGURES 7–12.</emphasis>
<taxonomicName id="4C6F4D09D572FF99FEAD45B32C0061E0" box="[345,588,1680,1703]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="willcocksorum" status="sp. nov.">
<emphasis id="B91BEA98D572FF99FEAD45B32C0061E0" box="[345,588,1680,1703]" italics="true" pageId="2" pageNumber="3">Wharia willcocksorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857E3D572FF99FDA045AC2C9561E1" box="[596,729,1679,1702]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
, hind femur, posterior, and hind femur, posterior, basal area. 7–8, specimen from Tree Trunk Gorge [LACM ENT 221624]. 9–10, specimen from Raglan [LACM ENT 223165]. 11–12, specimen from Desert Road at Oturere River [LACM ENT 221483].
</paragraph>
</caption>
<paragraph id="8BD0368AD572FF9FFF32443C2DC0645A" blockId="2.[151,1436,1823,2012]" lastBlockId="4.[151,1436,734,797]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<emphasis id="B91BEA98D572FF99FF32443C2F2F607E" bold="true" box="[198,355,1823,1849]" pageId="2" pageNumber="3">Recognition.</emphasis>
Based on the structure of the male genitalia, setation of the frons, and setation of the tibiae, this genus most closely resembles
<taxonomicName id="4C6F4D09D572FF99FDC7446B2D016025" authority="Schmitz" authorityName="Schmitz" box="[563,845,1864,1890]" class="Insecta" family="Phoridae" genus="Bothroprosopa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D572FF99FDC7446B2CAE6026" box="[563,738,1864,1889]" italics="true" pageId="2" pageNumber="3">Bothroprosopa</emphasis>
Schmitz
</taxonomicName>
, also known only from
<collectingCountry id="F378761AD572FF99FB9E446B2B406025" box="[1130,1292,1864,1890]" name="New Zealand" pageId="2" pageNumber="3">New Zealand</collectingCountry>
. In his revision of phorid genera, Brown (1992) placed
<taxonomicName id="4C6F4D09D572FF99FD6844532D0760CE" box="[668,843,1904,1929]" class="Insecta" family="Phoridae" genus="Bothroprosopa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D572FF99FD6844532D0760CE" box="[668,843,1904,1929]" italics="true" pageId="2" pageNumber="3">Bothroprosopa</emphasis>
</taxonomicName>
in his subfamily
<taxonomicName id="4C6F4D09D572FF99FBE044532AE160CD" box="[1044,1197,1904,1930]" class="Insecta" family="Phoridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Hypocerinae">Hypocerinae</taxonomicName>
, based in part on the strongly curved base of wing vein M1 (
<figureCitation id="13542A0FD572FF99FDAB44BB2CFA60F5" box="[607,694,1944,1970]" captionStart="FIGURE 17" captionStartId="4.[151,255,664,688]" captionTargetBox="[255,1328,209,611]" captionTargetId="figure@4.[235,1352,197,638]" captionTargetPageId="4" captionText="FIGURE 17. Bothroprosopa mirifica Schmitz, wing." httpUri="https://zenodo.org/record/184914/files/figure.png" pageId="2" pageNumber="3">Fig. 17</figureCitation>
), which is found throughout this group. Such a strongly curved vein is not found in
<taxonomicName id="4C6F4D09D572FF99FE7644E02F96609B" box="[386,474,1987,2012]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D572FF99FE7644E02F96609B" box="[386,474,1987,2012]" italics="true" pageId="2" pageNumber="3">Wharia</emphasis>
</taxonomicName>
(
<figureCitation id="13542A0FD572FF99FE1E44E12C0D609B" box="[490,577,1986,2012]" captionStart="FIGURES 13 – 16" captionStartId="3.[151,269,1926,1950]" captionTargetBox="[210,1367,187,1886]" captionTargetId="figure@3.[195,1382,159,1904]" captionTargetPageId="3" captionText="FIGURES 13 – 16. Wharia willcocksorum new species, male. 13, head, specimen from Rotopounamu [LACM ENT 223564]. 14, wing, specimen from Tree Trunk Gorge [LACM ENT 221624]. 15 – 16, male terminalia, right lateral and left lateral, specimen (molecular voucher) from Sandy Bay [LACM ENT 240024]." httpUri="https://zenodo.org/record/184913/files/figure.png" pageId="2" pageNumber="3">Fig. 14</figureCitation>
), whose placement among other phorid genera is unknown. We have extracted DNA from
<taxonomicName id="4C6F4D09D574FF9FFE6C41FD2FBC65B0" box="[408,496,734,759]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D574FF9FFE6C41FD2FBC65B0" box="[408,496,734,759]" italics="true" pageId="4" pageNumber="5">Wharia</emphasis>
</taxonomicName>
specimens, however, and they will be included in the molecular phylogenetic analysis of phorid genera (B. Brown &amp; P. Smith, in preparation).
</paragraph>
<caption id="DF106602D573FF98FF6344A52A4460A4" httpUri="https://zenodo.org/record/184913/files/figure.png" pageId="3" pageNumber="4" targetBox="[210,1367,187,1886]" targetPageId="3">
<paragraph id="8BD0368AD573FF98FF6344A52A4460A4" blockId="3.[151,1436,1926,2019]" pageId="3" pageNumber="4">
<emphasis id="B91BEA98D573FF98FF6344A52F2D60D9" bold="true" box="[151,353,1926,1950]" pageId="3" pageNumber="4">FIGURES 13–16.</emphasis>
<taxonomicName id="4C6F4D09D573FF98FE9844A42C2E60D9" box="[364,610,1927,1950]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="willcocksorum" status="sp. nov.">
<emphasis id="B91BEA98D573FF98FE9844A42C2E60D9" box="[364,610,1927,1950]" italics="true" pageId="3" pageNumber="4">Wharia willcocksorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857E3D573FF98FD9944A52CB860DA" box="[621,756,1926,1949]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
, male. 13, head, specimen from Rotopounamu [LACM ENT 223564]. 14, wing, specimen from Tree Trunk Gorge [LACM ENT 221624]. 15–16, male terminalia, right lateral and left lateral, specimen (molecular voucher) from Sandy Bay [LACM ENT 240024].
</paragraph>
</caption>
<caption id="DF106602D574FF9FFF6341BB2C9C65F7" box="[151,720,664,688]" httpUri="https://zenodo.org/record/184914/files/figure.png" pageId="4" pageNumber="5" targetBox="[255,1328,209,611]" targetPageId="4">
<paragraph id="8BD0368AD574FF9FFF6341BB2C9C65F7" blockId="4.[151,720,664,688]" box="[151,720,664,688]" pageId="4" pageNumber="5">
<emphasis id="B91BEA98D574FF9FFF6341BB2F6A65F7" bold="true" box="[151,294,664,688]" pageId="4" pageNumber="5">FIGURE 17.</emphasis>
<taxonomicName id="4C6F4D09D574FF9FFED941BA2CC465F7" authority="Schmitz" authorityName="Schmitz" box="[301,648,665,688]" class="Insecta" family="Phoridae" genus="Bothroprosopa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mirifica">
<emphasis id="B91BEA98D574FF9FFED941BA2C6B65F7" box="[301,551,665,688]" italics="true" pageId="4" pageNumber="5">Bothroprosopa mirifica</emphasis>
Schmitz
</taxonomicName>
, wing.
</paragraph>
</caption>
<paragraph id="8BD0368AD574FF9FFF3240682DBA64F2" blockId="4.[151,1436,843,949]" pageId="4" pageNumber="5">
In the latest keys to phorid genera (Disney, 1994), males of
<taxonomicName id="4C6F4D09D574FF9FFC6940682DB96423" box="[925,1013,843,868]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D574FF9FFC6940682DB96423" box="[925,1013,843,868]" italics="true" pageId="4" pageNumber="5">Wharia</emphasis>
</taxonomicName>
key to couplet 77 of the male key, which was revised by
<bibRefCitation id="EFFE4B7BD574FF9FFE5540502CF964CA" author="Brown" box="[417,693,883,909]" pageId="4" pageNumber="5" refString="Brown, B. V. &amp; Kung, G. (2004). Two new genera of Phoridae (Insecta: Diptera) from the Neotropical Region. Zootaxa, 554, 1 - 7." type="journal article" year="2004">Brown &amp; Kung (2004)</bibRefCitation>
. They do not key correctly to
<taxonomicName id="4C6F4D09D574FF9FFBE840502B0464CA" authority="Brown &amp; Kung" authorityName="Brown &amp; Kung" box="[1052,1352,883,909]" class="Insecta" family="Phoridae" genus="Tapantia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D574FF9FFBE840502ACF64CB" box="[1052,1155,883,908]" italics="true" pageId="4" pageNumber="5">Tapantia</emphasis>
Brown &amp; Kung
</taxonomicName>
, nor to lead 77A. To remedy this situation, we herein revise these couplets again:
</paragraph>
<paragraph id="8BD0368AD574FF9FFF6340C82BD86155" blockId="4.[151,1436,1003,1555]" pageId="4" pageNumber="5">
76. Fork of vein 3 complete [=R2+3 fully developed]
<emphasis id="B91BEA98D574FF9FFCFB40C82B006343" box="[783,1356,1003,1028]" italics="true" pageId="4" pageNumber="5">
..............................................................
<taxonomicName id="4C6F4D09D574FF9FFB2E40C82B006343" box="[1242,1356,1003,1028]" class="Insecta" family="Phoridae" genus="Triphleba" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Triphleba</taxonomicName>
</emphasis>
(part) - Inner branch of fork (R2+3) partly to completely absent.........................................................................77 77. At least left side, and often both sides of male terminalia posterodorsally elongate, forming surstyluslike lobe ...............................................................................................................................................77A - Both sides of male terminalia lacking surstylus-like lobes (although
<taxonomicName id="4C6F4D09D574FF9FFC0847B22A1863ED" box="[1020,1108,1169,1194]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D574FF9FFC0847B22A1863ED" box="[1020,1108,1169,1194]" italics="true" pageId="4" pageNumber="5">Wharia</emphasis>
</taxonomicName>
with median process) ....... .............................................................................................................................................................77B 77A. Restricted to
<collectingCountry id="F378761AD574FF9FFE8847C22C5563BC" box="[380,537,1249,1275]" name="New Zealand" pageId="4" pageNumber="5">New Zealand</collectingCountry>
<emphasis id="B91BEA98D574FF9FFDE947C22BD963BD" box="[541,1429,1249,1274]" italics="true" pageId="4" pageNumber="5">
..........................................................................................................
<taxonomicName id="4C6F4D09D574FF9FFAD947C22BD963BD" box="[1325,1429,1249,1274]" class="Insecta" family="Phoridae" genus="Kierania" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Kierania</taxonomicName>
</emphasis>
- Found in Holarctic and Neotropical Regions
<emphasis id="B91BEA98D574FF9FFD1F462A2B006265" box="[747,1356,1289,1314]" italics="true" pageId="4" pageNumber="5">
...................................................................
<taxonomicName id="4C6F4D09D574FF9FFB2E462A2B006265" box="[1242,1356,1289,1314]" class="Insecta" family="Phoridae" genus="Triphleba" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Triphleba</taxonomicName>
</emphasis>
(part) 77B. Cerci elongate, pointing dorsally; epandrium without posteromedial, ventrally-pointed, elongate process (
<bibRefCitation id="EFFE4B7BD574FF9FFEEB467A2CDB6234" author="Brown" box="[287,663,1369,1395]" pageId="4" pageNumber="5" refString="Brown, B. V. &amp; Kung, G. (2004). Two new genera of Phoridae (Insecta: Diptera) from the Neotropical Region. Zootaxa, 554, 1 - 7." type="journal article" year="2004">Brown &amp; Kung, 2004, figs. 3–4</bibRefCitation>
); costal setae short, numerous (
<bibRefCitation id="EFFE4B7BD574FF9FFBF0467A2B1F6234" author="Brown" box="[1028,1363,1369,1395]" pageId="4" pageNumber="5" refString="Brown, B. V. &amp; Kung, G. (2004). Two new genera of Phoridae (Insecta: Diptera) from the Neotropical Region. Zootaxa, 554, 1 - 7." type="journal article" year="2004">Brown &amp; Kung, 2004, fig. 2</bibRefCitation>
); posterior face of hind femur unmodified; Central
<collectingCountry id="F378761AD574FF9FFD0E46A22D1362DC" box="[762,863,1409,1435]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
<emphasis id="B91BEA98D574FF9FFC9446A22BD962DD" box="[864,1429,1409,1434]" italics="true" pageId="4" pageNumber="5">
..............................................................
<taxonomicName id="4C6F4D09D574FF9FFADA46A22BD962DD" box="[1326,1429,1409,1434]" class="Insecta" family="Phoridae" genus="Tapantia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Tapantia</taxonomicName>
</emphasis>
- Cerci short; epandrium with elongate, posteromedial, ventrally-pointed process (
<figureCitation id="13542A0FD574FF9FFB56468A2AB76284" box="[1186,1275,1449,1475]" captionStart="FIGURES 13 – 16" captionStartId="3.[151,269,1926,1950]" captionTargetBox="[210,1367,187,1886]" captionTargetId="figure@3.[195,1382,159,1904]" captionTargetPageId="3" captionText="FIGURES 13 – 16. Wharia willcocksorum new species, male. 13, head, specimen from Rotopounamu [LACM ENT 223564]. 14, wing, specimen from Tree Trunk Gorge [LACM ENT 221624]. 15 – 16, male terminalia, right lateral and left lateral, specimen (molecular voucher) from Sandy Bay [LACM ENT 240024]." httpUri="https://zenodo.org/record/184913/files/figure.png" pageId="4" pageNumber="5">Fig. 15</figureCitation>
); costal setae relatively longer and fewer (
<figureCitation id="13542A0FD574FF9FFDC646F22CC762AC" box="[562,651,1489,1515]" captionStart="FIGURES 13 – 16" captionStartId="3.[151,269,1926,1950]" captionTargetBox="[210,1367,187,1886]" captionTargetId="figure@3.[195,1382,159,1904]" captionTargetPageId="3" captionText="FIGURES 13 – 16. Wharia willcocksorum new species, male. 13, head, specimen from Rotopounamu [LACM ENT 223564]. 14, wing, specimen from Tree Trunk Gorge [LACM ENT 221624]. 15 – 16, male terminalia, right lateral and left lateral, specimen (molecular voucher) from Sandy Bay [LACM ENT 240024]." httpUri="https://zenodo.org/record/184913/files/figure.png" pageId="4" pageNumber="5">Fig. 14</figureCitation>
); posterior face of hind femur often with large, blunt, ventrobasal process (
<figureCitation id="13542A0FD574FF9FFEB346DA2F846154" box="[327,456,1529,1555]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 12" captionStartId-0="1.[151,269,1856,1880]" captionStartId-1="2.[151,270,1679,1703]" captionTargetBox-0="[151,1436,357,1822]" captionTargetBox-1="[151,1436,194,1651]" captionTargetId-0="figure@1.[151,1436,357,1830]" captionTargetId-1="figure@2.[151,1436,194,1655]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 6. Wharia willcocksorum new species, specimen from Rotopounamu [LACM ENT 223564]. 1, habitus, left lateral. 2, head and thorax, left lateral. 3, head, anterior. 4, thorax, dorsal. 5, hind femur, posterior. 6, hind femur, posterior, basal area." captionText-1="FIGURES 7 – 12. Wharia willcocksorum new species, hind femur, posterior, and hind femur, posterior, basal area. 7 – 8, specimen from Tree Trunk Gorge [LACM ENT 221624]. 9 – 10, specimen from Raglan [LACM ENT 223165]. 11 – 12, specimen from Desert Road at Oturere River [LACM ENT 221483]." httpUri-0="https://zenodo.org/record/184910/files/figure.png" httpUri-1="https://zenodo.org/record/184912/files/figure.png" pageId="4" pageNumber="5">Figs. 5–10</figureCitation>
);
<collectingCountry id="F378761AD574FF9FFE2946DA2C366154" box="[477,634,1529,1555]" name="New Zealand" pageId="4" pageNumber="5">New Zealand</collectingCountry>
<emphasis id="B91BEA98D574FF9FFD8946DA2BD86155" box="[637,1428,1529,1554]" italics="true" pageId="4" pageNumber="5">
................................................................................................
<taxonomicName id="4C6F4D09D574FF9FFAB446DA2BD86155" box="[1344,1428,1529,1554]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Wharia</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BD0368AD574FF9FFF32456A2ACB6199" blockId="4.[151,1436,1609,1878]" pageId="4" pageNumber="5">
Females of
<taxonomicName id="4C6F4D09D574FF9FFEA4456A2FE46125" box="[336,424,1609,1634]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D574FF9FFEA4456A2FE46125" box="[336,424,1609,1634]" italics="true" pageId="4" pageNumber="5">Wharia</emphasis>
</taxonomicName>
key to
<taxonomicName id="4C6F4D09D574FF9FFDF6456A2CFD6125" box="[514,689,1609,1634]" class="Insecta" family="Phoridae" genus="Bothroprosopa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D574FF9FFDF6456A2CFD6125" box="[514,689,1609,1634]" italics="true" pageId="4" pageNumber="5">Bothroprosopa</emphasis>
</taxonomicName>
at couplet
<quantity id="4C979B6FD574FF9FFCC1456A2DCC6124" box="[821,896,1609,1635]" metricMagnitude="0" metricUnit="m" metricValue="3.8608" pageId="4" pageNumber="5" unit="in" value="152.0">152 in</quantity>
the female key of Disney (1994). They differ in having a shorter costa, less curved vein M1, and a much shorter proboscis than in
<taxonomicName id="4C6F4D09D574FF9FFB9945522B5061CD" box="[1133,1308,1649,1674]" class="Insecta" family="Phoridae" genus="Bothroprosopa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D574FF9FFB9945522B5061CD" box="[1133,1308,1649,1674]" italics="true" pageId="4" pageNumber="5">Bothroprosopa</emphasis>
</taxonomicName>
. The other new genus described in this paper also keys to this couplet, so we offer modification to the key that allow identification of both new genera in the Recognition section for
<taxonomicName id="4C6F4D09D574FF9FFC7345E72DB0619A" box="[903,1020,1732,1757]" class="Insecta" family="Phoridae" genus="Minicosta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91BEA98D574FF9FFC7345E72DB0619A" box="[903,1020,1732,1757]" italics="true" pageId="4" pageNumber="5">Minicosta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857E3D574FF9FFBF745E72ACF6199" box="[1027,1155,1732,1758]" pageId="4" pageNumber="5" rank="genus">new genus</taxonomicNameLabel>
.
</paragraph>
<paragraph id="8BD0368AD574FF9FFF3245CF2CA76011" blockId="4.[151,1436,1609,1878]" pageId="4" pageNumber="5">
<emphasis id="B91BEA98D574FF9FFF3245CF2FF16041" bold="true" box="[198,445,1772,1798]" pageId="4" pageNumber="5">Derivation of name.</emphasis>
<taxonomicName id="4C6F4D09D574FF9FFE3045CF2C506042" box="[452,540,1772,1797]" class="Insecta" family="Phoridae" genus="Wharia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA98D574FF9FFE3045CF2C506042" box="[452,540,1772,1797]" italics="true" pageId="4" pageNumber="5">Wharia</emphasis>
</taxonomicName>
is an arbitrary combination of letters, although loosely based on the first two initials of the name of W. H. Robinson, who as been extremely helpful to both authors in the past. For Latinization purposes, we consider it feminine in gender.
</paragraph>
</subSubSection>
</treatment>
</document>
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