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<document id="6099F45063EB2F7653168740E6C81332" ID-DOI="10.1080/00222933.2018.1435833" ID-ISSN="1464-5262" ID-Zenodo-Dep="5174451" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628558511936" checkinUser="felipe" docAuthor="Wagner, Philipp, Greenbaum, Eli, Bauer, Aaron M., Kusamba, Chifundera &amp; Leaché, Adam D." docDate="2018" docId="03BFD058FF82FFB0158D89F2BA3BFDB5" docLanguage="en" docName="JNatHist.52.13-16.771-817.pdf" docOrigin="Journal of Natural History 52 (13 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2018.1435833" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Acanthocercus branchi Wagner, Greenbaum and Bauer 2012" docType="treatment" docVersion="3" lastPageNumber="797" masterDocId="FF86A820FF9BFFAB154A8901BE4DFF9E" masterDocTitle="Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae)" masterLastPageNumber="817" masterPageNumber="771" pageNumber="795" updateTime="1699046512504" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DB9F62109CC798E79AFDAFC4697CF81B">Lifting the blue-headed veil - integrative taxonomy of the Acanthocercus atricollis species complex (Squamata: Agamidae)</mods:title>
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<mods:namePart id="AC43D4964C597B580C0F0A838E8AF9A7">Greenbaum, Eli</mods:namePart>
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<mods:namePart id="D849F3A6CFCF367818A2F3CA6A4EF9B5">Bauer, Aaron M.</mods:namePart>
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<mods:namePart id="2B16D42002455CB89AE983F2EDD20656">Kusamba, Chifundera</mods:namePart>
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<mods:date id="7252C26203BBDDED2F65E70115501095">2018</mods:date>
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<mods:number id="F1E04FC75DD779281CC4411BC2C34880">13 - 16</mods:number>
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<paragraph id="8BA9614EFF82FFB2158D89F2BA11FE93" blockId="25.[199,1116,243,304]" box="[199,1116,243,269]" pageId="25" pageNumber="795">
<heading id="D0E1D622FF82FFB2158D89F2BA11FE93" box="[199,1116,243,269]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="795" reason="5">
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<treatmentCitation id="0AB7475FFF82FFB2158D89F2BD92FE93" author="Wagner P &amp; Greenbaum E &amp; Bauer AM" box="[199,991,243,269]" page="22" pageId="25" pageNumber="795" year="2012">
<taxonomicName id="4C161ACDFF82FFB2158D89F2BD92FE93" authority="Wagner, Greenbaum and Bauer, 2012 b: 22" authorityName="Wagner, Greenbaum and Bauer" authorityPageNumber="22" authorityYear="2012" box="[199,991,243,269]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="795" phylum="Chordata" rank="species" species="branchi">
<emphasis id="B962BD5CFF82FFB2158D89F2BF95FE93" bold="true" box="[199,472,243,269]" italics="true" pageId="25" pageNumber="795">Acanthocercus branchi</emphasis>
<bibRefCitation id="EF871CBFFF82FFB214AA89F5BD92FE93" author="Wagner P &amp; Greenbaum E &amp; Bauer AM" box="[480,991,244,269]" pageId="25" pageNumber="795" pagination="21 - 30" refId="ref21608" refString="Wagner P, Greenbaum E, Bauer AM. 2012 b. A new species of the Acanthocercus atricollis (Squamata: Agamidae) complex from Zambia. Salamandra. 48: 21 - 30." type="journal article" year="2012">Wagner, Greenbaum and Bauer, 2012b: 22</bibRefCitation>
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, figs. 12a
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</paragraph>
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<subSubSection id="C30C32C5FF82FFB2171B8816BF4CFE29" pageId="25" pageNumber="795" type="description">
<paragraph id="8BA9614EFF82FFB2171B8816BC9FFEAE" blockId="25.[199,1116,243,304]" box="[593,722,279,304]" pageId="25" pageNumber="795">
(
<figureCitation id="132D7DCBFF82FFB217138816BC87FEAE" box="[601,714,279,304]" captionStart="Figure 11" captionStartId="26.[160,225,1533,1556]" captionTargetBox="[178,1138,142,1502]" captionTargetId="figure-1@26.[178,1138,142,1502]" captionTargetPageId="26" captionText="Figure 11. Holotype (ZFMK 88682) of Acanthocercus branchi, from Chipata, Zambia." figureDoi="http://doi.org/10.5281/zenodo.5174475" httpUri="https://zenodo.org/record/5174475/files/figure.png" pageId="25" pageNumber="795">Figure 11</figureCitation>
)
</paragraph>
<paragraph id="8BA9614EFF82FFB215EA885BBF4CFE29" blockId="25.[160,1129,346,439]" pageId="25" pageNumber="795">
2012
<taxonomicName id="4C161ACDFF82FFB215AE885BBD38FEEC" authority="Wagner, Greenbaum &amp; Bauer, A" authorityName="Wagner, Greenbaum &amp; Bauer, A" authorityYear="2018" box="[228,885,346,370]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="795" phylum="Chordata" rank="species" species="branchi" status="sp. nov.">
<emphasis id="B962BD5CFF82FFB215AE885BBFA4FEEC" box="[228,489,346,370]" italics="true" pageId="25" pageNumber="795">Acanthocercus branchi</emphasis>
Wagner, Greenbaum &amp; Bauer, A
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<taxonomicNameLabel id="A2510027FF82FFB216CB885BBA5FFEEC" box="[897,1042,346,370]" pageId="25" pageNumber="795" rank="species">new species</taxonomicNameLabel>
of the
<taxonomicName id="4C161ACDFF82FFB215F4887CBFF5FE0B" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[190,440,381,405]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="795" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFF82FFB215F4887CBFF5FE0B" box="[190,440,381,405]" italics="true" pageId="25" pageNumber="795">Acanthocercus atricollis</emphasis>
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complex (
<taxonomicName id="4C161ACDFF82FFB21760887CBCD0FE0B" box="[554,669,381,405]" class="Reptilia" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="795" phylum="Chordata" rank="order">Squamata</taxonomicName>
:
<taxonomicName id="4C161ACDFF82FFB217EC887CBD55FE0B" authorityName="Gray" authorityYear="1827" box="[678,792,381,405]" class="Reptilia" family="Agamidae" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="795" phylum="Chordata" rank="family">Agamidae</taxonomicName>
) from
<collectingCountry id="F30121DEFF82FFB2162A887CBDF8FE0B" box="[864,949,381,405]" name="Zambia" pageId="25" pageNumber="795">Zambia</collectingCountry>
. Salamandra 48: 2130.
</paragraph>
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<subSubSection id="C30C32C5FF82FFB215EA88EFBF28FDD2" pageId="25" pageNumber="795" type="materials_examined">
<paragraph id="8BA9614EFF82FFB215EA88EFBF46FD99" blockId="25.[160,1155,494,588]" box="[160,267,494,519]" pageId="25" pageNumber="795">
<emphasis id="B962BD5CFF82FFB215EA88EFBF46FD99" bold="true" box="[160,267,494,519]" italics="true" pageId="25" pageNumber="795">
<typeStatus id="54ADDFECFF82FFB215EA88EFBF46FD99" box="[160,267,494,519]" pageId="25" pageNumber="795" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA9614EFF82FFB215EA8B10BF28FDD2" blockId="25.[160,1155,494,588]" pageId="25" pageNumber="795">
<materialsCitation id="3B7E6B13FF82FFB215EA8B10BF28FDD2" collectionCode="ZAWA" country="Zambia" location="Chipata'" pageId="25" pageNumber="795" specimenCode="ZFMK 88682" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB0C935FF82FFB215EA8B10BF78FDB7" box="[160,309,529,553]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="25" pageNumber="795" type="Museum">ZFMK 88682</specimenCode>
, from dense Miombo woodland close to the
<collectionCode id="ED07F98BFF82FFB216218B10BDFCFDB7" box="[875,945,529,553]" pageId="25" pageNumber="795">ZAWA</collectionCode>
head office near
<location id="8EC93795FF82FFB215EA8B35BEB6FDD2" LSID="urn:lsid:plazi:treatment:03BFD058FF82FFB0158D89F2BA3BFDB5:8EC93795FF82FFB215EA8B35BEB6FDD2" box="[160,251,564,588]" country="Zambia" name="Chipata'" pageId="25" pageNumber="795">Chipata</location>
,
<collectingCountry id="F30121DEFF82FFB214408B35BF2CFDD2" box="[266,353,564,588]" name="Zambia" pageId="25" pageNumber="795">Zambia</collectingCountry>
.
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<subSubSection id="C30C32C5FF82FFB215EA8B82BC9BFB58" pageId="25" pageNumber="795" type="description">
<paragraph id="8BA9614EFF82FFB215EA8B82BF6BFD02" blockId="25.[160,1156,643,1222]" box="[160,294,643,668]" pageId="25" pageNumber="795">
<emphasis id="B962BD5CFF82FFB215EA8B82BF6BFD02" bold="true" box="[160,294,643,668]" italics="true" pageId="25" pageNumber="795">Description</emphasis>
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A small to medium sized species with a total length up to
<quantity id="4CEECCABFF82FFB216048BA7BDE1FD20" box="[846,940,678,702]" metricMagnitude="-1" metricUnit="m" metricValue="3.13" pageId="25" pageNumber="795" unit="mm" value="313.0">313 mm</quantity>
(SVL:
<quantity id="4CEECCABFF82FFB216BC8BA7BACDFD20" box="[1014,1152,678,702]" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.31" metricValueMin="0.83" pageId="25" pageNumber="795" unit="mm" value="107.0" valueMax="131.0" valueMin="83.0">83131 mm</quantity>
, x =
<quantity id="4CEECCABFF82FFB215868BC8BF72FD7F" box="[204,319,713,737]" metricMagnitude="-1" metricUnit="m" metricValue="1.132" pageId="25" pageNumber="795" unit="mm" value="113.2">113.2 mm</quantity>
, n = 4), tail short, about 1.4 times longer than SVL (SVL/TL:
<quantity id="4CEECCABFF82FFB216958BC8BACDFD7F" box="[991,1152,712,737]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="25" pageNumber="795" unit="mm" value="0.7" valueMax="0.75" valueMin="0.65">0.650.75 mm</quantity>
, x =
<quantity id="4CEECCABFF82FFB215818BEABF7DFC9D" box="[203,304,747,771]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="25" pageNumber="795" unit="mm" value="0.71">0.71 mm</quantity>
, n = 4). Head distinctly broad in males, as in other species of the genus. Ear openings about the same size as the eyes, with the tympanum visible. Occipital scale lacking, but a parietal shield including a pineal window present. Nostril pear-shaped, slightly convex and below the canthus rostralis. Vertebral crest indistinct. Scales arranged in 103130 (x = 115.5, n = 4) rows around midbody, and 6781 (x = 71.2, n = 4) longitudinal rows along the vertebrae. Matrix scales small, usually smooth and scattered with pale, enlarged and keeled scales. Enlarged scales not organized into clusters, but indistinct transverse rows between the limbs present. Vertebral scales keeled, with a higher density of enlarged scales than on the flanks, sometimes giving an impression of a margin between the vertebral area and flanks. Gular scales flat, smooth, juxtaposed and becoming smaller towards the gular fold. Ventral scales small and smooth, in 7995 (x = 87.8, n = 4) longitudinal rows. Males usually with two continuous rows of 2125 (x = 22.8, n = 4) precloacal pores; it is assumed these are lacking in females, as is typical for most agamids.
</paragraph>
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<subSubSection id="C30C32C5FF82FFB015EA8DFDBA3BFDB5" lastPageId="27" lastPageNumber="797" pageId="25" pageNumber="795" type="distribution">
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<emphasis id="B962BD5CFF82FFB215EA8DFDBF61FA8B" bold="true" box="[160,300,1276,1301]" italics="true" pageId="25" pageNumber="795">Distribution</emphasis>
</paragraph>
<paragraph id="8BA9614EFF82FFB215EA8C21BACEFAC5" blockId="25.[160,1155,1276,1371]" pageId="25" pageNumber="795">
<materialsCitation id="3B7E6B13FF82FFB215EA8C21BC0DFAC5" country="Zambia" location="Lusaka" pageId="25" pageNumber="795" specimenCount="1" stateProvince="Lusaka">
In
<collectingCountry id="F30121DEFF82FFB2158E8C21BF55FAA6" box="[196,280,1312,1336]" name="Zambia" pageId="25" pageNumber="795">Zambia</collectingCountry>
the species is known from localities within the Luangwa (Chipata) and Zambezi (
<collectingRegion id="49D2AFACFF82FFB214458C42BF12FAC5" box="[271,351,1347,1371]" country="Zambia" name="Lusaka" pageId="25" pageNumber="795">Lusaka</collectingRegion>
) valleys (
<figureCitation id="132D7DCBFF82FFB214818C42BC7AFAC5" box="[459,567,1347,1371]" captionStart="Figure 10" captionStartId="24.[160,225,1538,1561]" captionTargetBox="[178,1138,172,1507]" captionTargetId="figure-2@24.[178,1138,172,1507]" captionTargetPageId="24" captionText="Figure 10. Distribution of Acanthocercus atricollis, A. branchi and A. cyanocephalus. The numbers refer to the specific localities." figureDoi="http://doi.org/10.5281/zenodo.5174473" httpUri="https://zenodo.org/record/5174473/files/figure.png" pageId="25" pageNumber="795">Figure 10</figureCitation>
)
</materialsCitation>
. In
<collectingRegion id="49D2AFACFF82FFB217278C42BCF1FAC5" box="[621,700,1347,1371]" country="Zambia" name="Lusaka" pageId="25" pageNumber="795">Lusaka</collectingRegion>
, it is known from three localities: Great
</paragraph>
<paragraph id="8BA9614EFF82FFB215EA8CCBBA70F92A" blockId="25.[160,1157,1482,1716]" pageId="25" pageNumber="795">
30 = Chongoni Forest Reserve; 31 =
<collectingRegion id="49D2AFACFF82FFB217588CCBBC29FA40" box="[530,612,1482,1502]" country="Malawi" name="Lilongwe" pageId="25" pageNumber="795">Lilongwe</collectingRegion>
; 32 =
<quantity id="4CEECCABFF82FFB217FA8CCBBD4DFA40" box="[688,768,1482,1502]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="25" pageNumber="795" unit="mi" value="30.0">30 miles</quantity>
NNW of Furancungo; 33 = Chifumbazi; 34 = Chikowa; 35 = Chipata; 36 = Katete; 37 =
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W of Katete; 38 = Petauke; 39 = Chakwenga River; 40 =
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E of
<collectingRegion id="49D2AFACFF82FFB2147F8F01BF3EF98A" box="[309,371,1536,1556]" country="Zambia" name="Lusaka" pageId="25" pageNumber="795">Lusaka</collectingRegion>
; 41 =
<collectingRegion id="49D2AFACFF82FFB214E08F01BFA5F98A" box="[426,488,1536,1556]" country="Zambia" name="Lusaka" pageId="25" pageNumber="795">Lusaka</collectingRegion>
; 42 = Ruwa near
<collectingRegion id="49D2AFACFF82FFB217CE8F01BC8DF98A" box="[644,704,1536,1556]" country="Zimbabwe" name="Harare" pageId="25" pageNumber="795">Harare</collectingRegion>
; 43 = Fern Valley; 44 = Mt. Sei, Melsetter; 45 = Mt. Silinda; 46 =
<collectingRegion id="49D2AFACFF82FFB2145D8F1BBF23F9B0" box="[279,366,1562,1582]" country="Zimbabwe" name="Bulawayo" pageId="25" pageNumber="795">Bulawayo</collectingRegion>
; 47 = Rhodes Grave, Matobo Hills; 48 = Krueger National Park; 49 = Hoedspruit Airforce base; 50 =
<quantity id="4CEECCABFF82FFB2145A8F34BF0CF9D7" box="[272,321,1589,1609]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="25" pageNumber="795" unit="km" value="7.0">7 km</quantity>
N of Chibuto; 51 =
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; 52 = Gabane; 53 = Otse; 54 =
<quantity id="4CEECCABFF82FFB216EC8F34BDACF9D7" box="[934,993,1589,1609]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="795" unit="km" value="12.0">12 km</quantity>
N of Lydenberg; 55 = Gerlachshoop [Mission Station]; 56 = Loskopdam; 57 = Waterval Onder; 58 = Barberton; 59 = Louwscreek; 60 = Delagoa Bay (
<collectingRegion id="49D2AFACFF82FFB214068F6BBFD8F9E0" box="[332,405,1642,1662]" country="Mozambique" name="Maputo" pageId="25" pageNumber="795">Maputo</collectingRegion>
); 61 = Ponta do Ouro; 62 = Mkuze Game Reserve; 63 = Somkele; 64 = Empangeni; 65 = Durban; 66 = Umfolozi; 67 = Tugela Estates [Farm]; 68 = Rustenberg [Farm]; 69 = Lunda; 70 = Malange; 71 = Lydenburg (=Mashishing); 72 = Wonderboom, near Pretoria; 73 = Woodbush; 74 = Komatipoort.
</paragraph>
<caption id="DF6931C6FF81FFB115EA8CFCBDA4F98A" ID-DOI="http://doi.org/10.5281/zenodo.5174475" ID-Zenodo-Dep="5174475" box="[160,1001,1532,1556]" httpUri="https://zenodo.org/record/5174475/files/figure.png" pageId="26" pageNumber="796" startId="26.[160,225,1533,1556]" targetBox="[178,1138,142,1502]" targetPageId="26">
<paragraph id="8BA9614EFF81FFB115EA8CFCBDA4F98A" blockId="26.[160,1001,1532,1556]" box="[160,1001,1532,1556]" pageId="26" pageNumber="796">
<emphasis id="B962BD5CFF81FFB115EA8CFCBF44F98A" bold="true" box="[160,265,1533,1556]" pageId="26" pageNumber="796">Figure 11.</emphasis>
Holotype (ZFMK 88682) of
<taxonomicName id="4C161ACDFF81FFB1176B8CFDBCB1F98D" authorityName="Wagner, Greenbaum &amp; Bauer, A" authorityYear="2018" box="[545,764,1532,1555]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="796" phylum="Chordata" rank="species" species="branchi">
<emphasis id="B962BD5CFF81FFB1176B8CFDBCB1F98D" box="[545,764,1532,1555]" italics="true" pageId="26" pageNumber="796">Acanthocercus branchi</emphasis>
</taxonomicName>
, from Chipata, Zambia.
</paragraph>
</caption>
<paragraph id="8BA9614EFF80FFB015EA898FBD6DFECA" blockId="27.[160,1156,142,340]" pageId="27" pageNumber="797">
East Road and Makishi Road, both within the town, and the Pioneer Camp about
<quantity id="4CEECCABFF80FFB01177898FBACEFF38" box="[1085,1155,142,166]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="27" pageNumber="797" unit="km" value="25.0">25 km</quantity>
outside Lusaka on the road to
<collectingCountry id="F30121DEFF80FFB014B189B0BC00FF57" box="[507,589,177,201]" name="Malawi" pageId="27" pageNumber="797">Malawi</collectingCountry>
. In
<collectingCountry id="F30121DEFF80FFB0173F89B0BC89FF57" box="[629,708,177,201]" name="Malawi" pageId="27" pageNumber="797">Malawi</collectingCountry>
it is recognized from
<collectingRegion id="49D2AFACFF80FFB016F789B0BA6EFF57" box="[957,1059,177,201]" country="Malawi" name="Lilongwe" pageId="27" pageNumber="797">Lilongwe</collectingRegion>
and the Chongoni Forest Reserve, south-east of
<collectingRegion id="49D2AFACFF80FFB017CF89D5BCA3FF72" box="[645,750,212,236]" country="Malawi" name="Lilongwe" pageId="27" pageNumber="797">Lilongwe</collectingRegion>
. Moreover, new records show its presence in
<collectingCountry id="F30121DEFF80FFB0147A89F7BFEAFE90" box="[304,423,246,270]" name="Zimbabwe" pageId="27" pageNumber="797">Zimbabwe</collectingCountry>
(Ruwa, near
<collectingRegion id="49D2AFACFF80FFB0170F89F7BCDCFE90" box="[581,657,246,270]" country="Zimbabwe" name="Harare" pageId="27" pageNumber="797">Harare</collectingRegion>
; P. Wagner pers. observation) and the Rift Valley in south-eastern
<collectingCountry id="F30121DEFF80FFB014FE8818BFA9FEAF" box="[436,484,281,305]" name="Democratic Republic of the Congo" pageId="27" pageNumber="797">DRC</collectingCountry>
(Manono,
<collectingRegion id="49D2AFACFF80FFB0172E8818BD60FEAF" box="[612,813,281,305]" country="Democratic Republic of the Congo" name="Katanga" pageId="27" pageNumber="797">Katanga Province</collectingRegion>
) where the species could be sympatric with other members of the
<taxonomicName id="4C161ACDFF80FFB0171F883DBC81FECA" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[597,716,316,340]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="797" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFF80FFB0171F883DBC81FECA" box="[597,716,316,340]" italics="true" pageId="27" pageNumber="797">A. atricollis</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="8BA9614EFF80FFB015EA8886BFD8FE3E" blockId="27.[160,1155,391,555]" box="[160,405,391,416]" pageId="27" pageNumber="797">
<emphasis id="B962BD5CFF80FFB015EA8886BFD8FE3E" bold="true" box="[160,405,391,416]" italics="true" pageId="27" pageNumber="797">Habitat and ecology</emphasis>
</paragraph>
<paragraph id="8BA9614EFF80FFB015EA88ABBA3BFDB5" blockId="27.[160,1155,391,555]" pageId="27" pageNumber="797">
This species is a tree dweller as is typical for the
<taxonomicName id="4C161ACDFF80FFB017A188ABBD08FE5C" baseAuthorityName="Smith" baseAuthorityYear="1849" box="[747,837,426,450]" class="Reptilia" family="Agamidae" genus="Acanthocercus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="797" phylum="Chordata" rank="species" species="atricollis">
<emphasis id="B962BD5CFF80FFB017A188ABBD08FE5C" box="[747,837,426,450]" italics="true" pageId="27" pageNumber="797">atricollis</emphasis>
</taxonomicName>
complex. In Chipata it was collected in dense Miombo woodland with large trees and a structured canopy. In
<collectingRegion id="49D2AFACFF80FFB015EA88F1BEA1FD96" box="[160,236,496,520]" country="Zambia" name="Lusaka" pageId="27" pageNumber="797">Lusaka</collectingRegion>
it was captured on a solitary tree close to the Great East Road, near Manda Hill Shopping Mall. At both localities, small colonies were observed (
<bibRefCitation id="EF871CBFFF80FFB016CB8B12BA27FDB5" author="Wagner P &amp; Greenbaum E &amp; Bauer AM" box="[897,1130,531,555]" pageId="27" pageNumber="797" pagination="21 - 30" refId="ref21608" refString="Wagner P, Greenbaum E, Bauer AM. 2012 b. A new species of the Acanthocercus atricollis (Squamata: Agamidae) complex from Zambia. Salamandra. 48: 21 - 30." type="journal article" year="2012">Wagner et al. 2012b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>