278 lines
30 KiB
XML
278 lines
30 KiB
XML
<document id="14370810B70222FA57DBC09A2E8399B0" ID-CLB-Dataset="36820" ID-DOI="10.11646/zootaxa.3829.1.1" ID-GBIF-Dataset="a4b69269-5a4c-4b5b-9f89-6b314d3ba284" ID-ISSN="1175-5326" ID-Zenodo-Dep="286556" ID-ZooBank="75210CDC-AC6A-4624-A6F1-1BC969BC7CAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452709399" checkinUser="plazi" docAuthor="Pyron, Robert Alexander & Wallach, Van" docDate="2014" docId="03B587DDC10EB179CFD7CCAEFCE3FBFA" docLanguage="en" docName="zt03829p081.pdf" docOrigin="Zootaxa 3829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Argyrophis Gray 1845" docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="FF8CFFA5C13AB14CCF40CA0EFFB9FFA2" masterDocTitle="Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence" masterLastPageNumber="81" masterPageNumber="1" pageNumber="53" updateTime="1698338434491" updateUser="ExternalLinkService">
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<mods:mods id="51ED723B04881A92A0C6DCD161E48CF5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E2BE2E98B7D3D8E491CA625C72BC3950">
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<mods:title id="4C84A73644FA5828D3295C5F5E8466B8">Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence</mods:title>
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<mods:name id="93A295FC15EDC1A87B9C84F61C7E0968" type="personal">
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<mods:roleTerm id="3F36E21C09FA9EAFB4AAE04165627947">Author</mods:roleTerm>
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<mods:namePart id="DC5556908277E229ACCC294255C9414D">Pyron, Robert Alexander</mods:namePart>
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</mods:name>
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<mods:name id="BC4DBB9A09C7C4170D5D18E782BF4BD5" type="personal">
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<mods:role id="8366314726384260EF18C7AD75D0CD94">
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<mods:roleTerm id="46B2C9C30D4999A573B1C554936F206F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BBA4F093CA8EB4034AC1ABC678DBB23E">Wallach, Van</mods:namePart>
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<mods:typeOfResource id="E7B767EF0AE9DAE13C969B3E2DDE1871">text</mods:typeOfResource>
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<mods:titleInfo id="4EA4DC9046EB51FFE608BE24495694EB">
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<mods:title id="7E12A2B58BEF6C181EC974CADDF01945">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B5AF09883CD554B65F95485C6FB7C493">
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<mods:date id="02DBF4C02F074D796E08BFB798EEF0D2">2014</mods:date>
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<mods:detail id="486D73A1AB247F3380A0F0C8AEFFEC2B" type="volume">
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<mods:number id="CFE5C6DBD4DE4944B452F3A562DC3599">3829</mods:number>
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</mods:detail>
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<mods:detail id="13D84BA121456438AE422B3BE8308E3C" type="issue">
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<mods:number id="4DC49B1B1557DA09A00D98666AB170AB">1</mods:number>
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<mods:classification id="3354241C590981DE621F9059873B49C1">journal article</mods:classification>
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<mods:identifier id="1384F4FD2A7200940792B99472C0E427" type="CLB-Dataset">36820</mods:identifier>
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<mods:identifier id="852E771DDC4AAB2E44350D4371E0BDDF" type="DOI">10.11646/zootaxa.3829.1.1</mods:identifier>
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<mods:identifier id="2893FE6DEB7DF90A09C1332496FFED81" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B587DDC10EB179CFD7CCAEFCE3FBFA" ID-DOI="http://doi.org/10.5281/zenodo.6127990" ID-GBIF-Taxon="127668675" ID-Zenodo-Dep="6127990" LSID="urn:lsid:plazi:treatment:03B587DDC10EB179CFD7CCAEFCE3FBFA" httpUri="http://treatment.plazi.org/id/03B587DDC10EB179CFD7CCAEFCE3FBFA" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
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<subSubSection id="C3066540C10EB178CFD7CCAEFE16F918" box="[151,431,1696,1722]" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="8BA336CBC10EB178CFD7CCAEFE16F918" blockId="52.[151,431,1696,1722]" box="[151,431,1696,1722]" pageId="52" pageNumber="53">
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<heading id="D0EB81A7C10EB178CFD7CCAEFE16F918" bold="true" box="[151,431,1696,1722]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
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<taxonomicName id="4C1C4D48C10EB178CFD7CCAEFE16F918" ID-CoL="32RJ" ID-ENA="1539852" authority="Gray, 1845" authorityName="Gray" authorityYear="1845" box="[151,431,1696,1722]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10EB178CFD7CCAEFE16F918" bold="true" box="[151,431,1696,1722]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CFD7CCAEFEA7F918" bold="true" box="[151,286,1696,1722]" italics="true" pageId="52" pageNumber="53">Argyrophis</emphasis>
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<bibRefCitation id="EF8D4B3AC10EB178CE65CCAEFE16F918" author="Gray" box="[293,431,1696,1722]" pageId="52" pageNumber="53" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. British Museum (Natural History), London, xxvii + 322 pp. http: // dx. doi. org / 10.5962 / bhl. title. 5499" type="book" year="1845">Gray, 1845</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10EB178CFD7CCE8FB05F8E4" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="8BA336CBC10EB178CFD7CCE8FD16F95D" blockId="52.[151,1437,1766,2007]" box="[151,687,1766,1791]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CFD7CCE8FE8BF95D" bold="true" box="[151,306,1766,1791]" pageId="52" pageNumber="53">
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<typeStatus id="54A78869C10EB178CFD7CCE8FF6BF95D" box="[151,210,1766,1791]" pageId="52" pageNumber="53">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C1C4D48C10EB178CE79CCE8FD16F95D" authority="Schlegel, 1839" authorityName="Schlegel" authorityYear="1839" box="[313,687,1766,1791]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="muelleri">
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<emphasis id="B968EAD9C10EB178CE79CCE8FDB8F95D" box="[313,513,1766,1791]" italics="true" pageId="52" pageNumber="53">Typhlops muelleri</emphasis>
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Schlegel, 1839
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA336CBC10EB178CF87CD04FB05F8E4" blockId="52.[151,1437,1766,2007]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CF87CD04FE3FF881" bold="true" box="[199,390,1802,1827]" pageId="52" pageNumber="53">Species content.</emphasis>
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<emphasis id="B968EAD9C10EB178CECFCD04FB05F8E4" italics="true" pageId="52" pageNumber="53">
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<taxonomicName id="4C1C4D48C10EB178CECFCD04FD69F881" box="[399,720,1802,1827]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="bothriorhynchus">Argyrophis bothriorhynchus</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CD9DCD05FCEAF881" box="[733,851,1802,1827]" class="Reptilia" family="Gerrhopilidae" genus="Asiatyphlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="diardii">Ar. diardii</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CC20CD05FC6BF881" box="[864,978,1803,1827]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="fuscus">Ar. fuscus</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CC9FCD05FB2EF881" box="[991,1175,1802,1827]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="giadinhensis">Ar. giadinhensis</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CBE5CD05FAC9F881" box="[1189,1392,1802,1827]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="hypsobothrius">Ar. hypsobothrius</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CA3CCD05FEBBF8E5" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="klemmeri">Ar. klemmeri</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CE4DCD21FE07F8E5" box="[269,446,1838,1863]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="koshunensis">Ar. koshunensis</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CE89CD21FDE9F8E5" box="[457,592,1838,1863]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="muelleri">Ar. muelleri</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CD1BCD21FD76F8E4" box="[603,719,1839,1862]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="oatesii">Ar. oatesii</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CD9BCD21FCD5F8E4" box="[731,876,1839,1862]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="roxaneae">Ar. roxaneae</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CC37CD21FBB7F8E4" box="[887,1038,1839,1862]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="siamensis">Ar. siamensis</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CB59CD21FB01F8E4" box="[1049,1208,1839,1862]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="trangensis">Ar. trangensis</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10EB179CF87CD5CFEABFDBA" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="diagnosis">
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<paragraph id="8BA336CBC10EB179CF87CD5CFE31FE0F" blockId="52.[151,1437,1766,2007]" lastBlockId="53.[151,1437,151,1112]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CF87CD5CFEF8F8C9" bold="true" box="[199,321,1874,1899]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
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<taxonomicName id="4C1C4D48C10EB178CE0BCD5CFE7EF8C9" box="[331,455,1874,1899]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10EB178CE0BCD5CFE7EF8C9" box="[331,455,1874,1899]" italics="true" pageId="52" pageNumber="53">Argyrophis</emphasis>
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</taxonomicName>
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can be distinguished from all other typhlopoids by the combination of the following characters: T-II, T-III, or T-V
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<collectingCountry id="F30B765BC10EB178CEA4CD79FDB4F82D" box="[484,525,1911,1935]" name="Cyprus" pageId="52" pageNumber="53">SIP</collectingCountry>
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and scale row reduction, and left lung present in
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<emphasis id="B968EAD9C10EB178CB79CD79FA87F82D" box="[1081,1342,1910,1935]" italics="true" pageId="52" pageNumber="53">
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<taxonomicName id="4C1C4D48C10EB178CB79CD79FB17F82D" box="[1081,1198,1910,1935]" class="Reptilia" family="Gerrhopilidae" genus="Asiatyphlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="diardii">Ar. diardii</taxonomicName>
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,
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<taxonomicName id="4C1C4D48C10EB178CBF9CD79FA87F82D" box="[1209,1342,1910,1935]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="muelleri">Ar. muelleri</taxonomicName>
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</emphasis>
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, and
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<taxonomicName id="4C1C4D48C10EB178CA3CCD79FEBDF811" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="siamensis">
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<emphasis id="B968EAD9C10EB178CA3CCD79FEBDF811" italics="true" pageId="52" pageNumber="53">Ar. siamensis</emphasis>
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</taxonomicName>
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. Small- to large-sized (total length
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<quantity id="4CE49B2EC10EB178CDE9CD94FC81F811" box="[681,824,1946,1971]" metricMagnitude="-1" metricUnit="m" metricValue="3.075" metricValueMax="5.4" metricValueMin="0.75" pageId="52" pageNumber="53" unit="mm" value="307.5" valueMax="540.0" valueMin="75.0">75–540 mm</quantity>
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), stout- to slender-bodied (length/width ratio 24–71) snakes with 20–30 scale rows (with reduction), 246–402 total middorsals, short to moderate tail (1.0–3.6% total length) with 5–26 subcaudals (length/width ratio 0.4–2.0), and apical spine small or thorn-like. Dorsal and lateral head profiles rounded, moderate rostral (0.25–0.40 head width), preocular in contact with second and third supralabials, eye moderate with distinct pupil, and postoculars 2–3. Lateral tongue papillae present; vestigial left lung present, tracheal, cardiac and right lungs multicameral (with 23–38 + 3–10 + 2–12 foramina, respectively); testes unsegmented; hemipenis eversible, lacking retrocloacal sacs, and moderate rectal caecum (1.2–5.0% SVL). Coloration brown, reddish-brown, purplish-black or black dorsally, transiting to a lighter venter (beige, cream or white) or bicolored (with a sharp midlateral demarcation between dark dorsum and light venter), snout, labials, and chin sometimes light.
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</paragraph>
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<paragraph id="8BA336CBC10FB179CF87CBB9FEABFDBA" blockId="53.[151,1437,151,1112]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CF87CBB9FE5FFE72" bold="true" box="[199,486,439,464]" pageId="53" pageNumber="54">Phylogenetic definition.</emphasis>
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Includes the MRCA of
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<taxonomicName id="4C1C4D48C10FB179CC4ECBB9FC58FE72" box="[782,993,439,464]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="diardii">
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<emphasis id="B968EAD9C10FB179CC4ECBB9FC58FE72" box="[782,993,439,464]" italics="true" pageId="53" pageNumber="54">Argyrophis diardii</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1C4D48C10FB179CB62CBB6FB12FE72" box="[1058,1195,439,464]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="muelleri">
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<emphasis id="B968EAD9C10FB179CB62CBB6FB12FE72" box="[1058,1195,439,464]" italics="true" pageId="53" pageNumber="54">Ar. muelleri</emphasis>
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</taxonomicName>
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and all descendants thereof, all species more closely related to
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<taxonomicName id="4C1C4D48C10FB179CDC4CBD2FCB2FE57" box="[644,779,476,501]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="muelleri">
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<emphasis id="B968EAD9C10FB179CDC4CBD2FCB2FE57" box="[644,779,476,501]" italics="true" pageId="53" pageNumber="54">Ar. muelleri</emphasis>
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</taxonomicName>
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than to the
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<typeStatus id="54A78869C10FB179CCD6CBD3FC7FFE57" box="[918,966,477,501]" pageId="53" pageNumber="54">type</typeStatus>
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species of the 15 other typhlopid genera listed here.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10FB179CF87C82AFD73FDC2" pageId="53" pageNumber="54" type="etymology">
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<paragraph id="8BA336CBC10FB179CF87C82AFD73FDC2" blockId="53.[151,1437,151,1112]" pageId="53" pageNumber="54">
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<emphasis id="B968EAD9C10FB179CF87C82AFEF7FD9F" bold="true" box="[199,334,548,573]" pageId="53" pageNumber="54">Etymology.</emphasis>
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Unclear. Many names erected by J.E. Gray were apparently chosen for euphony but lack any substantial etymology (A.M. Bauer, pers. comm.).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10FB179CF87C862FB00FD27" box="[199,1209,620,645]" pageId="53" pageNumber="54" type="distribution">
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||
<paragraph id="8BA336CBC10FB179CF87C862FB00FD27" blockId="53.[151,1437,151,1112]" box="[199,1209,620,645]" pageId="53" pageNumber="54">
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||
<emphasis id="B968EAD9C10FB179CF87C862FED9FD27" bold="true" box="[199,352,620,645]" pageId="53" pageNumber="54">Distribution.</emphasis>
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Occurs in Southeast Asia, primarily
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<collectingCountry id="F30B765BC10FB179CDBEC862FCD9FD27" box="[766,864,620,645]" name="Vietnam" pageId="53" pageNumber="54">Vietnam</collectingCountry>
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||
,
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<collectingCountry id="F30B765BC10FB179CC2BC862FC68FD27" box="[875,977,620,645]" name="Thailand" pageId="53" pageNumber="54">Thailand</collectingCountry>
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, and West
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<collectingCountry id="F30B765BC10FB179CB0BC862FB0CFD27" box="[1099,1205,620,645]" name="Malaysia" pageId="53" pageNumber="54">Malaysia</collectingCountry>
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.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066540C10FB179CF87C881FCE3FBFA" pageId="53" pageNumber="54" type="discussion">
|
||
<paragraph id="8BA336CBC10FB179CF87C881FCE3FBFA" blockId="53.[151,1437,151,1112]" pageId="53" pageNumber="54">
|
||
<emphasis id="B968EAD9C10FB179CF87C881FE82FD0A" bold="true" box="[199,315,655,680]" pageId="53" pageNumber="54">Remarks.</emphasis>
|
||
The species
|
||
<taxonomicName id="4C1C4D48C10FB179CE99C881FD1FFD0A" box="[473,678,655,680]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="fuscus">
|
||
<emphasis id="B968EAD9C10FB179CE99C881FD1FFD0A" box="[473,678,655,680]" italics="true" pageId="53" pageNumber="54">Argyrophis fuscus</emphasis>
|
||
</taxonomicName>
|
||
was considered
|
||
<taxonomicName id="4C1C4D48C10FB179CC2DC881FC40FD0A" box="[877,1017,655,680]" class="Reptilia" family="Typhlopidae" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="family">Typhlopidae</taxonomicName>
|
||
<emphasis id="B968EAD9C10FB179CB44C89FFB1AFD0A" box="[1028,1187,655,680]" italics="true" pageId="53" pageNumber="54">incertae sedis</emphasis>
|
||
by
|
||
<bibRefCitation id="EF8D4B3AC10FB179CB95C881FAD1FD0A" author="Hahn" box="[1237,1384,655,680]" pageId="53" pageNumber="54" refString="Hahn, D. E. (1980) Liste der rezenten Amphibien und Reptilien: Anomalepididae, Leptotyphlopidae, Typhlopidae. Das Tierreich, Berlin, 101, i - xii + 1 - 93. http: // dx. doi. org / 10.1515 / 9783111385556. fm" type="journal article" year="1980">Hahn (1980)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF8D4B3AC10FB179CFD7C8BAFE0DFD6E" box="[151,436,692,717]" pageId="53" pageNumber="54" refString="McDiarmid, R. W., Campbell, J. A. & Toure, T. S. A. (1999) Snake Species of the World: a Taxonomic and Geographic Reference (Volume 1). Herpetologists' League, Washington, DC, 512 pp." type="book">
|
||
McDiarmid
|
||
<emphasis id="B968EAD9C10FB179CE63C8BBFEE7FD6F" box="[291,350,692,717]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
|
||
(1999)
|
||
</bibRefCitation>
|
||
, who suspected the
|
||
<typeStatus id="54A78869C10FB179CDDEC8BBFD77FD6F" box="[670,718,693,717]" pageId="53" pageNumber="54">type</typeStatus>
|
||
locality of Java to be in error. The
|
||
<typeStatus id="54A78869C10FB179CB2EC8BBFB27FD6F" box="[1134,1182,693,717]" pageId="53" pageNumber="54">type</typeStatus>
|
||
(MNHN 1062) of
|
||
<taxonomicName id="4C1C4D48C10FB179CA3CC8BAFF64FD52" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="fuscus">
|
||
<emphasis id="B968EAD9C10FB179CA3CC8BAFF64FD52" italics="true" pageId="53" pageNumber="54">Ar. fuscus</emphasis>
|
||
</taxonomicName>
|
||
has been examined and reveals a relationship to the
|
||
<taxonomicName id="4C1C4D48C10FB179CC10C8D6FC7CFD52" box="[848,965,727,752]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="diardii">
|
||
<emphasis id="B968EAD9C10FB179CC10C8D6FC7CFD52" box="[848,965,727,752]" italics="true" pageId="53" pageNumber="54">Ar. diardii</emphasis>
|
||
</taxonomicName>
|
||
group (Table 2), so the locality may be correct. The species
|
||
<taxonomicName id="4C1C4D48C10FB179CECEC8F2FDE0FCB7" box="[398,601,764,789]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="hypsobothrius">
|
||
<emphasis id="B968EAD9C10FB179CECEC8F2FDE0FCB7" box="[398,601,764,789]" italics="true" pageId="53" pageNumber="54">Ar. hypsobothrius</emphasis>
|
||
</taxonomicName>
|
||
is known only from a briefly described and now lost ZMH
|
||
<typeStatus id="54A78869C10FB179CA7AC8F2FA25FCB7" box="[1338,1436,764,789]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
|
||
destroyed in
|
||
<date id="FFA2100BC10FB179CE6DC911FE24FC9A" box="[301,413,799,824]" pageId="53" pageNumber="54" value="1943-07">July 1943</date>
|
||
(during WW II;
|
||
<emphasis id="B968EAD9C10FB179CD25C911FD36FC9A" box="[613,655,799,824]" italics="true" pageId="53" pageNumber="54">fide</emphasis>
|
||
<bibRefCitation id="EF8D4B3AC10FB179CDD8C911FCD8FC9A" author="Hallermann" box="[664,865,799,824]" pageId="53" pageNumber="54" refString="Hallermann, J. (1998) Annotated catalogue of the type specimens of the herpetological collection in the Zoological Museum of the University of Hamburg. Mitteilungen aus dem Hamburgisches Zoologisches Museum und Institut, 95, 197 - 223." type="journal article" year="1998">Hallermann 1998</bibRefCitation>
|
||
), and
|
||
<taxonomicName id="4C1C4D48C10FB179CCEAC92EFBE2FC9A" box="[938,1115,799,824]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="koshunensis">
|
||
<emphasis id="B968EAD9C10FB179CCEAC92EFBE2FC9A" box="[938,1115,799,824]" italics="true" pageId="53" pageNumber="54">Ar. koshunensis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F30B765BC10FB179CBE4C911FB4FFC9A" box="[1188,1270,799,824]" name="Taiwan" pageId="53" pageNumber="54">Taiwan</collectingCountry>
|
||
is also poorly known. However, these species are all clearly allied with
|
||
<taxonomicName id="4C1C4D48C10FB179CC69C94AFC1CFCFF" box="[809,933,836,861]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10FB179CC69C94AFC1CFCFF" box="[809,933,836,861]" italics="true" pageId="53" pageNumber="54">Argyrophis</emphasis>
|
||
</taxonomicName>
|
||
on the basis of morphology (Table 2). Note that this genus is not included in our molecular phylogenetic analysis, but was analyzed by
|
||
<bibRefCitation id="EF8D4B3AC10FB179CBECC969FA2EFC22" box="[1196,1431,871,896]" pageId="53" pageNumber="54" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. & Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article">
|
||
Hedges
|
||
<emphasis id="B968EAD9C10FB179CA48C967FAFBFC22" box="[1288,1346,871,896]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
, who found weak support for a sister-group relationship between
|
||
<taxonomicName id="4C1C4D48C10FB179CC31C982FC54FC07" box="[881,1005,908,933]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10FB179CC31C982FC54FC07" box="[881,1005,908,933]" italics="true" pageId="53" pageNumber="54">Argyrophis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B968EAD9C10FB179CB65C982FB78FC07" box="[1061,1217,908,933]" italics="true" pageId="53" pageNumber="54">
|
||
<taxonomicName id="4C1C4D48C10FB179CB65C982FB04FC07" box="[1061,1213,908,933]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">Xerotyphlops</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Those authors also included
|
||
<taxonomicName id="4C1C4D48C10FB179CE40C9A1FDA3FC6A" box="[256,538,943,968]" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="leucomelas">
|
||
<emphasis id="B968EAD9C10FB179CE40C9A1FDA3FC6A" box="[256,538,943,968]" italics="true" pageId="53" pageNumber="54">Indotyphlops leucomelas</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D48C10FB179CD15C9BFFD5AFC6A" box="[597,739,943,968]" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="tenuicollis">
|
||
<emphasis id="B968EAD9C10FB179CD15C9BFFD5AFC6A" box="[597,739,943,968]" italics="true" pageId="53" pageNumber="54">I. tenuicollis</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<emphasis id="B968EAD9C10FB179CC4CC9A1FC24FC6A" box="[780,925,943,968]" italics="true" pageId="53" pageNumber="54">Asiatyphlops</emphasis>
|
||
(=
|
||
<taxonomicName id="4C1C4D48C10FB179CCFEC9A1FB83FC6A" box="[958,1082,943,968]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10FB179CCFEC9A1FB83FC6A" box="[958,1082,943,968]" italics="true" pageId="53" pageNumber="54">Argyrophis</emphasis>
|
||
</taxonomicName>
|
||
), but those species seem to be allied with
|
||
<taxonomicName id="4C1C4D48C10FB179CE58C9DAFE13FC4F" box="[280,426,980,1005]" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10FB179CE58C9DAFE13FC4F" box="[280,426,980,1005]" italics="true" pageId="53" pageNumber="54">Indotyphlops</emphasis>
|
||
</taxonomicName>
|
||
on the basis of a T-III
|
||
<collectingCountry id="F30B765BC10FB179CDF4C9DAFD65FC4E" box="[692,732,980,1004]" name="Cyprus" pageId="53" pageNumber="54">SIP</collectingCountry>
|
||
, caudal spine reduced to nubbin, and invisible eye. They also included
|
||
<taxonomicName id="4C1C4D48C10FB179CE40C9F6FE71FBB2" box="[256,456,1015,1040]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="hypsobothrius">
|
||
<emphasis id="B968EAD9C10FB179CE40C9F6FE71FBB2" box="[256,456,1015,1040]" italics="true" pageId="53" pageNumber="54">Ar. hypsobothrius</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C1C4D48C10FB179CEAFC9F9FD38FBB2" box="[495,641,1015,1040]" class="Reptilia" family="Typhlopidae" genus="Indotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10FB179CEAFC9F9FD38FBB2" box="[495,641,1015,1040]" italics="true" pageId="53" pageNumber="54">Indotyphlops</emphasis>
|
||
</taxonomicName>
|
||
, though it is likely allied with
|
||
<taxonomicName id="4C1C4D48C10FB179CCA6C9F9FBDBFBB2" box="[998,1122,1015,1040]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10FB179CCA6C9F9FBDBFBB2" box="[998,1122,1015,1040]" italics="true" pageId="53" pageNumber="54">Argyrophis</emphasis>
|
||
</taxonomicName>
|
||
on the basis of head shield pits or grooves as in
|
||
<emphasis id="B968EAD9C10FB179CEDFCE12FD34FB97" box="[415,653,1052,1077]" italics="true" pageId="53" pageNumber="54">
|
||
<taxonomicName id="4C1C4D48C10FB179CEDFCE12FD31FB97" box="[415,648,1052,1077]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="bothriorhynchus">Ar. bothriorhynchus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="F30B765BC10FB179CDD9CE12FD01FB96" box="[665,696,1052,1076]" name="American Samoa" pageId="53" pageNumber="54">As</collectingCountry>
|
||
noted above,
|
||
<taxonomicName id="4C1C4D48C10FB179CC29CE12FC5CFB97" box="[873,997,1052,1077]" class="Squamata" family="Typhlopidae" genus="Argyrophis" higherTaxonomySource="CoL" kingdom="Animalia" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10FB179CC29CE12FC5CFB97" box="[873,997,1052,1077]" italics="true" pageId="53" pageNumber="54">Argyrophis</emphasis>
|
||
</taxonomicName>
|
||
is the objective senior synonym for
|
||
<emphasis id="B968EAD9C10FB179CFD7CE31FE91FBFA" box="[151,296,1087,1112]" italics="true" pageId="53" pageNumber="54">Asiatyphlops</emphasis>
|
||
, as
|
||
<taxonomicName id="4C1C4D48C10FB179CE14CE4EFE61FBFA" box="[340,472,1087,1112]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="muelleri">
|
||
<emphasis id="B968EAD9C10FB179CE14CE4EFE61FBFA" box="[340,472,1087,1112]" italics="true" pageId="53" pageNumber="54">Ar. muelleri</emphasis>
|
||
</taxonomicName>
|
||
is the
|
||
<typeStatus id="54A78869C10FB179CD62CE4EFDEBFBFA" box="[546,594,1088,1112]" pageId="53" pageNumber="54">type</typeStatus>
|
||
species of both genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |