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<document id="6D3E2CC79E5C96D6078C66ABA83CC9D8" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F06CFFE7FF42F901FC37FB38" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Coprophanaeoides) hirticollis Olsoufieff 1924" docType="treatment" docVersion="10" lastPageNumber="17" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="15" updateTime="1698234910999" updateUser="plazi">
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<mods:title id="CF2AD294C3EC220D224FC751B4FEA228">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
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<mods:namePart id="395B76D6C51C32ACDF468765818F0CF4">François Génier</mods:namePart>
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<mods:namePart id="48988E739C652502B76991C303F4716D">Patrick Arnaud</mods:namePart>
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<mods:date id="252077443C80EAD6E7ABFB50167D5317">2016</mods:date>
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<mods:identifier id="9F852A243EF8FADAE1A766DDCC904D8E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C68782F06CFFE7FF42F901FC37FB38" ID-DOI="http://doi.org/10.5281/zenodo.5780262" ID-GBIF-Taxon="127631978" ID-Zenodo-Dep="5780262" LSID="urn:lsid:plazi:treatment:03C68782F06CFFE7FF42F901FC37FB38" httpUri="http://treatment.plazi.org/id/03C68782F06CFFE7FF42F901FC37FB38" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="C375651FF06CFFF9FF42F901FC52F8D6" box="[151,948,1772,1798]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BD03694F06CFFF9FF42F901FC52F8D6" blockId="14.[151,948,1772,1831]" box="[151,948,1772,1798]" pageId="14" pageNumber="15">
<heading id="D09881F8F06CFFF9FF42F901FC52F8D6" bold="true" box="[151,948,1772,1798]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B91BEA86F06CFFF9FF42F901FC52F8D6" bold="true" box="[151,948,1772,1798]" pageId="14" pageNumber="15">
6.
<taxonomicName id="4C6F4D17F06CFFF9FF61F901FC52F8D6" ID-CoL="6CLCS" authority="Olsoufieff, 1924" authorityName="Olsoufieff" authorityYear="1924" box="[180,948,1772,1798]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hirticollis" subGenus="Coprophanaeoides">
<emphasis id="B91BEA86F06CFFF9FF61F901FD0EF8D6" bold="true" box="[180,744,1772,1798]" italics="true" pageId="14" pageNumber="15">Dendropaemon (Coprophanaeoides) hirticollis</emphasis>
<bibRefCitation id="EFFE4B65F06CFFF9FD3AF901FC52F8D6" author="Olsoufieff" box="[751,948,1772,1798]" pageId="14" pageNumber="15" refString="Olsoufieff, G. d' (1924) Les Phanaeides (Coleoptera-Lamellicornia), Famille Scarabaeidae-Tr. Coprini. Insecta, 13, 4 - 172, 15 pls." type="journal article" year="1924">Olsoufieff, 1924</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06CFFF9FF42F8E3FE5CF8F7" box="[151,442,1806,1831]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BD03694F06CFFF9FF42F8E3FE5CF8F7" blockId="14.[151,948,1772,1831]" box="[151,442,1806,1831]" pageId="14" pageNumber="15">
(
<figureCitation id="13542A11F06CFFF9FF4AF8E3FF08F8F7" box="[159,238,1806,1831]" captionStart="FIGURES 1 – 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 – 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="14" pageNumber="15">Figs. 6</figureCitation>
, 50–51, 134, 156)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06CFFF8FF42F8BBFBBCFE2F" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BD03694F06CFFF9FF42F8BBFC2FF8BC" blockId="14.[151,1218,1878,2023]" box="[151,969,1878,1901]" pageId="14" pageNumber="15">
<taxonomicName id="4C6F4D17F06CFFF9FF42F8BBFDA5F8BC" ID-CoL="6CLCS" authority="Olsoufieff 1924" authorityName="Olsoufieff" authorityYear="1924" box="[151,579,1878,1900]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F06CFFF9FF42F8BBFE7BF8BC" box="[151,413,1878,1900]" italics="true" pageId="14" pageNumber="15">Dendropaemon hirticollis</emphasis>
<bibRefCitation id="EFFE4B65F06CFFF9FE77F8BBFDA5F8BC" author="Olsoufieff" box="[418,579,1878,1900]" pageId="14" pageNumber="15" refString="Olsoufieff, G. d' (1924) Les Phanaeides (Coleoptera-Lamellicornia), Famille Scarabaeidae-Tr. Coprini. Insecta, 13, 4 - 172, 15 pls." type="journal article" year="1924">
Olsoufieff
<emphasis id="B91BEA86F06CFFF9FDC5F8BAFDA5F8BC" bold="true" box="[528,579,1879,1900]" pageId="14" pageNumber="15">1924</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06CFFF9FD85F8B5FD7FF8BD" box="[592,665,1880,1901]" italics="true" pageId="14" pageNumber="15">Insecta</emphasis>
13: 128 (original description)
</paragraph>
<paragraph id="8BD03694F06CFFF9FF42F898FC51F818" blockId="14.[151,1218,1878,2023]" pageId="14" pageNumber="15">
<taxonomicName id="4C6F4D17F06CFFF9FF42F898FD9CF85A" ID-CoL="6CLCS" authority="Pessoa &amp; Lane 1936" authorityName="Pessoa &amp; Lane" authorityYear="1936" box="[151,634,1909,1931]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F06CFFF9FF42F898FE7BF85B" box="[151,413,1909,1931]" italics="true" pageId="14" pageNumber="15">Dendropaemon hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06CFFF9FE7CF89BFD9CF85A" author="Pessoa" box="[425,634,1909,1931]" pageId="14" pageNumber="15" refString="Pessoa, S. B. &amp; Lane, F. (1936) Sobre os coleopteros do genero Dendropaemon Perty, de S. Paulo e regioes visinhas. Revista de Biologia e Hygiene, 7 (2), 89 - 93." type="journal article" year="1936">
Pessôa &amp; Lane
<emphasis id="B91BEA86F06CFFF9FD9DF898FD9CF85A" bold="true" box="[584,634,1909,1930]" pageId="14" pageNumber="15">1936</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06CFFF9FD52F89BFCA4F85B" box="[647,834,1909,1931]" italics="true" pageId="14" pageNumber="15">Rev. Biol. Hygiene</emphasis>
7: 90 (identification key, comment)
<taxonomicName id="4C6F4D17F06CFFF9FF42F879FDB0F879" ID-CoL="34R3H" authority="Pessoa &amp; Lane 1936" authorityName="Pessoa &amp; Lane" authorityYear="1936" box="[151,598,1940,1962]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="montei">
<emphasis id="B91BEA86F06CFFF9FF42F879FE99F87A" box="[151,383,1940,1962]" italics="true" pageId="14" pageNumber="15">Dendropaemon montei</emphasis>
<bibRefCitation id="EFFE4B65F06CFFF9FE50F879FDB0F879" author="Pessoa" box="[389,598,1940,1962]" pageId="14" pageNumber="15" refString="Pessoa, S. B. &amp; Lane, F. (1936) Sobre os coleopteros do genero Dendropaemon Perty, de S. Paulo e regioes visinhas. Revista de Biologia e Hygiene, 7 (2), 89 - 93." type="journal article" year="1936">
Pessôa &amp; Lane
<emphasis id="B91BEA86F06CFFF9FDF6F879FDB0F879" bold="true" box="[547,598,1940,1961]" pageId="14" pageNumber="15">1936</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06CFFF9FDB7F878FCF8F87A" box="[610,798,1940,1962]" italics="true" pageId="14" pageNumber="15">Rev. Biol. Hygiene</emphasis>
7: 90 (original description)
<taxonomicName id="4C6F4D17F06CFFF9FF42F85FFDEBF817" ID-CoL="6CLCS" authority="Blut 1939" authorityName="Blut" authorityYear="1939" box="[151,525,1970,1992]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F06CFFF9FF42F85FFE7BF818" box="[151,413,1970,1992]" italics="true" pageId="14" pageNumber="15">Dendropaemon hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06CFFF9FE7CF85FFDEBF817" author="Blut" box="[425,525,1970,1992]" pageId="14" pageNumber="15" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">
Blut
<emphasis id="B91BEA86F06CFFF9FE0EF85FFDEBF817" bold="true" box="[475,525,1970,1991]" pageId="14" pageNumber="15">1939</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06CFFF9FDCEF85FFD03F818" box="[539,741,1970,1992]" italics="true" pageId="14" pageNumber="15">Arch. Naturg. (N.F.)</emphasis>
8: 273 (monograph)
</paragraph>
<paragraph id="8BD03694F06CFFF8FF42F83CFC1EFF1C" blockId="14.[151,1218,1878,2023]" lastBlockId="15.[151,1339,152,511]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<taxonomicName id="4C6F4D17F06CFFF9FF42F83CFD9CF836" ID-CoL="6CLCS" authority="Pessoa &amp; Lane 1941" authorityName="Pessoa &amp; Lane" authorityYear="1941" box="[151,634,2001,2023]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F06CFFF9FF42F83CFE7BF837" box="[151,413,2001,2023]" italics="true" pageId="14" pageNumber="15">Dendropaemon hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06CFFF9FE7CF83FFD9CF836" author="Pessoa" box="[425,634,2001,2023]" pageId="14" pageNumber="15" refString="Pessoa, S. B. &amp; Lane, F. (1941) Coleopteros necrophagos de interesse medico-legal. Ensaio monografico sobre a familia Scarabaeidae de Sao Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo, 2, 389 - 504." type="journal article" year="1941">
Pessôa &amp; Lane
<emphasis id="B91BEA86F06CFFF9FD9DF83CFD9CF836" bold="true" box="[584,634,2001,2022]" pageId="14" pageNumber="15">1941</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06CFFF9FD52F83FFCAFF837" box="[647,841,2001,2023]" italics="true" pageId="14" pageNumber="15">Arq. Zool. S. Paulo</emphasis>
2: 491 (identification key, comment)
<taxonomicName id="4C6F4D17F06DFFF8FF42FF75FDBBFF7D" ID-CoL="34R3H" authority="Pessoa &amp; Lane 1941" authorityName="Pessoa &amp; Lane" authorityYear="1941" box="[151,605,152,174]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="montei">
<emphasis id="B91BEA86F06DFFF8FF42FF75FE99FF7E" box="[151,383,152,174]" italics="true" pageId="15" pageNumber="16">Dendropaemon montei</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FE59FF75FDBBFF7D" author="Pessoa" box="[396,605,152,174]" pageId="15" pageNumber="16" refString="Pessoa, S. B. &amp; Lane, F. (1941) Coleopteros necrophagos de interesse medico-legal. Ensaio monografico sobre a familia Scarabaeidae de Sao Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo, 2, 389 - 504." type="journal article" year="1941">
Pessôa &amp; Lane
<emphasis id="B91BEA86F06DFFF8FDFFFF75FDBBFF7D" bold="true" box="[554,605,152,173]" pageId="15" pageNumber="16">1941</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FDBCFF74FCCAFF7E" box="[617,812,152,174]" italics="true" pageId="15" pageNumber="16">Arq. Zool. S. Paulo</emphasis>
2: 491 (identification key, redescription)
<taxonomicName id="4C6F4D17F06DFFF8FF42FF5BFDB6FF1C" authority="Blackwelder 1944" authorityName="Blackwelder" authorityYear="1944" box="[151,592,182,204]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hirticolle">
<emphasis id="B91BEA86F06DFFF8FF42FF5BFE6FFF1C" box="[151,393,182,204]" italics="true" pageId="15" pageNumber="16">Dendropemon hirticolle</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FE42FF5BFDB6FF1C" author="Blackwelder" box="[407,592,182,204]" pageId="15" pageNumber="16" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 341. http: // dx. doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">
Blackwelder
<emphasis id="B91BEA86F06DFFF8FDC8FF5AFDB6FF1C" bold="true" box="[541,592,183,204]" pageId="15" pageNumber="16">1944</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FD89FF5AFCCFFF1C" box="[604,809,182,205]" italics="true" pageId="15" pageNumber="16">
<collectingCountry id="F3787604F06DFFF8FD89FF5AFD6FFF1C" box="[604,649,183,204]" name="United States of America" pageId="15" pageNumber="16">U.S.</collectingCountry>
<taxonomicName id="4C6F4D17F06DFFF8FD5BFF55FD50FF1D" box="[654,694,184,205]" class="Insecta" family="Coccinellidae" genus="Nat" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Nat</taxonomicName>
. Mus. Bull.
</emphasis>
185: 211 (checklist)
</paragraph>
<paragraph id="8BD03694F06DFFF8FF42FF38FC07FF3B" blockId="15.[151,1339,152,511]" box="[151,993,213,235]" pageId="15" pageNumber="16">
<taxonomicName id="4C6F4D17F06DFFF8FF42FF38FDDEFF3A" authority="Blackwelder 1944" authorityName="Blackwelder" authorityYear="1944" box="[151,568,213,235]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="montei">
<emphasis id="B91BEA86F06DFFF8FF42FF38FE94FF3B" box="[151,370,213,235]" italics="true" pageId="15" pageNumber="16">Dendropemon montei</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FEAAFF38FDDEFF3A" author="Blackwelder" box="[383,568,213,235]" pageId="15" pageNumber="16" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 341. http: // dx. doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">
Blackwelder
<emphasis id="B91BEA86F06DFFF8FDD0FF38FDDEFF3A" bold="true" box="[517,568,213,234]" pageId="15" pageNumber="16">1944</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FD90FF3BFCF7FF3B" box="[581,785,213,235]" italics="true" pageId="15" pageNumber="16">
<collectingCountry id="F3787604F06DFFF8FD90FF3BFD97FF3B" box="[581,625,214,235]" name="United States of America" pageId="15" pageNumber="16">U.S.</collectingCountry>
<taxonomicName id="4C6F4D17F06DFFF8FDA2FF3BFD79FF3B" box="[631,671,214,235]" class="Insecta" family="Coccinellidae" genus="Nat" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Nat</taxonomicName>
. Mus. Bull.
</emphasis>
185: 211 (checklist)
</paragraph>
<paragraph id="8BD03694F06DFFF8FF42FF19FC13FEF8" blockId="15.[151,1339,152,511]" pageId="15" pageNumber="16">
<taxonomicName id="4C6F4D17F06DFFF8FF42FF19FD7FFED9" authority="Pereira &amp; Martinez 1956" authorityName="Pereira &amp; Martinez" authorityYear="1956" box="[151,665,244,266]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F06DFFF8FF42FF19FE69FEDA" box="[151,399,244,266]" italics="true" pageId="15" pageNumber="16">Dendropemon hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FE48FF19FD7FFED9" author="Pereira" box="[413,665,244,266]" pageId="15" pageNumber="16" refString="Pereira, F. S. &amp; Martinez, A. (1956) Algunas notas sinonimicas en Phanaeini (Col. Scarabaeidae, Coprinae). Revista Brasileira de Entomologia, 5, 229 - 240." type="journal article" year="1956">
Pereira &amp; Martínez
<emphasis id="B91BEA86F06DFFF8FDB3FF19FD7FFED9" bold="true" box="[614,665,244,265]" pageId="15" pageNumber="16">1956</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FD73FF18FCADFEDA" box="[678,843,244,266]" italics="true" pageId="15" pageNumber="16">
Rev.
<collectingCountry id="F3787604F06DFFF8FD03FF19FCF1FEDA" box="[726,791,244,266]" name="Brazil" pageId="15" pageNumber="16">Brasil</collectingCountry>
. Ent.
</emphasis>
5: 237 (comment taxonomy)
<taxonomicName id="4C6F4D17F06DFFF8FF42FEFFFD9AFEF7" authority="Pereira &amp; Martinez 1956" authorityName="Pereira &amp; Martinez" authorityYear="1956" box="[151,636,274,296]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="montei">
<emphasis id="B91BEA86F06DFFF8FF42FEFFFE94FEF8" box="[151,370,274,296]" italics="true" pageId="15" pageNumber="16">Dendropemon montei</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FEAAFEFFFD9AFEF7" author="Pereira" box="[383,636,274,296]" pageId="15" pageNumber="16" refString="Pereira, F. S. &amp; Martinez, A. (1956) Algunas notas sinonimicas en Phanaeini (Col. Scarabaeidae, Coprinae). Revista Brasileira de Entomologia, 5, 229 - 240." type="journal article" year="1956">
Pereira &amp; Martínez
<emphasis id="B91BEA86F06DFFF8FD9CFEFFFD9AFEF7" bold="true" box="[585,636,274,295]" pageId="15" pageNumber="16">1956</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FD5DFEFEFCC8FEF8" box="[648,814,274,296]" italics="true" pageId="15" pageNumber="16">
Rev.
<collectingCountry id="F3787604F06DFFF8FD6CFEFFFD1CFEF8" box="[697,762,274,296]" name="Brazil" pageId="15" pageNumber="16">Brasil</collectingCountry>
. Ent.
</emphasis>
5: 237 (synonymy)
</paragraph>
<paragraph id="8BD03694F06DFFF8FF42FEDCFC13FEB6" blockId="15.[151,1339,152,511]" pageId="15" pageNumber="16">
<taxonomicName id="4C6F4D17F06DFFF8FF42FEDCFD7FFE96" authority="Pereira &amp; Martinez 1960" authorityName="Pereira &amp; Martinez" authorityYear="1960" box="[151,665,305,327]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F06DFFF8FF42FEDCFE69FE97" box="[151,399,305,327]" italics="true" pageId="15" pageNumber="16">Dendropemon hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FE48FEDCFD7FFE96" author="Pereira" box="[413,665,305,327]" pageId="15" pageNumber="16" refString="Martinez, A. &amp; Pereira, F. S. (1960) Algunos interesantes coprini neotropicales (Col. Scarabaeidae). Revista de la Sociedad Entomologica Argentina, 22 (1 - 4), 77 - 84." type="journal article" year="1960">
Pereira &amp; Martínez
<emphasis id="B91BEA86F06DFFF8FDB3FEDCFD7FFE96" bold="true" box="[614,665,305,326]" pageId="15" pageNumber="16">1960</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FD73FEDFFCADFE97" box="[678,843,305,327]" italics="true" pageId="15" pageNumber="16">
Rev.
<collectingCountry id="F3787604F06DFFF8FD03FEDCFCF1FE97" box="[726,791,305,327]" name="Brazil" pageId="15" pageNumber="16">Brasil</collectingCountry>
. Ent.
</emphasis>
9: 55 (comment taxonomy)
<taxonomicName id="4C6F4D17F06DFFF8FF42FEBDFD61FEB5" ID-CoL="34R3H" authority="Pereira &amp; Martinez 1960" authorityName="Pereira &amp; Martinez" authorityYear="1960" box="[151,647,336,358]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="montei">
<emphasis id="B91BEA86F06DFFF8FF42FEBDFE99FEB6" box="[151,383,336,358]" italics="true" pageId="15" pageNumber="16">Dendropaemon montei</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FE59FEBDFD61FEB5" author="Pereira" box="[396,647,336,358]" pageId="15" pageNumber="16" refString="Martinez, A. &amp; Pereira, F. S. (1960) Algunos interesantes coprini neotropicales (Col. Scarabaeidae). Revista de la Sociedad Entomologica Argentina, 22 (1 - 4), 77 - 84." type="journal article" year="1960">
Pereira &amp; Martínez
<emphasis id="B91BEA86F06DFFF8FD80FEBDFD61FEB5" bold="true" box="[597,647,336,357]" pageId="15" pageNumber="16">1960</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FD40FEBCFCDCFEB6" box="[661,826,336,358]" italics="true" pageId="15" pageNumber="16">
Rev.
<collectingCountry id="F3787604F06DFFF8FD13FEBDFCE1FEB6" box="[710,775,336,358]" name="Brazil" pageId="15" pageNumber="16">Brasil</collectingCountry>
. Ent.
</emphasis>
9: 55 (synonymy)
</paragraph>
<paragraph id="8BD03694F06DFFF8FF42FE83FC6DFE73" blockId="15.[151,1339,152,511]" pageId="15" pageNumber="16">
<taxonomicName id="4C6F4D17F06DFFF8FF42FE83FCFEFE54" ID-CoL="6CLCS" authority="Edmonds 1972" authorityName="Edmonds" authorityYear="1972" box="[151,792,366,388]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hirticollis" subGenus="Coprophanaeoides">
<emphasis id="B91BEA86F06DFFF8FF42FE83FD95FE54" box="[151,627,366,388]" italics="true" pageId="15" pageNumber="16">Dendropaemon (Coprophanaeoides) hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FD55FE83FCFEFE54" author="Edmonds" box="[640,792,366,388]" pageId="15" pageNumber="16" refString="Edmonds, W. D. (1972) Comparative Skeletal Morphology, Systematics and Evolution of the Phanaeine Dung Beetles (Coleoptera: Scarabaeidae). University of Kansas Science Bulletin, 49 (11), 731 - 874." type="journal article" year="1972">
Edmonds
<emphasis id="B91BEA86F06DFFF8FD33FE82FCFEFE54" bold="true" box="[742,792,367,388]" pageId="15" pageNumber="16">1972</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FCF0FE9DFBEEFE54" box="[805,1032,366,389]" italics="true" pageId="15" pageNumber="16">Univ. Kansas Sc. Bull.</emphasis>
49: 850 (comment taxonomy)
<taxonomicName id="4C6F4D17F06DFFF8FF42FE60FDC8FE72" ID-CoL="6CLCS" authority="Arnaud 1982" authorityName="Arnaud" authorityYear="1982" box="[151,558,397,419]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F06DFFF8FF42FE60FE7BFE73" box="[151,413,397,419]" italics="true" pageId="15" pageNumber="16">Dendropaemon hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FE7CFE60FDC8FE72" author="Arnaud" box="[425,558,397,419]" pageId="15" pageNumber="16" refString="Arnaud, P. (1982) Liste des types de Phanaeini du Museum national d'Histoire naturelle de Paris, suivi d'une note synonymique pour quatre especes decrite par G. d'Olsoufieff (Coleoptera, Scarabaeidae). Revue Francaise d'Entomologie, Nouvelle Serie, 4 (3), 113 - 118." type="journal article" year="1982">
Arnaud
<emphasis id="B91BEA86F06DFFF8FE29FE60FDC8FE72" bold="true" box="[508,558,397,418]" pageId="15" pageNumber="16">1982</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FDEEFE63FD1FFE73" box="[571,761,398,419]" italics="true" pageId="15" pageNumber="16">Rev. Fr. Ent. (N.S.)</emphasis>
4: 117 (notes)
</paragraph>
<paragraph id="8BD03694F06DFFF8FF42FE41FBFDFE12" blockId="15.[151,1339,152,511]" box="[151,1051,428,450]" pageId="15" pageNumber="16">
<taxonomicName id="4C6F4D17F06DFFF8FF42FE41FD4FFE11" ID-CoL="6CLCS" authority="Martinez &amp; Clavijo 1990" authorityName="Martinez &amp; Clavijo" authorityYear="1990" box="[151,681,428,450]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F06DFFF8FF42FE41FE7BFE12" box="[151,413,428,450]" italics="true" pageId="15" pageNumber="16">Dendropaemon hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FE7CFE41FD4FFE11" author="Martinez" box="[425,681,428,450]" pageId="15" pageNumber="16" refString="Martinez, A. &amp; Clavijo, J. (1990) Notas sobre Phanaeina Venezolanos, con descripcion de una nueva subespecie de Diabroctis (Coleoptera, Scarabaeidae, Coprini). Boletin de Entomologia Venezolana, Nueva Seria, 5 (20) 147 - 157." type="journal article" year="1990">
Martínez &amp; Clavijo
<emphasis id="B91BEA86F06DFFF8FDA3FE41FD4FFE11" bold="true" box="[630,681,428,449]" pageId="15" pageNumber="16">1990</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FD60FE41FC97FE11" box="[693,881,428,450]" italics="true" pageId="15" pageNumber="16">Bol. Ent. Ven. N.S.</emphasis>
5: 155 (biology)
</paragraph>
<paragraph id="8BD03694F06DFFF8FF42FE27FC3EFE30" blockId="15.[151,1339,152,511]" box="[151,984,458,480]" pageId="15" pageNumber="16">
<taxonomicName id="4C6F4D17F06DFFF8FF42FE27FE71FE30" box="[151,407,458,480]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hirticolle">
<emphasis id="B91BEA86F06DFFF8FF42FE27FE71FE30" box="[151,407,458,480]" italics="true" pageId="15" pageNumber="16">Dendropaemon hirticolle</emphasis>
</taxonomicName>
: Vaz-de-Mello
<emphasis id="B91BEA86F06DFFF8FDE0FE27FD81FE0F" bold="true" box="[565,615,458,479]" pageId="15" pageNumber="16">2000</emphasis>
,
<emphasis id="B91BEA86F06DFFF8FDA1FE26FCD0FE30" box="[628,822,459,480]" italics="true" pageId="15" pageNumber="16">Hac. Proy. CYTED</emphasis>
: 192 (faunistic)
</paragraph>
<paragraph id="8BD03694F06DFFF8FF42FE04FBBCFE2F" blockId="15.[151,1339,152,511]" box="[151,1114,489,511]" pageId="15" pageNumber="16">
<taxonomicName id="4C6F4D17F06DFFF8FF42FE04FCE0FE2E" ID-CoL="6CLCS" authority="Arnaud 2002" authorityName="Arnaud" authorityYear="2002" box="[151,774,489,511]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hirticollis" subGenus="Coprophanaeoides">
<emphasis id="B91BEA86F06DFFF8FF42FE04FD95FE2F" box="[151,627,489,511]" italics="true" pageId="15" pageNumber="16">Dendropaemon (Coprophanaeoides) hirticollis</emphasis>
:
<bibRefCitation id="EFFE4B65F06DFFF8FD55FE04FCE0FE2E" author="Arnaud" box="[640,774,489,511]" pageId="15" pageNumber="16" refString="Arnaud, P. (2002) Phanaeini, Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Les Coleopteres du Monde, 28. Hillside Books, Canterbury, 151 pp." type="book" year="2002">
Arnaud
<emphasis id="B91BEA86F06DFFF8FD06FE04FCE0FE2E" bold="true" box="[723,774,489,510]" pageId="15" pageNumber="16">2002</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F06DFFF8FCC7FE04FC6FFE2F" box="[786,905,489,511]" italics="true" pageId="15" pageNumber="16">Col. Monde</emphasis>
28: 15 (monograph)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06DFFF8FF42FDC1FE10FD95" box="[151,502,556,581]" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BD03694F06DFFF8FF42FDC1FE10FD95" blockId="15.[151,1437,556,2021]" box="[151,502,556,581]" pageId="15" pageNumber="16">
<emphasis id="B91BEA86F06DFFF8FF42FDC1FECBFD95" bold="true" box="[151,301,556,581]" pageId="15" pageNumber="16">
<typeStatus id="54D48836F06DFFF8FF42FDC1FF34FD95" box="[151,210,556,581]" pageId="15" pageNumber="16">Type</typeStatus>
locality
</emphasis>
. No
<typeStatus id="54D48836F06DFFF8FEB0FDC0FE73FD95" box="[357,405,557,581]" pageId="15" pageNumber="16">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06DFFF8FF12FDA2FC02FD05" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BD03694F06DFFF8FF12FDA2FC02FD05" blockId="15.[151,1437,556,2021]" pageId="15" pageNumber="16">
<emphasis id="B91BEA86F06DFFF8FF12FDA2FEA7FDB7" bold="true" box="[199,321,591,616]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
The acutely notched external side of each clypeal tooth will place this species in
<taxonomicName id="4C6F4D17F06DFFF8FF42FD99FE8DFD5D" box="[151,363,628,653]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F06DFFF8FF42FD99FE8DFD5D" box="[151,363,628,653]" italics="true" pageId="15" pageNumber="16">Coprophanaeoides</emphasis>
</taxonomicName>
. Within
<taxonomicName id="4C6F4D17F06DFFF8FE03FD99FD4CFD5D" box="[470,682,628,653]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F06DFFF8FE03FD99FD4CFD5D" box="[470,682,628,653]" italics="true" pageId="15" pageNumber="16">Coprophanaeoides</emphasis>
</taxonomicName>
, the lack of basal elytral margin will separate it from all other species except its sister species
<taxonomicName id="4C6F4D17F06DFFF8FDF6FD74FCE1FD60" box="[547,775,665,688]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="compressipennis">
<emphasis id="B91BEA86F06DFFF8FDF6FD74FCE1FD60" box="[547,775,665,688]" italics="true" pageId="15" pageNumber="16">D. compressipennis</emphasis>
</taxonomicName>
from which it differs in having a less dorsoventrally compressed body and in lacking a well-defined metasternal v-shaped ridge.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06DFFE7FF12FD32FCFBFD54" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="8BD03694F06DFFE7FF12FD32FD73FE00" blockId="15.[151,1437,556,2021]" lastBlockId="16.[151,1437,151,1256]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B91BEA86F06DFFF8FF12FD32FEB7FD28" bold="true" box="[199,337,735,760]" pageId="15" pageNumber="16">Description</emphasis>
. Female
<typeStatus id="54D48836F06DFFF8FE68FD32FDF9FD28" box="[445,543,735,760]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
(
<figureCitation id="13542A11F06DFFF8FDE4FD32FD91FD28" box="[561,631,735,760]" captionStart="FIGURES 1 – 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 – 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
).
<emphasis id="B91BEA86F06DFFF8FD5BFD32FD2DFD28" bold="true" box="[654,715,735,760]" pageId="15" pageNumber="16">Body</emphasis>
. Body moderately large, length 13.0 mm, maximum width 7.0 mm; body subrectangular in dorsal view; dorsum narrowly flat.
<emphasis id="B91BEA86F06DFFF8FCA8FCE9FC58FCCD" bold="true" box="[893,958,772,797]" pageId="15" pageNumber="16">Color</emphasis>
. Dorsal surface black, glossy, with green metallic sheen; head with green metallic sheen on genae and frons; pronotum with green metallic sheen except for anteromedian carina, on anterior portion of disc and surface adjacent to lateral fossae; elytra with uniform green metallic sheen; ventrum with faint greenish and coppery metallic sheen; pygidium with green metallic sheen; legs with coppery and greenish metallic sheen on femora and tibiae.
<emphasis id="B91BEA86F06DFFF8FC89FC79FC7AFC7D" bold="true" box="[860,924,916,941]" pageId="15" pageNumber="16">Head</emphasis>
. Clypeus broadly arcuate laterally, straight on a short distance laterally to clypeal teeth, anterior portion upturned; clypeal teeth acutely triangular; clypeal median emargination narrowly v-shaped, clypeal edge acutely notched on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin ill-defined, bordered posteriorly by a more or less regular row of punctures, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with small irregular tubercles, lacking distinct transverse carina, simply convex; clypeofrontal carina rather low, approximately 4 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge straight in frontal view; eyes large in dorsal view, interocular ratio 3.5.
<emphasis id="B91BEA86F06DFFF8FF42FB59FEF6FB1D" bold="true" box="[151,272,1204,1229]" pageId="15" pageNumber="16">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.7; disc of pronotum minutely punctate with some scattered large setiferous puncture medially and irregular setiferous rough punctation anterolaterally, with an ill-defined shallow longitudinal depression on posterior half; pronotal anterior margin wider and flat lateral to eyes; anterior portion with a simple anteriorly arcuate carina; anterior angles surface with rough setiferous punctures, similar to lateral margin along posterior edge of anterior margin; lateral fossae oval, simple; lateral portions strongly explanate; pronotal basal fossae ill-defined, slightly concave; posterior margin well-defined laterally, crenulate, with few long setae.
<emphasis id="B91BEA86F06DFFF8FD26FA61FCD9FA75" bold="true" box="[755,831,1420,1445]" pageId="15" pageNumber="16">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.1; elytral base lacking distinct margin, simply convex; elytral striae 1–4 fine and well-defined, evenly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures fine, setiferous and scattered, adjacent strial edge encroaching on interval, stria 1 weakly impressed apically, going straight to elytral apical margin; interstriae slightly convex, minutely punctate and with few larger setiferous punctures along striae, surface glossy.
<emphasis id="B91BEA86F06DFFF8FE4CF9D2FDBCF988" bold="true" box="[409,602,1599,1624]" pageId="15" pageNumber="16">Thoracic sterna</emphasis>
. Proepisternal carina absent; metasternal median lobe bluntly angularly produced anteromedially, ventral ridge absent.
<emphasis id="B91BEA86F06DFFF8FD11F989FD1AF9AD" bold="true" box="[708,764,1636,1661]" pageId="15" pageNumber="16">Legs</emphasis>
. Profemur posterior surface flat and glabrous internally, posterointernal margin rather thin, evenly developed, internal edge rather wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle bluntly lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with some ill-defined irregular punctures externally to median carina, surface glossy between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge straight in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less flat, bordered anteriorly and posteriorly by an almost complete setal row. Mesotarsus similar in shape to metatarsus, 3- segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur internal edge nearly straight and lateral edge arcuate, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on more than half the length. Metatibia moderately slender, slightly widening toward apex in anterior view, anterior surface with distinct row of setae, surface glossy, metatibial posterior surface flat between longitudinal row of setae and lateral edge, with ill-defined microsculpture. Metatarsus 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge.
<emphasis id="B91BEA86F072FFE7FB4BFEA1FA72FEB5" bold="true" box="[1182,1428,332,357]" pageId="16" pageNumber="17">Abdominal sternites</emphasis>
. Sternites 3–6 longitudinally flat; sternites 4–6 with more than three unaligned row of setae laterally, narrowly glabrous medially; sternite 7 approximately longitudinally flat medially, subequal in length to segment 6 along midline; pygidium minutely punctate on disc.
</paragraph>
<paragraph id="8BD03694F072FFE7FF12FE31FB5EFE24" blockId="16.[151,1437,151,1256]" box="[199,1208,476,501]" pageId="16" pageNumber="17">
<emphasis id="B91BEA86F072FFE7FF12FE31FE90FE25" bold="true" box="[199,374,476,501]" pageId="16" pageNumber="17">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF072FFE7FE5DFE31FE32FE25" box="[392,468,476,501]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
,
<specimenCount id="9D69FD1DF072FFE7FE35FE30FDA8FE25" box="[480,590,476,501]" pageId="16" pageNumber="17" type="female">7 females</specimenCount>
). Length: male 15.0, female 13.0–15.0 (13.6±0.7) mm.
</paragraph>
<paragraph id="8BD03694F072FFE7FF12FE12FCFBFD54" blockId="16.[151,1437,151,1256]" pageId="16" pageNumber="17">
<emphasis id="B91BEA86F072FFE7FF12FE12FE47FDC8" bold="true" box="[199,417,511,536]" pageId="16" pageNumber="17">
Primary
<typeStatus id="54D48836F072FFE7FEE6FE12FE80FDC8" box="[307,358,511,536]" pageId="16" pageNumber="17">type</typeStatus>
data
</emphasis>
(
<figureCitation id="13542A11F072FFE7FE65FE12FDF4FDC8" box="[432,530,511,536]" captionStart="FIGURES 134 – 154" captionStartId="115.[151,264,1903,1926]" captionTargetBox="[216,1361,200,1865]" captionTargetId="figure@115.[210,1372,193,1876]" captionTargetPageId="115" captionText="FIGURES 134 – 154. Primary type labels. 134. D. hirticollis; 135. D. pauliani; 136. D. renatii; 137. D. ater; 138. D. piceus; 139: D. telephus; 140. D. viridis; 141. D. viridipennis. 142. D. convexus; 143. D. fredericki; 144. D. angustipennis; 145. D. bahianus; 146. D. amyntas (= D. attalus); 147. D. ganglbaueri; 148. D. refulgens; 149. D. fractipes; 150. D. fasces; 151. D. haroldi; 152. D. nitidicollis; 153. D. quadratus; 154. D. similis; 155. D. denticollis." httpUri="https://zenodo.org/record/268285/files/figure.png" pageId="16" pageNumber="17">Fig. 134</figureCitation>
).
<typeStatus id="54D48836F072FFE7FDF0FE12FD6BFDC8" box="[549,653,511,536]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
female (MNHN): [Collection/ Mniszech]; [Dendrop.
<taxonomicName id="4C6F4D17F072FFE7FB32FE12FAB4FDC8" box="[1255,1362,511,536]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hirticollis" status="sp. nov.">hirticollis</taxonomicName>
<taxonomicNameLabel id="A22857FDF072FFE7FA8CFDEDFA75FDC8" box="[1369,1427,512,536]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
/ det. G. OLSOUFIEFF] partly handwritten; [
<taxonomicName id="4C6F4D17F072FFE7FDABFDC9FCC8FDED" box="[638,814,548,573]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
♀/
<taxonomicName id="4C6F4D17F072FFE7FC8DFDC9FC11FDED" authority="Ols." authorityName="Ols." box="[856,1015,548,573]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hirticollis">hirticollis Ols.</taxonomicName>
/
<typeStatus id="54D48836F072FFE7FBD2FDC9FB7CFDEC" box="[1031,1178,548,572]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
/ P. Arnaud DET 1982] partly handwritten, red border; [WORLD/ SCARAB./ DATABASE/ WSD00016487]; [
<taxonomicName id="4C6F4D17F072FFE7FB68FDAAFA8BFDB0" box="[1213,1389,583,608]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
♀/
<taxonomicName id="4C6F4D17F072FFE7FF42FD81FEE4FD55" box="[151,258,620,645]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hirticollis">hirticollis</taxonomicName>
/
<bibRefCitation id="EFFE4B65F072FFE7FEC4FD81FE2FFD54" author="Olsoufieff" box="[273,457,620,645]" pageId="16" pageNumber="17" refString="Olsoufieff, G. d' (1924) Les Phanaeides (Coleoptera-Lamellicornia), Famille Scarabaeidae-Tr. Coprini. Insecta, 13, 4 - 172, 15 pls." type="journal article" year="1924">Olsoufieff, 1924</bibRefCitation>
/ vid. Génier &amp; Arnaud, 2009]
</paragraph>
</subSubSection>
<subSubSection id="C375651FF072FFE7FF12FD62FDC7FC3D" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD03694F072FFE7FF12FD62FAF1FC50" blockId="16.[151,1437,151,1256]" pageId="16" pageNumber="17">
<emphasis id="B91BEA86F072FFE7FF12FD62FE53FD78" bold="true" box="[199,437,655,680]" pageId="16" pageNumber="17">Material examined</emphasis>
.
<collectingCountry id="F3787604F072FFE7FE1AFD62FD9CFD78" box="[463,634,655,680]" name="Argentina" pageId="16" pageNumber="17">
<emphasis id="B91BEA86F072FFE7FE1AFD62FD9CFD78" bold="true" box="[463,634,655,680]" pageId="16" pageNumber="17">ARGENTINA</emphasis>
</collectingCountry>
: MISIONES, Departamento Concepción, Santa María, (
<geoCoordinate id="EE5B5053F072FFE7FAE9FD62FA71FD78" box="[1340,1431,655,680]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="-27.9">27°54'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F072FFE7FF42FD59FF1BFD1D" box="[151,253,692,717]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="-55.4">55°24'W</geoCoordinate>
),
<date id="FFD11054F072FFE7FEC5FD59FEB9FD1D" box="[272,351,692,717]" pageId="16" pageNumber="17" value="1960-05">v.1960</date>
, coll. M.J. Viana—
<specimenCount id="9D69FD1DF072FFE7FDE9FD59FD64FD1D" box="[572,642,692,717]" pageId="16" pageNumber="17" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F072FFE7FD44FD59FD03FD1C" box="[657,741,692,716]" pageId="16" pageNumber="17">BDGC</collectionCode>
); same locality,
<date id="FFD11054F072FFE7FC4DFD59FC09FD1D" box="[920,1007,692,717]" pageId="16" pageNumber="17" value="1958-11">xi.1958</date>
, coll. M.J. Viana—
<specimenCount id="9D69FD1DF072FFE7FB18FD59FACFFD1D" box="[1229,1321,692,717]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F072FFE7FAE2FD59FA76FD1C" box="[1335,1424,692,716]" pageId="16" pageNumber="17">CMNC</collectionCode>
); same locality,
<date id="FFD11054F072FFE7FE94FD3AFE46FD20" box="[321,416,727,752]" pageId="16" pageNumber="17" value="1948-12">xii.1948</date>
, coll. M.J. Viana—
<specimenCount id="9D69FD1DF072FFE7FD58FD3AFD0BFD20" box="[653,749,727,752]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F072FFE7FCD5FD35FCBFFD20" box="[768,857,728,752]" pageId="16" pageNumber="17">CMNC</collectionCode>
);
<collectingCountry id="F3787604F072FFE7FCA4FD35FC3CFD20" box="[881,986,728,752]" name="Brazil" pageId="16" pageNumber="17">BRAZIL</collectingCountry>
: [unspecified locality], [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF072FFE7FE99FD11FE4AFCC5" box="[332,428,764,789]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F072FFE7FE15FD11FDF1FCC4" box="[448,535,764,788]" pageId="16" pageNumber="17">ZMHB</collectionCode>
);
<collectionCode id="ED7EAE51F072FFE7FDFAFD11FD61FCC5" box="[559,647,764,789]" pageId="16" pageNumber="17">GOIÁS</collectionCode>
, Fazenda Monjolinho, Corumbá, (
<geoCoordinate id="EE5B5053F072FFE7FBCEFD11FB93FCC5" box="[1051,1141,764,789]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="925" value="-15.916667">15°55'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F072FFE7FB50FD11FB0AFCC4" box="[1157,1260,764,788]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="925" value="-48.8">48°48'W</geoCoordinate>
),
<date id="FFD11054F072FFE7FAD1FD11FABDFCC5" box="[1284,1371,764,789]" pageId="16" pageNumber="17" value="1945-11">xi.1945</date>
, coll. Barreto—
<specimenCount id="9D69FD1DF072FFE7FED9FCCDFE81FCE8" box="[268,359,799,824]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F072FFE7FEA3FCCDFE29FCE8" box="[374,463,800,824]" pageId="16" pageNumber="17">WDEC</collectionCode>
);
<collectionCode id="ED7EAE51F072FFE7FE37FCCDFDDAFCE8" box="[482,572,800,824]" pageId="16" pageNumber="17">MINAS</collectionCode>
GERAIS, Cupí,
<date id="FFD11054F072FFE7FD2FFCF2FC8BFCE8" box="[762,877,799,824]" pageId="16" pageNumber="17" value="1982-07-03">3.vii.1982</date>
, coll. P. Moret—
<specimenCount id="9D69FD1DF072FFE7FBE4FCCDFB6AFCE8" box="[1073,1164,799,824]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F072FFE7FB4FFCCDFB03FCE8" box="[1178,1253,800,824]" pageId="16" pageNumber="17">CPFA</collectionCode>
); Santa Bárbara, (
<geoCoordinate id="EE5B5053F072FFE7FF4AFCA9FEC6FC8D" box="[159,288,836,861]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="-19.958889">19°57'32''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F072FFE7FEE5FCA9FE5BFC8D" box="[304,445,836,861]" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="15" value="-43.415">43°24'54''W</geoCoordinate>
),
<date id="FFD11054F072FFE7FE01FCA9FDDEFC8C" box="[468,568,836,860]" pageId="16" pageNumber="17" value="1993-10-08">8.x.1993</date>
, coll. Zanuncio—
<specimenCount id="9D69FD1DF072FFE7FCDAFCA9FC88FC8D" box="[783,878,836,861]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F072FFE7FC54FCA9FC33FC8C" box="[897,981,836,860]" pageId="16" pageNumber="17">CEMT</collectionCode>
); [unspecified locality], [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF072FFE7FE99FC85FE4EFC50" box="[332,424,871,896]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F072FFE7FE62FC85FDF7FC50" box="[439,529,872,896]" pageId="16" pageNumber="17">BMNH</collectionCode>
); [NO
<collectionCode id="ED7EAE51F072FFE7FD88FC85FD4BFC50" box="[605,685,872,896]" pageId="16" pageNumber="17">DATA</collectionCode>
]: -., coll. [anonymous]—
<specimenCount id="9D69FD1DF072FFE7FC1CFC85FBC2FC50" box="[969,1060,871,896]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(
<typeStatus id="54D48836F072FFE7FBE6FC8AFB7FFC50" box="[1075,1177,871,896]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
) (
<collectionCode id="ED7EAE51F072FFE7FB7AFC85FAEDFC50" LSID="urn:lsid:biocol.org:col:34988" box="[1199,1291,872,896]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8BD03694F072FFE7FF12FC61FDC7FC3D" blockId="16.[151,1437,151,1256]" pageId="16" pageNumber="17">
<emphasis id="B91BEA86F072FFE7FF12FC61FE63FC75" bold="true" box="[199,389,908,933]" pageId="16" pageNumber="17">Natural history</emphasis>
.
<bibRefCitation id="EFFE4B65F072FFE7FE4FFC61FD3AFC74" author="Martinez" box="[410,732,908,933]" pageId="16" pageNumber="17" refString="Martinez, A. &amp; Clavijo, J. (1990) Notas sobre Phanaeina Venezolanos, con descripcion de una nueva subespecie de Diabroctis (Coleoptera, Scarabaeidae, Coprini). Boletin de Entomologia Venezolana, Nueva Seria, 5 (20) 147 - 157." type="journal article" year="1990">Martínez &amp; Clavijo (1990)</bibRefCitation>
report that M.J. Viana caught numerous specimens of
<taxonomicName id="4C6F4D17F072FFE7FA54FC60FEE2FC18" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="B91BEA86F072FFE7FA54FC60FEE2FC18" italics="true" pageId="16" pageNumber="17">D. hirticollis</emphasis>
</taxonomicName>
leaving a termite nest during a rain in Misiones,
<collectingCountry id="F3787604F072FFE7FCF1FC42FC71FC18" box="[804,919,943,968]" name="Argentina" pageId="16" pageNumber="17">Argentina</collectingCountry>
. A specimen from Minas Gerais was collected in a spider web under a street light.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF072FFE7FF12FC1AFC37FB38" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BD03694F072FFE7FF12FC1AFC37FB38" blockId="16.[151,1437,151,1256]" pageId="16" pageNumber="17">
<emphasis id="B91BEA86F072FFE7FF12FC1AFED2FBC0" bold="true" box="[199,308,1015,1040]" pageId="16" pageNumber="17">Remarks</emphasis>
. A single male of this species is known and differs from the females in having the anterior pronotal carina wider and somewhat atrophied medially in addition to having the abdominal sternites 5–7 almost completely setose. The parameres are simply rounded in dorsal view and the surface finely irregular apically (Figs. 50–51). Variation, as usual, the extent of the green metallic markings on head and pronotum varies. The Argentinean specimens differs in having the anterolateral surface of pronotum more simply convex and the pronotal disc less heavily sculptured between setose punctures as well as having the elytral apex more evenly convex with striae less impressed. For the moment this variation will be considered intraspecific.
</paragraph>
</subSubSection>
</treatment>
</document>
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