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<document id="C21E2FDABDDB3891C95E976B9BD421E6" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F03AFFAFFF42F9F9FD68F844" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Glaphyropaemon) Genier &amp; Arnaud" docType="treatment" docVersion="10" lastPageNumber="89" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="89" updateTime="1698234910999" updateUser="plazi">
<mods:mods id="8804C3B567C19F697D93E557F888C815" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BB57CEB60CA71DE8257C2A766E08A6ED">
<mods:title id="AED5591D0921490D6DD39371414835F1">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
</mods:titleInfo>
<mods:name id="D255607424E7D09252E3BBEC943C46EC" type="personal">
<mods:role id="1D25FD8F784981C706887538442F50C9">
<mods:roleTerm id="904D1672D567448B0ABA89427D7D494A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F4D9FD066473D4CCED0E4675E82C47A">François Génier</mods:namePart>
</mods:name>
<mods:name id="472391D06C33F930EAFD751EB2B62F43" type="personal">
<mods:role id="BCCE9C5170124BC74ABF3CB82658A089">
<mods:roleTerm id="658C56D8BEA04FE9343E8093521FEA9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C7E20DBBEADCFA11FCAC786A5C14D0C">Patrick Arnaud</mods:namePart>
</mods:name>
<mods:typeOfResource id="628A27288AD03145C0005369876264FC">text</mods:typeOfResource>
<mods:relatedItem id="D68D0B7A3B3EEA25704712F4F523C0AE" type="host">
<mods:titleInfo id="BD3E7B9F77A0616B7CA09C639573C8D9">
<mods:title id="415E9DAA3C20200E5B079D9369D088DF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CE06971338132500FBD8528A471F3D28">
<mods:date id="86359CD3CFE236F4CBFDFF94CDCB751F">2016</mods:date>
<mods:detail id="65F7EAC6972C1760E0D0207CDB480491" type="volume">
<mods:number id="CAB8B8FA0B3A8C9E0AFBFA7555BEF677">4099</mods:number>
</mods:detail>
<mods:detail id="79A8C59E713C53483AC582C2C1DD315F" type="issue">
<mods:number id="7C4152C4FEF2293CB58718394A31840E">1</mods:number>
</mods:detail>
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<mods:classification id="544E8728DFAA4EDE194A6225BE7E5F02">journal volume</mods:classification>
<mods:identifier id="1A0AF9B13AC60D8110BFC32498E495E7" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
<mods:identifier id="1032CF571C250C1CE712330B45DE2241" type="GBIF-Dataset">5d7eab55-66e2-43ed-b764-bac0c6e5b6d6</mods:identifier>
<mods:identifier id="1B273BA5022F7DA2E050FEF1C36A06C0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5174640CE0517EA65856E2D62AD0210E" type="Zenodo-Dep">268274</mods:identifier>
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<treatment id="03C68782F03AFFAFFF42F9F9FD68F844" ID-DOI="http://doi.org/10.5281/zenodo.5780394" ID-GBIF-Taxon="127631943" ID-Zenodo-Dep="5780394" LSID="urn:lsid:plazi:treatment:03C68782F03AFFAFFF42F9F9FD68F844" httpUri="http://treatment.plazi.org/id/03C68782F03AFFAFFF42F9F9FD68F844" lastPageNumber="89" pageId="88" pageNumber="89">
<subSubSection id="C375651FF03AFFAFFF42F9F9FB85F980" pageId="88" pageNumber="89" type="nomenclature">
<paragraph id="8BD03694F03AFFAFFF42F9F9FC59F9FD" blockId="88.[151,1155,1556,1941]" box="[151,959,1556,1581]" pageId="88" pageNumber="89">
<heading id="D09881F8F03AFFAFFF42F9F9FC59F9FD" bold="true" box="[151,959,1556,1581]" fontSize="10" level="2" pageId="88" pageNumber="89" reason="2">
<emphasis id="B91BEA86F03AFFAFFF42F9F9FC59F9FD" bold="true" box="[151,959,1556,1581]" pageId="88" pageNumber="89">
<taxonomicName id="4C6F4D17F03AFFAFFF42F9F9FCEAF9FD" ID-CoL="8CWN" authority="Genier &amp; Arnaud" authorityName="Genier &amp; Arnaud" box="[151,780,1556,1581]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="subGenus" subGenus="Glaphyropaemon">
<emphasis id="B91BEA86F03AFFAFFF42F9F9FEACF9FD" bold="true" box="[151,330,1556,1581]" italics="true" pageId="88" pageNumber="89">Dendropaemon</emphasis>
(
<emphasis id="B91BEA86F03AFFAFFE8FF9F9FDC5F9FD" bold="true" box="[346,547,1556,1581]" italics="true" pageId="88" pageNumber="89">Glaphyropaemon</emphasis>
) Génier &amp; Arnaud
</taxonomicName>
; new subgenus
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD03694F03AFFAFFF42F9D5FB85F980" blockId="88.[151,1155,1556,1941]" box="[151,1123,1591,1616]" pageId="88" pageNumber="89">
23.
<taxonomicName id="4C6F4D17F03AFFAFFF14F9DAFC6DF980" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" box="[193,907,1591,1616]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="angustipennis" subGenus="Glaphyropaemon">
<emphasis id="B91BEA86F03AFFAFFF14F9DAFE89F980" box="[193,367,1591,1616]" italics="true" pageId="88" pageNumber="89">Dendropaemon</emphasis>
(
<emphasis id="B91BEA86F03AFFAFFEABF9DAFDA7F980" box="[382,577,1591,1616]" italics="true" pageId="88" pageNumber="89">Glaphyropaemon</emphasis>
)
<emphasis id="B91BEA86F03AFFAFFD84F9D4FD0BF980" box="[593,749,1593,1616]" italics="true" pageId="88" pageNumber="89">angustipennis</emphasis>
Harold, 1869
</taxonomicName>
;
<emphasis id="B91BEA86F03AFFAFFC43F9DAFB85F980" bold="true" box="[918,1123,1591,1616]" pageId="88" pageNumber="89">new combination</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF03AFFAFFF12F9B1FCF2F9A5" box="[199,788,1628,1653]" pageId="88" pageNumber="89" type="reference_group">
<paragraph id="8BD03694F03AFFAFFF12F9B1FCF2F9A5" blockId="88.[151,1155,1556,1941]" box="[199,788,1628,1653]" pageId="88" pageNumber="89">
<taxonomicName id="4C6F4D17F03AFFAFFF12F9B1FDBEF9A4" ID-CoL="34R3L" authority="Blut, 1939" authorityName="Blut" authorityYear="1939" box="[199,600,1628,1653]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="silvanus">
<emphasis id="B91BEA86F03AFFAFFF12F9B1FE31F9A5" box="[199,471,1628,1653]" italics="true" pageId="88" pageNumber="89">Dendropaemon silvanus</emphasis>
<bibRefCitation id="EFFE4B65F03AFFAFFE0BF9B1FDBEF9A4" author="Blut" box="[478,600,1628,1653]" pageId="88" pageNumber="89" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">Blut, 1939</bibRefCitation>
</taxonomicName>
;
<emphasis id="B91BEA86F03AFFAFFDB6F9B1FCF2F9A5" bold="true" box="[611,788,1628,1653]" pageId="88" pageNumber="89">new synonymy</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF03AFFAFFF12F992FBC8F96D" pageId="88" pageNumber="89" type="distribution">
<paragraph id="8BD03694F03AFFAFFF12F992FBC8F96D" blockId="88.[151,1155,1556,1941]" pageId="88" pageNumber="89">
Distribution (
<figureCitation id="13542A11F03AFFAFFE88F992FE26F948" box="[349,448,1663,1688]" captionStart="FIGURE 158" captionStartId="118.[151,250,1248,1270]" captionTargetBox="[150,1436,193,1226]" captionTargetId="figure@118.[150,1436,193,1227]" captionTargetPageId="118" captionText="FIGURE 158. Known distribution of D. convexus, D. fredericki, D. angustipennis, D. bahianus, D. inemarginatus, D. nigritulus." httpUri="https://zenodo.org/record/268288/files/figure.png" pageId="88" pageNumber="89">Fig. 158</figureCitation>
):
<collectingCountry id="F3787604F03AFFAFFE01F992FDCFF948" box="[468,553,1663,1688]" name="Bolivia" pageId="88" pageNumber="89">Bolivia</collectingCountry>
,
<collectingCountry id="F3787604F03AFFAFFDE0F992FD9FF948" box="[565,633,1663,1688]" name="Brazil" pageId="88" pageNumber="89">Brazil</collectingCountry>
(AC, AM, MG, PA),
<collectingCountry id="F3787604F03AFFAFFCBCF992FC3CF948" box="[873,986,1663,1688]" name="Colombia" pageId="88" pageNumber="89">Colombia</collectingCountry>
,
<collectingCountry id="F3787604F03AFFAFFC33F992FBA3F948" box="[998,1093,1663,1688]" name="Ecuador" pageId="88" pageNumber="89">Ecuador</collectingCountry>
,
<collectingCountry id="F3787604F03AFFAFFB85F96DFB65F947" box="[1104,1155,1664,1687]" name="Peru" pageId="88" pageNumber="89">Peru</collectingCountry>
24.
<taxonomicName id="4C6F4D17F03AFFAFFF14F949FCB0F96D" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[193,854,1700,1725]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="bahianus" subGenus="Glaphyropaemon">
<emphasis id="B91BEA86F03AFFAFFF14F949FE89F96D" box="[193,367,1700,1725]" italics="true" pageId="88" pageNumber="89">Dendropaemon</emphasis>
(
<emphasis id="B91BEA86F03AFFAFFEABF949FDA7F96D" box="[382,577,1700,1725]" italics="true" pageId="88" pageNumber="89">Glaphyropaemon</emphasis>
)
<emphasis id="B91BEA86F03AFFAFFD84F949FD51F96D" box="[593,695,1700,1725]" italics="true" pageId="88" pageNumber="89">bahianus</emphasis>
Harold, 1868
</taxonomicName>
;
<emphasis id="B91BEA86F03AFFAFFCB7F949FBC8F96D" bold="true" box="[866,1070,1700,1725]" pageId="88" pageNumber="89">new combination</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF03AFFAFFF12F92AFCD0F8F8" pageId="88" pageNumber="89" type="reference_group">
<paragraph id="8BD03694F03AFFAFFF12F92AFCB8F930" blockId="88.[151,1155,1556,1941]" box="[199,862,1735,1760]" pageId="88" pageNumber="89">
<taxonomicName id="4C6F4D17F03AFFAFFF12F92AFD47F930" ID-CoL="34R9D" authority="Waterhouse, 1891" authorityName="Waterhouse" authorityYear="1891" box="[199,673,1735,1760]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="lobatus">
<emphasis id="B91BEA86F03AFFAFFF12F92AFE28F930" box="[199,462,1735,1760]" italics="true" pageId="88" pageNumber="89">Dendropaemon lobatus</emphasis>
<bibRefCitation id="EFFE4B65F03AFFAFFE01F92AFD47F930" author="Waterhouse" box="[468,673,1735,1760]" pageId="88" pageNumber="89" refString="Waterhouse, C. O. (1891) New Scarabaeidae in the British Museum: a Fifth Contribution. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology, Sixth Series, 8 (43), 53 - 61. http: // dx. doi. org / 10.1080 / 00222939109460388" type="journal article" year="1891">Waterhouse, 1891</bibRefCitation>
</taxonomicName>
;
<emphasis id="B91BEA86F03AFFAFFD79F92AFCB8F930" bold="true" box="[684,862,1735,1760]" pageId="88" pageNumber="89">new synonymy</emphasis>
</paragraph>
<paragraph id="8BD03694F03AFFAFFF12F901FCB5F8D5" blockId="88.[151,1155,1556,1941]" box="[199,851,1772,1797]" pageId="88" pageNumber="89">
<taxonomicName id="4C6F4D17F03AFFAFFF12F901FD71F8D4" ID-CoL="34R3P" authority="Felsche, 1909" authorityName="Felsche" authorityYear="1909" box="[199,663,1772,1797]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="tenuitarsis">
<emphasis id="B91BEA86F03AFFAFFF12F901FE17F8D4" box="[199,497,1772,1797]" italics="true" pageId="88" pageNumber="89">Dendropaemon tenuitarsis</emphasis>
<bibRefCitation id="EFFE4B65F03AFFAFFE2DF901FD71F8D4" author="Felsche" box="[504,663,1772,1797]" pageId="88" pageNumber="89" refString="Felsche, C. (1909) Neue und alte coprophage Scarabaeiden. (Col.). Deutsche Entomologische Zeitschrift, 1909 (6), 751 - 765. http: // dx. doi. org / 10.1002 / mmnd. 48019090608" type="journal article" year="1909">Felsche, 1909</bibRefCitation>
</taxonomicName>
;
<emphasis id="B91BEA86F03AFFAFFD77F901FCB5F8D5" bold="true" box="[674,851,1772,1797]" pageId="88" pageNumber="89">new synonymy</emphasis>
</paragraph>
<paragraph id="8BD03694F03AFFAFFF12F8E2FCD0F8F8" blockId="88.[151,1155,1556,1941]" box="[199,822,1807,1832]" pageId="88" pageNumber="89">
<taxonomicName id="4C6F4D17F03AFFAFFF12F8E2FD9FF8F8" ID-CoL="6CLCF" authority="Blut, 1939" authorityName="Blut" authorityYear="1939" box="[199,633,1807,1832]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="batrachites">
<emphasis id="B91BEA86F03AFFAFFF12F8E2FE1FF8F8" box="[199,505,1807,1832]" italics="true" pageId="88" pageNumber="89">Dendropaemon batrachites</emphasis>
<bibRefCitation id="EFFE4B65F03AFFAFFDD5F8E2FD9FF8F8" author="Blut" box="[512,633,1807,1832]" pageId="88" pageNumber="89" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">Blut, 1939</bibRefCitation>
</taxonomicName>
;
<emphasis id="B91BEA86F03AFFAFFD50F8E2FCD0F8F8" bold="true" box="[645,822,1807,1832]" pageId="88" pageNumber="89">new synonymy</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF03AFFAFFF12F8D9FD68F844" pageId="88" pageNumber="89" type="distribution">
<paragraph id="8BD03694F03AFFAFFF12F8D9FBB3F8A0" blockId="88.[151,1155,1556,1941]" pageId="88" pageNumber="89">
Distribution (
<figureCitation id="13542A11F03AFFAFFE88F8D9FE26F89D" box="[349,448,1844,1869]" captionStart="FIGURE 158" captionStartId="118.[151,250,1248,1270]" captionTargetBox="[150,1436,193,1226]" captionTargetId="figure@118.[150,1436,193,1227]" captionTargetPageId="118" captionText="FIGURE 158. Known distribution of D. convexus, D. fredericki, D. angustipennis, D. bahianus, D. inemarginatus, D. nigritulus." httpUri="https://zenodo.org/record/268288/files/figure.png" pageId="88" pageNumber="89">Fig. 158</figureCitation>
):
<collectingCountry id="F3787604F03AFFAFFE01F8D9FDFEF89D" box="[468,536,1844,1869]" name="Brazil" pageId="88" pageNumber="89">Brazil</collectingCountry>
(BA, DF, GO, MG, MT, PB, RJ),
<collectingCountry id="F3787604F03AFFAFFC43F8D8FC1BF89D" box="[918,1021,1845,1869]" name="Paraguay" pageId="88" pageNumber="89">Paraguay</collectingCountry>
25.
<taxonomicName id="4C6F4D17F03AFFAFFF14F8BAFC26F8A0" authority="Genier &amp; Arnaud" authorityName="Genier &amp; Arnaud" box="[193,960,1879,1904]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="inemarginatus" subGenus="Glaphyropaemon">
<emphasis id="B91BEA86F03AFFAFFF14F8BAFE89F8A0" box="[193,367,1879,1904]" italics="true" pageId="88" pageNumber="89">Dendropaemon</emphasis>
(
<emphasis id="B91BEA86F03AFFAFFEABF8BAFDA7F8A0" box="[382,577,1879,1904]" italics="true" pageId="88" pageNumber="89">Glaphyropaemon</emphasis>
)
<emphasis id="B91BEA86F03AFFAFFD84F8B4FD14F8A0" box="[593,754,1881,1904]" italics="true" pageId="88" pageNumber="89">inemarginatus</emphasis>
Génier &amp; Arnaud
</taxonomicName>
;
<emphasis id="B91BEA86F03AFFAFFC19F8BAFBB3F8A0" bold="true" box="[972,1109,1879,1904]" pageId="88" pageNumber="89">new species</emphasis>
</paragraph>
<paragraph id="8BD03694F03AFFAFFF12F891FD68F844" blockId="88.[151,1155,1556,1941]" box="[199,654,1916,1941]" pageId="88" pageNumber="89">
Distribution (
<figureCitation id="13542A11F03AFFAFFE88F891FE26F845" box="[349,448,1916,1941]" captionStart="FIGURE 158" captionStartId="118.[151,250,1248,1270]" captionTargetBox="[150,1436,193,1226]" captionTargetId="figure@118.[150,1436,193,1227]" captionTargetPageId="118" captionText="FIGURE 158. Known distribution of D. convexus, D. fredericki, D. angustipennis, D. bahianus, D. inemarginatus, D. nigritulus." httpUri="https://zenodo.org/record/268288/files/figure.png" pageId="88" pageNumber="89">Fig. 158</figureCitation>
):
<collectingCountry id="F3787604F03AFFAFFE01F891FDA1F845" box="[468,583,1916,1941]" name="Venezuela" pageId="88" pageNumber="89">Venezuela</collectingCountry>
(AM)
</paragraph>
</subSubSection>
</treatment>
</document>
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