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<document id="62D045E228ECCDA7A4E906BE5FA3522A" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F05AFFCDFF42F949FD0EFA6F" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Onthoecus) attalus Genier &amp; Arnaud" docType="treatment" docVersion="10" lastPageNumber="59" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="57" updateTime="1698234910999" updateUser="plazi">
<mods:mods id="C0FBCC782DAC13C7DC0FAB1C42C7973F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E5502A183EDDF9B87FCFF780469410CF">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
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<mods:name id="DEADC3409FC139D86474F02916341D36" type="personal">
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<mods:roleTerm id="B4C32300D742922B56D962DEB366DFEC">Author</mods:roleTerm>
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<mods:namePart id="42FDC1BA7E1A5E27EC40FCF224ECD04F">François Génier</mods:namePart>
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<mods:namePart id="C607C0AC61DF3C2D04DB1CBE47E8CC44">Patrick Arnaud</mods:namePart>
</mods:name>
<mods:typeOfResource id="DD91366ADC84970F9303741E239C591F">text</mods:typeOfResource>
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<mods:title id="63D7F23B84EF864B2806D308C8509CB2">Zootaxa</mods:title>
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<mods:part id="92BC421A8319CA435A643BC4353D1AB1">
<mods:date id="8EBFB5DD5D700036943E45D2F3932F1C">2016</mods:date>
<mods:detail id="11C9272C9A63063DB7787589FF56E13C" type="volume">
<mods:number id="D979161C42BCF89EEA6CF0D30298BD6D">4099</mods:number>
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<mods:detail id="CD512020A3ED2C94758954BF8F578F2D" type="issue">
<mods:number id="2518006D5234CB71EB41E43674517BAE">1</mods:number>
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<mods:classification id="8109545686B9C800CB95219AA19FB2ED">journal volume</mods:classification>
<mods:identifier id="BEB65E7D910B824201B0A92C2127EB93" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
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<mods:identifier id="B82E0E4DC6B0CA71B6CAD35B56483226" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A9C237173A2C74DCDA7EAAD256703E86" type="Zenodo-Dep">268274</mods:identifier>
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<treatment id="03C68782F05AFFCDFF42F949FD0EFA6F" ID-DOI="http://doi.org/10.5281/zenodo.5780324" ID-GBIF-Taxon="127631973" ID-Zenodo-Dep="5780324" LSID="urn:lsid:plazi:treatment:03C68782F05AFFCDFF42F949FD0EFA6F" httpUri="http://treatment.plazi.org/id/03C68782F05AFFCDFF42F949FD0EFA6F" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57">
<subSubSection id="C375651FF05AFFCFFF42F949FBFDF96E" box="[151,1051,1700,1726]" pageId="56" pageNumber="57" type="nomenclature">
<paragraph id="8BD03694F05AFFCFFF42F949FBFDF96E" blockId="56.[151,1051,1700,1759]" box="[151,1051,1700,1726]" pageId="56" pageNumber="57">
<heading id="D09881F8F05AFFCFFF42F949FBFDF96E" bold="true" box="[151,1051,1700,1726]" fontSize="11" level="1" pageId="56" pageNumber="57" reason="1">
<emphasis id="B91BEA86F05AFFCFFF42F949FBFDF96E" bold="true" box="[151,1051,1700,1726]" pageId="56" pageNumber="57">
28.
<taxonomicName id="4C6F4D17F05AFFCFFF16F949FCBDF96E" ID-CoL="34R2L" authority="Genier &amp; Arnaud" authorityName="Genier &amp; Arnaud" box="[195,859,1700,1726]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="attalus" subGenus="Onthoecus">
<emphasis id="B91BEA86F05AFFCFFF16F949FD95F96E" bold="true" box="[195,627,1700,1726]" italics="true" pageId="56" pageNumber="57">Dendropaemon (Onthoecus) attalus</emphasis>
Génier &amp; Arnaud
</taxonomicName>
, nomen novum
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05AFFCFFF42F92BFCCCF8B2" pageId="56" pageNumber="57" type="description">
<paragraph id="8BD03694F05AFFCFFF42F92BFE21F90F" blockId="56.[151,1051,1700,1759]" box="[151,455,1734,1759]" pageId="56" pageNumber="57">
(
<figureCitation id="13542A11F05AFFCFFF4AF92BFF1AF90F" box="[159,252,1734,1759]" captionStart="FIGURES 25 – 30" captionStartId="103.[151,264,1962,1985]" captionTargetBox="[269,1325,201,1926]" captionTargetId="figure@103.[251,1336,193,1941]" captionTargetPageId="103" captionText="FIGURES 25 – 30. Dorsal habitus. 25. D. inemarginatus, female holotype; 26. D. nigritulus, male holotype; 27. D. amyntas, male; 28. D. attalus, male holotype; 29. D. lydiae, male holotype; 30. D. morettoi, male holotype." httpUri="https://zenodo.org/record/268279/files/figure.png" pageId="56" pageNumber="57">Figs. 28</figureCitation>
, 86–87, 123, 159)
</paragraph>
<paragraph id="8BD03694F05AFFCFFF42F8E3FC73F8F4" blockId="56.[151,1158,1806,2013]" box="[151,917,1806,1828]" pageId="56" pageNumber="57">
<taxonomicName id="4C6F4D17F05AFFCFFF42F8E3FDECF8F4" authority="Amyntas Harold 1868" authorityName="Amyntas Harold" authorityYear="1868" box="[151,522,1806,1828]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F05AFFCFFF42F8E3FE62F8F4" box="[151,388,1806,1828]" italics="true" pageId="56" pageNumber="57">Dendropemon Amyntas</emphasis>
Harold
<emphasis id="B91BEA86F05AFFCFFE02F8E2FDECF8F4" bold="true" box="[471,522,1807,1828]" pageId="56" pageNumber="57">1868</emphasis>
</taxonomicName>
,
<emphasis id="B91BEA86F05AFFCFFDC3F8E3FD99F8F4" box="[534,639,1806,1828]" italics="true" pageId="56" pageNumber="57">Col. Hefte</emphasis>
4: 83 (original description)
</paragraph>
<paragraph id="8BD03694F05AFFCFFF42F8C0FCB7F893" blockId="56.[151,1158,1806,2013]" box="[151,849,1837,1859]" pageId="56" pageNumber="57">
<taxonomicName id="4C6F4D17F05AFFCFFF42F8C0FE62F893" authority="Amyntas" authorityName="Amyntas" box="[151,388,1837,1859]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F05AFFCFFF42F8C0FE62F893" box="[151,388,1837,1859]" italics="true" pageId="56" pageNumber="57">Dendropemon Amyntas</emphasis>
</taxonomicName>
: Harold
<emphasis id="B91BEA86F05AFFCFFE0BF8C0FDF7F892" bold="true" box="[478,529,1837,1858]" pageId="56" pageNumber="57">1869</emphasis>
,
<emphasis id="B91BEA86F05AFFCFFDC8F8C3FD9CF893" box="[541,634,1837,1859]" italics="true" pageId="56" pageNumber="57">Cat. Col.</emphasis>
IV: 1020 (catalogue)
</paragraph>
<paragraph id="8BD03694F05AFFCFFF42F8A1FCCCF8B2" blockId="56.[151,1158,1806,2013]" box="[151,810,1868,1890]" pageId="56" pageNumber="57">
<taxonomicName id="4C6F4D17F05AFFCFFF42F8A1FE62F8B1" authority="Amyntas" authorityName="Amyntas" box="[151,388,1868,1890]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F05AFFCFFF42F8A1FE62F8B1" box="[151,388,1868,1890]" italics="true" pageId="56" pageNumber="57">Dendropemon Amyntas</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFFE4B65F05AFFCFFE47F8A1FDE5F8B1" author="Gillet" box="[402,515,1868,1890]" pageId="56" pageNumber="57" refString="Gillet, J. J. E. (1911) Coleopterorum Catalogus, Scarabaeidae: Coprinae I. Vol. 38. W Junk, Berlin, 263 pp." type="book" year="1911">
Gillet
<emphasis id="B91BEA86F05AFFCFFE07F8A1FDE5F8B1" bold="true" box="[466,515,1868,1889]" pageId="56" pageNumber="57">1911</emphasis>
</bibRefCitation>
,
<emphasis id="B91BEA86F05AFFCFFDDAF8A1FD8AF8B1" box="[527,620,1868,1890]" italics="true" pageId="56" pageNumber="57">Col. Cat.</emphasis>
38: 88 (catalogue)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05AFFCEFF42F887FBE2FF3B" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="reference_group">
<paragraph id="8BD03694F05AFFCFFF42F887FC9DF850" blockId="56.[151,1158,1806,2013]" box="[151,891,1898,1920]" pageId="56" pageNumber="57">
<taxonomicName id="4C6F4D17F05AFFCFFF42F887FDAFF8AF" authority="Olsoufieff 1924" authorityName="Olsoufieff" authorityYear="1924" box="[151,585,1898,1920]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="amynthas">
<emphasis id="B91BEA86F05AFFCFFF42F887FE7DF850" box="[151,411,1898,1920]" italics="true" pageId="56" pageNumber="57">Dendropaemon amynthas</emphasis>
:
<bibRefCitation id="EFFE4B65F05AFFCFFE7DF887FDAFF8AF" author="Olsoufieff" box="[424,585,1898,1920]" pageId="56" pageNumber="57" refString="Olsoufieff, G. d' (1924) Les Phanaeides (Coleoptera-Lamellicornia), Famille Scarabaeidae-Tr. Coprini. Insecta, 13, 4 - 172, 15 pls." type="journal article" year="1924">
Olsoufieff
<emphasis id="B91BEA86F05AFFCFFDC3F887FDAFF8AF" bold="true" box="[534,585,1898,1919]" pageId="56" pageNumber="57">1924</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05AFFCFFD80F886FD78F850" box="[597,670,1899,1920]" italics="true" pageId="56" pageNumber="57">Insecta</emphasis>
13: 122 (monograph)
</paragraph>
<paragraph id="8BD03694F05AFFCEFF42F864FBE2FF3B" blockId="56.[151,1158,1806,2013]" lastBlockId="57.[151,1066,152,235]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
<taxonomicName id="4C6F4D17F05AFFCFFF42F864FD92F84E" authority="Blut 1939" authorityName="Blut" authorityYear="1939" box="[151,628,1929,1951]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="subSpecies" species="amynthas" subSpecies="amynthas">
<emphasis id="B91BEA86F05AFFCFFF42F864FDE4F84F" box="[151,514,1929,1951]" italics="true" pageId="56" pageNumber="57">Dendropaemon amynthas amynthas</emphasis>
:
<bibRefCitation id="EFFE4B65F05AFFCFFDDAF864FD92F84E" author="Blut" box="[527,628,1929,1951]" pageId="56" pageNumber="57" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">
Blut
<emphasis id="B91BEA86F05AFFCFFD94F864FD92F84E" bold="true" box="[577,628,1929,1950]" pageId="56" pageNumber="57">1939</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05AFFCFFD54F864FCADF84F" box="[641,843,1929,1951]" italics="true" pageId="56" pageNumber="57">Arch. Naturg. (N.F.)</emphasis>
8: 267 (monograph)
<taxonomicName id="4C6F4D17F05AFFCFFF42F845FDAEF86D" authority="Blackwelder 1944" authorityName="Blackwelder" authorityYear="1944" box="[151,584,1960,1982]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F05AFFCFFF42F845FE67F86E" box="[151,385,1960,1982]" italics="true" pageId="56" pageNumber="57">Dendropemon amyntas</emphasis>
:
<bibRefCitation id="EFFE4B65F05AFFCFFE5AF845FDAEF86D" author="Blackwelder" box="[399,584,1960,1982]" pageId="56" pageNumber="57" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 341. http: // dx. doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">
Blackwelder
<emphasis id="B91BEA86F05AFFCFFDC0F845FDAEF86D" bold="true" box="[533,584,1960,1981]" pageId="56" pageNumber="57">1944</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05AFFCFFD81F844FCC7F86E" box="[596,801,1960,1982]" italics="true" pageId="56" pageNumber="57">
<collectingCountry id="F3787604F05AFFCFFD81F844FD66F86E" box="[596,640,1961,1982]" name="United States of America" pageId="56" pageNumber="57">U.S.</collectingCountry>
<taxonomicName id="4C6F4D17F05AFFCFFD53F844FD48F86E" box="[646,686,1961,1982]" class="Insecta" family="Coccinellidae" genus="Nat" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">Nat</taxonomicName>
. Mus. Bull.
</emphasis>
185: 210 (checklist)
<taxonomicName id="4C6F4D17F05AFFCFFF42F82BFD82F80C" ID-CoL="34R2K" authority="Edmonds 1972" authorityName="Edmonds" authorityYear="1972" box="[151,612,1990,2013]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="amyntas" subGenus="Dendropaemon">
<emphasis id="B91BEA86F05AFFCFFF42F82BFE5BF80D" box="[151,445,1990,2013]" italics="true" pageId="56" pageNumber="57">Dendropaemon (D.) amyntas</emphasis>
:
<bibRefCitation id="EFFE4B65F05AFFCFFE1EF82BFD82F80C" author="Edmonds" box="[459,612,1990,2012]" pageId="56" pageNumber="57" refString="Edmonds, W. D. (1972) Comparative Skeletal Morphology, Systematics and Evolution of the Phanaeine Dung Beetles (Coleoptera: Scarabaeidae). University of Kansas Science Bulletin, 49 (11), 731 - 874." type="journal article" year="1972">
Edmonds
<emphasis id="B91BEA86F05AFFCFFDE4F82AFD82F80C" bold="true" box="[561,612,1991,2012]" pageId="56" pageNumber="57">1972</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05AFFCFFDA5F825FCB5F80C" box="[624,851,1990,2013]" italics="true" pageId="56" pageNumber="57">Univ. Kansas Sc. Bull.</emphasis>
49: 850 (comment taxonomy)
<taxonomicName id="4C6F4D17F05BFFCEFF42FF75FDC6FF7D" ID-CoL="34R2K" authority="Arnaud 1982" authorityName="Arnaud" authorityYear="1982" box="[151,544,152,174]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F05BFFCEFF42FF75FE68FF7E" box="[151,398,152,174]" italics="true" pageId="57" pageNumber="58">Dendropaemon amyntas</emphasis>
:
<bibRefCitation id="EFFE4B65F05BFFCEFE4EFF75FDC6FF7D" author="Arnaud" box="[411,544,152,174]" pageId="57" pageNumber="58" refString="Arnaud, P. (1982) Liste des types de Phanaeini du Museum national d'Histoire naturelle de Paris, suivi d'une note synonymique pour quatre especes decrite par G. d'Olsoufieff (Coleoptera, Scarabaeidae). Revue Francaise d'Entomologie, Nouvelle Serie, 4 (3), 113 - 118." type="journal article" year="1982">
Arnaud
<emphasis id="B91BEA86F05BFFCEFE38FF75FDC6FF7D" bold="true" box="[493,544,152,173]" pageId="57" pageNumber="58">1982</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05BFFCEFDF8FF74FD0DFF7E" box="[557,747,153,174]" italics="true" pageId="57" pageNumber="58">Rev. Fr. Ent. (N.S.)</emphasis>
4: 115 (
<typeStatus id="54D48836F05BFFCEFCEBFF75FC7AFF7E" box="[830,924,152,174]" pageId="57" pageNumber="58" type="lectotype">lectotype</typeStatus>
designation)
<taxonomicName id="4C6F4D17F05BFFCEFF42FF5BFE0EFF1D" ID-CoL="34R2K" box="[151,488,182,205]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="amyntas" subSpecies="amyntas">
<emphasis id="B91BEA86F05BFFCEFF42FF5BFE0EFF1D" box="[151,488,182,205]" italics="true" pageId="57" pageNumber="58">Dendropaemon amyntas amyntas</emphasis>
</taxonomicName>
: Vaz-de-Mello
<emphasis id="B91BEA86F05BFFCEFD52FF5AFD5FFF1C" bold="true" box="[647,697,183,204]" pageId="57" pageNumber="58">2000</emphasis>
,
<emphasis id="B91BEA86F05BFFCEFD12FF55FC6EFF1C" box="[711,904,183,205]" italics="true" pageId="57" pageNumber="58">Hac. Proy. CYTED</emphasis>
: 192 (faunistic)
<taxonomicName id="4C6F4D17F05BFFCEFF42FF38FD64FF3A" ID-CoL="34R2K" authority="Philips et al. 2004" authorityName="Philips et al." authorityYear="2004" box="[151,642,213,235]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="amyntas" subGenus="Dendropaemon">
<emphasis id="B91BEA86F05BFFCEFF42FF38FE5BFF3B" box="[151,445,213,235]" italics="true" pageId="57" pageNumber="58">Dendropaemon (D.) amyntas</emphasis>
:
<bibRefCitation id="EFFE4B65F05BFFCEFE1EFF38FD64FF3A" author="Philips" box="[459,642,213,235]" pageId="57" pageNumber="58" refString="Philips, T. K., Edmonds, W. D. &amp; Scholtz, C. H. (2004) A phylogenetic analysis of the New World tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae): Hypotheses on relationships and origins. Insect Systematics &amp; Evolution, 35, 43 - 63. http: // dx. doi. org / 10.1163 / 187631204788964664" type="journal article" year="2004">
Philips et al.
<emphasis id="B91BEA86F05BFFCEFD9AFF38FD64FF3A" bold="true" box="[591,642,213,234]" pageId="57" pageNumber="58">2004</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F05BFFCEFD5BFF3BFCDAFF3B" box="[654,828,213,235]" italics="true" pageId="57" pageNumber="58">Insect Syst. Evol.</emphasis>
35: 51 (phylogeny)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05BFFCEFF42FEFAFE54FEE0" box="[151,434,279,304]" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8BD03694F05BFFCEFF42FEFAFE54FEE0" blockId="57.[151,1437,279,1997]" box="[151,434,279,304]" pageId="57" pageNumber="58">
<emphasis id="B91BEA86F05BFFCEFF42FEFAFECBFEE0" bold="true" box="[151,301,279,304]" pageId="57" pageNumber="58">
<typeStatus id="54D48836F05BFFCEFF42FEFAFF34FEE0" box="[151,210,279,304]" pageId="57" pageNumber="58">Type</typeStatus>
locality
</emphasis>
. Cayennae.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05BFFCEFF12FED1FF33FE4D" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BD03694F05BFFCEFF12FED1FF33FE4D" blockId="57.[151,1437,279,1997]" pageId="57" pageNumber="58">
<emphasis id="B91BEA86F05BFFCEFF12FED1FEA7FE83" bold="true" box="[199,321,316,341]" pageId="57" pageNumber="58">Diagnosis.</emphasis>
The large size, rather thick body combined with the dorsum dark metallic blue sheen will separate
<taxonomicName id="4C6F4D17F05BFFCEFF42FE8CFEE3FEA8" box="[151,261,351,376]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="attalus">
<emphasis id="B91BEA86F05BFFCEFF42FE8CFEE3FEA8" box="[151,261,351,376]" italics="true" pageId="57" pageNumber="58">D. attalus</emphasis>
</taxonomicName>
from all other species. It can be separated from its sister species,
<taxonomicName id="4C6F4D17F05BFFCEFC36FE8CFB84FEA8" box="[995,1122,353,376]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F05BFFCEFC36FE8CFB84FEA8" box="[995,1122,353,376]" italics="true" pageId="57" pageNumber="58">D. amyntas</emphasis>
</taxonomicName>
, by the shape of the clypeal teeth.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05BFFCDFF12FE4AFC0CFEB5" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="description">
<paragraph id="8BD03694F05BFFCEFF12FE4AFF08F81D" blockId="57.[151,1437,279,1997]" pageId="57" pageNumber="58">
<emphasis id="B91BEA86F05BFFCEFF12FE4AFEB7FE10" bold="true" box="[199,337,423,448]" pageId="57" pageNumber="58">Description</emphasis>
. Undamaged female specimen matching abraded male
<typeStatus id="54D48836F05BFFCEFC09FE4AFBA5FE10" box="[988,1091,423,448]" pageId="57" pageNumber="58" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="13542A11F05BFFCEFB83FE4AFB49FE10" box="[1110,1199,423,448]" captionStart="FIGURES 25 – 30" captionStartId="103.[151,264,1962,1985]" captionTargetBox="[269,1325,201,1926]" captionTargetId="figure@103.[251,1336,193,1941]" captionTargetPageId="103" captionText="FIGURES 25 – 30. Dorsal habitus. 25. D. inemarginatus, female holotype; 26. D. nigritulus, male holotype; 27. D. amyntas, male; 28. D. attalus, male holotype; 29. D. lydiae, male holotype; 30. D. morettoi, male holotype." httpUri="https://zenodo.org/record/268279/files/figure.png" pageId="57" pageNumber="58">Fig. 28</figureCitation>
).
<emphasis id="B91BEA86F05BFFCEFB13FE4AFAE5FE10" bold="true" box="[1222,1283,423,448]" pageId="57" pageNumber="58">Body</emphasis>
. Body large, length 22.0 mm, maximum width 12.0 mm; body subrectangular; dorsum narrowly flat.
<emphasis id="B91BEA86F05BFFCEFB77FE21FB05FE35" bold="true" box="[1186,1251,460,485]" pageId="57" pageNumber="58">Color</emphasis>
. Dorsal surface black, glossy, with metallic sheen; head with greenish to coppery metallic sheen adjacent to the eyes; pronotum with faint bluish metallic sheen along the border; elytra with uniform blue metallic sheen; ventrum dark brown to black; pygidium with greenish metallic sheen; legs black.
<emphasis id="B91BEA86F05BFFCEFC89FDDAFC7DFD80" bold="true" box="[860,923,567,592]" pageId="57" pageNumber="58">Head</emphasis>
. Clypeus broadly arcuate, anterior portion upturned; clypeal teeth triangular; clypeal median emargination broadly v-shaped, clypeal edge lacking emargination on external side of each clypeal tooth, clypeal teeth ventral surface with a fine v-shaped carina, clypeal margin well-defined and sharply carinate posteriorly, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with small and coarse rugulae anteriorly and simply punctate posteriorly, transversely tumescent; clypeofrontal carina low, more than 6 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge obtusely angular medially in frontal view; eyes small in dorsal view, interocular ratio 5.8.
<emphasis id="B91BEA86F05BFFCEFCB8FCD9FC03FC9D" bold="true" box="[877,997,820,845]" pageId="57" pageNumber="58">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.6; disc of pronotum finely punctate basally, punctures changing into fine rough rugulae anteriorly, with an ill-defined shallow longitudinal depression on posterior half; pronotal anterior margin slightly wider and concave lateral to eye; anterior portion with a tri-sinuous carina; anterior angles surface with fine rugulae, similar to lateral margin along posterior edge of anterior margin; lateral fossae simply rounded, concave; lateral portions strongly explanate; pronotal basal fossae well-defined, concave; posterior margin well-defined, lacking crenulation and setae.
<emphasis id="B91BEA86F05BFFCEFE3CFBE1FDD3FBF5" bold="true" box="[489,565,1036,1061]" pageId="57" pageNumber="58">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/ length ratio 1.2; elytral base lacking distinct margin, simply convex; elytral striae 1–4 fine and well-defined, evenly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures fine, not encroaching on interval, stria 1 well-defined apically, connecting to marginal stria; interstriae slightly convex, minutely punctate throughout, surface glossy.
<emphasis id="B91BEA86F05BFFCEFE9BFB71FDF4FB65" bold="true" box="[334,530,1180,1205]" pageId="57" pageNumber="58">Thoracic sterna</emphasis>
. Proepisternal carina complete, extending laterally; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, v-shaped.
<emphasis id="B91BEA86F05BFFCEFC19FB52FBE2FB08" bold="true" box="[972,1028,1215,1240]" pageId="57" pageNumber="58">Legs</emphasis>
. Profemur posterior surface convex, glabrous and punctate internally, posterointernal margin rather thick, evenly developed, internal edge rather wide, with a brush of long setae along anterointernal edge, remaining surface irregularly punctate. Protibia with four teeth on lateral edge; internal basal angle bluntly lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with coarse irregular punctures externally to median carina, surface between punctures finely and irregularly microsculptured, with a single interrupted setal row along lateral teeth. Mesofemur unmodified on anterointernal edge apically. Mesotibia rather slender, gradually widening toward apex in anterior view; anteroapical edge straight in anterior view, anteroapical row of setae incomplete, irregular; apicoanterior edge obliquely truncated internally; external edge rounded basally, more or less flat apically, surface with irregular punctures and microsculpture. Mesotarsus similar in shape to metatarsus, 3-segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur elongate, internal and lateral edges mostly parallel in ventral view, more than twice as long as wide, depressed anterointernally before apex, apicoposterior edge lobate beyond tibial insertion, surface coarsely microsculptured, anterior surface with a row of irregular punctures, punctures not bordered anteriorly by a sharp and well-define sulcus. Metatibia robust, short, slightly widening toward apex in anterior view, anterior surface completely covered with irregular sculpturing and punctures, metatibial posterior surface concave between longitudinal row of setae and lateral edge, surface dull and irregularly punctate. Metatarsus 3-segmented (
<figureCitation id="13542A11F05BFFCEFABAF912FF22F8EC" captionStart="FIGURES 118 – 125" captionStartId="113.[151,264,1924,1947]" captionTargetBox="[217,1363,200,1895]" captionTargetId="figure@113.[211,1376,193,1903]" captionTargetPageId="113" captionText="FIGURES 118 – 125. Metatarsus, ventral view. 118. D. fredericki, holotype; 119. D. angustipennis; 120. D. bahianus; 121. D. nigritulus, paratype; 122. D. amyntas; 123. D. attalus; 124. D. lydiae, paratype; 125. D. morettoi, paratype." httpUri="https://zenodo.org/record/268283/files/figure.png" pageId="57" pageNumber="58">Fig. 123</figureCitation>
), first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge.
<emphasis id="B91BEA86F05BFFCEFCC3F8AAFBEBF8B0" bold="true" box="[790,1037,1863,1888]" pageId="57" pageNumber="58">Abdominal sternites</emphasis>
. Sternites 3–6 longitudinally flat; sternites 4–6 with more than three unaligned row of setae laterally, glabrous medially; sternite 7 approximately longitudinally flat medially, shorter than segment 6 along midline; pygidium with mixed minute and fine punctures on disc.
</paragraph>
<paragraph id="8BD03694F058FFCDFF12FF7AFA7DFF7F" blockId="58.[151,1437,151,1472]" box="[199,1435,151,176]" pageId="58" pageNumber="59">
<emphasis id="B91BEA86F058FFCDFF12FF7AFE90FF60" bold="true" box="[199,374,151,176]" pageId="58" pageNumber="59">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF058FFCDFE53FF75FE0BFF60" box="[390,493,151,176]" pageId="58" pageNumber="59" type="male">10 males</specimenCount>
,
<specimenCount id="9D69FD1DF058FFCDFE2CFF75FD93FF60" box="[505,629,151,176]" pageId="58" pageNumber="59" type="female">24 females</specimenCount>
). Length: male 16.0–20.0 (18.7±1.3), female 16.0–22.5 (19.5±1.5) mm.
</paragraph>
<paragraph id="8BD03694F058FFCDFF12FF51FC0CFEB5" blockId="58.[151,1437,151,1472]" pageId="58" pageNumber="59">
<emphasis id="B91BEA86F058FFCDFF12FF51FE41FF05" bold="true" box="[199,423,188,213]" pageId="58" pageNumber="59">
Primary
<typeStatus id="54D48836F058FFCDFEE0FF51FE8EFF05" box="[309,360,188,213]" pageId="58" pageNumber="59">type</typeStatus>
data
</emphasis>
(
<figureCitation id="13542A11F058FFCDFE6DFF51FDFBFF05" box="[440,541,188,213]" captionStart="FIGURES 134 – 154" captionStartId="115.[151,264,1903,1926]" captionTargetBox="[216,1361,200,1865]" captionTargetId="figure@115.[210,1372,193,1876]" captionTargetPageId="115" captionText="FIGURES 134 – 154. Primary type labels. 134. D. hirticollis; 135. D. pauliani; 136. D. renatii; 137. D. ater; 138. D. piceus; 139: D. telephus; 140. D. viridis; 141. D. viridipennis. 142. D. convexus; 143. D. fredericki; 144. D. angustipennis; 145. D. bahianus; 146. D. amyntas (= D. attalus); 147. D. ganglbaueri; 148. D. refulgens; 149. D. fractipes; 150. D. fasces; 151. D. haroldi; 152. D. nitidicollis; 153. D. quadratus; 154. D. similis; 155. D. denticollis." httpUri="https://zenodo.org/record/268285/files/figure.png" pageId="58" pageNumber="59">Fig. 146</figureCitation>
).
<typeStatus id="54D48836F058FFCDFDE7FF50FD44FF05" box="[562,674,189,213]" pageId="58" pageNumber="59" type="lectotype">Lectotype</typeStatus>
female (MNHN): [Cayenae/ D./
<taxonomicName id="4C6F4D17F058FFCDFBF4FF51FB60FF04" box="[1057,1158,188,212]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="120" phylum="Arthropoda" rank="species" species="amyntas">Amyntas</taxonomicName>
/ t. Harold] handwritten, red border; [Ex-Musaeo/ E.Harold]; [
<taxonomicName id="4C6F4D17F058FFCDFD84FF32FC40FF28" authority="Har" authorityName="Har" box="[593,934,223,248]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="amyntas">Dendropaemon amyntas Har</taxonomicName>
/ det. G. OLSOUFIEFF] partly handwritten; [Museum Paris/ ex Coll./ R.Oberthur] green card; [WORLD/ SCARAB./ DATABASE/ WSD00016598]; [
<taxonomicName id="4C6F4D17F058FFCDFF48FECAFEAAFE90" box="[157,332,295,320]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F058FFCDFEBCFEC5FDE2FE90" authority="Har" authorityName="Har" box="[361,516,296,320]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="amyntas">amyntas Har</taxonomicName>
/
<typeStatus id="54D48836F058FFCDFDF4FEC5FD26FE90" box="[545,704,296,320]" pageId="58" pageNumber="59" type="lectotype">LECTOTYPE</typeStatus>
/ P.ARNAUD DET 1982] partly handwritten, red border; [
<taxonomicName id="4C6F4D17F058FFCDFF75FEA1FEA8FEB5" box="[160,334,332,357]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F058FFCDFF75FEA1FEA8FEB5" box="[160,334,332,357]" italics="true" pageId="58" pageNumber="59">Dendropaemon</emphasis>
</taxonomicName>
♀/
<taxonomicName id="4C6F4D17F058FFCDFEADFEA1FE22FEB5" box="[376,452,332,357]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="attalus">
<emphasis id="B91BEA86F058FFCDFEADFEA1FE22FEB5" box="[376,452,332,357]" italics="true" pageId="58" pageNumber="59">attalus</emphasis>
</taxonomicName>
/ Génier &amp; Arnaud/ dét. Génier &amp; Arnaud,2012].
</paragraph>
</subSubSection>
<subSubSection id="C375651FF058FFCDFF12FE82FBD9FBAC" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="8BD03694F058FFCDFF12FE82FBFCFB88" blockId="58.[151,1437,151,1472]" pageId="58" pageNumber="59">
<emphasis id="B91BEA86F058FFCDFF12FE82FE57FE58" bold="true" box="[199,433,367,392]" pageId="58" pageNumber="59">Material examined</emphasis>
.
<collectingCountry id="F3787604F058FFCDFE12FE82FDD4FE58" box="[455,562,367,392]" name="Brazil" pageId="58" pageNumber="59">
<emphasis id="B91BEA86F058FFCDFE12FE82FDD4FE58" bold="true" box="[455,562,367,392]" pageId="58" pageNumber="59">BRAZIL</emphasis>
</collectingCountry>
: [unspecified locality], [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF058FFCDFB1BFE9DFAD3FE58" box="[1230,1333,367,392]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
,
<specimenCount id="9D69FD1DF058FFCDFA9DFE9DFA7DFE58" box="[1352,1435,367,392]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFF4BFE79FF11FE7C" box="[158,247,404,428]" pageId="58" pageNumber="59">BMNH</collectionCode>
,
<collectionCode id="ED7EAE51F058FFCDFED0FE79FEA8FE7C" box="[261,334,404,428]" pageId="58" pageNumber="59">CPFA</collectionCode>
); AMAZONAS, Municipio Barcelos, Pico Tamacuari,
<quantity id="4C979B71F058FFCDFC1FFE79FBFEFE7C" box="[970,1048,404,428]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="58" pageNumber="59" unit="m" value="2700.0">2700m</quantity>
NW Missão Marari, elev.
<quantity id="4C979B71F058FFCDFA87FE79FA7AFE7C" box="[1362,1436,404,428]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="58" pageNumber="59" unit="m" value="340.0">340 m</quantity>
(
<geoCoordinate id="EE5B5053F058FFCDFF4AFE5AFECAFE00" box="[159,300,439,464]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="1" value="1.2073333">1°12'26.4''N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFEE9FE5AFE38FE00" box="[316,478,439,464]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="-64.78836">64°47'18.1''W</geoCoordinate>
),
<date id="FFD11054F058FFCDFE22FE5AFD55FE00" box="[503,691,439,464]" pageId="58" pageNumber="59" value="2004-04-29" valueMax="2004-05-10" valueMin="2004-04-29">29.iv.–10.v.2004</date>
, coll. U. Caramaschi—
<specimenCount id="9D69FD1DF058FFCDFC1BFE5AFBFFFE00" box="[974,1049,439,464]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFBF8FE55FB67FE00" box="[1069,1153,440,464]" pageId="58" pageNumber="59">CEMT</collectionCode>
); Rio Parauary, (
<geoCoordinate id="EE5B5053F058FFCDFA9EFE5AFA7EFE00" box="[1355,1432,439,464]" direction="south" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="-4.6">4°36'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFF42FE31FF1BFE25" box="[151,253,476,501]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="-57.783333">57°47'W</geoCoordinate>
),
<date id="FFD11054F058FFCDFEC1FE31FE76FE25" box="[276,400,476,501]" pageId="58" pageNumber="59" value="1937-03-15">15.iii.1937</date>
, coll. Zellibor-Hauff—
<specimenCount id="9D69FD1DF058FFCDFD74FE31FCE6FE25" box="[673,768,476,501]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFCC7FE31FC8DFE24" box="[786,875,476,500]" pageId="58" pageNumber="59">CMNC</collectionCode>
);
<collectionCode id="ED7EAE51F058FFCDFC57FE31FC37FE24" box="[898,977,476,500]" pageId="58" pageNumber="59">MATO</collectionCode>
GROSSO, Fazenda Paraíso, Municipio Tangara da Serra, elev.
<quantity id="4C979B71F058FFCDFE78FE12FE1EFDC7" box="[429,504,511,536]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="58" pageNumber="59" unit="m" value="500.0">500 m</quantity>
(
<geoCoordinate id="EE5B5053F058FFCDFDDEFDEDFD98FDC8" box="[523,638,512,536]" direction="south" orientation="latitude" pageId="58" pageNumber="59" precision="15" value="-14.667223">14°40'2''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFD5BFE12FCEBFDC8" box="[654,781,511,536]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="15" value="-57.385834">57°23'9''W</geoCoordinate>
),
<date id="FFD11054F058FFCDFCF1FE12FC00FDC8" box="[804,998,511,536]" pageId="58" pageNumber="59" value="2008-11-27" valueMax="2008-12-04" valueMin="2008-11-27">27.xi.–4.xii.2008</date>
, coll. L.R. Silva—
<specimenCount id="9D69FD1DF058FFCDFB1CFE12FACFFDC8" box="[1225,1321,511,536]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFAE9FDEDFA76FDC8" box="[1340,1424,512,536]" pageId="58" pageNumber="59">CEMT</collectionCode>
);
<collectionCode id="ED7EAE51F058FFCDFF42FDC9FF04FDED" box="[151,226,548,573]" pageId="58" pageNumber="59">PARÁ</collectionCode>
, [unspecified locality], [no date], coll. S. Sieber—
<specimenCount id="9D69FD1DF058FFCDFCF2FDC9FC62FDED" box="[807,900,548,573]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFC43FDC9FC0BFDEC" box="[918,1005,548,572]" pageId="58" pageNumber="59">ZMHB</collectionCode>
); RONDÔNIA, Barragem de Samuel, Candeias do Jamari, (
<geoCoordinate id="EE5B5053F058FFCDFE4AFDAAFE0DFDB0" box="[415,491,583,608]" direction="south" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="-8.766666">8°46'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFE2BFDAAFD82FDB0" box="[510,612,583,608]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="-63.466667">63°28'W</geoCoordinate>
),
<date id="FFD11054F058FFCDFD55FDAAFD5FFDB0" box="[640,697,583,608]" pageId="58" pageNumber="59" value="1989-04">iv.89</date>
, coll. Dégallier—
<specimenCount id="9D69FD1DF058FFCDFC40FDAAFC1EFDB0" box="[917,1016,583,608]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFBDBFDA5FBBFFDB0" box="[1038,1113,584,608]" pageId="58" pageNumber="59">CPFA</collectionCode>
); Fazenda estrela de Davi, Guajará-Mirim, elev.
<quantity id="4C979B71F058FFCDFE42FD81FE03FD54" box="[407,485,620,645]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="58" pageNumber="59" unit="m" value="165.0">165 m</quantity>
(
<geoCoordinate id="EE5B5053F058FFCDFE2EFD81FD77FD55" box="[507,657,620,645]" direction="south" orientation="latitude" pageId="58" pageNumber="59" precision="1" value="-10.743084">10°44'35.1''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFD71FD81FCB2FD55" box="[676,852,620,645]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="-65.2996">65°17'58.57''W</geoCoordinate>
),
<date id="FFD11054F058FFCDFCBBFD81FC04FD55" box="[878,994,620,645]" pageId="58" pageNumber="59" value="2010-02-18">18.ii.2010</date>
, coll. F. Coletti—
<specimenCount id="9D69FD1DF058FFCDFB16FD81FAC0FD55" box="[1219,1318,620,645]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFAE9FD81FA76FD54" box="[1340,1424,620,644]" pageId="58" pageNumber="59">CEMT</collectionCode>
);
<collectingCountry id="F3787604F058FFCDFF42FD7DFEF4FD78" box="[151,274,656,680]" name="Guyana" pageId="58" pageNumber="59">GUYANA</collectingCountry>
: [unspecified locality], [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF058FFCDFC49FD7DFC18FD78" box="[924,1022,655,680]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFBC6FD7DFB89FD78" LSID="urn:lsid:biocol.org:col:34988" box="[1043,1135,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="59">MNHN</collectionCode>
); CUYUNI-MAZARUNI, Kartabo, (
<geoCoordinate id="EE5B5053F058FFCDFEDCFD59FE66FD1D" box="[265,384,692,717]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="15" value="6.2825003">6°16'57''N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFE5BFD59FDEAFD1D" box="[398,524,692,717]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="15" value="-58.585556">58°35'8''W</geoCoordinate>
),
<date id="FFD11054F058FFCDFDF4FD59FD7DFD1D" box="[545,667,692,717]" pageId="58" pageNumber="59" value="1925-06-23">23.vi.1925</date>
, coll. [anonymous]—
<specimenCount id="9D69FD1DF058FFCDFC42FD59FC12FD1D" box="[919,1012,692,717]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFBD0FD59FBB8FD1C" box="[1029,1118,692,716]" pageId="58" pageNumber="59">CMNC</collectionCode>
);
<emphasis id="B91BEA86F058FFCDFBA7FD59FA72FD1D" bold="true" box="[1138,1428,692,717]" pageId="58" pageNumber="59">
<collectingCountry id="F3787604F058FFCDFBA7FD59FB0FFD1D" box="[1138,1257,692,717]" name="French Guiana" pageId="58" pageNumber="59">GUYANE</collectingCountry>
FRANÇAISE
</emphasis>
: Cacao, (
<geoCoordinate id="EE5B5053F058FFCDFF26FD3AFEA2FD20" box="[243,324,727,752]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="4.5833335">4°35'N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFE9AFD3AFE50FD20" box="[335,438,727,752]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="-52.466667">52°28'W</geoCoordinate>
),
<date id="FFD11054F058FFCDFE1CFD3AFDC4FD20" box="[457,546,727,752]" pageId="58" pageNumber="59" value="2007-03">iii.2007</date>
, coll. [anonymous]—
<specimenCount id="9D69FD1DF058FFCDFCC1FD35FC99FD20" box="[788,895,727,752]" pageId="58" pageNumber="59" type="female">2 females</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFC5BFD35FC3FFD20" box="[910,985,728,752]" pageId="58" pageNumber="59">CPFA</collectionCode>
); Cayenne, (
<geoCoordinate id="EE5B5053F058FFCDFBB6FD3AFB2AFD20" box="[1123,1228,727,752]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="15" value="4.9008336">4°54'3''N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFB02FD3AFA85FD20" box="[1239,1379,727,752]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="15" value="-52.303333">52°18'12''W</geoCoordinate>
), [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF058FFCDFE0FFD11FDA9FCC5" box="[474,591,764,789]" pageId="58" pageNumber="59" type="female">5 females</specimenCount>
(incl.
<typeStatus id="54D48836F058FFCDFD79FD11FCF6FCC5" box="[684,784,764,789]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
) (
<collectionCode id="ED7EAE51F058FFCDFCE5FD11FC9FFCC4" box="[816,889,764,788]" pageId="58" pageNumber="59">CPFA</collectionCode>
,
<collectionCode id="ED7EAE51F058FFCDFC58FD11FC04FCC4" box="[909,994,764,788]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="58" pageNumber="59">IRSNB</collectionCode>
,
<collectionCode id="ED7EAE51F058FFCDFC2DFD11FBB5FCC4" LSID="urn:lsid:biocol.org:col:34988" box="[1016,1107,764,788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="59">MNHN</collectionCode>
); Crique Blanche, R.N. 2, (
<geoCoordinate id="EE5B5053F058FFCDFF4AFCCDFEF1FCE8" box="[159,279,800,824]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="15" value="4.5608335">4°33'39''N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFEF2FCF2FE52FCE8" box="[295,436,799,824]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="15" value="-52.397224">52°23'50''W</geoCoordinate>
),
<date id="FFD11054F058FFCDFE19FCF2FDFBFCE8" box="[460,541,799,824]" pageId="58" pageNumber="59" value="2009-02">ii.2009</date>
, coll. Giuglaris—
<specimenCount id="9D69FD1DF058FFCDFD3AFCF2FCB8FCE8" box="[751,862,799,824]" pageId="58" pageNumber="59" type="female">2 females</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFCA7FCCDFC5BFCE8" box="[882,957,800,824]" pageId="58" pageNumber="59">CPFA</collectionCode>
); environs de Cacao, (
<geoCoordinate id="EE5B5053F058FFCDFB12FCF2FAFFFCE8" box="[1223,1305,799,824]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="4.5833335">4°35'N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFAFCFCF2FA69FCE8" box="[1321,1423,799,824]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="-52.466667">52°28'W</geoCoordinate>
),
<date id="FFD11054F058FFCDFF42FCA9FF16FC8D" box="[151,240,836,861]" pageId="58" pageNumber="59" value="2007-03">iii.2007</date>
, coll. E. Degrad &amp; P. Bonin—
<specimenCount id="9D69FD1DF058FFCDFD9EFCA9FD4FFC8D" box="[587,681,836,861]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
,
<specimenCount id="9D69FD1DF058FFCDFD60FCA9FCE6FC8D" box="[693,768,836,861]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFCC5FCA9FCBDFC8C" box="[784,859,836,860]" pageId="58" pageNumber="59">CPFA</collectionCode>
); Paramaca (Kourou), (
<geoCoordinate id="EE5B5053F058FFCDFBB4FCA9FB2DFC8D" box="[1121,1227,836,861]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="15" value="5.148333">5°8'54''N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFB03FCA9FA85FC8D" box="[1238,1379,836,861]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="15" value="-52.680836">52°40'51''W</geoCoordinate>
), [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF058FFCDFE11FC85FDD4FC50" box="[452,562,871,896]" pageId="58" pageNumber="59" type="female">2 females</specimenCount>
,
<specimenCount id="9D69FD1DF058FFCDFDE8FC85FD75FC50" box="[573,659,871,896]" pageId="58" pageNumber="59" type="male">2 males</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFD77FC85FD00FC50" box="[674,742,872,896]" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="58" pageNumber="59">MTD</collectionCode>
); PK 125, Route N2 (Cayenne-Saint-Georges-de-L’Oyapock), (
<geoCoordinate id="EE5B5053F058FFCDFF4AFC61FEF1FC74" box="[159,279,908,932]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="15" value="4.2225">4°13'21''N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFEF0FC61FE42FC75" box="[293,420,908,933]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="15" value="-52.12083">52°7'15''W</geoCoordinate>
),
<date id="FFD11054F058FFCDFE6EFC61FDCEFC75" box="[443,552,908,933]" pageId="58" pageNumber="59" value="2008-03-06">6.iii.2008</date>
, coll. J.L. Giuglaris—
<specimenCount id="9D69FD1DF058FFCDFCFAFC61FC68FC75" box="[815,910,908,933]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFC75FC61FC0DFC74" box="[928,1003,908,932]" pageId="58" pageNumber="59">CPFA</collectionCode>
); same locality,
<date id="FFD11054F058FFCDFB72FC61FACFFC75" box="[1191,1321,908,933]" pageId="58" pageNumber="59" value="2008-07-20">20.vii.2008</date>
, coll. J.L. Giuglaris—
<specimenCount id="9D69FD1DF058FFCDFEF4FC42FE9BFC18" box="[289,381,943,968]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFE58FC5DFE3EFC18" box="[397,472,944,968]" pageId="58" pageNumber="59">CPFA</collectionCode>
); PK 5, Piste KM 25, Route Régina-Saint-Georges, (
<geoCoordinate id="EE5B5053F058FFCDFBFAFC42FB7FFC18" box="[1071,1177,943,968]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="15" value="4.115">4°6'54''N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFB70FC42FAC2FC18" box="[1189,1316,943,968]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="15" value="-52.12111">52°7'16''W</geoCoordinate>
),
<date id="FFD11054F058FFCDFAEDFC42FA71FC18" box="[1336,1431,943,968]" pageId="58" pageNumber="59" value="2008-07">vii.2008</date>
, coll. J.L. Giuglaris—
<specimenCount id="9D69FD1DF058FFCDFE46FC39FE15FC3D" box="[403,499,980,1005]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFDD2FC39FDB1FC3C" box="[519,599,980,1004]" pageId="58" pageNumber="59">ATHC</collectionCode>
); same locality,
<date id="FFD11054F058FFCDFCCDFC39FC96FC3D" box="[792,880,980,1005]" pageId="58" pageNumber="59" value="2008-09">ix.2008</date>
, coll. J.L. Giuglaris—
<specimenCount id="9D69FD1DF058FFCDFBA9FC39FB20FC3D" box="[1148,1222,980,1005]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFB0FFC39FACCFC3C" box="[1242,1322,980,1004]" pageId="58" pageNumber="59">ATHC</collectionCode>
); Réserve naturelle des Nouragues, (
<geoCoordinate id="EE5B5053F058FFCDFE15FC15FDF4FBC0" box="[448,530,1016,1040]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="4.3166666">4°19'N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFDCAFC1AFD63FBC0" box="[543,645,1015,1040]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="-52.366665">52°22'W</geoCoordinate>
),
<date id="FFD11054F058FFCDFD4FFC1AFCE8FBC0" box="[666,782,1015,1040]" pageId="58" pageNumber="59" value="2008-07-02">2.vii.2008</date>
, coll. F. Feer—
<specimenCount id="9D69FD1DF058FFCDFC11FC15FBEDFBC0" box="[964,1035,1015,1040]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFBCEFC15FB6CFBC0" box="[1051,1162,1016,1040]" pageId="58" pageNumber="59">MNHNB</collectionCode>
); Savane Matiti, (
<geoCoordinate id="EE5B5053F058FFCDFA80FC1AFA7EFBC0" box="[1365,1432,1015,1040]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="925" value="5.0833335">5°5'N</geoCoordinate>
,
<geoCoordinate id="EE5B5053F058FFCDFF42FBF1FF1BFBE5" box="[151,253,1052,1077]" direction="west" orientation="longitude" pageId="58" pageNumber="59" precision="925" value="-52.616665">52°37'W</geoCoordinate>
),
<date id="FFD11054F058FFCDFEC1FBF1FEBAFBE5" box="[276,348,1052,1077]" pageId="58" pageNumber="59" value="2011-01">i.2011</date>
, coll. [anonymous]—
<specimenCount id="9D69FD1DF058FFCDFD8EFBF1FD5BFBE5" box="[603,701,1052,1077]" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
,
<specimenCount id="9D69FD1DF058FFCDFD19FBF1FCFCFBE5" box="[716,794,1052,1077]" pageId="58" pageNumber="59" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFCF9FBF1FC91FBE4" box="[812,887,1052,1076]" pageId="58" pageNumber="59">CPFA</collectionCode>
); same locality,
<date id="FFD11054F058FFCDFBE1FBF1FB6AFBE5" box="[1076,1164,1052,1077]" pageId="58" pageNumber="59" value="2009-09">ix.2009</date>
, coll. J.L. Giuglaris—
<specimenCount id="9D69FD1DF058FFCDFA41FBF1FF05FB88" pageId="58" pageNumber="59" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFF27FBADFEAEFB88" box="[242,328,1088,1112]" pageId="58" pageNumber="59">PMOC</collectionCode>
); [
<emphasis id="B91BEA86F058FFCDFEB1FBD2FE06FB88" bold="true" box="[356,480,1087,1112]" pageId="58" pageNumber="59">
NO
<collectionCode id="ED7EAE51F058FFCDFE40FBD2FE06FB88" box="[405,480,1087,1112]" pageId="58" pageNumber="59">DATA</collectionCode>
</emphasis>
]: -., coll. [anonymous]—
<specimenCount id="9D69FD1DF058FFCDFCD4FBADFCB3FB88" box="[769,853,1087,1112]" pageId="58" pageNumber="59" type="male">3 males</specimenCount>
(
<collectionCode id="ED7EAE51F058FFCDFCB6FBADFC4AFB88" box="[867,940,1088,1112]" pageId="58" pageNumber="59">CPFA</collectionCode>
,
<collectionCode id="ED7EAE51F058FFCDFC62FBADFBE8FB88" box="[951,1038,1088,1112]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="58" pageNumber="59">IRSNB</collectionCode>
).
</paragraph>
<paragraph id="8BD03694F058FFCDFF12FB89FBD9FBAC" blockId="58.[151,1437,151,1472]" box="[199,1087,1124,1149]" pageId="58" pageNumber="59">
<emphasis id="B91BEA86F058FFCDFF12FB89FE9BFBAD" bold="true" box="[199,381,1124,1149]" pageId="58" pageNumber="59">Natural history</emphasis>
. Some specimens were collected with flight interception traps.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF058FFCDFF12FB6AFD0EFA6F" pageId="58" pageNumber="59" type="discussion">
<paragraph id="8BD03694F058FFCDFF12FB6AFE5BFAE0" blockId="58.[151,1437,151,1472]" pageId="58" pageNumber="59">
<emphasis id="B91BEA86F058FFCDFF12FB6AFED2FB70" bold="true" box="[199,308,1159,1184]" pageId="58" pageNumber="59">Remarks</emphasis>
. Meso and hyperthelic males have the clypeofrontal carina trapezoidal in front view, the pronotal anterior declivity has a concave impression on each side of mid line and the anterior transverse carina projecting forward and emarginate medially in dorsal view. Parameres (Figs. 86–87) feebly dented in dorsal view, lacking distinct raspy tubercles apically. Variation. Little except for size and a single specimen from the state of Rondônia with green metallic sheen.
</paragraph>
<paragraph id="8BD03694F058FFCDFF12FAD1FC7DFAA7" blockId="58.[151,1437,151,1472]" pageId="58" pageNumber="59">
<emphasis id="B91BEA86F058FFCDFF12FAD1FDD1FA85" bold="true" box="[199,567,1340,1365]" pageId="58" pageNumber="59">Nomenclature and taxonomy</emphasis>
.
<taxonomicName id="4C6F4D17F058FFCDFD84FAD1FB79FA84" authority="Genier &amp; Arnaud, 2014" authorityName="Genier &amp; Arnaud" authorityYear="2014" box="[593,1183,1340,1365]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="attalus">
<emphasis id="B91BEA86F058FFCDFD84FAD1FCB8FA85" box="[593,862,1340,1365]" italics="true" pageId="58" pageNumber="59">Dendropaemon attalus</emphasis>
Génier &amp; Arnaud, 2014
</taxonomicName>
<emphasis id="B91BEA86F058FFCDFB67FAD1FA8FFA85" bold="true" box="[1202,1385,1340,1365]" pageId="58" pageNumber="59">nomen novum</emphasis>
for
<emphasis id="B91BEA86F058FFCDFF42FAB2FEBCFAA8" box="[151,346,1375,1400]" italics="true" pageId="58" pageNumber="59">Dendrop[a]emon</emphasis>
<taxonomicName id="4C6F4D17F058FFCDFEB4FA8CFDBFFAA8" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[353,601,1375,1400]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F058FFCDFEB4FA8CFE58FAA8" box="[353,446,1377,1400]" italics="true" pageId="58" pageNumber="59">amyntas</emphasis>
Harold, 1868
</taxonomicName>
<emphasis id="B91BEA86F058FFCDFDB5FAB2FC72FAA8" bold="true" box="[608,916,1375,1400]" pageId="58" pageNumber="59">primary junior homonym</emphasis>
.
</paragraph>
<paragraph id="8BD03694F058FFCDFF42FA69FD0EFA6F" blockId="58.[151,1437,151,1472]" pageId="58" pageNumber="59">
A replacement name is necessary for
<taxonomicName id="4C6F4D17F058FFCDFD95FA68FCBBFA4D" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[576,861,1412,1437]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F058FFCDFD95FA68FDB2FA4C" box="[576,596,1413,1436]" italics="true" pageId="58" pageNumber="59">D</emphasis>
.
<emphasis id="B91BEA86F058FFCDFDB1FA68FD27FA4C" box="[612,705,1413,1436]" italics="true" pageId="58" pageNumber="59">amyntas</emphasis>
Harold, 1868
</taxonomicName>
because of the previous description of
<taxonomicName id="4C6F4D17F058FFCDFACEFA68FEBFFA10" authority="Lacordaire, 1856" authorityName="Lacordaire" authorityYear="1856" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F058FFCDFACEFA68FAC9FA4C" box="[1307,1327,1413,1436]" italics="true" pageId="58" pageNumber="59">D</emphasis>
.
<emphasis id="B91BEA86F058FFCDFAEBFA68FA7DFA4C" box="[1342,1435,1413,1436]" italics="true" pageId="58" pageNumber="59">amyntas</emphasis>
Lacordaire, 1856
</taxonomicName>
, which represents a different taxon.
</paragraph>
</subSubSection>
</treatment>
</document>
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