217 lines
25 KiB
XML
217 lines
25 KiB
XML
<document id="80470543B0D363DD3687FAC593EC11D9" ID-CLB-Dataset="36820" ID-DOI="10.11646/zootaxa.3829.1.1" ID-GBIF-Dataset="a4b69269-5a4c-4b5b-9f89-6b314d3ba284" ID-ISSN="1175-5326" ID-Zenodo-Dep="286556" ID-ZooBank="75210CDC-AC6A-4624-A6F1-1BC969BC7CAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452709399" checkinUser="plazi" docAuthor="Pyron, Robert Alexander & Wallach, Van" docDate="2014" docId="03B587DDC10CB17ACFD7CA96FB9AF919" docLanguage="en" docName="zt03829p081.pdf" docOrigin="Zootaxa 3829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lemuriatyphlops Pyron & Wallach, 2014, gen. nov." docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="FF8CFFA5C13AB14CCF40CA0EFFB9FFA2" masterDocTitle="Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence" masterLastPageNumber="81" masterPageNumber="1" pageNumber="55" updateTime="1698338434491" updateUser="ExternalLinkService">
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<mods:mods id="2FDB837EEE2469D80C59FC403F614A45" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A301D1A6A19B11D810C509395708E567">
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<mods:title id="CE9B506B203DFB395CE42B5532D2401D">Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence</mods:title>
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<mods:name id="BA6C1E65F7999D268AFEE2B7B1D08656" type="personal">
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<mods:namePart id="740BA4E889E0D94E83901B6A894FE98D">Pyron, Robert Alexander</mods:namePart>
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<mods:name id="ABBE3E5A1ED9670DE9568254959D21EA" type="personal">
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<mods:roleTerm id="1E1B498D2EA6EC7DFE6A7E50066513B2">Author</mods:roleTerm>
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<mods:namePart id="68C8E13B71F4A3883E70F0F8A705FB0C">Wallach, Van</mods:namePart>
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<mods:title id="8FFC7DE0864A360F214CA13F5330B811">Zootaxa</mods:title>
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<mods:part id="F5222BA79017F56296F18E7C67AADC15">
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<mods:date id="D45FA9ADBED2960C59A04811604AC8B4">2014</mods:date>
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<mods:number id="AA9179A95ED84878D20CBB035DF72A67">3829</mods:number>
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<mods:number id="5BC238B6EFD2569332B07B32A33CDF32">1</mods:number>
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<mods:identifier id="8DDBF2F71A35E994A8B8D9F3FB2093FE" type="DOI">10.11646/zootaxa.3829.1.1</mods:identifier>
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<mods:identifier id="711551FD6051A31878D0EB16A50C8A41" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="257AB236B3BA9CDB558D63C690F09080" type="Zenodo-Dep">286556</mods:identifier>
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<treatment id="03B587DDC10CB17ACFD7CA96FB9AF919" ID-DOI="http://doi.org/10.5281/zenodo.6127994" ID-GBIF-Taxon="127668693" ID-Zenodo-Dep="6127994" LSID="urn:lsid:plazi:treatment:03B587DDC10CB17ACFD7CA96FB9AF919" httpUri="http://treatment.plazi.org/id/03B587DDC10CB17ACFD7CA96FB9AF919" lastPageNumber="55" pageId="54" pageNumber="55">
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<subSubSection id="C3066540C10CB17ACFD7CA96FE60FF10" box="[151,473,152,178]" pageId="54" pageNumber="55" type="nomenclature">
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<paragraph id="8BA336CBC10CB17ACFD7CA96FE60FF10" blockId="54.[151,473,152,178]" box="[151,473,152,178]" pageId="54" pageNumber="55">
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<heading id="D0EB81A7C10CB17ACFD7CA96FE60FF10" bold="true" box="[151,473,152,178]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
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<emphasis id="B968EAD9C10CB17ACFD7CA96FE60FF10" bold="true" box="[151,473,152,178]" pageId="54" pageNumber="55">
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<taxonomicName id="4C1C4D48C10CB17ACFD7CA96FEDDFF10" box="[151,356,152,178]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B968EAD9C10CB17ACFD7CA96FEDDFF10" bold="true" box="[151,356,152,178]" italics="true" pageId="54" pageNumber="55">Lemuriatyphlops</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57A2C10CB17ACE2BCA96FE60FF10" box="[363,473,152,178]" pageId="54" pageNumber="55" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10CB17ACFD7CAD0FAFAFEB9" pageId="54" pageNumber="55" type="materials_examined">
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<paragraph id="8BA336CBC10CB17ACFD7CAD0FD4BFF55" blockId="54.[151,1437,222,1723]" box="[151,754,222,247]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACFD7CAD0FE8BFF55" bold="true" box="[151,306,222,247]" pageId="54" pageNumber="55">
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<typeStatus id="54A78869C10CB17ACFD7CAD0FF6BFF55" box="[151,210,222,247]" pageId="54" pageNumber="55">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C1C4D48C10CB17ACE79CAD0FD4BFF55" authority="Werner, 1909" authorityName="Werner" authorityYear="1909" box="[313,754,222,247]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="microcephalus">
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<emphasis id="B968EAD9C10CB17ACE79CAD0FDF1FF55" box="[313,584,222,247]" italics="true" pageId="54" pageNumber="55">Typhlops microcephalus</emphasis>
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<bibRefCitation id="EF8D4B3AC10CB17ACD0FCAD1FD4BFF55" author="Werner" box="[591,754,223,247]" pageId="54" pageNumber="55" refString="Werner, F. (1909 a) Beschreibung neuer Reptilien aus dem Kgl. Naturalienkabinett in Stuttgart. Jahreshefte des Vereins fur vaterlandische Naturkunde Wurttemberg, 65, 55 - 63." type="journal article" year="1909" yearSuffix="a">Werner, 1909a</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA336CBC10CB17ACF87CB0CFAFAFEB9" blockId="54.[151,1437,222,1723]" box="[199,1347,258,283]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACF87CB0CFE3CFEB9" bold="true" box="[199,389,258,283]" pageId="54" pageNumber="55">Species content.</emphasis>
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<emphasis id="B968EAD9C10CB17ACECCCB0CFBC2FEB9" box="[396,1147,258,283]" italics="true" pageId="54" pageNumber="55">
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<taxonomicName id="4C1C4D48C10CB17ACECCCB0CFD47FEB9" authority="Lem." authorityName="Lem." box="[396,766,258,283]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="albanalis">Lemuriatyphlops albanalis, Lem.</taxonomicName>
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<taxonomicName id="4C1C4D48C10CB17ACC45CB0CFC7DFEB8" authority="Lem." authorityName="Lem." box="[773,964,258,283]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="domerguei">
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domerguei,
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<emphasis id="B968EAD9C10CB17ACCC9CB0DFC7DFEB8" bold="true" box="[905,964,259,282]" italics="true" pageId="54" pageNumber="55">Lem.</emphasis>
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</taxonomicName>
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<taxonomicName id="4C1C4D48C10CB17ACC8BCB0CFBCDFEB9" box="[971,1140,258,283]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="microcephalus">microcephalus</taxonomicName>
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,
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</emphasis>
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and
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<emphasis id="B968EAD9C10CB17ACBF2CB0AFAFAFEB9" box="[1202,1347,260,283]" italics="true" pageId="54" pageNumber="55">Lem. reuteri.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10CB17ACF87CB28FC83FD4D" pageId="54" pageNumber="55" type="diagnosis">
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<paragraph id="8BA336CBC10CB17ACF87CB28FBE4FD05" blockId="54.[151,1437,222,1723]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACF87CB28FEF8FE9D" bold="true" box="[199,321,294,319]" pageId="54" pageNumber="55">Diagnosis.</emphasis>
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<taxonomicName id="4C1C4D48C10CB17ACE0ACB28FDB1FE9D" box="[330,520,294,319]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10CB17ACE0ACB28FDB1FE9D" box="[330,520,294,319]" italics="true" pageId="54" pageNumber="55">Lemuriatyphlops</emphasis>
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</taxonomicName>
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can be distinguished from all other typhlopoids by lack of scale row reduction, and by the combination of the following characters: small-to moderate-sized (total length
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<quantity id="4CE49B2EC10CB17ACBDCCB44FA90FEC1" box="[1180,1321,330,355]" metricMagnitude="-1" metricUnit="m" metricValue="1.765" metricValueMax="2.76" metricValueMin="0.77" pageId="54" pageNumber="55" unit="mm" value="176.5" valueMax="276.0" valueMin="77.0">77–276 mm</quantity>
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), stout- to slender-bodied (length/width ratio 34–94) snakes with 20–22 scale rows (without reduction), 252–520 total middorsals, short to moderate tail (1.6–4.0% total length) with 6–16 subcaudals (length/width ratio 0.9–2.2), and apical spine small or nubbin. Dorsal and lateral head profiles rounded, narrow to moderate rostral (0.26–0.61 head width), inferior nasal suture in contact with second supralabial, preocular in contact with second and third supralabials, eye small with distinct pupil or eyespot, T-III or T-V
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<collectingCountry id="F30B765BC10CB17ACC3ACBF1FC1BFDB5" box="[890,930,511,535]" name="Cyprus" pageId="54" pageNumber="55">SIP</collectingCountry>
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, and postoculars 1–3. Lateral tongue papillae present; left lung absent; tracheal lung multicameral (with 11–32 chambers), cardiac lung multicameral (with 2–4 chambers) or unicameral, and right lung multicameral (with 2–4 chambers); testes unsegmented; hemipenis eversible, lacking retrocloacal sacs; and rectal caecum small (0.8–2.5% SVL). Coloration light brown to black, venter lighter; and ventral snout, chin, cloacal region, and subcaudals yellow or white.
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</paragraph>
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<paragraph id="8BA336CBC10CB17ACF87C8BCFC83FD4D" blockId="54.[151,1437,222,1723]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACF87C8BCFE5BFD69" bold="true" box="[199,482,690,715]" pageId="54" pageNumber="55">Phylogenetic definition.</emphasis>
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Includes all species more closely related to
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<taxonomicName id="4C1C4D48C10CB17ACC9BC8BCFAFCFD69" box="[987,1349,690,715]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="microcephalus">
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<emphasis id="B968EAD9C10CB17ACC9BC8BCFAFCFD69" box="[987,1349,690,715]" italics="true" pageId="54" pageNumber="55">Lemuriatyphlops microcephalus</emphasis>
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</taxonomicName>
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than to the
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<typeStatus id="54A78869C10CB17ACF80C8D9FF49FD4D" box="[192,240,727,751]" pageId="54" pageNumber="55">type</typeStatus>
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species of the 15 other typhlopid genera listed here.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10CB17ACF87C8F4FDACFC95" pageId="54" pageNumber="55" type="etymology">
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<paragraph id="8BA336CBC10CB17ACF87C8F4FDACFC95" blockId="54.[151,1437,222,1723]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACF87C8F4FEF7FCB1" bold="true" box="[199,334,762,787]" pageId="54" pageNumber="55">Etymology.</emphasis>
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Name is a masculine noun, and refers to the mythical lost continent of Lemuria, which connected
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<collectingCountry id="F30B765BC10CB17ACFD7C910FEA7FC95" box="[151,286,798,823]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
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to
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<collectingCountry id="F30B765BC10CB17ACE02C910FEC2FC95" box="[322,379,798,823]" name="India" pageId="54" pageNumber="55">India</collectingCountry>
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and Oceania.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10CB17ACF87C94CFD5BFCDD" pageId="54" pageNumber="55" type="distribution">
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<paragraph id="8BA336CBC10CB17ACF87C94CFD5BFCDD" blockId="54.[151,1437,222,1723]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACF87C94CFED9FCF9" bold="true" box="[199,352,834,859]" pageId="54" pageNumber="55">Distribution.</emphasis>
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Available data suggest this genus is endemic to
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<collectingCountry id="F30B765BC10CB17ACCD6C94CFB98FCF9" box="[918,1057,834,859]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
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;
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<taxonomicName id="4C1C4D48C10CB17ACB6FC94CFAE7FCF9" box="[1071,1374,834,859]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="albanalis">
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<emphasis id="B968EAD9C10CB17ACB6FC94CFAE7FCF9" box="[1071,1374,834,859]" italics="true" pageId="54" pageNumber="55">Lemuriatyphlops albanalis</emphasis>
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</taxonomicName>
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is of uncertain origin, though
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<collectingCountry id="F30B765BC10CB17ACEECC968FD8AFCDD" box="[428,563,870,895]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
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is a possibility.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10CB17ACF87C984FB9AF919" pageId="54" pageNumber="55" type="discussion">
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<paragraph id="8BA336CBC10CB17ACF87C984FB9AF919" blockId="54.[151,1437,222,1723]" pageId="54" pageNumber="55">
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<emphasis id="B968EAD9C10CB17ACF87C984FE82FC01" bold="true" box="[199,315,906,931]" pageId="54" pageNumber="55">Remarks.</emphasis>
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Note that we also include
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<taxonomicName id="4C1C4D48C10CB17ACD37C984FCF4FC01" box="[631,845,906,931]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="albanalis">
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<emphasis id="B968EAD9C10CB17ACD37C984FCF4FC01" box="[631,845,906,931]" italics="true" pageId="54" pageNumber="55">Typhlops albanalis</emphasis>
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</taxonomicName>
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in
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<taxonomicName id="4C1C4D48C10CB17ACC37C984FB8CFC01" box="[887,1077,906,931]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10CB17ACC37C984FB8CFC01" box="[887,1077,906,931]" italics="true" pageId="54" pageNumber="55">Lemuriatyphlops</emphasis>
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</taxonomicName>
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here, based on 20 dorsal scale rows lacking reduction (
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<bibRefCitation id="EF8D4B3AC10CB17ACEF4C9A0FDE1FC65" author="Rendahl" box="[436,600,942,967]" pageId="54" pageNumber="55" refString="Rendahl, H. (1918) Zwei neue Arten der Gattung Typhlops aus dem Kaplande. Arkiv for Zoologi, Stockholm, 11, 1 - 6." type="journal article" year="1918">Rendahl 1918</bibRefCitation>
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). The species was synonymized with
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<taxonomicName id="4C1C4D48C10CB17ACB5BC9A0FA9AFC65" box="[1051,1315,942,967]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="ocularis">
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<emphasis id="B968EAD9C10CB17ACB5BC9A0FA9AFC65" box="[1051,1315,942,967]" italics="true" pageId="54" pageNumber="55">Madatyphlops ocularis</emphasis>
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</taxonomicName>
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by Roux-
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<bibRefCitation id="EF8D4B3AC10CB17ACFD7C9DDFE8CFC49" author="Esteve" box="[151,309,979,1003]" pageId="54" pageNumber="55" refString="Roux-Esteve, R. (1974) Revision systematique des Typhlopidae d'Afrique, Reptilia-Serpentes. Memoires du Museum National d'Histoire Naturelle, Paris, Serie 2, 87 A, 1 - 313." type="journal article" year="1974">Estève (1974)</bibRefCitation>
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, but examination of the
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<typeStatus id="54A78869C10CB17ACD04C9DDFDC6FC49" box="[580,639,979,1003]" pageId="54" pageNumber="55">types</typeStatus>
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(NHR 3351 and 2574) by V.W. found it to be distinct from that species based upon numerous external and internal characters (Tables 2, 3), including the following (
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<taxonomicName id="4C1C4D48C10CB17ACBE6C9F9FA95FBAD" box="[1190,1324,1014,1039]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="albanalis">
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<emphasis id="B968EAD9C10CB17ACBE6C9F9FA95FBAD" box="[1190,1324,1014,1039]" italics="true" pageId="54" pageNumber="55">L. albanalis</emphasis>
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</taxonomicName>
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data first, followed by
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||
<taxonomicName id="4C1C4D48C10CB17ACE6CCE12FE09FB91" box="[300,432,1050,1075]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="ocularis">
|
||
<emphasis id="B968EAD9C10CB17ACE6CCE12FE09FB91" box="[300,432,1050,1075]" italics="true" pageId="54" pageNumber="55">M. ocularis</emphasis>
|
||
</taxonomicName>
|
||
data): dorsal head profile (tapered vs. ogival), lateral head profile (blunt vs. pointed), rostral shape (sagittate vs. parallel), anterior rostral (convex vs. concave with anterodorsal pocket behind dorsally projecting point), superior nasal suture (vertical, parallel to rostral vs. oblique), pupil of eye (invisible vs. visible), occipital shields (absent vs. present), color pattern (bicolored vs. uniform), length/width ratio (80–92 vs. 55–67), cardiac and right lungs (unicameral vs. multicameral), and significant differences between the lengths and midpoints of the liver, gonads, adrenals, kidneys and lungs plus most of the gaps and intervals between all the viscera. Therefore we ressurect the former species here under the combination of
|
||
<taxonomicName id="4C1C4D48C10CB17ACB01CEFCFAD6FAA9" box="[1089,1391,1266,1291]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="albanalis">
|
||
<emphasis id="B968EAD9C10CB17ACB01CEFCFAD6FAA9" box="[1089,1391,1266,1291]" italics="true" pageId="54" pageNumber="55">Lemuriatyphlops albanalis</emphasis>
|
||
</taxonomicName>
|
||
. Its distribution is unknown (purported to be from
|
||
<collectingCountry id="F30B765BC10CB17ACDE3CF18FC8AFA8D" box="[675,819,1302,1327]" name="South Africa" pageId="54" pageNumber="55">South Africa</collectingCountry>
|
||
or
|
||
<collectingCountry id="F30B765BC10CB17ACC19CF18FC5FFA8D" box="[857,998,1302,1327]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
|
||
). This species was originally described from “Kapland” (=
|
||
<collectingCountry id="F30B765BC10CB17ACE2BCF34FE46FAF1" box="[363,511,1338,1363]" name="South Africa" pageId="54" pageNumber="55">South Africa</collectingCountry>
|
||
) but was supposedly corrected by
|
||
<bibRefCitation id="EF8D4B3AC10CB17ACCC4CF34FBEFFAF1" author="FitzSimons" box="[900,1110,1338,1363]" pageId="54" pageNumber="55" refString="FitzSimons, V. F. M. (1962) Snakes of southern Africa. MacDonald, London, 423 pp." type="book" year="1962">FitzSimons (1962)</bibRefCitation>
|
||
to
|
||
<collectingCountry id="F30B765BC10CB17ACB3ACF34FAB8FAF1" box="[1146,1281,1338,1363]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
|
||
based upon a label by W. Kaudern from
|
||
<date id="FFA2100BC10CB17ACE85CF50FDC4FAD5" box="[453,637,1374,1399]" pageId="54" pageNumber="55" value="1906-11">November 1906</date>
|
||
. However, this species was not mentioned by either
|
||
<bibRefCitation id="EF8D4B3AC10CB17ACB8DCF50FA2EFAD5" author="Andersson" box="[1229,1431,1374,1399]" pageId="54" pageNumber="55" refString="Andersson, L. G. (1911) Reptiles and batrachians from the north-west of Madagascar collected by W. Kaudern, 1906 - 1907. Arkiv for Zoologi, 7, 1 - 15." type="journal article" year="1911">Andersson (1911)</bibRefCitation>
|
||
, who reported on Kaudern’s
|
||
<collectingCountry id="F30B765BC10CB17ACE90CF8CFD84FA39" box="[464,573,1410,1435]" name="Madagascar" pageId="54" pageNumber="55">Malagasy</collectingCountry>
|
||
snake collection of
|
||
<date id="FFA2100BC10CB17ACC60CF8CFC19FA39" box="[800,928,1410,1435]" pageId="54" pageNumber="55" value="1906" valueMax="1907">1906–1907</date>
|
||
, nor by
|
||
<bibRefCitation id="EF8D4B3AC10CB17ACCBCCF8CFB09FA39" author="Kaudern" box="[1020,1200,1410,1435]" pageId="54" pageNumber="55" refString="Kaudern, W. (1922) Sauropsiden aus Madagascar. Zoologischer Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 45, 395 - 458." type="journal article" year="1922">Kaudern (1922)</bibRefCitation>
|
||
, who reported on his own collections of
|
||
<date id="FFA2100BC10CB17ACEC3CFA8FDB8FA1D" box="[387,513,1446,1471]" pageId="54" pageNumber="55" value="1906" valueMax="1907">1906–1907</date>
|
||
and
|
||
<date id="FFA2100BC10CB17ACD07CFA9FD71FA1D" box="[583,712,1447,1471]" pageId="54" pageNumber="55" value="1911" valueMax="1912">1911–1912</date>
|
||
. Similar to other
|
||
<emphasis id="B968EAD9C10CB17ACCE7CFA8FBD2FA1D" box="[935,1131,1446,1471]" italics="true" pageId="54" pageNumber="55">
|
||
<taxonomicName id="4C1C4D48C10CB17ACCE7CFA8FBD1FA1D" box="[935,1128,1446,1471]" class="Reptilia" family="Typhlopidae" genus="Lemuriatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">Lemuriatyphlops</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
this species is small- to moderate-sized (total length
|
||
<quantity id="4CE49B2EC10CB17ACEA7CFC4FD3DFA41" box="[487,644,1482,1507]" metricMagnitude="-1" metricUnit="m" metricValue="2.295" metricValueMax="2.76" metricValueMin="1.83" pageId="54" pageNumber="55" unit="mm" value="229.5" valueMax="276.0" valueMin="183.0">183–276 mm</quantity>
|
||
) and slender-bodied (length/width ratio 78–94) with 20 scale rows throughout, 499–520 total middorsals, short to moderate tail (1.8–2.2% total length) with 15–16 subcaudals (length/width ratio 2.0–2.2), broad oval rostral (0.54–0.61 head width), inferior nasal in contact with second supralabial, T-III
|
||
<collectingCountry id="F30B765BC10CB17ACE1CCC39FE3DF9ED" box="[348,388,1591,1615]" name="Cyprus" pageId="54" pageNumber="55">SIP</collectingCountry>
|
||
, postoculars two, tracheal lung multicameral, cardiac and right lungs unicameral, and a large rectal caecum. Note that these species were all included in
|
||
<taxonomicName id="4C1C4D48C10CB17ACC66CC54FC7FF9D1" box="[806,966,1626,1651]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10CB17ACC66CC54FC7FF9D1" box="[806,966,1626,1651]" italics="true" pageId="54" pageNumber="55">Madatyphlops</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8D4B3AC10CB17ACCAFCC54FB6AF9D1" box="[1007,1235,1626,1651]" pageId="54" pageNumber="55" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. & Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article">
|
||
Hedges
|
||
<emphasis id="B968EAD9C10CB17ACB0ACC52FB3BF9D1" box="[1098,1154,1626,1651]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
without inclusion in their phylogenies, but are clearly morphologically distinct (Tables 2, 3), and strongly supported outside of
|
||
<taxonomicName id="4C1C4D48C10CB17ACFD7CCACFE8EF919" box="[151,311,1698,1723]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10CB17ACFD7CCACFE8EF919" box="[151,311,1698,1723]" italics="true" pageId="54" pageNumber="55">Madatyphlops</emphasis>
|
||
</taxonomicName>
|
||
in the phylogeny, in a separate Palearctic and Asian clade (
|
||
<figureCitation id="13272A4EC10CB17ACC91CCACFBABF919" box="[977,1042,1698,1723]" captionStart="FIGURE 1" captionStartId="39.[151,249,1923,1945]" captionTargetBox="[212,1348,195,1894]" captionTargetId="figure@39.[208,1364,193,1894]" captionTargetPageId="39" captionText="FIGURE 1. Results from a molecular phylogenetic analysis of 95 of the 275 known, extant species of typhlopoid blindsnakes. Tree represents the ML estimate from a concatenated matrix of 4 mitochondrial and 6 nuclear genes (6290 bp total), inferred using 200 independent searches in RAxMLv 7.2.8, with support estimated from 1000 non-parametric BS replicates (> 50 % shown)." httpUri="https://zenodo.org/record/286557/files/figure.png" pageId="54" pageNumber="55">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |