treatments-xml/data/03/B8/CF/03B8CF7BB960FF94B1D2F8B8FD69F804.xml

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<mods:title id="92E892762781589FB1843BEC771A1160">Ngan mayla gen. et sp. nov, a new blind potamid cave crab from Borneo, Kalimantan (Crustacea: Decapoda: Brachyura: Potamoidea, Potamidae), and three other unidentified cave crabs from the same region</mods:title>
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<mods:namePart id="32754B7B5F38F6E85CDFF0C3FD04AA52">Guinot, Danièle</mods:namePart>
<mods:affiliation id="8047F938487AB6BCBF0B282B886ED6BB">Institut de Systématique, Evolution, Biodiversité (ISYEB), UMR 7205 CNRS, MNHN, EPHE</mods:affiliation>
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<mods:namePart id="02A062FB8E381EF8EE655B671655D559">Rodríguez Moreno, Paula A.</mods:namePart>
<mods:affiliation id="C59168239B567C250A86577C8B0ECF7E">Direction Générale Déléguée aux Collections, Muséum national d’Histoire naturelle, MNHN,</mods:affiliation>
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<subSubSection id="C30B2DE6B960FF95B1D2F8B8FC01F8A5" pageId="2" pageNumber="117" type="nomenclature">
<paragraph id="8BAE7E6DB960FF95B1D2F8B8FEF7F881" blockId="2.[151,953,1854,1916]" box="[151,335,1854,1880]" pageId="2" pageNumber="117">
<heading id="D0E6C901B960FF95B1D2F8B8FEF7F881" bold="true" box="[151,335,1854,1880]" fontSize="11" level="1" pageId="2" pageNumber="117" reason="1">
<emphasis id="B965A27FB960FF95B1D2F8B8FEF7F881" bold="true" box="[151,335,1854,1880]" pageId="2" pageNumber="117">
<taxonomicName id="4C1105EEB960FF95B1D2F8B8FF62F881" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[151,218,1854,1880]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="117" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB960FF95B1D2F8B8FF62F881" bold="true" box="[151,218,1854,1880]" italics="true" pageId="2" pageNumber="117">Ngan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2561F04B960FF95B1A4F8B8FEF7F881" box="[225,335,1854,1880]" pageId="2" pageNumber="117" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="C30B2DE6B960FF95B1D2F82CFCA5F81D" box="[151,797,1961,1988]" pageId="2" pageNumber="117" type="type_taxon">
<paragraph id="8BAE7E6DB960FF95B1D2F82CFCA5F81D" blockId="2.[151,797,1961,1988]" box="[151,797,1961,1988]" pageId="2" pageNumber="117">
<emphasis id="B965A27FB960FF95B1D2F82CFE93F81A" bold="true" box="[151,299,1961,1988]" pageId="2" pageNumber="117">
<typeStatus id="54AAC0CFB960FF95B1D2F82CFF6AF81D" box="[151,210,1962,1988]" pageId="2" pageNumber="117">Type</typeStatus>
species
</emphasis>
: 
<taxonomicName id="4C1105EEB960FF95B07FF82CFE7AF81D" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[314,450,1962,1988]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="mayla" status="sp. nov.">
<emphasis id="B965A27FB960FF95B07FF82CFE7AF81D" box="[314,450,1962,1988]" italics="true" pageId="2" pageNumber="117">Ngan mayla</emphasis>
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<taxonomicNameLabel id="A2561F04B960FF95B08CF82CFD9BF81D" box="[457,547,1962,1988]" pageId="2" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(present designation).
</paragraph>
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<subSubSection id="C30B2DE6B961FF94B1D2FF10FDBBFCE0" pageId="3" pageNumber="118" type="diagnosis">
<paragraph id="8BAE7E6DB961FF94B1D2FF10FDBBFCE0" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<emphasis id="B965A27FB961FF94B1D2FF10FEB2FF69" bold="true" box="[151,266,150,176]" pageId="3" pageNumber="118">Diagnosis</emphasis>
. Carapace widest anteriorly, narrowing posteriorly, flat; each postorbital and epigastric ridge consist of 1 row of granules, each ridge separated from each other, epigastric ridge transverse, placed mesioposterior to postorbital ridge, postorbital ridge short, oblique, well separated from anterolateral margin; anterolateral margin long, without teeth or epibranchial incision, forming a finely granular, even finely serrated carina, joining orbital margin anteriorly, this carina posteriorly curving onto dorsal surface of carapace; posterolateral margin longer, straight, with transverse curved ridges formed by rows of granules; front flat but slightly concave medially, directed somewhat down, with very small frontal triangle (as enlarged proepistome); cervical groove not visible in median area; orbits large, orbital margin entire. Eyes strongly reduced, filling less than half the orbit, bullet-shaped, with only tiny pale brown pigment. Antenna very reduced. Mandibular palp three-articulated; last article simple, not bifurcated, oval. Third maxilipeds filling most of oral cavity; exopod without flagellum. Epistome with straight anterior margin; its concave posterior margin ending medially in triangular apex and, at either side of this triangle, two concavities separated by sharp point, one small (for reception of the palp of third maxilliped), the other rounded (forming anterior margin of efferent openings). Efferent openings very large, circular, reaching lateral margin of oral field. Chelipeds remarkably spiny, with numerous acute teeth and spines (some horn-tipped), and short simple setae or spine-like setae on all articles, may be in groups of two or three, some teeth bearing short setae at tips; fingers closing over full length and ending in sharp curved tips, and bearing short horny spines and simple setae. Pereiopods extremely long and thin, slender with short teeth and simple setae; dactyli with numerous longer simple setae on both sides. Vulvae on thoracic sternite 6, at margin with sternite 5, with wide sternal cover, and occupying two-thirds of width of sternite 6.
</paragraph>
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<subSubSection id="C30B2DE6B961FF94B182FCC5FC67FBC8" pageId="3" pageNumber="118" type="etymology">
<paragraph id="8BAE7E6DB961FF94B182FCC5FC67FBC8" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<emphasis id="B965A27FB961FF94B182FCC5FEF6FC84" bold="true" box="[199,334,835,861]" pageId="3" pageNumber="118">Etymology.</emphasis>
The genus is dedicated to the memory of 
<taxonomicName id="4C1105EEB961FF94B273FCC5FC63FC84" authority="Kee NG" authorityName="Kee NG" box="[822,987,835,861]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">Ngan Kee NG</taxonomicName>
(1966 
<emphasis id="B965A27FB961FF94B563FCC5FB8CFC84" bold="true" box="[1062,1076,835,861]" pageId="3" pageNumber="118">–</emphasis>
2022). She leaves a void in our scientific community, where she had a special place. She was more than a colleague, a very dear friend. She liked to stay in our Muséum, in Paris, and among her last works are those done here in winter over the last decade, notably the 
<emphasis id="B965A27FB961FF94B185FC29FB0BFC11" box="[192,1203,943,969]" italics="true" pageId="3" pageNumber="118">
Annotated type-catalogue of 
<taxonomicName id="4C1105EEB961FF94B344FC29FDC1FC10" box="[513,633,943,969]" class="Reptilia" family="Colubridae" genus="Brachyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="118" phylum="Chordata" rank="genus">Brachyura</taxonomicName>
of the Muséum national d’Histoire naturelle, Paris
</emphasis>
, the first one dealing with 
<taxonomicName id="4C1105EEB961FF94B197FC55FEDEFC34" authorityName="H. Milne Edwards" authorityYear="1837" box="[210,358,979,1005]" class="Malacostraca" family="Gecarcinidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Gecarcinidae</taxonomicName>
and 
<taxonomicName id="4C1105EEB961FF94B0E5FC55FCADFC34" authority="(N. K. Ng et al. 2019)" baseAuthorityName="N. K. Ng" baseAuthorityYear="2019" box="[416,789,979,1005]" class="Malacostraca" family="Grapsidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">
Grapsidae (N.K. 
<bibRefCitation id="EF80039CB961FF94B326FC55FCB5FC34" author="Ng, N. K. &amp; Rodriguez Moreno, P. A. &amp; Naruse, T. &amp; Guinot, D. &amp; Mollaret, N." box="[611,781,979,1005]" pageId="3" pageNumber="118" pagination="91 - 130" refId="ref15859" refString="Ng, N. K., Rodriguez Moreno, P. A., Naruse, T., Guinot, D. &amp; Mollaret, N. (2019). Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Museum national d'Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors. Zoosystema, 41 (7), 91 - 130. https: // doi. org / 10.5252 / zoosystema 2019 v 41 a 7" type="journal article" year="2019">
Ng 
<emphasis id="B965A27FB961FF94B3CBFC55FD79FC34" box="[654,705,979,1005]" italics="true" pageId="3" pageNumber="118">et al</emphasis>
. 2019
</bibRefCitation>
)
</taxonomicName>
, then 
<taxonomicName id="4C1105EEB961FF94B21EFC55FC64FC34" authorityName="Dana" authorityYear="1851" box="[859,988,979,1005]" class="Malacostraca" family="Plagusiidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Plagusiidae</taxonomicName>
and 
<taxonomicName id="4C1105EEB961FF94B552FC55FB30FC34" authorityName="Stevcic" authorityYear="2005" box="[1047,1160,979,1005]" class="Malacostraca" family="Percnidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Percnidae</taxonomicName>
, but the last one, on the 
<taxonomicName id="4C1105EEB961FF94B1D2FC71FEB4FBC8" authorityName="H. Milne Edwards" authorityYear="1853" box="[151,268,1015,1041]" class="Malacostraca" family="Varunidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Varunidae</taxonomicName>
, her favourite group, has not been completed. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="C30B2DE6B961FF94B182FB9DFD69F804" pageId="3" pageNumber="118" type="discussion">
<paragraph id="8BAE7E6DB961FF94B182FB9DFCBCFAA0" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<emphasis id="B965A27FB961FF94B182FB9DFE83FBEC" bold="true" box="[199,315,1051,1077]" pageId="3" pageNumber="118">Remarks.</emphasis>
The shape of the three-articulated mandibular palp ending by a single terminal lobe (
<figureCitation id="132A62E8B961FF94B5BAFB9DFAEBFBEC" box="[1279,1363,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1670,1694]" captionTargetBox="[151,1435,181,1646]" captionTargetId="figure-31@5.[151,1435,181,1646]" captionTargetPageId="5" captionText="FIGURE 3. Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710: A, frontal view; B, left cheliped; C, D, carapace and chelipeds; E, ventral view; F, ventral anterior view. Scale bars: A–E = 5 mm; F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731237" httpUri="https://zenodo.org/record/12731237/files/figure.png" pageId="3" pageNumber="118">Fig. 3F</figureCitation>
) (
<bibRefCitation id="EF80039CB961FF94B42FFB9DFF6BFB80" author="Bott, R." pageId="3" pageNumber="118" pagination="1 - 338" refId="ref13140" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammes-geschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." type="journal article" year="1970">Bott 1970</bibRefCitation>
: fig. 133; 
<bibRefCitation id="EF80039CB961FF94B000FBB9FE13FB80" author="Ng, P. K. L." box="[325,427,1087,1113]" pageId="3" pageNumber="118" refId="ref16006" refString="Ng, P. K. L. (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, viii + 156 pp., figs. 1 - 63, 4 colour pls." type="book" year="1988">Ng 1988</bibRefCitation>
: fig. 2A; 
<bibRefCitation id="EF80039CB961FF94B350FBB9FDC3FB80" author="Ng, P. K. L." box="[533,635,1087,1113]" pageId="3" pageNumber="118" pagination="311 - 336" refId="ref16224" refString="Ng, P. K. L. (2004) Crustacea: Decapoda, Brachyura. Freshwater Invertebrates of the Malaysian Region. In: Yule, C. M. &amp; Sen, Y. H. (Eds.), Freshwater Invertebrates of the Malaysian Region. Academy of Science Malaysia, Kuala Lumpur, pp. 311 - 336." type="book chapter" year="2004">Ng 2004</bibRefCitation>
: fig. 5I, J) indicates that 
<taxonomicName id="4C1105EEB961FF94B2D7FBB9FC77FB81" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[914,975,1087,1112]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B2D7FBB9FC77FB81" box="[914,975,1087,1112]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B293FBB9FBF8FB80" bold="true" box="[982,1088,1087,1113]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B293FBB9FBF8FB80" box="[982,1088,1087,1113]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
belongs to the 
<taxonomicName id="4C1105EEB961FF94B5A8FBB9FADFFB80" authorityName="Ortmann" authorityYear="1896" box="[1261,1383,1087,1113]" class="Malacostraca" family="Potamidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="family">Potamidae</taxonomicName>
. The closest genus is 
<taxonomicName id="4C1105EEB961FF94B009FBE5FDC8FBA4" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[332,624,1123,1149]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B009FBE5FE7AFBA4" box="[332,450,1123,1149]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
<bibRefCitation id="EF80039CB961FF94B08FFBE5FDC8FBA4" author="Holthuis, L. B." box="[458,624,1123,1149]" pageId="3" pageNumber="118" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
that consists of two species, both from the Gunung Mulu National Park, north 
<collectingCountry id="F3063EFDB961FF94B19DFB01FE83FB78" box="[216,315,1159,1185]" name="Malaysia" pageId="3" pageNumber="118">Sarawak</collectingCountry>
, the Malaysian part of Borneo and close to the southern border of 
<collectingCountry id="F3063EFDB961FF94B54EFB01FBE4FB78" box="[1035,1116,1159,1185]" name="Brunei" pageId="3" pageNumber="118">Brunei</collectingCountry>
: 
<taxonomicName id="4C1105EEB961FF94B521FB01FAD6FB78" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[1124,1390,1159,1185]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB961FF94B521FB01FB7CFB79" box="[1124,1220,1159,1185]" italics="true" pageId="3" pageNumber="118">C. caeca</emphasis>
<bibRefCitation id="EF80039CB961FF94B58FFB01FAD6FB78" author="Holthuis, L. B." box="[1226,1390,1159,1185]" pageId="3" pageNumber="118" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C1105EEB961FF94B1D2FB2DFE11FB1C" authority="Holthuis, 1979" authorityName="Holthuis" authorityYear="1979" box="[151,425,1195,1221]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB961FF94B1D2FB2DFF41FB1C" box="[151,249,1195,1221]" italics="true" pageId="3" pageNumber="118">C. tipula</emphasis>
<bibRefCitation id="EF80039CB961FF94B045FB2DFE11FB1C" author="Holthuis, L. B." box="[256,425,1195,1221]" pageId="3" pageNumber="118" pagination="1 - 47" refId="ref14725" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapods crustacean from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 1 - 47." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
</taxonomicName>
. The area explored by Lips’ team (
<bibRefCitation id="EF80039CB961FF94B26BFB2DFC58FB1C" author="Lips, B. &amp; Lips, J. &amp; Robert, X. &amp; Senecal, P. &amp; Placido, C. &amp; Geneau, A." box="[814,992,1195,1221]" pageId="3" pageNumber="118" refId="ref15541" refString="Lips, B., Lips, J., Robert, X., Senecal, P., Placido, C. &amp; Geneau, A. (2010) Borneo 2010 Kalimantan (Indonesie) 28 juillet au 30 aout 2010. Groupe Speleologique Vulcain. Federation Francaise de Speleologie, Lyon, 66 pp." type="book" year="2010">
Lips 
<emphasis id="B965A27FB961FF94B222FB2DFC20FB1C" box="[871,920,1195,1221]" italics="true" pageId="3" pageNumber="118">et al</emphasis>
. 2010
</bibRefCitation>
) in East Kalimantan, where the Lubang Gedung Cave (
<figureCitation id="132A62E8B961FF94B004FB49FE2EFB30" box="[321,406,1231,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1567,1591]" captionTargetBox="[201,1386,179,1541]" captionTargetId="figure-23@2.[201,1386,179,1541]" captionTargetPageId="2" captionText="FIGURE 1. A, Borneo Island and Kalimantan; B, Karst of Merabu, East Kalimantan, northernmost limestone massif of the Mangkalihat Peninsula; C, D, Lubang Gedung Cave, two views. Courtesy, J. and B. Lips." figureDoi="http://doi.org/10.5281/zenodo.12731231" httpUri="https://zenodo.org/record/12731231/files/figure.png" pageId="3" pageNumber="118">Fig. 1B</figureCitation>
) is located, is about 
<quantity id="4CE9D388B961FF94B3C5FB49FD6DFB30" box="[640,725,1231,1257]" metricMagnitude="5" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="118" unit="km" value="370.0">370 km</quantity>
southeast of the Gunung Mulu National Park. Like 
<taxonomicName id="4C1105EEB961FF94B45AFB49FA2DFB30" authorityName="Holthuis" authorityYear="1979" box="[1311,1429,1231,1257]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B45AFB49FA2DFB30" box="[1311,1429,1231,1257]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1105EEB961FF94B1D2FB75FF6CFAD5" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[151,212,1267,1292]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B1D2FB75FF6CFAD5" box="[151,212,1267,1292]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B19BFB75FEF2FAD4" bold="true" box="[222,330,1267,1293]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B19BFB75FEF2FAD4" box="[222,330,1267,1293]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
has the exopod of the third maxillipeds without a flagellum. Both species of 
<taxonomicName id="4C1105EEB961FF94B594FB75FAFFFAD4" authorityName="Holthuis" authorityYear="1979" box="[1233,1351,1267,1293]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B594FB75FAFFFAD4" box="[1233,1351,1267,1293]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
have a similar arrangement of the anterior part of the carapace, a similar front and probably a similar proepistome, a similar ornamentation of the carinate and non-incised anterolateral margin, and the posterolateral portion of carapace with several striae; the rounded vulvae are similar (
<figureCitation id="132A62E8B961FF94B3D8FAD9FD4FFAA0" box="[669,759,1375,1401]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BAE7E6DB961FF94B182FA05FBC6F988" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
Differences between 
<taxonomicName id="4C1105EEB961FF94B0FDFA05FE4DFA45" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[440,501,1411,1436]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B0FDFA05FE4DFA45" box="[440,501,1411,1436]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B0B8FA05FDD0FA44" bold="true" box="[509,616,1411,1437]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B0B8FA05FDD0FA44" box="[509,616,1411,1437]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C1105EEB961FF94B3E7FA05FCA0FA44" authorityName="Holthuis" authorityYear="1979" box="[674,792,1411,1437]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B3E7FA05FCA0FA44" box="[674,792,1411,1437]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
concern the shape of the carapace, subquadrate in 
<taxonomicName id="4C1105EEB961FF94B425FA05FA25FA45" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1376,1437,1411,1436]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B425FA05FA25FA45" box="[1376,1437,1411,1436]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2561F04B961FF94B1D2FA21FF44FA18" box="[151,252,1447,1473]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
(trapezoidal in 
<taxonomicName id="4C1105EEB961FF94B0ECFA21FDA7FA18" authorityName="Holthuis" authorityYear="1979" box="[425,543,1447,1473]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B0ECFA21FDA7FA18" box="[425,543,1447,1473]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
), the postorbital and epigastric ridges marked in 
<taxonomicName id="4C1105EEB961FF94B57EFA21FBC0FA19" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1083,1144,1447,1472]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B57EFA21FBC0FA19" box="[1083,1144,1447,1472]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B53BFA21FB5EFA18" bold="true" box="[1150,1254,1447,1473]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B53BFA21FB5EFA18" box="[1150,1254,1447,1473]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(inconspicuous, even absent or weak in 
<taxonomicName id="4C1105EEB961FF94B0E2FA4DFDA5FA3C" authorityName="Holthuis" authorityYear="1979" box="[423,541,1483,1509]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B0E2FA4DFDA5FA3C" box="[423,541,1483,1509]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
), the chelipeds with numerous spines and setae in 
<taxonomicName id="4C1105EEB961FF94B521FA4DFB19FA3D" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1124,1185,1483,1508]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B521FA4DFB19FA3D" box="[1124,1185,1483,1508]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B5ECFA4DFAACFA3C" bold="true" box="[1193,1300,1483,1509]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B5ECFA4DFAACFA3C" box="[1193,1300,1483,1509]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(practically absent in 
<taxonomicName id="4C1105EEB961FF94B046FA69FEC1F9D0" authorityName="Holthuis" authorityYear="1979" box="[259,377,1519,1545]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B046FA69FEC1F9D0" box="[259,377,1519,1545]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
). In 
<taxonomicName id="4C1105EEB961FF94B0F5FA69FD80F9D0" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[432,568,1519,1545]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="mayla" status="sp. nov.">
<emphasis id="B965A27FB961FF94B0F5FA69FD80F9D0" box="[432,568,1519,1545]" italics="true" pageId="3" pageNumber="118">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B37AFA69FD21F9D0" bold="true" box="[575,665,1519,1545]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B37AFA69FD21F9D0" box="[575,665,1519,1545]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the presence on the chelipeds of horny spines accompanied by setae and of simple setae without setules (spinelike setae), which are usually chemoreceptive (
<bibRefCitation id="EF80039CB961FF94B5D4F995FA97F9F4" author="Watling, L." box="[1169,1327,1555,1581]" pageId="3" pageNumber="118" refId="ref17629" refString="Watling, L. (1989) A classification system for crustacean setae based on the homology concept. In: Felgenhauer, B., Watling, L. &amp; Thistle, A. B. (Eds.), Functional morphology of feeding and grooming in Crustacea. Crustacean Issues. Vol. 6. A. A. Balkema, Rotterdam, pp. 15 - 26. https: // doi. org / 10.1201 / 9781003079354 - 2" type="book" year="1989">Watling 1989</bibRefCitation>
; 
<bibRefCitation id="EF80039CB961FF94B478F995FE8AF988" author="Garm, A. &amp; Watling, L." pageId="3" pageNumber="118" refId="ref13976" refString="Garm, A. &amp; Watling, L. (2013) The crustacean integument: setae, setules, and other ornamentation. In: Watling, L. &amp; Thiel, M. (Eds.), Functional morphology and diversity. The natural history of the Crustacea. Vol. 1. Oxford University Press, New York, New York, pp. 167 - 198. https: // doi. org / 10.1093 / acprof: osobl / 9780195398038.003.0006" type="book" year="2013">Garm &amp; Watling 2013</bibRefCitation>
), is probably the sign of a high degree of troglomorphism (see Discussion).
</paragraph>
<paragraph id="8BAE7E6DB961FF94B182F9DDFD69F804" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="118">
<taxonomicName id="4C1105EEB961FF94B182F9DDFE29F9AC" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[199,401,1626,1653]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B182F9DDFEA5F9AC" box="[199,285,1627,1653]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
<bibRefCitation id="EF80039CB961FF94B060F9DDFE29F9AC" author="Ng, P. K. L." box="[293,401,1626,1653]" pageId="3" pageNumber="118" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995</bibRefCitation>
</taxonomicName>
, an unmodified potamid genus from Borneo, not cavernicolous but probably semiterrestrial and known by two species, 
<taxonomicName id="4C1105EEB961FF94B096F9F9FD53F940" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[467,747,1662,1689]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="aethes">
<emphasis id="B965A27FB961FF94B096F9F9FDC1F940" box="[467,633,1663,1689]" italics="true" pageId="3" pageNumber="118">Ibanum aethes</emphasis>
<bibRefCitation id="EF80039CB961FF94B3C4F9F9FD53F940" author="Ng, P. K. L." box="[641,747,1662,1689]" pageId="3" pageNumber="118" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF80039CB961FF94B3BEF9F9FCDBF940" author="Ng, P. K. L." box="[763,867,1662,1689]" pageId="3" pageNumber="118" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng 1995</bibRefCitation>
: figs. 4 
<emphasis id="B965A27FB961FF94B2F2F9F9FC7DF940" bold="true" box="[951,965,1663,1689]" pageId="3" pageNumber="118">–</emphasis>
6), the 
<typeStatus id="54AAC0CFB961FF94B553F9F9FBFEF940" box="[1046,1094,1663,1689]" pageId="3" pageNumber="118">type</typeStatus>
species from Lanjak-Entimau Wildlife Sanctuary, 
<collectingCountry id="F3063EFDB961FF94B03CF925FE64F964" box="[377,476,1699,1725]" name="Malaysia" pageId="3" pageNumber="118">Sarawak</collectingCountry>
, and 
<taxonomicName id="4C1105EEB961FF94B35FF925FCC8F964" authority="(De Man, 1899)" baseAuthorityName="De Man" baseAuthorityYear="1899" box="[538,880,1699,1725]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="bicristatum">
<emphasis id="B965A27FB961FF94B35FF925FD0AF964" box="[538,690,1699,1725]" italics="true" pageId="3" pageNumber="118">I. bicristatum</emphasis>
(De Man, 1899)
</taxonomicName>
(
<bibRefCitation id="EF80039CB961FF94B2C5F925FC51F964" author="Ng, P. K. L." box="[896,1001,1698,1725]" pageId="3" pageNumber="118" pagination="57 - 72" refId="ref16169" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng 1995</bibRefCitation>
: figs. 1 
<emphasis id="B965A27FB961FF94B579F925FBF2F964" bold="true" box="[1084,1098,1699,1725]" pageId="3" pageNumber="118">–</emphasis>
3), from Kapuas, Kalimantan, is close phylogenetically to 
<taxonomicName id="4C1105EEB961FF94B095F941FDB5F939" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[464,525,1735,1760]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B095F941FDB5F939" box="[464,525,1735,1760]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B351F941FDC5F938" bold="true" box="[532,637,1735,1761]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B351F941FDC5F938" box="[532,637,1735,1761]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C1105EEB961FF94B3F6F941FC91F938" authorityName="Holthuis" authorityYear="1979" box="[691,809,1735,1761]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B3F6F941FC91F938" box="[691,809,1735,1761]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
(P.K.L. Ng, pers. comm.). The three genera share many similarities: carapace widest anteriorly; antero-lateral margin short, arcuate, carinate, granulated to gently serrated, without teeth or epibranchial incision, with the carina curving onto the carapace dorsal surface; front flat and rather straight; and posterolateral margins long, with transverse ridges. Such as 
<taxonomicName id="4C1105EEB961FF94B289F8B5FBB1F895" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[972,1033,1843,1868]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B289F8B5FBB1F895" box="[972,1033,1843,1868]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B555F8B5FBC1F894" bold="true" box="[1040,1145,1843,1869]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B555F8B5FBC1F894" box="[1040,1145,1843,1869]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C1105EEB961FF94B5CDF8B5FB66F894" authorityName="Ng" authorityYear="1995" box="[1160,1246,1843,1869]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B5CDF8B5FB66F894" box="[1160,1246,1843,1869]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
</taxonomicName>
has well defined postorbital cristae. 
<taxonomicName id="4C1105EEB961FF94B02AF8D1FE7DF8A8" authorityName="Ng" authorityYear="1995" box="[367,453,1879,1905]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B02AF8D1FE7DF8A8" box="[367,453,1879,1905]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
</taxonomicName>
distinguishes from both 
<taxonomicName id="4C1105EEB961FF94B39AF8D1FCA4F8A9" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[735,796,1879,1904]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B965A27FB961FF94B39AF8D1FCA4F8A9" box="[735,796,1879,1904]" italics="true" pageId="3" pageNumber="118">Ngan</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B261F8D1FC36F8A8" bold="true" box="[804,910,1879,1905]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B261F8D1FC36F8A8" box="[804,910,1879,1905]" pageId="3" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C1105EEB961FF94B282F8D1FB85F8A8" authorityName="Holthuis" authorityYear="1979" box="[967,1085,1879,1905]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B282F8D1FB85F8A8" box="[967,1085,1879,1905]" italics="true" pageId="3" pageNumber="118">Cerberusa</emphasis>
</taxonomicName>
by the presence of a flagellum on the exopod of the third maxilliped (P.K.L. Ng, pers. comm.). The vulvae of 
<taxonomicName id="4C1105EEB961FF94B54AF8FDFBDDF84C" authorityName="Ng" authorityYear="1995" box="[1039,1125,1915,1941]" class="Malacostraca" family="Potamidae" genus="Ibanum" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B965A27FB961FF94B54AF8FDFBDDF84C" box="[1039,1125,1915,1941]" italics="true" pageId="3" pageNumber="118">Ibanum</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B531F8FDFB74F84C" box="[1140,1228,1915,1941]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6D</figureCitation>
) are smaller, oval, distinctly obliquely positioned on sternite 6, instead of rounded and central in 
<taxonomicName id="4C1105EEB961FF94B569F819FB0FF860" authorityName="Guinot &amp; Rodríguez Moreno" authorityYear="2024" box="[1068,1207,1951,1977]" class="Malacostraca" family="Potamidae" genus="Ngan" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="mayla" status="sp. nov.">
<emphasis id="B965A27FB961FF94B569F819FB0FF860" box="[1068,1207,1951,1977]" italics="true" pageId="3" pageNumber="118">Ngan mayla</emphasis>
</taxonomicName>
<emphasis id="B965A27FB961FF94B587F819FAA7F860" bold="true" box="[1218,1311,1951,1977]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="A2561F04B961FF94B587F819FAA7F860" box="[1218,1311,1951,1977]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132A62E8B961FF94B477F819FA37F860" box="[1330,1423,1951,1977]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6A</figureCitation>
), 
<taxonomicName id="4C1105EEB961FF94B1D2F845FEEDF805" authorityName="Holthuis" authorityYear="1979" box="[151,341,1987,2013]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B965A27FB961FF94B1D2F845FEEDF805" box="[151,341,1987,2013]" italics="true" pageId="3" pageNumber="118">Cerberusa caeca</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B020F845FE03F804" box="[357,443,1987,2013]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6B</figureCitation>
) and 
<taxonomicName id="4C1105EEB961FF94B0BFF845FDE3F804" authorityName="Holthuis" authorityYear="1979" box="[506,603,1987,2013]" class="Malacostraca" family="Potamidae" genus="Cerberusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="tipula">
<emphasis id="B965A27FB961FF94B0BFF845FDB5F804" box="[506,525,1987,2013]" italics="true" pageId="3" pageNumber="118">C</emphasis>
. 
<emphasis id="B965A27FB961FF94B35FF845FDE3F804" box="[538,603,1987,2013]" italics="true" pageId="3" pageNumber="118">tipula</emphasis>
</taxonomicName>
(
<figureCitation id="132A62E8B961FF94B32EF845FD7CF804" box="[619,708,1987,2013]" captionStart="FIGURE 6" captionStartId="8.[151,250,1684,1708]" captionTargetBox="[164,1423,885,1661]" captionTargetId="figure-401@8.[164,1423,885,1661]" captionTargetPageId="8" captionText="FIGURE 6. Vulvae: A, Ngan mayla gen. et sp. nov., holotype female, 12.2 × 15.3 mm, MZB Cru 5710; B, Cerberusa caeca Holthuis, 1979, female, 9.9 × 13.3 mm, ZRC 2013.1220; C, C. tipula Holthuis, 1979, female, 13.0 × 17.6 mm, ZRC 2013.1219; D, Ibanum aethes Ng, 1995, female, 10.8 × 14.7 mm, ZRC 2019.0141. B–D, courtesy P.K.L. Ng." figureDoi="http://doi.org/10.5281/zenodo.12731252" httpUri="https://zenodo.org/record/12731252/files/figure.png" pageId="3" pageNumber="118">Fig. 6C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>