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<document id="71D9F734A43E3A621404502D3A50EB3E" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F06AFFFDFF42F92AFCE6FADD" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Coprophanaeoides) carinifer Genier &amp; Arnaud, new species" docType="treatment" docVersion="10" lastPageNumber="11" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="9" updateTime="1698234910999" updateUser="plazi">
<mods:mods id="66B7D49F9922190C82ED740B67A6915D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="36E4E6EEA201A62BC3033BE1BBFF6B9D">
<mods:title id="2AFF48086131040DC35B0BFF8B39E19C">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
</mods:titleInfo>
<mods:name id="94920FFC807CA139784F6299472AB2D9" type="personal">
<mods:role id="C310F5F2BD4F2F6C7B2402D5D60C3563">
<mods:roleTerm id="BDD059B328FDBA670A64D967B2D9AABF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D1C6E4A8388A46C94D65A4967F893327">François Génier</mods:namePart>
</mods:name>
<mods:name id="432B395D82A28347D2C7F5EBE89CEDBF" type="personal">
<mods:role id="9C6238D475A540E2BEC4F7F4985C626C">
<mods:roleTerm id="24B33D40E2FA990703AE7ECE7FB06865">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02C04A00F80AE2B94BFE4E6BE98F5AAD">Patrick Arnaud</mods:namePart>
</mods:name>
<mods:typeOfResource id="6FBD98DB2093DA23F2B62DC1A54F91FE">text</mods:typeOfResource>
<mods:relatedItem id="29DB1CE25939A82F8410525D135EC9F1" type="host">
<mods:titleInfo id="056E8B77AC82CA14C5B3EBFE073F4D55">
<mods:title id="15AC3C62EFA432EF32EA1C3840B78402">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6727B9EC57D863CE2E92724BB591C2A4">
<mods:date id="AF532A6A2C06A0CFE9BF179B04730B45">2016</mods:date>
<mods:detail id="A5B6D4B7AA14DC32234D35873090C398" type="volume">
<mods:number id="F49EE17A32266A8971DFC1843546443A">4099</mods:number>
</mods:detail>
<mods:detail id="D06B234AED841559BEF7C8B717EA8A85" type="issue">
<mods:number id="4DDE1FFB7216ABDDF733887DDD5A449A">1</mods:number>
</mods:detail>
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<mods:classification id="80F0CDAA5696C3C7B8D91D933008E5BA">journal volume</mods:classification>
<mods:identifier id="4714765E46E2F617F73D57D192A76F22" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
<mods:identifier id="6106FD46EB2586831EA34CA4B29F1FF8" type="GBIF-Dataset">5d7eab55-66e2-43ed-b764-bac0c6e5b6d6</mods:identifier>
<mods:identifier id="296D50F5DFB00DF541B896C6CCFA4CFC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C7072E1B0EB773143EB4CDD61F6F3710" type="Zenodo-Dep">268274</mods:identifier>
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</mods:mods>
<treatment id="03C68782F06AFFFDFF42F92AFCE6FADD" ID-DOI="http://doi.org/10.5281/zenodo.5780254" ID-GBIF-Taxon="127631979" ID-Zenodo-Dep="5780254" LSID="urn:lsid:plazi:treatment:03C68782F06AFFFDFF42F92AFCE6FADD" httpUri="http://treatment.plazi.org/id/03C68782F06AFFFDFF42F92AFCE6FADD" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="C375651FF06AFFFFFF42F92AFBDEF930" box="[151,1080,1735,1760]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BD03694F06AFFFFFF42F92AFBDEF930" blockId="8.[151,1080,1735,1797]" box="[151,1080,1735,1760]" pageId="8" pageNumber="9">
<heading id="D09881F8F06AFFFFFF42F92AFBDEF930" bold="true" box="[151,1080,1735,1760]" fontSize="10" level="2" pageId="8" pageNumber="9" reason="2">
<emphasis id="B91BEA86F06AFFFFFF42F92AFBDEF930" bold="true" box="[151,1080,1735,1760]" pageId="8" pageNumber="9">
2.
<taxonomicName id="4C6F4D17F06AFFFFFF66F92AFC47F930" ID-CoL="6CLD3" authority="Genier &amp; Arnaud" authorityName="Genier &amp; Arnaud" box="[179,929,1735,1760]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="carinifer" status="sp. nov." subGenus="Coprophanaeoides">
<emphasis id="B91BEA86F06AFFFFFF66F92AFD25F930" bold="true" box="[179,707,1735,1760]" italics="true" pageId="8" pageNumber="9">Dendropaemon (Coprophanaeoides) carinifer</emphasis>
Génier &amp; Arnaud
</taxonomicName>
,
<taxonomicNameLabel id="A22857FDF06AFFFFFC7BF92AFBDEF930" box="[942,1080,1735,1760]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06AFFFFFF42F901FE5EF8D5" box="[151,440,1772,1797]" pageId="8" pageNumber="9" type="description">
<paragraph id="8BD03694F06AFFFFFF42F901FE5EF8D5" blockId="8.[151,1080,1735,1797]" box="[151,440,1772,1797]" pageId="8" pageNumber="9">
(
<figureCitation id="13542A11F06AFFFFFF4AF901FF08F8D4" box="[159,238,1772,1797]" captionStart="FIGURES 1 – 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 – 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="8" pageNumber="9">Figs. 2</figureCitation>
, 42–43, 111, 156)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06AFFFFFF42F8D9FD02F89D" box="[151,740,1844,1869]" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BD03694F06AFFFFFF42F8D9FD02F89D" blockId="8.[151,1436,1844,2013]" box="[151,740,1844,1869]" pageId="8" pageNumber="9">
<emphasis id="B91BEA86F06AFFFFFF42F8D9FECBF89D" bold="true" box="[151,301,1844,1869]" pageId="8" pageNumber="9">
<typeStatus id="54D48836F06AFFFFFF42F8D9FF34F89D" box="[151,210,1844,1869]" pageId="8" pageNumber="9">Type</typeStatus>
locality
</emphasis>
. MA-10, Pedrinhas, Maranhão,
<collectingCountry id="F3787604F06AFFFFFD4DF8D9FD06F89D" box="[664,736,1844,1869]" name="Brazil" pageId="8" pageNumber="9">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06AFFFEFF12F8BAFBDDFEB4" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BD03694F06AFFFEFF12F8BAFBDDFEB4" blockId="8.[151,1436,1844,2013]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B91BEA86F06AFFFFFF12F8BAFEA7F8BF" bold="true" box="[199,321,1879,1904]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
The long elytral pilosity combined with the sharply carinate lateral edge of the pronotal lateral depressions will separate
<taxonomicName id="4C6F4D17F06AFFFFFE6EF890FDA6F844" box="[443,576,1916,1940]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="carinifer">
<emphasis id="B91BEA86F06AFFFFFE6EF890FDA6F844" box="[443,576,1916,1940]" italics="true" pageId="8" pageNumber="9">D. carinifer</emphasis>
</taxonomicName>
from most other in the genus. The much less pilose dorsum, especially the glabrous eighth elytral interval will separate it from
<taxonomicName id="4C6F4D17F06AFFFFFD3BF84CFC7DF869" box="[750,923,1952,1977]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pilosissimus">
<emphasis id="B91BEA86F06AFFFFFD3BF84CFC7DF869" box="[750,923,1952,1977]" italics="true" pageId="8" pageNumber="9">D. pilosissimus</emphasis>
</taxonomicName>
and the simply broadly arcuate clypeal edge will distinguish it from
<taxonomicName id="4C6F4D17F06AFFFFFE61F828FDD0F80D" box="[436,566,1988,2013]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="furtadoi">
<emphasis id="B91BEA86F06AFFFFFE61F828FDD0F80D" box="[436,566,1988,2013]" italics="true" pageId="8" pageNumber="9">D. furtadoi</emphasis>
</taxonomicName>
. Finally, it can be separated from its sibling species
<taxonomicName id="4C6F4D17F06AFFFFFB60F828FAA3F80D" box="[1205,1349,1988,2013]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cribrosus">
<emphasis id="B91BEA86F06AFFFFFB60F828FAA3F80D" box="[1205,1349,1988,2013]" italics="true" pageId="8" pageNumber="9">D. cribrosus</emphasis>
</taxonomicName>
by the distinctly less punctate pronotal disc. In
<taxonomicName id="4C6F4D17F06BFFFEFDB8FF74FD12FF60" box="[621,756,152,176]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="carinifer">
<emphasis id="B91BEA86F06BFFFEFDB8FF74FD12FF60" box="[621,756,152,176]" italics="true" pageId="9" pageNumber="10">D. carinifer</emphasis>
</taxonomicName>
there is at most 25 (usually 10 to 12) non confluent large setiferous punctures on disc as in
<taxonomicName id="4C6F4D17F06BFFFEFDC2FF50FD42FF05" box="[535,676,188,213]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cribrosus">
<emphasis id="B91BEA86F06BFFFEFDC2FF50FD42FF05" box="[535,676,188,213]" italics="true" pageId="9" pageNumber="10">D. cribrosus</emphasis>
</taxonomicName>
there is at least twice that many confluent punctures. Additionally, the clypeal pilosity is much reduced in
<taxonomicName id="4C6F4D17F06BFFFEFD8DFF0CFD3BFF28" box="[600,733,224,248]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="carinifer">
<emphasis id="B91BEA86F06BFFFEFD8DFF0CFD3BFF28" box="[600,733,224,248]" italics="true" pageId="9" pageNumber="10">D. carinifer</emphasis>
</taxonomicName>
and in most cases completely absent, when present, only few setae are set on each side of midline in front of the clypeofrontal tubercle. In
<taxonomicName id="4C6F4D17F06BFFFEFC26FEE8FB9BFECD" box="[1011,1149,260,285]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cribrosus">
<emphasis id="B91BEA86F06BFFFEFC26FEE8FB9BFECD" box="[1011,1149,260,285]" italics="true" pageId="9" pageNumber="10">D. cribrosus</emphasis>
</taxonomicName>
, the pilosity is present on most of the surface and the surface anterior to the clypeofrontal tubercle present minute setiferous granules as opposed to large transverse rugulae similar to the remaining surface in
<taxonomicName id="4C6F4D17F06BFFFEFC67FEA0FBD2FEB4" box="[946,1076,332,356]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="carinifer">
<emphasis id="B91BEA86F06BFFFEFC67FEA0FBD2FEB4" box="[946,1076,332,356]" italics="true" pageId="9" pageNumber="10">D. carinifer</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF06BFFFDFF12FE82FC9AFF28" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="8BD03694F06BFFFEFF12FE82FC0EFCC4" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
<emphasis id="B91BEA86F06BFFFEFF12FE82FEB7FE58" bold="true" box="[199,337,367,392]" pageId="9" pageNumber="10">Description</emphasis>
.
<collectingCountry id="F3787604F06BFFFEFE8AFE82FE7FFE58" box="[351,409,367,392]" name="Malta" pageId="9" pageNumber="10">Male</collectingCountry>
<typeStatus id="54D48836F06BFFFEFE4AFE82FDE7FE58" box="[415,513,367,392]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
(
<figureCitation id="13542A11F06BFFFEFDC4FE82FDB0FE58" box="[529,598,367,392]" captionStart="FIGURES 1 – 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 – 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="9" pageNumber="10">Fig. 2</figureCitation>
).
<emphasis id="B91BEA86F06BFFFEFDBFFE82FD41FE58" bold="true" box="[618,679,367,392]" pageId="9" pageNumber="10">Body</emphasis>
. Body moderately large, length
<quantity id="4C979B71F06BFFFEFBDAFE82FB92FE57" box="[1039,1140,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="9" pageNumber="10" unit="mm" value="11.5">11.5 mm</quantity>
, maximum width
<quantity id="4C979B71F06BFFFEFAEAFE82FA7EFE57" box="[1343,1432,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="10" unit="mm" value="6.5">6.5 mm</quantity>
; body subrectangular in dorsal view; dorsum slightly convex.
<emphasis id="B91BEA86F06BFFFEFC82FE79FC7EFE7D" bold="true" box="[855,920,404,429]" pageId="9" pageNumber="10">Color</emphasis>
. Dorsal surface dark brown to black, glossy, with green metallic sheen; head black along anterior edge of clypeus, metallic green on remaining surface; pronotum with green metallic sheen; elytra with uniform green metallic sheen; ventrum with faint greenish metallic sheen; pygidium with green metallic sheen; legs with coppery and greenish metallic sheen on femora and tibiae.
<emphasis id="B91BEA86F06BFFFEFF42FDC9FF31FDED" bold="true" box="[151,215,548,573]" pageId="9" pageNumber="10">Head</emphasis>
. Clypeus broadly arcuate, anterior portion slightly upturned; clypeal teeth acutely triangular; clypeal median emargination v-shaped, clypeal edge acutely notched on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin ill-defined, lacking sharp carina posteriorly, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with small and blunt irregular tubercles, lacking distinct transverse carina, simply convex; clypeofrontal carina rather low, approximately 4 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge straight in frontal view; eyes large in dorsal view, interocular ratio 3.1.
</paragraph>
<paragraph id="8BD03694F06BFFFEFF12FCF2FC59F869" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="10">
<emphasis id="B91BEA86F06BFFFEFF12FCF2FEA6FCE8" bold="true" box="[199,320,799,824]" pageId="9" pageNumber="10">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.6; disc of pronotum minutely punctate basally with some scattered large setiferous puncture medially, with dense squamose punctation anteriorly, with an ill-defined shallow longitudinal depression on posterior half; pronotal anterior margin unmodified lateral to eye; anterior portion with a slightly tri-sinuous carina notched medially; anterior angles surface finely granulate, similar to lateral margin along posterior edge of anterior margin; lateral fossae oval, bordered laterally by a sharp carina and anteriorly by a blunt tubercle; lateral portions strongly explanate; pronotal basal fossae ill-defined, slightly concave; posterior margin well-defined and crenulate, with several long setae.
<emphasis id="B91BEA86F06BFFFEFF42FBF1FF05FBE5" bold="true" box="[151,227,1052,1077]" pageId="9" pageNumber="10">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.2; elytral base distinctly marginate; elytral striae 1–4 moderately wide basally and fine apically, evenly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures fine, well-defined and setiferous, adjacent strial edge encroaching on interval, stria 1 weakly impressed apically, going straight to elytral apical margin; interstriae slightly convex, minutely punctate and with few larger setiferous punctures along striae, surface glossy.
<emphasis id="B91BEA86F06BFFFEFAE4FB41FF04FB38" bold="true" pageId="9" pageNumber="10">Thoracic sterna</emphasis>
. Proepisternal carina absent; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, y-shaped.
<emphasis id="B91BEA86F06BFFFEFE70FB19FE3BFADD" bold="true" box="[421,477,1268,1293]" pageId="9" pageNumber="10">Legs</emphasis>
. Profemur posterior surface flat and glabrous internally, posterointernal margin rather thin, uneven, internal edge rather wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle bluntly lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with some ill-defined irregular punctures externally to median carina, surface glossy between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge slightly sinuate in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less rounded, with several large elongate setiferous punctures. Mesotarsus similar in shape to metatarsus, 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur elongate, internal and lateral edges mostly parallel in ventral view, more than twice as long as wide, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with an irregular fine sulcus medially. Metatibia rather slender, gradually widening toward apex in anterior view, anterior surface with distinct row of setae, surface glossy, metatibial posterior surface flat between longitudinal row of setae and lateral edge, with ill-defined microsculpture. Metatarsus 3-segmented (
<figureCitation id="13542A11F06BFFFEFB57F901FB0FF8D4" box="[1154,1257,1772,1797]" captionStart="FIGURES 110 – 117" captionStartId="112.[151,264,1933,1956]" captionTargetBox="[219,1362,202,1901]" captionTargetId="figure@112.[212,1375,193,1913]" captionTargetPageId="112" captionText="FIGURES 110 – 117. Metatarsus, ventral view. 110. D. bluti; 111. D. carinifer; 112. D. aenigmaticus; 113. D. amazonicus; 114. D. angustulus; 115. D. viridis; 116. D. viridipennis; 117. D. convexus." httpUri="https://zenodo.org/record/268282/files/figure.png" pageId="9" pageNumber="10">Fig. 111</figureCitation>
), first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge.
<emphasis id="B91BEA86F06BFFFEFE22F8D9FD09F89D" bold="true" box="[503,751,1844,1869]" pageId="9" pageNumber="10">Abdominal sternites</emphasis>
. Sternites 3–6 longitudinally flat; sternites 4–6 with 1–3 unaligned rows of setae laterally, narrowly glabrous medially; sternite 7 approximately longitudinally flat medially, shorter than segment 6 along midline; pygidium finely punctate on disc.
<emphasis id="B91BEA86F06BFFFEFC1BF891FB9FF845" bold="true" box="[974,1145,1916,1941]" pageId="9" pageNumber="10">
<collectingCountry id="F3787604F06BFFFEFC1BF891FBECF845" box="[974,1034,1916,1941]" name="Malta" pageId="9" pageNumber="10">Male</collectingCountry>
genitalia
</emphasis>
(Figs. 42–43). Parameres simply rounded apically in dorsal view; surface smooth, glossy apically.
</paragraph>
<paragraph id="8BD03694F06BFFFEFF12F829FA6DF80C" blockId="9.[151,1437,151,2013]" box="[199,1419,1988,2013]" pageId="9" pageNumber="10">
<emphasis id="B91BEA86F06BFFFEFF12F829FE90F80D" bold="true" box="[199,374,1988,2013]" pageId="9" pageNumber="10">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF06BFFFEFE53F829FE0BF80D" box="[390,493,1988,2013]" pageId="9" pageNumber="10" type="male">10 males</specimenCount>
,
<specimenCount id="9D69FD1DF06BFFFEFE2CF829FD9FF80D" box="[505,633,1988,2013]" pageId="9" pageNumber="10" type="female">12 females</specimenCount>
). Length: male 11.0–12.5 (11.7±.5), female 10.5–13.5 (12.2±0.9) mm.
</paragraph>
<paragraph id="8BD03694F068FFFDFF12FF7AFC9AFF28" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="11">
<emphasis id="B91BEA86F068FFFDFF12FF7AFE4EFF60" bold="true" box="[199,424,151,176]" pageId="10" pageNumber="11">
Primary
<typeStatus id="54D48836F068FFFDFEE3FF7AFE8FFF60" box="[310,361,151,176]" pageId="10" pageNumber="11">type</typeStatus>
data
</emphasis>
.
<typeStatus id="54D48836F068FFFDFE6CFF7AFDC7FF60" box="[441,545,151,176]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
male (CEMT): [
<date id="FFD11054F068FFFDFD31FF75FC81FF60" box="[740,871,152,176]" pageId="10" pageNumber="11" value="1987-09-02">02/IX/1987</date>
/ BR-MA-Pedrinhas/ armadilha luninosa/ Ma-10/ Bergmann, E col.] handwritten; WORLD/ SCARAB./ DATABASE/ WSD00017508]; [
<typeStatus id="54D48836F068FFFDFB28FF51FA75FF04" box="[1277,1427,188,212]" pageId="10" pageNumber="11" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C6F4D17F068FFFDFF42FF32FEA2FF28" box="[151,324,223,248]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F068FFFDFF42FF32FEA2FF28" box="[151,324,223,248]" italics="true" pageId="10" pageNumber="11">Dendropaemon</emphasis>
</taxonomicName>
/
<taxonomicName id="4C6F4D17F068FFFDFE86FF0DFE53FF28" box="[339,437,224,248]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="carinifer" status="sp. nov.">
<emphasis id="B91BEA86F068FFFDFE86FF0DFE53FF28" box="[339,437,224,248]" italics="true" pageId="10" pageNumber="11">carinifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22857FDF068FFFDFE69FF0DFE08FF28" box="[444,494,224,248]" pageId="10" pageNumber="11" rank="species">n.sp.</taxonomicNameLabel>
/ Génier &amp; Arnaud, 2014] red card.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF068FFFDFF12FEE9FB34FC50" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BD03694F068FFFDFF12FEE9FB34FC50" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="11">
<emphasis id="B91BEA86F068FFFDFF12FEE9FE52FECD" bold="true" box="[199,436,260,285]" pageId="10" pageNumber="11">Material examined</emphasis>
.
<collectingCountry id="F3787604F068FFFDFE19FEE9FDD1FECD" box="[460,567,260,285]" name="Brazil" pageId="10" pageNumber="11">
<emphasis id="B91BEA86F068FFFDFE19FEE9FDD1FECD" bold="true" box="[460,567,260,285]" pageId="10" pageNumber="11">BRAZIL</emphasis>
</collectingCountry>
:
<collectionCode id="ED7EAE51F068FFFDFD84FEE9FD4BFECC" box="[593,685,260,284]" pageId="10" pageNumber="11">BAHIA</collectionCode>
, Cândido Sales, (
<geoCoordinate id="EE5B5053F068FFFDFC5AFEE9FBF6FECD" box="[911,1040,260,285]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="15" value="-15.505">15°30'18''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F068FFFDFBF3FEE9FB57FECC" box="[1062,1201,260,284]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="15" value="-41.238888">41°14'20''W</geoCoordinate>
),
<date id="FFD11054F068FFFDFB05FEE9FAD6FECD" box="[1232,1328,260,285]" pageId="10" pageNumber="11" value="2012-12">xii.2012</date>
, coll. P. Wagner—
<specimenCount id="9D69FD1DF068FFFDFEDAFEC5FE97FE90" box="[271,369,295,320]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(
<typeStatus id="54D48836F068FFFDFE53FEC5FE0CFE90" box="[390,490,296,320]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFDD2FEC5FDBDFE90" box="[519,603,296,320]" pageId="10" pageNumber="11">CEMT</collectionCode>
);
<collectionCode id="ED7EAE51F068FFFDFDA0FECAFD31FE90" box="[629,727,295,320]" pageId="10" pageNumber="11">CEARÁ</collectionCode>
, Chapada do Araripe, (
<geoCoordinate id="EE5B5053F068FFFDFC21FEC5FBA7FE90" box="[1012,1089,296,320]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="-7.3333335">7°20'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F068FFFDFB86FEC5FB4DFE90" box="[1107,1195,296,320]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="-40.0">40°0'W</geoCoordinate>
),
<date id="FFD11054F068FFFDFB13FECAFAC9FE90" box="[1222,1327,295,320]" pageId="10" pageNumber="11" value="1962-04-05">5.iv.1962</date>
, coll. D. Zajciw—
<specimenCount id="9D69FD1DF068FFFDFED6FEA1FE96FEB5" box="[259,368,332,357]" pageId="10" pageNumber="11" type="female">3 females</specimenCount>
(incl. 3
<typeStatus id="54D48836F068FFFDFE0FFEA1FDA1FEB4" box="[474,583,332,356]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFDB4FEA1FD54FEB4" box="[609,690,332,356]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="10" pageNumber="11">MNRJ</collectionCode>
); Parque Botânico do Ceará, Caucaia, (
<geoCoordinate id="EE5B5053F068FFFDFB54FEA1FB15FEB4" box="[1153,1267,332,356]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="15" value="-3.7119446">3°42'43''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F068FFFDFAD6FEA1FA69FEB5" box="[1283,1423,332,357]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="15" value="-38.645836">38°38'45''W</geoCoordinate>
),
<date id="FFD11054F068FFFDFF42FE82FEEDFE58" box="[151,267,367,392]" pageId="10" pageNumber="11" value="2006-02-27">27.ii.2006</date>
, coll. F.A. Nunes—
<specimenCount id="9D69FD1DF068FFFDFE27FE9DFDAFFE58" box="[498,585,367,392]" pageId="10" pageNumber="11" type="male">2 males</specimenCount>
(incl. 2
<typeStatus id="54D48836F068FFFDFD7AFE9DFCFAFE58" box="[687,796,368,392]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
) (MEFEIS); MARANHÃO, Base da Geraldina, Parque Estadual Mirador, Mirador, (
<geoCoordinate id="EE5B5053F068FFFDFE22FE79FDBFFE7D" box="[503,601,404,429]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="92" value="-6.37">6°22.2'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F068FFFDFDBAFE79FD0DFE7C" box="[623,747,404,428]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="92" value="-44.363335">44°21.8'W</geoCoordinate>
),
<date id="FFD11054F068FFFDFCDCFE79FC53FE7D" box="[777,949,404,429]" pageId="10" pageNumber="11" value="2006-12-20" valueMax="2006-12-24" valueMin="2006-12-20">20–24.xii.2006</date>
, coll. F. Limeira-de-Oliveira—
<specimenCount id="9D69FD1DF068FFFDFAE3FE79FA7AFE7D" box="[1334,1436,404,429]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(
<typeStatus id="54D48836F068FFFDFF4BFE55FEE4FE00" box="[158,258,440,464]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFECBFE55FE94FE00" box="[286,370,440,464]" pageId="10" pageNumber="11">CEMT</collectionCode>
); same locality,
<date id="FFD11054F068FFFDFDE0FE5AFD31FE00" box="[565,727,439,464]" pageId="10" pageNumber="11" value="2007-04-20" valueMax="2007-04-22" valueMin="2007-04-20">20–22.iv.2007</date>
, coll. F. Limeira-de-Oliveira—
<specimenCount id="9D69FD1DF068FFFDFB9DFE5AFB4FFE00" box="[1096,1193,439,464]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(
<typeStatus id="54D48836F068FFFDFB69FE55FAC6FE00" box="[1212,1312,440,464]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFAE9FE55FA76FE00" box="[1340,1424,440,464]" pageId="10" pageNumber="11">CEMT</collectionCode>
); same locality,
<date id="FFD11054F068FFFDFEE3FE31FE0AFE25" box="[310,492,476,501]" pageId="10" pageNumber="11" value="2009-02-22" valueMax="2009-03-01" valueMin="2009-02-22">22.ii.–1.iii.2009</date>
, coll. M.B. Aguilar-Neto &amp; J.A. Holanda—
<specimenCount id="9D69FD1DF068FFFDFC0CFE31FBF8FE25" box="[985,1054,476,501]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(
<typeStatus id="54D48836F068FFFDFBF9FE31FB76FE24" box="[1068,1168,476,500]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFB73FE31FB17FE24" box="[1190,1265,476,500]" pageId="10" pageNumber="11">CPFA</collectionCode>
); same locality,
<date id="FFD11054F068FFFDFF42FE12FE81FDC8" box="[151,359,511,536]" pageId="10" pageNumber="11" value="2011-07-30" valueMax="2011-08-06" valueMin="2011-07-30">30.vii.–6.viii.2011</date>
, coll. F. Limeira-de-Oliveira, T.T.A. Silva &amp; A.A. Santos—
<specimenCount id="9D69FD1DF068FFFDFBC0FDEDFB94FDC8" box="[1045,1138,511,536]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(
<typeStatus id="54D48836F068FFFDFB54FDEDFB03FDC8" box="[1153,1253,512,536]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFB28FDEDFAB7FDC8" box="[1277,1361,512,536]" pageId="10" pageNumber="11">CEMT</collectionCode>
); Base do Mosquito, Parque Estadual Mirador, Mirador,
<date id="FFD11054F068FFFDFD01FDC9FCB3FDED" box="[724,853,548,573]" pageId="10" pageNumber="11" value="2011-02-04" valueMax="2011-02-08" valueMin="2011-02-04">4–8.ii.2011</date>
, coll. F. Limeira-de-Oliveira—
<specimenCount id="9D69FD1DF068FFFDFB15FDC9FAF9FDED" box="[1216,1311,548,573]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(
<typeStatus id="54D48836F068FFFDFAE4FDC9FA73FDEC" box="[1329,1429,548,572]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFF4AFDA5FF15FDB0" box="[159,243,584,608]" pageId="10" pageNumber="11">CEMT</collectionCode>
); Pedrinhas, (
<geoCoordinate id="EE5B5053F068FFFDFE47FDAAFDE2FDB0" box="[402,516,583,608]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="15" value="-2.6236112">2°37'25''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F068FFFDFDC6FDA5FD79FDB0" box="[531,671,584,608]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="15" value="-44.224445">44°13'28''W</geoCoordinate>
),
<date id="FFD11054F068FFFDFD62FDAAFCD7FDB0" box="[695,817,583,608]" pageId="10" pageNumber="11" value="1987-03-11">11.iii.1987</date>
, coll. E.C. Bergmann—
<specimenCount id="9D69FD1DF068FFFDFB98FDA5FB70FDB0" box="[1101,1174,583,608]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(
<typeStatus id="54D48836F068FFFDFB7DFDA5FAEAFDB0" box="[1192,1292,584,608]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (MEFEIS); same locality,
<date id="FFD11054F068FFFDFEEBFD81FE4CFD55" box="[318,426,620,645]" pageId="10" pageNumber="11" value="1987-09-02">2.ix.1987</date>
, coll. E. Bergmann—
<specimenCount id="9D69FD1DF068FFFDFD78FD81FD10FD55" box="[685,758,620,645]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(
<typeStatus id="54D48836F068FFFDFCDDFD81FC8BFD55" box="[776,877,620,645]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFC52FD81FC3DFD54" box="[903,987,620,644]" pageId="10" pageNumber="11">CEMT</collectionCode>
); Pedrinhas,
<collectingCountry id="F3787604F068FFFDFBA7FD81FB7CFD55" box="[1138,1178,620,645]" name="Iceland" pageId="10" pageNumber="11">Isla</collectingCountry>
São Luís, (
<geoCoordinate id="EE5B5053F068FFFDFAF0FD81FA71FD55" box="[1317,1431,620,645]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="15" value="-2.6236112">2°37'25''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F068FFFDFF42FD7DFEC5FD78" box="[151,291,656,680]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="15" value="-44.224445">44°13'28''W</geoCoordinate>
),
<date id="FFD11054F068FFFDFEE2FD62FE5FFD78" box="[311,441,655,680]" pageId="10" pageNumber="11" value="1984-07-20">20.vii.1984</date>
, coll. E.C. Bergmann—
<specimenCount id="9D69FD1DF068FFFDFD19FD7DFCF4FD78" box="[716,786,655,680]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(
<typeStatus id="54D48836F068FFFDFCF4FD7DFC63FD78" box="[801,901,656,680]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFC49FD7DFC16FD78" box="[924,1008,656,680]" pageId="10" pageNumber="11">CEMT</collectionCode>
); same locality,
<date id="FFD11054F068FFFDFB71FD62FAC1FD78" box="[1188,1319,655,680]" pageId="10" pageNumber="11" value="1984-07-25">25.vii.1984</date>
, coll. E.C. Bergmann—
<specimenCount id="9D69FD1DF068FFFDFEF9FD59FE6CFD1D" box="[300,394,692,717]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
allotype (MEFEIS); same locality,
<date id="FFD11054F068FFFDFCF4FD59FC4DFD1D" box="[801,939,692,717]" pageId="10" pageNumber="11" value="1984-08-28">28.viii.1984</date>
, coll. E.C. Bergmann—
<specimenCount id="9D69FD1DF068FFFDFB16FD59FAC6FD1D" box="[1219,1312,692,717]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(
<typeStatus id="54D48836F068FFFDFAE4FD59FA73FD1C" box="[1329,1429,692,716]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (MEFEIS); same locality,
<date id="FFD11054F068FFFDFE11FD35FDC1FD20" box="[452,551,728,752]" pageId="10" pageNumber="11" value="1987-05-07">7.v.1987</date>
, coll. E.C. Bergmann—
<specimenCount id="9D69FD1DF068FFFDFC94FD35FC6FFD20" box="[833,905,727,752]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(
<typeStatus id="54D48836F068FFFDFC4EFD35FC19FD20" box="[923,1023,728,752]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFBC2FD35FB8DFD20" box="[1047,1131,728,752]" pageId="10" pageNumber="11">CEMT</collectionCode>
); Posto Avançado do Mel, Parque Estadual Mirador, Mirador, (
<geoCoordinate id="EE5B5053F068FFFDFDFEFD11FD78FCC5" box="[555,670,764,789]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="15" value="-6.7305555">6°43'50''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F068FFFDFD7CFD11FCD0FCC5" box="[681,822,764,789]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="15" value="-44.983055">44°58'59''W</geoCoordinate>
),
<date id="FFD11054F068FFFDFC9CFD11FC28FCC5" box="[841,974,764,789]" pageId="10" pageNumber="11" value="2011-04-02" valueMax="2011-04-08" valueMin="2011-04-02">2–8.iv.2011</date>
, coll. F. Limeira-de-Oliveira,G.A. Reis &amp; M.S. Oliveira—
<specimenCount id="9D69FD1DF068FFFDFE87FCF2FE7EFCE8" box="[338,408,799,824]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(
<typeStatus id="54D48836F068FFFDFE72FCCDFDEDFCE8" box="[423,523,800,824]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFDF7FCCDFD90FCE8" box="[546,630,800,824]" pageId="10" pageNumber="11">CEMT</collectionCode>
); same locality,
<date id="FFD11054F068FFFDFCFEFCCDFC22FCE8" box="[811,964,800,824]" pageId="10" pageNumber="11" value="2011-05-30" valueMax="2011-05-31" valueMin="2011-05-30">30–31.v.2011</date>
, coll. F. Limeira-de-Oliveira, A.A. Santos &amp; T.T.A. Silva—
<specimenCount id="9D69FD1DF068FFFDFE8CFCA9FE21FC8D" box="[345,455,836,861]" pageId="10" pageNumber="11" type="female">2 females</specimenCount>
,
<specimenCount id="9D69FD1DF068FFFDFE07FCA9FDCFFC8D" box="[466,553,836,861]" pageId="10" pageNumber="11" type="male">4 males</specimenCount>
(incl. 6
<typeStatus id="54D48836F068FFFDFD52FCA9FD12FC8C" box="[647,756,836,860]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFCDEFCA9FCB9FC8C" box="[779,863,836,860]" pageId="10" pageNumber="11">CEMT</collectionCode>
); Reserva Ecológica do Inhamum, Caxias, (
<geoCoordinate id="EE5B5053F068FFFDFA9EFCA9FA71FC8D" box="[1355,1431,836,861]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="-4.866667">4°52'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F068FFFDFF42FC85FF1BFC50" box="[151,253,872,896]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="-43.366665">43°22'W</geoCoordinate>
),
<date id="FFD11054F068FFFDFEC4FC8AFE7FFC50" box="[273,409,871,896]" pageId="10" pageNumber="11" value="2009-06-05" valueMax="2009-06-07" valueMin="2009-06-05">5–7.vi.2009</date>
, coll. E.A. Barbosa &amp; M.B. Aguliar-Neto—
<specimenCount id="9D69FD1DF068FFFDFC5BFC8AFC0FFC50" box="[910,1001,871,896]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(
<typeStatus id="54D48836F068FFFDFC22FC85FBBDFC50" box="[1015,1115,872,896]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
) (
<collectionCode id="ED7EAE51F068FFFDFBA7FC85FB20FC50" box="[1138,1222,872,896]" pageId="10" pageNumber="11">CEMT</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C375651FF068FFFDFF12FC61FD2EFBE4" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BD03694F068FFFDFF12FC61FEC7FC18" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="11">
<emphasis id="B91BEA86F068FFFDFF12FC61FEA1FC75" bold="true" box="[199,327,908,933]" pageId="10" pageNumber="11">Etymology</emphasis>
.
<taxonomicName id="4C6F4D17F068FFFDFE89FC61FE25FC74" box="[348,451,908,932]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="carinifer">
<emphasis id="B91BEA86F068FFFDFE89FC61FE25FC74" box="[348,451,908,932]" italics="true" pageId="10" pageNumber="11">Carinifer</emphasis>
</taxonomicName>
, an adjective relating to the sharply carinate lateral edge of the pronotal lateral depressions.
</paragraph>
<paragraph id="8BD03694F068FFFDFF12FC39FD2EFBE4" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="11">
<emphasis id="B91BEA86F068FFFDFF12FC39FE98FC3D" bold="true" box="[199,382,980,1005]" pageId="10" pageNumber="11">Natural history</emphasis>
. Half of the specimens were collected using black light traps. Two specimens were collected in closed tree forest (mata tabuleiro) and two others in mangrove areas. A single specimen was collected in a pitfall trap baited with decaying fish in a mangrove area.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF068FFFDFF12FBD2FCE6FADD" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BD03694F068FFFDFF12FBD2FCE6FADD" blockId="10.[151,1437,151,1293]" pageId="10" pageNumber="11">
<emphasis id="B91BEA86F068FFFDFF12FBD2FED2FB88" bold="true" box="[199,308,1087,1112]" pageId="10" pageNumber="11">Remarks</emphasis>
. Slight variation aside the extent of the black marking on the pronotum. Some individuals only have the marginal area with green metallic sheen. A female present some coppery sheen along the median longitudinal pronotal sulcus. Females can be separated from males by their slightly wider anteromedian pronotal carina which lack the medially notched transverse tubercle and the more widely glabrous sternites 4–6. Two female specimens from Chapada de Aripe (Ceará) have wider and straighter carina, in these specimens the edges of the anterior pronotal carina in line with the lateral edge of the eyes.
</paragraph>
</subSubSection>
</treatment>
</document>
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