treatments-xml/data/03/B5/87/03B587C01C4EA511FF680EC6FD75AFA1.xml

<document id="E823B91723CEF8B4E15F454A0413E5B2" ID-CLB-Dataset="20239" ID-DOI="10.11646/zootaxa.5115.2.4" ID-GBIF-Dataset="65327e15-2216-4e6c-9681-d92f5bf3aff5" ID-ISSN="1175-5326" ID-Zenodo-Dep="6352402" ID-ZooBank="85A2F1F0-BC45-4ED0-AC0A-41757CCCB41B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647259108896" checkinUser="plazi" docAuthor="Lee, Jonathan J. &amp; Baumann, Richard W." docDate="2022" docId="03B587C01C4EA511FF680EC6FD75AFA1" docLanguage="en" docName="zootaxa.5115.2.4.pdf" docOrigin="Zootaxa 5115 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dracoleuctra Lee &amp; Baumann 2022, new genus" docType="treatment" docVersion="7" lastPageNumber="265" masterDocId="FF8CFFB81C4EA516FFFF095EFF84AB48" masterDocTitle="Dracoleuctra, a new wingless stonefly genus from southern Oregon, U. S. A. (Plecoptera: Leuctridae)" masterLastPageNumber="266" masterPageNumber="258" pageNumber="258" updateTime="1699318007820" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DFB192C07C8EC0EE5FA0CA9AB092A356">Dracoleuctra, a new wingless stonefly genus from southern Oregon, U. S. A. (Plecoptera: Leuctridae)</mods:title>
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<mods:namePart id="D7743AEC09BDEF8E1F7F3BFA0F660F79">Lee, Jonathan J.</mods:namePart>
<mods:affiliation id="53B9C3A4AAB4B2294E5D62E6A0E589DE">2337 15 th Street, Eureka, CA, U. S. A. 95501. jonleebugs @ gmail. com</mods:affiliation>
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<mods:namePart id="A33DF553362708AF1D2EC7FD35A8A8CF">Baumann, Richard W.</mods:namePart>
<mods:affiliation id="CE3714F172A4C54B05CF21D89BC5904E">Department of Biology and Monte L. Bean Life Science Museum, Brigham Young University, Provo, UT, U. S. A. 84602. richard _ baumann @ byu. edu</mods:affiliation>
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<treatment id="03B587C01C4EA511FF680EC6FD75AFA1" ID-DOI="http://doi.org/10.5281/zenodo.6354789" ID-GBIF-Taxon="194020412" ID-Zenodo-Dep="6354789" LSID="urn:lsid:plazi:treatment:03B587C01C4EA511FF680EC6FD75AFA1" httpUri="http://treatment.plazi.org/id/03B587C01C4EA511FF680EC6FD75AFA1" lastPageId="7" lastPageNumber="265" pageId="0" pageNumber="258">
<subSubSection id="C306655D1C4EA516FF680EC6FE43ACFB" box="[151,455,1944,1971]" pageId="0" pageNumber="258" type="nomenclature">
<paragraph id="8BA336D61C4EA516FF680EC6FE43ACFB" blockId="0.[151,455,1944,1971]" box="[151,455,1944,1971]" pageId="0" pageNumber="258">
<heading id="D0EB81BA1C4EA516FF680EC6FE43ACFB" bold="true" box="[151,455,1944,1971]" fontSize="11" level="1" pageId="0" pageNumber="258" reason="1">
<emphasis id="B968EAC41C4EA516FF680EC6FE43ACFB" bold="true" box="[151,455,1944,1971]" pageId="0" pageNumber="258">
<taxonomicName id="4C1C4D551C4EA516FF680EC6FEB3ACFB" ID-CoL="9HZZF" authority="Lee &amp; Baumann, 2022" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[151,311,1944,1971]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="258" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B968EAC41C4EA516FF680EC6FEB3ACFB" bold="true" box="[151,311,1944,1971]" italics="true" pageId="0" pageNumber="258">Dracoleuctra</emphasis>
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, 
<taxonomicNameLabel id="A25B57BF1C4EA516FEB90EC7FE43ACFB" box="[326,455,1945,1971]" pageId="0" pageNumber="258" rank="genus">new genus</taxonomicNameLabel>
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<paragraph id="8BA336D61C4EA516FF680E80FCA8ACBE" blockId="0.[151,812,2013,2038]" box="[151,812,2013,2038]" pageId="0" pageNumber="258">
<typeStatus id="54A788741C4EA516FF680E80FF4DACBE" box="[151,201,2014,2038]" pageId="0" pageNumber="258">Type</typeStatus>
species:. 
<taxonomicName id="4C1C4D551C4EA516FED20E83FD89ACBD" authority="Lee &amp; Baumann, 2022" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[301,525,2013,2037]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="258" phylum="Arthropoda" rank="species" species="siskiyou" status="sp. nov.">
<emphasis id="B968EAC41C4EA516FED20E83FD89ACBD" box="[301,525,2013,2037]" italics="true" pageId="0" pageNumber="258">Dracoleuctra siskiyou</emphasis>
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, 
<taxonomicNameLabel id="A25B57BF1C4EA516FDE50E80FD12ACBD" box="[538,662,2013,2037]" pageId="0" pageNumber="258" rank="species">new species</taxonomicNameLabel>
, by monotypy.
</paragraph>
<paragraph id="8BA336D61C4FA517FF6809C9FADDAB9D" blockId="1.[151,1437,151,861]" pageId="1" pageNumber="259">
<emphasis id="B968EAC41C4FA517FF6809C9FEAAABF9" bold="true" box="[151,302,151,177]" pageId="1" pageNumber="259">Morphology.</emphasis>
<emphasis id="B968EAC41C4FA517FECB09C9FEFEABF9" box="[308,378,151,177]" italics="true" pageId="1" pageNumber="259">Adults</emphasis>
. Apterous (
<figureCitation id="13272A531C4FA517FE0809C9FDB2ABF9" box="[503,566,151,177]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[151,1437,885,1827]" captionTargetId="figure-309@1.[151,1437,885,1827]" captionTargetPageId="1" captionText="FIGURES 1–4. Dracoleuctra siskiyou: 1. Thorax, dorsal. 2. Thorax, ventral. 3. Female abdomen, dorsal. 4. Female terminalia, ventral. Bs = probasisternum; Fs = profurcasternum; MBs = mesobasisternum; MeBs = Metabasisternum; MePrs = Metapresternum; MFs = mesofurcasternum; Mg = marginal groove of pronotum; MPfs = Mesopostfurcasternum; MPrs = Mesopresternum; MSs1 = Mesospinasternum 1; MSs2 = Mesospinasternum 2; Pfs = propostfurcasternum; Pn = central disc of pronotum; Psc = mesoprescutum; Pscl = mesopostscutellum; Prs = propresternum; Sct = mesoscutum." figureDoi="http://doi.org/10.5281/zenodo.6352404" httpUri="https://zenodo.org/record/6352404/files/figure.png" pageId="1" pageNumber="259">Fig. 1</figureCitation>
), head brown, lacking ocelli, antennae elongate, with at least 34 segments, body bearing numerous dark and light clothing hairs, sclerotized parts brown to dark brown, membranous areas pale.
</paragraph>
<paragraph id="8BA336D61C4FA517FF380981FE22A815" blockId="1.[151,1437,151,861]" pageId="1" pageNumber="259">
Prothoracic tergum with large central pronotum wider than long, central disc brown with dark brown middorsal line, dark brown rugosities lateral to middorsal line (
<figureCitation id="13272A531C4FA517FD06085DFCB8AA55" box="[761,828,259,285]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[151,1437,885,1827]" captionTargetId="figure-309@1.[151,1437,885,1827]" captionTargetPageId="1" captionText="FIGURES 1–4. Dracoleuctra siskiyou: 1. Thorax, dorsal. 2. Thorax, ventral. 3. Female abdomen, dorsal. 4. Female terminalia, ventral. Bs = probasisternum; Fs = profurcasternum; MBs = mesobasisternum; MeBs = Metabasisternum; MePrs = Metapresternum; MFs = mesofurcasternum; Mg = marginal groove of pronotum; MPfs = Mesopostfurcasternum; MPrs = Mesopresternum; MSs1 = Mesospinasternum 1; MSs2 = Mesospinasternum 2; Pfs = propostfurcasternum; Pn = central disc of pronotum; Psc = mesoprescutum; Pscl = mesopostscutellum; Prs = propresternum; Sct = mesoscutum." figureDoi="http://doi.org/10.5281/zenodo.6352404" httpUri="https://zenodo.org/record/6352404/files/figure.png" pageId="1" pageNumber="259">Fig. 1</figureCitation>
), light brown marginal groove (
<bibRefCitation id="EF8D4B271C4FA517FB4E085DFADBAA55" author="Claassen, P. W." box="[1201,1375,259,285]" pageId="1" pageNumber="259" pagination="1 - 199" refId="ref3717" refString="Claassen, P. W. (1931) Plecoptera nymphs of America (north of Mexico). Entomological Society of America, Thomas Say Foundation, 3, 1 - 199." type="journal article" year="1931">Claassen, 1931</bibRefCitation>
) surrounding central disc; anterior and posterior margins with three shallow longitudinal furrows not reaching central disc, demarking four anterior and four posterior rectangular areas, larger in posterior row. Mesonotal anterior prescutum narrow, transverse, anteriorly curved, projecting slightly under pronotum, separated from scutum by only a shallow groove; scutum widest anteriorly projecting laterally forward of anepisternum, narrowing then widening slightly posteriorly, rugosities lateral to middorsal line; scutellum not apparent; postscutellum widest anteriorly, gradually narrowing posteriorly, middorsal suture line reaching scutum, demarked from scutum by transverse light brown line, suture not apparent. Metanotum similar to mesonotum but smaller and not as narrow posteriorly. Prothoracic basisternum and presternum fused, spade-shaped (
<figureCitation id="13272A531C4FA517FCDC0B7DFCE2A975" box="[803,870,547,573]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[151,1437,885,1827]" captionTargetId="figure-309@1.[151,1437,885,1827]" captionTargetPageId="1" captionText="FIGURES 1–4. Dracoleuctra siskiyou: 1. Thorax, dorsal. 2. Thorax, ventral. 3. Female abdomen, dorsal. 4. Female terminalia, ventral. Bs = probasisternum; Fs = profurcasternum; MBs = mesobasisternum; MeBs = Metabasisternum; MePrs = Metapresternum; MFs = mesofurcasternum; Mg = marginal groove of pronotum; MPfs = Mesopostfurcasternum; MPrs = Mesopresternum; MSs1 = Mesospinasternum 1; MSs2 = Mesospinasternum 2; Pfs = propostfurcasternum; Pn = central disc of pronotum; Psc = mesoprescutum; Pscl = mesopostscutellum; Prs = propresternum; Sct = mesoscutum." figureDoi="http://doi.org/10.5281/zenodo.6352404" httpUri="https://zenodo.org/record/6352404/files/figure.png" pageId="1" pageNumber="259">Fig. 2</figureCitation>
), furcasternum narrow mesally, expanded laterally, somewhat bowtie-shaped, separate from basisternum; postfurcasternum large, transversely oval, midventral line membranous. Mesosternal spinasternum 1 with dark oval mesal depression, lateral sclerotized bands shorter than prothoracic furcasternal width; presternum heart-shaped, basisternum large, ray-shaped, “wings” extending laterally past the lateral margin of coxae, bearing numerous clothing hairs, with posteromesal depression where furcasternum is embedded, suture lines not apparent; furcasternum round-oval, furcasternal pits short, arms not apparent; spinasternum 2 dark, reaching metathoracic presternum, postfurcasterna oval, separate from spinasternum 2. Metasternal presternum transversely oval, basisternum similar in shape to mesobasisternum but truncate posteriorly where furcasternum is embedded.
</paragraph>
<caption id="DF63665E1C4FA517FF680E65FC72ACAA" ID-DOI="http://doi.org/10.5281/zenodo.6352404" ID-Zenodo-Dep="6352404" httpUri="https://zenodo.org/record/6352404/files/figure.png" pageId="1" pageNumber="259" startId="1.[151,264,1851,1875]" targetBox="[151,1437,885,1827]" targetPageId="1">
<paragraph id="8BA336D61C4FA517FF680E65FC72ACAA" blockId="1.[151,1437,1851,2019]" pageId="1" pageNumber="259">
<emphasis id="B968EAC41C4FA517FF680E65FEBEAC1B" bold="true" box="[151,314,1851,1875]" pageId="1" pageNumber="259">FIGURES 1–4.</emphasis>
<taxonomicName id="4C1C4D551C4FA517FEBF0E65FDA5AC1B" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[320,545,1851,1875]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="259" phylum="Arthropoda" rank="species" species="siskiyou">
<emphasis id="B968EAC41C4FA517FEBF0E65FDA5AC1B" box="[320,545,1851,1875]" italics="true" pageId="1" pageNumber="259">Dracoleuctra siskiyou</emphasis>
</taxonomicName>
: 1. Thorax, dorsal. 2. Thorax, ventral. 3. Female abdomen, dorsal. 4. Female terminalia, ventral. Bs = probasisternum; Fs = profurcasternum; MBs = mesobasisternum; MeBs = Metabasisternum; MePrs = Metapresternum; MFs = mesofurcasternum; Mg = marginal groove of pronotum; MPfs = Mesopostfurcasternum; MPrs = Mesopresternum; MSs1 = Mesospinasternum 1; MSs2 = Mesospinasternum 2; Pfs = propostfurcasternum; Pn = central disc of pronotum; Psc = mesoprescutum; Pscl = mesopostscutellum; Prs = propresternum; Sct = mesoscutum.
</paragraph>
</caption>
<caption id="DF63665E1C4CA514FF680F80FF73AC52" ID-DOI="http://doi.org/10.5281/zenodo.6352406" ID-Zenodo-Dep="6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="2" pageNumber="260" startId="2.[151,264,1758,1782]" targetBox="[277,1311,182,1719]" targetPageId="2">
<paragraph id="8BA336D61C4CA514FF680F80FF73AC52" blockId="2.[151,1437,1758,1818]" pageId="2" pageNumber="260">
<emphasis id="B968EAC41C4CA514FF680F80FEBFADBE" bold="true" box="[151,315,1758,1782]" pageId="2" pageNumber="260">FIGURES 5–6.</emphasis>
<taxonomicName id="4C1C4D551C4CA514FEBD0F80FDA0ADBE" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[322,548,1758,1782]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="260" phylum="Arthropoda" rank="species" species="siskiyou">
<emphasis id="B968EAC41C4CA514FEBD0F80FDA0ADBE" box="[322,548,1758,1782]" italics="true" pageId="2" pageNumber="260">Dracoleuctra siskiyou</emphasis>
</taxonomicName>
: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed.
</paragraph>
</caption>
<paragraph id="8BA336D61C4CA515FF380E16FEE7A9E2" blockId="2.[151,1437,1864,2035]" lastBlockId="3.[151,1437,151,682]" lastPageId="3" lastPageNumber="261" pageId="2" pageNumber="260">
<emphasis id="B968EAC41C4CA514FF380E16FE85AC2A" box="[199,257,1864,1890]" italics="true" pageId="2" pageNumber="260">Male</emphasis>
. Length 4.5–6.0 mm, mean length 
<quantity id="4CE49B331C4CA514FD740E16FD65AC2B" box="[651,737,1864,1891]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="260" unit="mm" value="5.3">5.3 mm</quantity>
(n = 8). Abdominal segments sclerotized, dark to light brown. Abdominal segment 9 lacking vesicle (
<figureCitation id="13272A531C4CA514FDB70E32FD1DACCE" box="[584,665,1900,1926]" captionStart="FIGURES 9–14" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[155,1432,182,1805]" captionTargetId="figure-17@4.[155,1432,181,1806]" captionTargetPageId="4" captionText="FIGURES 9–14. Dracoleuctra siskiyou: 9. Male terminalia, dorsal. 10. Male terminalia, ventral. 11. Male terminalia, caudal. 12. Paraprocts, frontal. 13. Paraproct tips, lateral. 14. Female terminalia, ventral." figureDoi="http://doi.org/10.5281/zenodo.6352410" httpUri="https://zenodo.org/record/6352410/files/figure.png" pageId="2" pageNumber="260">Fig. 10</figureCitation>
), heavily sclerotized, uniformly dark brown, tergum narrow and widening laterally, sternum broad, cup-shaped then narrowing posteriorly to a dorsally produced subgenital process (we prefer this term to subgenital plate), process apically slightly emarginate; inner portion a membrane lined furrow where the subanal probe lies, membranous layer thick apically (
<figureCitation id="13272A531C4CA514FC920E86FC37ACBB" box="[877,947,2008,2035]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="2" pageNumber="260">Fig. 5</figureCitation>
, 
<figureCitation id="13272A531C4CA514FC3F0E86FC59ACBA" box="[960,989,2008,2034]" captionStart="FIGURES 9–14" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[155,1432,182,1805]" captionTargetId="figure-17@4.[155,1432,181,1806]" captionTargetPageId="4" captionText="FIGURES 9–14. Dracoleuctra siskiyou: 9. Male terminalia, dorsal. 10. Male terminalia, ventral. 11. Male terminalia, caudal. 12. Paraprocts, frontal. 13. Paraproct tips, lateral. 14. Female terminalia, ventral." figureDoi="http://doi.org/10.5281/zenodo.6352410" httpUri="https://zenodo.org/record/6352410/files/figure.png" pageId="2" pageNumber="260">11</figureCitation>
). Segment 10 forming sclerotized band under cerci and articulating with suspensory bars of paraprocts (
<figureCitation id="13272A531C4DA515FCA409C9FC1AABFA" box="[859,926,151,178]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Fig. 6</figureCitation>
), tergite with mesal longitudinal membranous line, mesoposterior border slightly produced posteriorly (
<figureCitation id="13272A531C4DA515FCDE09E5FCF6AB9D" box="[801,882,186,213]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Figs. 5</figureCitation>
, 
<figureCitation id="13272A531C4DA515FC7F09E5FC14AB9D" box="[896,912,187,213]" captionStart="FIGURES 9–14" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[155,1432,182,1805]" captionTargetId="figure-17@4.[155,1432,181,1806]" captionTargetPageId="4" captionText="FIGURES 9–14. Dracoleuctra siskiyou: 9. Male terminalia, dorsal. 10. Male terminalia, ventral. 11. Male terminalia, caudal. 12. Paraprocts, frontal. 13. Paraproct tips, lateral. 14. Female terminalia, ventral." figureDoi="http://doi.org/10.5281/zenodo.6352410" httpUri="https://zenodo.org/record/6352410/files/figure.png" pageId="3" pageNumber="261">9</figureCitation>
). Lateral sclerites narrowly contacting tergum 10 anteriorly, bases of cerci laterally, and epiproct sclerite posteromesally (
<figureCitation id="13272A531C4DA515FC220981FBA5ABB1" box="[989,1057,223,249]" captionStart="FIGURES 7–8" captionStartId="3.[151,264,1169,1193]" captionTargetBox="[206,1416,706,1146]" captionTargetId="figure-265@3.[171,1416,705,1146]" captionTargetPageId="3" captionText="FIGURES 7–8. Dracoleuctra siskiyou: 7. Male terminalia, dorsal, partially cleared, segment nine removed. 8. Male paraprocts, ventral, partially cleared." figureDoi="http://doi.org/10.5281/zenodo.6352408" httpUri="https://zenodo.org/record/6352408/files/figure.png" pageId="3" pageNumber="261">Fig. 7</figureCitation>
); epiproct sclerites fused mesally, extending posterolaterally, dorsally projecting epiproct lacking (
<figureCitation id="13272A531C4DA515FC97085DFC3CAA55" box="[872,952,258,285]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Figs. 5</figureCitation>
, 
<figureCitation id="13272A531C4DA515FC3A085DFC51AA55" box="[965,981,259,285]" captionStart="FIGURES 7–8" captionStartId="3.[151,264,1169,1193]" captionTargetBox="[206,1416,706,1146]" captionTargetId="figure-265@3.[171,1416,705,1146]" captionTargetPageId="3" captionText="FIGURES 7–8. Dracoleuctra siskiyou: 7. Male terminalia, dorsal, partially cleared, segment nine removed. 8. Male paraprocts, ventral, partially cleared." figureDoi="http://doi.org/10.5281/zenodo.6352408" httpUri="https://zenodo.org/record/6352408/files/figure.png" pageId="3" pageNumber="261">7</figureCitation>
); setose membranous pad present posterior to epiproct sclerite. Outer lobes of paraproct (suspensory bars) broad basally extending a short distance laterally then directed anteriorly as narrow sclerotized straps that articulate with posterior base of cerci (
<figureCitation id="13272A531C4DA515FB390815FA8FAA2E" box="[1222,1291,331,358]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Fig. 6</figureCitation>
), lateral portion with large distoposterior knob that articulates with posteroventral portion of segment 10. Basal sclerotization extends mesally along subanal probe as paired sclerotized rods that appear to be unsclerotized on mesal margin, dorsoapical sclerotization acuminate (
<figureCitation id="13272A531C4DA515FDBB08E9FD11AA99" box="[580,661,439,465]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Figs. 6</figureCitation>
, 
<figureCitation id="13272A531C4DA515FD5D08E9FD35AA99" box="[674,689,439,465]" captionStart="FIGURES 7–8" captionStartId="3.[151,264,1169,1193]" captionTargetBox="[206,1416,706,1146]" captionTargetId="figure-265@3.[171,1416,705,1146]" captionTargetPageId="3" captionText="FIGURES 7–8. Dracoleuctra siskiyou: 7. Male terminalia, dorsal, partially cleared, segment nine removed. 8. Male paraprocts, ventral, partially cleared." figureDoi="http://doi.org/10.5281/zenodo.6352408" httpUri="https://zenodo.org/record/6352408/files/figure.png" pageId="3" pageNumber="261">8</figureCitation>
, 
<figureCitation id="13272A531C4DA515FD3F08E9FC8DAA99" box="[704,777,439,465]" captionStart="FIGURES 9–14" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[155,1432,182,1805]" captionTargetId="figure-17@4.[155,1432,181,1806]" captionTargetPageId="4" captionText="FIGURES 9–14. Dracoleuctra siskiyou: 9. Male terminalia, dorsal. 10. Male terminalia, ventral. 11. Male terminalia, caudal. 12. Paraprocts, frontal. 13. Paraproct tips, lateral. 14. Female terminalia, ventral." figureDoi="http://doi.org/10.5281/zenodo.6352410" httpUri="https://zenodo.org/record/6352410/files/figure.png" pageId="3" pageNumber="261">12–13</figureCitation>
), ventral sclerotization ends subapically in a membranous structure (
<figureCitation id="13272A531C4DA515FEF60885FECBAABD" box="[265,335,474,501]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Fig. 5</figureCitation>
). Anterior to base of subanal probe is an interior membranous sac, at the mesoanterior end of sac lies the retractoral plate (
<figureCitation id="13272A531C4DA515FE4C08A1FD87A951" box="[435,515,511,537]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Figs. 6</figureCitation>
, 
<figureCitation id="13272A531C4DA515FDEE08A1FDA4A951" box="[529,544,511,537]" captionStart="FIGURES 7–8" captionStartId="3.[151,264,1169,1193]" captionTargetBox="[206,1416,706,1146]" captionTargetId="figure-265@3.[171,1416,705,1146]" captionTargetPageId="3" captionText="FIGURES 7–8. Dracoleuctra siskiyou: 7. Male terminalia, dorsal, partially cleared, segment nine removed. 8. Male paraprocts, ventral, partially cleared." figureDoi="http://doi.org/10.5281/zenodo.6352408" httpUri="https://zenodo.org/record/6352408/files/figure.png" pageId="3" pageNumber="261">8</figureCitation>
), a bluntly triangular sclerotized process (flat in lateral view), a pair of sclerotized straps (ducts?) extend from its base and are directed to the base of the subanal probe. Cerci large (
<figureCitation id="13272A531C4DA515FAE90B7DFAE1A975" box="[1302,1381,546,573]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Figs. 5</figureCitation>
, 
<figureCitation id="13272A531C4DA515FA8E0B7DFA0AA975" box="[1393,1422,547,573]" captionStart="FIGURES 9–14" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[155,1432,182,1805]" captionTargetId="figure-17@4.[155,1432,181,1806]" captionTargetPageId="4" captionText="FIGURES 9–14. Dracoleuctra siskiyou: 9. Male terminalia, dorsal. 10. Male terminalia, ventral. 11. Male terminalia, caudal. 12. Paraprocts, frontal. 13. Paraproct tips, lateral. 14. Female terminalia, ventral." figureDoi="http://doi.org/10.5281/zenodo.6352410" httpUri="https://zenodo.org/record/6352410/files/figure.png" pageId="3" pageNumber="261">11</figureCitation>
), heavily sclerotized, dark brown, horn shaped, directed posteriorly, curving mesally and slightly dorsally, apex acute with a subapical ventral tubercle, mesobasal margin with belt-like extension articulating with suspensory bars of paraproct (
<figureCitation id="13272A531C4DA515FEEF0BD1FED1A9E2" box="[272,341,655,682]" captionStart="FIGURES 5–6" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[277,1311,182,1719]" captionTargetId="figure-17@2.[276,1311,181,1734]" captionTargetPageId="2" captionText="FIGURES 5–6. Dracoleuctra siskiyou: 5. Male terminalia, dorsal. 6. Male terminalia, ventral, partially cleared, segment nine removed." figureDoi="http://doi.org/10.5281/zenodo.6352406" httpUri="https://zenodo.org/record/6352406/files/figure.png" pageId="3" pageNumber="261">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF63665E1C4DA515FF680DCFFE11AF86" ID-DOI="http://doi.org/10.5281/zenodo.6352408" ID-Zenodo-Dep="6352408" httpUri="https://zenodo.org/record/6352408/files/figure.png" pageId="3" pageNumber="261" startId="3.[151,264,1169,1193]" targetBox="[206,1416,706,1146]" targetPageId="3">
<paragraph id="8BA336D61C4DA515FF680DCFFE11AF86" blockId="3.[151,1437,1169,1230]" pageId="3" pageNumber="261">
<emphasis id="B968EAC41C4DA515FF680DCFFEBEAFE2" bold="true" box="[151,314,1169,1194]" pageId="3" pageNumber="261">FIGURES 7–8.</emphasis>
<taxonomicName id="4C1C4D551C4DA515FEC00DCCFDA4AFE2" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[319,544,1170,1194]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="261" phylum="Arthropoda" rank="species" species="siskiyou">
<emphasis id="B968EAC41C4DA515FEC00DCCFDA4AFE2" box="[319,544,1170,1194]" italics="true" pageId="3" pageNumber="261">Dracoleuctra siskiyou</emphasis>
</taxonomicName>
: 7. Male terminalia, dorsal, partially cleared, segment nine removed. 8. Male paraprocts, ventral, partially cleared.
</paragraph>
</caption>
<paragraph id="8BA336D61C4DA515FF380DA2FAABAD36" blockId="3.[151,1437,1276,1986]" pageId="3" pageNumber="261">
<emphasis id="B968EAC41C4DA515FF380DA2FE9FAE5E" box="[199,283,1276,1302]" italics="true" pageId="3" pageNumber="261">Female</emphasis>
. Length 
<quantity id="4CE49B331C4DA515FE800DA2FE5AAE5E" box="[383,478,1276,1302]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="3" pageNumber="261" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
, mean length 
<quantity id="4CE49B331C4DA515FD840DA2FD55AE5E" box="[635,721,1276,1303]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="261" unit="mm" value="5.7">5.7 mm</quantity>
(n = 19). Abdominal tergum 1 with large sclerite covering most of tergum; tergum 2 mostly membranous with large central sclerite and a few small sclerites laterally; terga 3–7 membranous with numerous small sclerites scattered over surface (
<figureCitation id="13272A531C4DA515FC7E0C1AFC40AE16" box="[897,964,1348,1374]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[151,1437,885,1827]" captionTargetId="figure-309@1.[151,1437,885,1827]" captionTargetPageId="1" captionText="FIGURES 1–4. Dracoleuctra siskiyou: 1. Thorax, dorsal. 2. Thorax, ventral. 3. Female abdomen, dorsal. 4. Female terminalia, ventral. Bs = probasisternum; Fs = profurcasternum; MBs = mesobasisternum; MeBs = Metabasisternum; MePrs = Metapresternum; MFs = mesofurcasternum; Mg = marginal groove of pronotum; MPfs = Mesopostfurcasternum; MPrs = Mesopresternum; MSs1 = Mesospinasternum 1; MSs2 = Mesospinasternum 2; Pfs = propostfurcasternum; Pn = central disc of pronotum; Psc = mesoprescutum; Pscl = mesopostscutellum; Prs = propresternum; Sct = mesoscutum." figureDoi="http://doi.org/10.5281/zenodo.6352404" httpUri="https://zenodo.org/record/6352404/files/figure.png" pageId="3" pageNumber="261">Fig. 3</figureCitation>
), each sclerite bearing one large dark seta; some sclerites longitudinally elongate and bearing 2–3 dark setae, these appear to be coalesced smaller sclerites. Tergum 8 mostly sclerotized with submesal membranous area wide anteriorly, narrowing posteriorly, bearing few small sclerites; terga 9 and 10 sclerotized. Abdominal sterna 1–6 with mesal sclerite surrounded by membranous area, sclerites increase in size with sternal number; sternum 7 mostly sclerotized, membranous anteriorly at pleural fold; sternum 8 (
<figureCitation id="13272A531C4DA515FEAC0CA6FE26AD5A" box="[339,418,1528,1554]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[151,1437,885,1827]" captionTargetId="figure-309@1.[151,1437,885,1827]" captionTargetPageId="1" captionText="FIGURES 1–4. Dracoleuctra siskiyou: 1. Thorax, dorsal. 2. Thorax, ventral. 3. Female abdomen, dorsal. 4. Female terminalia, ventral. Bs = probasisternum; Fs = profurcasternum; MBs = mesobasisternum; MeBs = Metabasisternum; MePrs = Metapresternum; MFs = mesofurcasternum; Mg = marginal groove of pronotum; MPfs = Mesopostfurcasternum; MPrs = Mesopresternum; MSs1 = Mesospinasternum 1; MSs2 = Mesospinasternum 2; Pfs = propostfurcasternum; Pn = central disc of pronotum; Psc = mesoprescutum; Pscl = mesopostscutellum; Prs = propresternum; Sct = mesoscutum." figureDoi="http://doi.org/10.5281/zenodo.6352404" httpUri="https://zenodo.org/record/6352404/files/figure.png" pageId="3" pageNumber="261">Figs. 4</figureCitation>
, 
<figureCitation id="13272A531C4DA515FE500CA6FE48AD5A" box="[431,460,1528,1554]" captionStart="FIGURES 9–14" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[155,1432,182,1805]" captionTargetId="figure-17@4.[155,1432,181,1806]" captionTargetPageId="4" captionText="FIGURES 9–14. Dracoleuctra siskiyou: 9. Male terminalia, dorsal. 10. Male terminalia, ventral. 11. Male terminalia, caudal. 12. Paraprocts, frontal. 13. Paraproct tips, lateral. 14. Female terminalia, ventral." figureDoi="http://doi.org/10.5281/zenodo.6352410" httpUri="https://zenodo.org/record/6352410/files/figure.png" pageId="3" pageNumber="261">14</figureCitation>
) with darkly sclerotized subgenital plate about two-thirds width and one-half length of sternite, posterior margin curving anteriorly in a symmetrical gentle arc or with a slight mesal posterior projection, posterior one-half of sternite membranous; sternum 9 sclerotized with a short anteromesal projection; sternum 10 membranous mesally; paraprocts roughly triangular, posterior margin blunt; cerci one segmented, setose.
</paragraph>
<paragraph id="8BA336D61C4DA515FF380FD7FE0AADEA" blockId="3.[151,1437,1276,1986]" box="[199,398,1672,1698]" pageId="3" pageNumber="261">
<emphasis id="B968EAC41C4DA515FF380FD7FE8EADEA" box="[199,266,1673,1698]" italics="true" pageId="3" pageNumber="261">Larva</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C306655D1C4DA513FF380FF2FB00A8ED" lastPageId="5" lastPageNumber="263" pageId="3" pageNumber="261" type="diagnosis">
<paragraph id="8BA336D61C4DA515FF380FF2FDC4AC8A" blockId="3.[151,1437,1276,1986]" pageId="3" pageNumber="261">
<emphasis id="B968EAC41C4DA515FF380FF2FEC6AD8E" bold="true" box="[199,322,1708,1734]" pageId="3" pageNumber="261">Diagnosis.</emphasis>
<emphasis id="B968EAC41C4DA515FEB30FF2FE58AD8E" box="[332,476,1708,1734]" italics="true" pageId="3" pageNumber="261">Morphology.</emphasis>
Both males and females of 
<taxonomicName id="4C1C4D551C4DA515FCD70FF2FC3BAD8E" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[808,959,1708,1734]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4DA515FCD70FF2FC3BAD8E" box="[808,959,1708,1734]" italics="true" pageId="3" pageNumber="261">Dracoleuctra</emphasis>
</taxonomicName>
can be recognized (even in the field) by the lack of wings and the one-segmented cerci. The following combination of characters are diagnostic for males: segment 9 heavily sclerotized, subgenital process produced dorsally with a membrane–lined interior furrow, inner apical portion with a thick membranous layer, vesicle lacking, heavily sclerotized horn-shaped cerci, lack of a dorsally projecting epiproct but with the lateral sclerites and epiproct sclerite together butterfly-shaped, and the membranous, bluntly cone shaped apex of the subanal probe pressed to inner membranous portion of subgenital process. Females can be recognized by the membranous abdominal tergites bearing numerous small, setose sclerites, and the size and shape of the subgenital plate.
</paragraph>
<caption id="DF63665E1C4AA512FF680E78FC4CAC2A" ID-DOI="http://doi.org/10.5281/zenodo.6352410" ID-Zenodo-Dep="6352410" httpUri="https://zenodo.org/record/6352410/files/figure.png" pageId="4" pageNumber="262" startId="4.[151,264,1830,1854]" targetBox="[155,1432,182,1805]" targetPageId="4">
<paragraph id="8BA336D61C4AA512FF680E78FC4CAC2A" blockId="4.[151,1437,1830,1890]" pageId="4" pageNumber="262">
<emphasis id="B968EAC41C4AA512FF680E78FECCAC76" bold="true" box="[151,328,1830,1854]" pageId="4" pageNumber="262">FIGURES 9–14.</emphasis>
<taxonomicName id="4C1C4D551C4AA512FEB00E78FDB5AC76" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[335,561,1830,1854]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="262" phylum="Arthropoda" rank="species" species="siskiyou">
<emphasis id="B968EAC41C4AA512FEB00E78FDB5AC76" box="[335,561,1830,1854]" italics="true" pageId="4" pageNumber="262">Dracoleuctra siskiyou</emphasis>
</taxonomicName>
: 9. Male terminalia, dorsal. 10. Male terminalia, ventral. 11. Male terminalia, caudal. 12. Paraprocts, frontal. 13. Paraproct tips, lateral. 14. Female terminalia, ventral.
</paragraph>
</caption>
<paragraph id="8BA336D61C4BA513FF3809C9FB00A8ED" blockId="5.[151,1437,150,1977]" pageId="5" pageNumber="263">
<bibRefCitation id="EF8D4B271C4BA513FF3809C9FE5CABFA" author="Ricker, W. E. &amp; Ross, H. H." box="[199,472,151,178]" pageId="5" pageNumber="263" pagination="1113 - 1127" refId="ref3995" refString="Ricker, W. E. &amp; Ross, H. H. (1969) The genus Zealeuctra and its position in the family Leuctridae (Plecoptera, Insecta). Canadian Journal of Zoology, 47, 1113 - 1127. https: // doi. org / 10.1139 / z 69 - 175" type="journal article" year="1969">Ricker and Ross (1969)</bibRefCitation>
, 
<bibRefCitation id="EF8D4B271C4BA513FE1A09C9FC9DABF9" author="Nelson, C. H. &amp; Hanson, J. F." box="[485,793,151,177]" pageId="5" pageNumber="263" pagination="187 - 199" refId="ref3957" refString="Nelson, C. H. &amp; Hanson, J. F. (1973) The genus Perlomyia (Plecoptera: Leuctridae). Journal of the Kansas Entomological Society, 46, 187 - 199." type="journal article" year="1973">Nelson and Hanson (1973)</bibRefCitation>
, 
<bibRefCitation id="EF8D4B271C4BA513FCD809C9FC48ABF9" author="Nelson, C. H." box="[807,972,151,177]" pageId="5" pageNumber="263" pagination="235 - 240" refId="ref3924" refString="Nelson, C. H. (1977) Position of &quot; Leuctra &quot; divisa Hitchcock within the family Leuctridae (Plecoptera). Entomological News, 88, 235 - 240." type="journal article" year="1977">Nelson (1977)</bibRefCitation>
, and 
<bibRefCitation id="EF8D4B271C4BA513FBF509C9FADBABF9" author="Shepard, W. D. &amp; Baumann, R. W." box="[1034,1375,150,177]" pageId="5" pageNumber="263" pagination="124 - 134" refId="ref4080" refString="Shepard, W. D. &amp; Baumann, R. W. (1995) Calileuctra, a new genus, and two new species of stoneflies from California (Plecoptera: Leuctridae). Great Basin Naturalist, 55, 124 - 134. https: // doi. org / 10.5962 / bhl. part. 22805" type="journal article" year="1995">Shepard and Baumann (1995)</bibRefCitation>
have proposed phylogenies for 
<taxonomicName id="4C1C4D551C4BA513FE4809E5FDB6AB9D" authorityName="Klapalek" authorityYear="1836" box="[439,562,187,213]" class="Insecta" family="Leuctridae" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="family">Leuctridae</taxonomicName>
. Although we do not propose a possible 
<taxonomicName id="4C1C4D551C4BA513FC1309E5FBE0AB9D" authorityName="Klapalek" authorityYear="1836" box="[1004,1124,187,213]" class="Insecta" family="Leuctridae" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="family">Leuctridae</taxonomicName>
phylogeny, we have studied these papers and examined leuctrid specimens we have in hand, including the western North American species of the following genera: 
<taxonomicName id="4C1C4D551C4BA513FE6E085DFCD5AA55" authority="Shepard &amp; Baumann, 1995" authorityName="Shepard &amp; Baumann" authorityYear="1995" box="[401,849,258,285]" class="Insecta" family="Leuctridae" genus="Calileuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FE6E085DFD95AA55" box="[401,529,259,285]" italics="true" pageId="5" pageNumber="263">Calileuctra</emphasis>
<bibRefCitation id="EF8D4B271C4BA513FDE6085DFCD5AA55" author="Shepard, W. D. &amp; Baumann, R. W." box="[537,849,258,285]" pageId="5" pageNumber="263" pagination="124 - 134" refId="ref4080" refString="Shepard, W. D. &amp; Baumann, R. W. (1995) Calileuctra, a new genus, and two new species of stoneflies from California (Plecoptera: Leuctridae). Great Basin Naturalist, 55, 124 - 134. https: // doi. org / 10.5962 / bhl. part. 22805" type="journal article" year="1995">Shepard &amp; Baumann, 1995</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C1C4D551C4BA513FCA2085DFBE7AA55" authority="Ricker, 1943" authorityName="Ricker" authorityYear="1943" box="[861,1123,259,285]" class="Insecta" family="Leuctridae" genus="Despaxia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FCA2085DFC43AA54" box="[861,967,259,284]" italics="true" pageId="5" pageNumber="263">Despaxia</emphasis>
Ricker, 1943
</taxonomicName>
, 
<taxonomicName id="4C1C4D551C4BA513FB90085DFA1CAA55" authority="Neave, 1934" authorityName="Neave" authorityYear="1934" box="[1135,1432,259,285]" class="Insecta" family="Leuctridae" genus="Megaleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FB90085DFB7AAA55" box="[1135,1278,259,285]" italics="true" pageId="5" pageNumber="263">Megaleuctra</emphasis>
Neave, 1934
</taxonomicName>
, 
<taxonomicName id="4C1C4D551C4BA513FF680879FE0EAA09" authority="Ricker, 1943" authorityName="Ricker" authorityYear="1943" box="[151,394,295,321]" class="Insecta" family="Leuctridae" genus="Moselia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FF680879FF75AA09" box="[151,241,295,321]" italics="true" pageId="5" pageNumber="263">Moselia</emphasis>
Ricker, 1943
</taxonomicName>
, 
<taxonomicName id="4C1C4D551C4BA513FE6A0879FD44AA09" authority="Hanson, 1941" authorityName="Hanson" authorityYear="1941" box="[405,704,295,321]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FE6A0879FD98AA09" box="[405,540,295,321]" italics="true" pageId="5" pageNumber="263">Paraleuctra</emphasis>
Hanson, 1941
</taxonomicName>
, 
<taxonomicName id="4C1C4D551C4BA513FD330879FC52AA09" authority="Banks, 1906" authorityName="Banks" authorityYear="1906" box="[716,982,295,321]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FD330879FCC5AA09" box="[716,833,295,321]" italics="true" pageId="5" pageNumber="263">Perlomyia</emphasis>
Banks, 1906
</taxonomicName>
, 
<taxonomicName id="4C1C4D551C4BA513FC1E0879FAE7AA09" authority="Stark &amp; Kyzar, 2001" authorityName="Stark &amp; Kyzar" authorityYear="2001" box="[993,1379,295,321]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FC1E0879FBF6AA09" box="[993,1138,295,321]" italics="true" pageId="5" pageNumber="263">Pomoleuctra</emphasis>
<bibRefCitation id="EF8D4B271C4BA513FB870879FAE7AA09" author="Stark, B. P. &amp; Kyzar, J. W." box="[1144,1379,295,321]" pageId="5" pageNumber="263" pagination="119 - 135" refId="ref4187" refString="Stark, B. P. &amp; Kyzar, J. W. (2001) Systematics of Nearctic Paraleuctra with description of a new genus (Plecoptera: Leuctridae). Tijdschrift voor Entomologie, 144, 119 - 135. https: // doi. org / 10.1163 / 22119434 - 99900060" type="journal article" year="2001">Stark &amp; Kyzar, 2001</bibRefCitation>
</taxonomicName>
, plus the apterous 
<taxonomicName id="4C1C4D551C4BA513FED90815FD6AAA2D" authority="Kacanski &amp; Zwick, 1970" authorityName="Kacanski &amp; Zwick" authorityYear="1970" box="[294,750,331,357]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="species" species="aptera">
<emphasis id="B968EAC41C4BA513FED90815FE49AA2C" box="[294,461,331,356]" italics="true" pageId="5" pageNumber="263">Leuctra aptera</emphasis>
<bibRefCitation id="EF8D4B271C4BA513FE2B0815FD6AAA2D" author="Kacanski, D. &amp; Zwick, P." box="[468,750,331,357]" pageId="5" pageNumber="263" pagination="1 - 16" refId="ref3830" refString="Kacanski, D. &amp; Zwick, P. (1970) Neue und wenig bekannte Plecopteren aus Jugoslawien. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 43, 1 - 16." type="journal article" year="1970">Kaćanski &amp; Zwick, 1970</bibRefCitation>
</taxonomicName>
from Europe and 
<taxonomicName id="4C1C4D551C4BA513FC460815FADBAA2D" authority="Shimizu, 2000" authorityName="Shimizu" authorityYear="2000" box="[953,1375,331,357]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="species" species="ambulans">
<emphasis id="B968EAC41C4BA513FC460815FB31AA2D" box="[953,1205,331,357]" italics="true" pageId="5" pageNumber="263">Paraleuctra ambulans</emphasis>
<bibRefCitation id="EF8D4B271C4BA513FB430815FADBAA2D" author="Shimizu, T." box="[1212,1375,331,357]" pageId="5" pageNumber="263" pagination="285 - 303" refId="ref4140" refString="Shimizu, T. (2000) Paraleuctra (Insecta: Plecoptera: Leuctridae) from Japan, with taxonomic notes on the Japanese Leuctridae. Species Diversity, 5, 285 - 303. https: // doi. org / 10.12782 / specdiv. 5.285" type="journal article" year="2000">Shimizu, 2000</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F30B76461C4BA513FF680831FF5EAAC1" box="[151,218,367,393]" name="Japan" pageId="5" pageNumber="263">Japan</collectingCountry>
. With input from Charles Nelson (personal communication, 2020), we feel 
<taxonomicName id="4C1C4D551C4BA513FBCF0831FB43AAC1" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[1072,1223,367,393]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FBCF0831FB43AAC1" box="[1072,1223,367,393]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
is closest to 
<taxonomicName id="4C1C4D551C4BA513FAA40831FF62AAE5" authorityName="Hanson" authorityYear="1941" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FAA40831FF62AAE5" italics="true" pageId="5" pageNumber="263">Paraleuctra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1C4D551C4BA513FEE508CDFE2FAAE5" authorityName="Stark &amp; Kyzar" authorityYear="2001" box="[282,427,403,429]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FEE508CDFE2FAAE5" box="[282,427,403,429]" italics="true" pageId="5" pageNumber="263">Pomoleuctra</emphasis>
</taxonomicName>
based on the general structure of the paraproct and paraproct lobes. 
<bibRefCitation id="EF8D4B271C4BA513FB6608CDFA18AAE5" author="Stark, B. P. &amp; Kyzar, J. W." box="[1177,1436,403,429]" pageId="5" pageNumber="263" pagination="119 - 135" refId="ref4187" refString="Stark, B. P. &amp; Kyzar, J. W. (2001) Systematics of Nearctic Paraleuctra with description of a new genus (Plecoptera: Leuctridae). Tijdschrift voor Entomologie, 144, 119 - 135. https: // doi. org / 10.1163 / 22119434 - 99900060" type="journal article" year="2001">Stark and Kyzar (2001)</bibRefCitation>
list nine shared features uniting species in 
<taxonomicName id="4C1C4D551C4BA513FD8908E9FD79AA99" authorityName="Hanson" authorityYear="1941" box="[630,765,439,465]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FD8908E9FD79AA99" box="[630,765,439,465]" italics="true" pageId="5" pageNumber="263">Paraleuctra</emphasis>
</taxonomicName>
. Two of the features are wing vein characters, so obviously these do not pertain to 
<taxonomicName id="4C1C4D551C4BA513FE680885FDAAAABD" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[407,558,475,501]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FE680885FDAAAABD" box="[407,558,475,501]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
. Additional unshared characters include basisternum and furcasternum of prothorax fused (not fused in 
<taxonomicName id="4C1C4D551C4BA513FE4508A1FDD5A951" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[442,593,511,537]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FE4508A1FDD5A951" box="[442,593,511,537]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
), epiproct reduced to a small, curved hook (dorsally projecting curved hook lacking in 
<taxonomicName id="4C1C4D551C4BA513FEEF0B7DFE23A975" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[272,423,547,573]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FEEF0B7DFE23A975" box="[272,423,547,573]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
), and female subgenital plate usually bilobed (not bilobed in 
<taxonomicName id="4C1C4D551C4BA513FB9E0B7DFB7CA975" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[1121,1272,547,573]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FB9E0B7DFB7CA975" box="[1121,1272,547,573]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
and only onehalf length of sternum 8). These 
<taxonomicName id="4C1C4D551C4BA513FDF80B19FD1AA929" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[519,670,583,609]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FDF80B19FD1AA929" box="[519,670,583,609]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
characters are also not shared with 
<taxonomicName id="4C1C4D551C4BA513FBCD0B19FB47A929" authorityName="Stark &amp; Kyzar" authorityYear="2001" box="[1074,1219,583,609]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FBCD0B19FB47A929" box="[1074,1219,583,609]" italics="true" pageId="5" pageNumber="263">Pomoleuctra</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C1C4D551C4BA513FB2E0B19FAECA929" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[1233,1384,583,609]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FB2E0B19FAECA929" box="[1233,1384,583,609]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
also differs from 
<taxonomicName id="4C1C4D551C4BA513FEDC0B35FE2EA9CD" authorityName="Hanson" authorityYear="1941" box="[291,426,619,645]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FEDC0B35FE2EA9CD" box="[291,426,619,645]" italics="true" pageId="5" pageNumber="263">Paraleuctra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1C4D551C4BA513FE200B35FDF4A9CD" authorityName="Stark &amp; Kyzar" authorityYear="2001" box="[479,624,619,645]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FE200B35FDF4A9CD" box="[479,624,619,645]" italics="true" pageId="5" pageNumber="263">Pomoleuctra</emphasis>
</taxonomicName>
in the lack of a vesicle, and from 
<taxonomicName id="4C1C4D551C4BA513FC180B35FBEAA9CD" authorityName="Hanson" authorityYear="1941" box="[999,1134,619,645]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FC180B35FBEAA9CD" box="[999,1134,619,645]" italics="true" pageId="5" pageNumber="263">Paraleuctra</emphasis>
</taxonomicName>
, having a robust sternum 9 with a posteriorly and dorsally produced subgenital process harboring the subanal probe. The western North American 
<taxonomicName id="4C1C4D551C4BA513FF380BEDFECAA985" authorityName="Hanson" authorityYear="1941" box="[199,334,691,717]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FF380BEDFECAA985" box="[199,334,691,717]" italics="true" pageId="5" pageNumber="263">Paraleuctra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1C4D551C4BA513FE770BEDFD9DA985" authorityName="Stark &amp; Kyzar" authorityYear="2001" box="[392,537,691,717]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FE770BEDFD9DA985" box="[392,537,691,717]" italics="true" pageId="5" pageNumber="263">Pomoleuctra</emphasis>
</taxonomicName>
we have examined also have furrows in sternum 9 demarcating the subgenital process, this feature is lacking in 
<taxonomicName id="4C1C4D551C4BA513FDF20B89FD20A9B9" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[525,676,727,753]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FDF20B89FD20A9B9" box="[525,676,727,753]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
. Additionally, the female abdominal tergum, membranous but bearing numerous small setose sclerites, appears to be unique to 
<taxonomicName id="4C1C4D551C4BA513FCC00BA5FC52A85D" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[831,982,763,789]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FCC00BA5FC52A85D" box="[831,982,763,789]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
. We have avoided using meso-metasternal characters for diagnosis since 
<bibRefCitation id="EF8D4B271C4BA513FDEB0A41FD2AA871" author="Zwick, P." box="[532,686,799,825]" pageId="5" pageNumber="263" pagination="709 - 746" refId="ref4317" refString="Zwick, P. (2000) Phylogenetic system and zoogeography of the Plecoptera. Annual Review of Entomology, 45, 709 - 746. https: // doi. org / 10.1146 / annurev. ento. 45.1.709" type="journal article" year="2000">Zwick (2000)</bibRefCitation>
questions how variable flight ability affects use of thoracic sternal characters for generic correlation. 
<taxonomicName id="4C1C4D551C4BA513FDE40A1DFD36A815" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[539,690,835,861]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FDE40A1DFD36A815" box="[539,690,835,861]" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
does have meso-metasternal characters similar to the descriptions and figures of apterous 
<taxonomicName id="4C1C4D551C4BA513FE600A39FE72A8C8" box="[415,502,871,896]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FE600A39FE72A8C8" box="[415,502,871,896]" italics="true" pageId="5" pageNumber="263">Leuctra</emphasis>
</taxonomicName>
from the Caucasus, 
<collectingCountry id="F30B76461C4BA513FD240A39FCB1A8C9" box="[731,821,871,897]" name="Georgia" pageId="5" pageNumber="263">Georgia</collectingCountry>
(
<bibRefCitation id="EF8D4B271C4BA513FCBC0A39FBB6A8C9" author="Teslenko, V. A. &amp; Palatov, D. M. &amp; Semenchenko, A. A." box="[835,1074,871,897]" pageId="5" pageNumber="263" pagination="546 - 560" refId="ref4241" refString="Teslenko, V. A., Palatov, D. M. &amp; Semenchenko, A. A. (2019) Description of new apterous winter species of Leuctra (Plecoptera: Leuctridae) based morphology and DNA barcoding and further records to stonefly fauna of the Caucasus, Georgia. Zootaxa, 4585 (3), 546 - 560. https: // doi. org / 10.11646 / zootaxa. 4585.3.9" type="journal article" year="2019">Teslenko et al., 2019</bibRefCitation>
). We are not sure if flight ability would affect prosternal sclerites, but we do include prosternal characters in our diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="C306655D1C4BA513FF380AF1FC09AF35" pageId="5" pageNumber="263" type="materials_examined">
<paragraph id="8BA336D61C4BA513FF380AF1FC09AF35" blockId="5.[151,1437,150,1977]" pageId="5" pageNumber="263">
<emphasis id="B968EAC41C4BA513FF380AF1FE0FA880" bold="true" box="[199,395,942,969]" pageId="5" pageNumber="263">Type specimens.</emphasis>
<materialsCitation id="3B743C8B1C4BA513FE6C0AF1FBC4AF59" ID-GBIF-Occurrence="3704872302" collectingDate="2020-05-06" collectionCode="CA, CAS" collectorName="J. Lee. To &amp; San Francisco" country="United States of America" county="Josephine County" elevation="1440" latitude="42.00162" location="Happy Camp Road" longLatPrecision="1" longitude="-123.53785" municipality="Academy of Sciences" pageId="5" pageNumber="263" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
<typeStatus id="54A788741C4BA513FE6C0AF1FE7FA881" box="[403,507,943,969]" pageId="5" pageNumber="263" type="holotype">Holotype</typeStatus>
male: 
<collectingRegion id="49D8F8341C4BA513FDB40AF1FD26A881" box="[587,674,943,969]" country="United States of America" name="Oregon" pageId="5" pageNumber="263">Oregon</collectingRegion>
, 
<collectingCounty id="62C24E5A1C4BA513FD510AF1FCFDA881" box="[686,889,943,969]" pageId="5" pageNumber="263">Josephine County</collectingCounty>
, 
<location id="8EC3600D1C4BA513FC7A0AF1FBE6A881" LSID="urn:lsid:plazi:treatment:03B587C01C4EA511FF680EC6FD75AFA1:8EC3600D1C4BA513FC7A0AF1FBE6A881" box="[901,1122,943,969]" country="United States of America" county="Josephine County" latitude="42.00162" longLatPrecision="1" longitude="-123.53785" municipality="Academy of Sciences" name="Happy Camp Road" pageId="5" pageNumber="263" stateProvince="Oregon">Happy Camp Road</location>
, 
<location id="8EC3600D1C4BA513FB910AF1FAA2A881" LSID="urn:lsid:plazi:treatment:03B587C01C4EA511FF680EC6FD75AFA1:8EC3600D1C4BA513FB910AF1FAA2A881" box="[1134,1318,943,969]" country="United States of America" county="Josephine County" latitude="42.00162" longLatPrecision="1" longitude="-123.53785" municipality="Academy of Sciences" name="Siskiyou Spring" pageId="5" pageNumber="263" stateProvince="Oregon">Siskiyou Spring</location>
, 
<geoCoordinate id="EE2850111C4BA513FACC0AF1FA18A881" box="[1331,1436,943,969]" degrees="42.00162" direction="north" orientation="latitude" pageId="5" pageNumber="263" precision="1" value="42.00162">42.00162</geoCoordinate>
<geoCoordinate id="EE2850111C4BA513FF680A8CFE9FA8A5" box="[151,283,978,1005]" degrees="123.53785" direction="west" orientation="longitude" pageId="5" pageNumber="263" precision="1" value="-123.53785">-123.53785</geoCoordinate>
, ~ 
<quantity id="4CE49B331C4BA513FECC0A8DFE0CA8A5" box="[307,392,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="5" pageNumber="263" unit="m" value="1440.0">
<elevation id="0031D1E51C4BA513FECC0A8DFE0CA8A5" box="[307,392,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="5" pageNumber="263" unit="m" value="1440.0">1440 m</elevation>
</quantity>
elevation, 
<location id="8EC3600D1C4BA513FDFC0A8DFCD4A8A5" LSID="urn:lsid:plazi:treatment:03B587C01C4EA511FF680EC6FD75AFA1:8EC3600D1C4BA513FDFC0A8DFCD4A8A5" box="[515,848,979,1005]" country="United States of America" county="Josephine County" latitude="42.00162" longLatPrecision="1" longitude="-123.53785" municipality="Academy of Sciences" name="Klamath-Siskiyou Mountains" pageId="5" pageNumber="263" stateProvince="Oregon">Klamath-Siskiyou Mountains</location>
, 
<date id="FFA210161C4BA513FCA50A8DFC67A8A5" box="[858,995,979,1005]" pageId="5" pageNumber="263" value="2020-05-06">
<collectingDate id="EFE6E9FE1C4BA513FCA50A8DFC67A8A5" box="[858,995,979,1005]" pageId="5" pageNumber="263" value="2020-05-06">6 May 2020</collectingDate>
</date>
, 
<collectorName id="26E953001C4BA513FC120A8DFBDDA8A5" box="[1005,1113,979,1005]" pageId="5" pageNumber="263">J. Lee. To</collectorName>
be deposited in the entomology collection at 
<collectingRegion id="49D8F8341C4BA513FEA50AA9FE48AF59" box="[346,460,1015,1041]" country="United States of America" name="California" pageId="5" pageNumber="263">California</collectingRegion>
<collectingMunicipality id="6BC7ACAC1C4BA513FE2D0AA9FD4CAF59" box="[466,712,1015,1041]" pageId="5" pageNumber="263">Academy of Sciences</collectingMunicipality>
, 
<collectorName id="26E953001C4BA513FD2B0AA9FCF1AF59" box="[724,885,1015,1041]" pageId="5" pageNumber="263">San Francisco</collectorName>
, 
<collectionCode id="ED0DAE131C4BA513FC7F0AA9FC2EAF59" box="[896,938,1015,1041]" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="263" type="Museum">CA</collectionCode>
, 
<collectingCountry id="F30B76461C4BA513FC490AA9FC68AF59" box="[950,1004,1015,1041]" name="United States of America" pageId="5" pageNumber="263">USA</collectingCountry>
(
<collectionCode id="ED0DAE131C4BA513FC030AA9FBBCAF59" box="[1020,1080,1015,1041]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="5" pageNumber="263" type="Museum">CAS</collectionCode>
)
</materialsCitation>
. 
<typeStatus id="54A788741C4BA513FBB40AA9FB38AF59" box="[1099,1212,1015,1041]" pageId="5" pageNumber="263" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D1AFD5F1C4BA513FB380AA9FAA5AF59" box="[1223,1313,1015,1041]" pageId="5" pageNumber="263" type="male">4 males</specimenCount>
, 
<materialsCitation id="3B743C8B1C4BA513FAD20AA9FE9FAF7D" ID-GBIF-Occurrence="3704872304" collectingDate="2020-05-06" collectionCode="CA, CAS" collectorName="J. Lee. To &amp; San Francisco" country="United States of America" county="Josephine County" elevation="1440" latitude="42.00162" location="Happy Camp Road" longLatPrecision="1" longitude="-123.53785" municipality="Academy of Sciences" pageId="5" pageNumber="263" specimenCount="1" stateProvince="Oregon" typeStatus="paratype">same data as holotype</materialsCitation>
; 
<specimenCount id="9D1AFD5F1C4BA513FEDA0D45FE17AF7D" box="[293,403,1051,1077]" pageId="5" pageNumber="263" type="female">9 females</specimenCount>
, 
<materialsCitation id="3B743C8B1C4BA513FE610D45FCFFAF11" ID-GBIF-Occurrence="3704872301" collectingDate="2020-05-01" collectionCode="CAS, BYUC" collectorName="Brigham Young University Collection" country="United States of America" county="Josephine County" elevation="1440" latitude="42.00162" location="To" longLatPrecision="1" longitude="-123.53785" municipality="Provo" pageId="5" pageNumber="263" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Utah" typeStatus="paratype">
same data as holotype but 
<date id="FFA210161C4BA513FD3C0D45FCC9AF7D" box="[707,845,1051,1077]" pageId="5" pageNumber="263" value="2020-05-01">
<collectingDate id="EFE6E9FE1C4BA513FD3C0D45FCC9AF7D" box="[707,845,1051,1077]" pageId="5" pageNumber="263" value="2020-05-01">1 May 2020</collectingDate>
</date>
. 
<location id="8EC3600D1C4BA513FCA90D45FCF7AF7D" LSID="urn:lsid:plazi:treatment:03B587C01C4EA511FF680EC6FD75AFA1:8EC3600D1C4BA513FCA90D45FCF7AF7D" box="[854,883,1051,1077]" country="United States of America" municipality="Provo" name="To" pageId="5" pageNumber="263" stateProvince="Utah">To</location>
be deposited at 
<collectionCode id="ED0DAE131C4BA513FBD90D45FBD8AF7D" box="[1062,1116,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="5" pageNumber="263" type="Museum">CAS</collectionCode>
(
<specimenCount id="9D1AFD5F1C4BA513FB940D45FB42AF7D" box="[1131,1222,1051,1077]" pageId="5" pageNumber="263" type="male">2 males</specimenCount>
; 
<specimenCount id="9D1AFD5F1C4BA513FB2F0D44FAB9AF7E" box="[1232,1341,1050,1078]" pageId="5" pageNumber="263" type="female">5 females</specimenCount>
) and the 
<collectorName id="26E953001C4BA513FF680D61FDC5AF11" box="[151,577,1087,1113]" pageId="5" pageNumber="263">Brigham Young University Collection</collectorName>
, 
<collectingMunicipality id="6BC7ACAC1C4BA513FDB40D61FD15AF11" box="[587,657,1087,1113]" pageId="5" pageNumber="263">Provo</collectingMunicipality>
, 
<collectingRegion id="49D8F8341C4BA513FD650D61FD57AF11" box="[666,723,1087,1113]" country="United States of America" name="Utah" pageId="5" pageNumber="263">Utah</collectingRegion>
, 
<collectingCountry id="F30B76461C4BA513FD210D61FC90AF11" box="[734,788,1087,1113]" name="United States of America" pageId="5" pageNumber="263">USA</collectingCountry>
(
<collectionCode id="ED0DAE131C4BA513FCDD0D61FCF7AF11" box="[802,883,1087,1113]" pageId="5" pageNumber="263">BYUC</collectionCode>
)
</materialsCitation>
<specimenCount id="9D1AFD5F1C4BA513FC7F0D61FC5DAF11" box="[896,985,1087,1113]" pageId="5" pageNumber="263" type="male">2 males</specimenCount>
; 
<specimenCount id="9D1AFD5F1C4BA513FC1C0D61FBD6AF11" box="[995,1106,1087,1113]" pageId="5" pageNumber="263" type="female">4 females</specimenCount>
.Additional material: 
<specimenCount id="9D1AFD5F1C4BA513FABF0D61FA1CAF11" box="[1344,1432,1087,1113]" pageId="5" pageNumber="263" type="male">3 males</specimenCount>
, 
<specimenCount id="9D1AFD5F1C4BA513FF680D3DFE87AF35" box="[151,259,1123,1149]" pageId="5" pageNumber="263" type="female">8 females</specimenCount>
(Jonathan J. Lee Collection); 
<materialsCitation id="3B743C8B1C4BA513FDAA0D3DFC0DAF35" ID-GBIF-Occurrence="3704872303" box="[597,905,1123,1149]" collectionCode="BYUC" country="United States of America" pageId="5" pageNumber="263" specimenCount="11" specimenCount-female="8" specimenCount-male="3" stateProvince="Utah">
<specimenCount id="9D1AFD5F1C4BA513FDAA0D3DFD2FAF35" box="[597,683,1123,1149]" pageId="5" pageNumber="263" type="male">3 males</specimenCount>
<specimenCount id="9D1AFD5F1C4BA513FD4D0D3DFC9AAF35" box="[690,798,1123,1149]" pageId="5" pageNumber="263" type="female">8 females</specimenCount>
(
<collectionCode id="ED0DAE131C4BA513FCD20D3DFC05AF35" box="[813,897,1123,1149]" pageId="5" pageNumber="263">BYUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C306655D1C4BA513FF380DD9FA02AE45" pageId="5" pageNumber="263" type="etymology">
<paragraph id="8BA336D61C4BA513FF380DD9FA02AE45" blockId="5.[151,1437,150,1977]" pageId="5" pageNumber="263">
<emphasis id="B968EAC41C4BA513FF380DD9FECAAFE9" bold="true" box="[199,334,1159,1185]" pageId="5" pageNumber="263">Etymology.</emphasis>
The prefix of the genus name 
<emphasis id="B968EAC41C4BA513FD520DD9FD71AFE8" box="[685,757,1159,1184]" italics="true" pageId="5" pageNumber="263">Draco</emphasis>
(Latin for lizard or dragon) was selected because the males had a reptilian quality running across the beating sheet. The suffix name was selected to indicate placement of the genus within the family 
<taxonomicName id="4C1C4D551C4BA513FE5D0D91FD99AFA1" authorityName="Klapalek" authorityYear="1836" box="[418,541,1231,1257]" class="Insecta" family="Leuctridae" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="family">Leuctridae</taxonomicName>
. Gender of the name is neuter. The trivial name 
<taxonomicName id="4C1C4D551C4BA513FBD40D91FB01AFA1" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[1067,1157,1231,1257]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="species" species="siskiyou">
<emphasis id="B968EAC41C4BA513FBD40D91FB01AFA1" box="[1067,1157,1231,1257]" italics="true" pageId="5" pageNumber="263">siskiyou</emphasis>
</taxonomicName>
was selected because the specimens were collected from Siskiyou Spring in the Siskiyou Mountains subregion of the Klamath Mountains.
</paragraph>
</subSubSection>
<subSubSection id="C306655D1C4BA511FF380C48FD75AFA1" lastPageId="7" lastPageNumber="265" pageId="5" pageNumber="263" type="biology_ecology">
<paragraph id="8BA336D61C4BA513FF380C48FE7FADA9" blockId="5.[151,1437,150,1977]" pageId="5" pageNumber="263">
<emphasis id="B968EAC41C4BA513FF380C48FEA3AE78" bold="true" box="[199,295,1302,1328]" pageId="5" pageNumber="263">Biology.</emphasis>
Siskiyou Spring is located in the Siskiyou Mountains subregion of the Klamath Mountains and is a headwater tributary of Althouse Creek in the Rogue River drainage. The spring emerges (
<figureCitation id="13272A531C4BA513FB6C0C65FB61AE1D" box="[1171,1253,1338,1365]" captionStart="FIGURES 15–20" captionStartId="6.[151,264,1920,1944]" captionTargetBox="[165,1423,182,1896]" captionTargetId="figure-17@6.[165,1423,181,1896]" captionTargetPageId="6" captionText="FIGURES 15–20. Siskiyou Spring: 15. Spring head. 16. Spring creek/seep. 17. Spring creek. 18. Spring creek and Sitka Alders. 19. Sitka Alder stand covering wetted area, 28 April 2020. 20. Sitka Alder stand covering wetted area, 2 July 2020." figureDoi="http://doi.org/10.5281/zenodo.6352412" httpUri="https://zenodo.org/record/6352412/files/figure.png" pageId="5" pageNumber="263">Fig. 15</figureCitation>
) at an elevation of about 
<quantity id="4CE49B331C4BA513FF020C00FED7AE31" box="[253,339,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="5" pageNumber="263" unit="m" value="1450.0">1450 m</quantity>
on a steep hillside as small creeks and seeps creating a wetted area several meters wide. At least two small creeks flow within the wetted area; between them seeps are moss and forb covered (
<figureCitation id="13272A531C4BA513FB2E0CDDFAD8AED6" box="[1233,1372,1411,1438]" captionStart="FIGURES 15–20" captionStartId="6.[151,264,1920,1944]" captionTargetBox="[165,1423,182,1896]" captionTargetId="figure-17@6.[165,1423,181,1896]" captionTargetPageId="6" captionText="FIGURES 15–20. Siskiyou Spring: 15. Spring head. 16. Spring creek/seep. 17. Spring creek. 18. Spring creek and Sitka Alders. 19. Sitka Alder stand covering wetted area, 28 April 2020. 20. Sitka Alder stand covering wetted area, 2 July 2020." figureDoi="http://doi.org/10.5281/zenodo.6352412" httpUri="https://zenodo.org/record/6352412/files/figure.png" pageId="5" pageNumber="263">Figs. 16–17</figureCitation>
). The longitudinal distance between the springhead and road is &lt;
<specimenCount id="9D1AFD5F1C4BA513FCBF0CF9FC14AE89" box="[832,912,1447,1473]" pageId="5" pageNumber="263" type="generic">
<quantity id="4CE49B331C4BA513FCBF0CF9FC08AE89" box="[832,908,1447,1473]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="263" unit="m" value="100.0">100 m</quantity>
.
</specimenCount>
The entire wetted area is covered by a nearly impenetrable stand of Sitka Alder (
<taxonomicName id="4C1C4D551C4BA513FDDF0C95FCBCAEAC" box="[544,824,1483,1509]" class="Insecta" family="Leuctridae" genus="Alnus" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="subSpecies" species="crispa" subSpecies="sinuata">
<emphasis id="B968EAC41C4BA513FDDF0C95FD29AEAC" box="[544,685,1483,1509]" italics="true" pageId="5" pageNumber="263">Alnus crispa</emphasis>
ssp. 
<emphasis id="B968EAC41C4BA513FD190C95FCBCAEAC" box="[742,824,1483,1508]" italics="true" pageId="5" pageNumber="263">sinuata</emphasis>
</taxonomicName>
) (
<figureCitation id="13272A531C4BA513FCAD0C95FC58AEAD" box="[850,988,1483,1509]" captionStart="FIGURES 15–20" captionStartId="6.[151,264,1920,1944]" captionTargetBox="[165,1423,182,1896]" captionTargetId="figure-17@6.[165,1423,181,1896]" captionTargetPageId="6" captionText="FIGURES 15–20. Siskiyou Spring: 15. Spring head. 16. Spring creek/seep. 17. Spring creek. 18. Spring creek and Sitka Alders. 19. Sitka Alder stand covering wetted area, 28 April 2020. 20. Sitka Alder stand covering wetted area, 2 July 2020." figureDoi="http://doi.org/10.5281/zenodo.6352412" httpUri="https://zenodo.org/record/6352412/files/figure.png" pageId="5" pageNumber="263">Figs. 18–20</figureCitation>
). Adult stoneflies collected with 
<taxonomicName id="4C1C4D551C4BA513FAB40C95FF62AD41" authorityName="Lee &amp; Baumann" authorityYear="2022" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FAB40C95FF62AD41" italics="true" pageId="5" pageNumber="263">Dracoleuctra</emphasis>
</taxonomicName>
were 
<taxonomicName id="4C1C4D551C4BA513FECE0CB1FD53AD41" authority="(Frison, 1937)" baseAuthorityName="Frison" baseAuthorityYear="1937" box="[305,727,1519,1545]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="species" species="forcipata">
<emphasis id="B968EAC41C4BA513FECE0CB1FDACAD41" box="[305,552,1519,1545]" italics="true" pageId="5" pageNumber="263">Paraleuctra forcipata</emphasis>
(Frison, 1937)
</taxonomicName>
, 
<taxonomicName id="4C1C4D551C4BA513FD190CB1FAB2AD41" authority="(Harper &amp; Wildman, 1985)" baseAuthorityName="Harper &amp; Wildman" baseAuthorityYear="1985" box="[742,1334,1518,1545]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis id="B968EAC41C4BA513FD190CB1FC75AD41" box="[742,1009,1519,1545]" italics="true" pageId="5" pageNumber="263">Pomoleuctra andersoni</emphasis>
(Harper &amp; Wildman, 1985)
</taxonomicName>
, females of an undescribed apterous 
<taxonomicName id="4C1C4D551C4BA513FE2B0F4DFDCCAD65" box="[468,584,1555,1581]" class="Insecta" family="Capniidae" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="family">Capniidae</taxonomicName>
, 
<emphasis id="B968EAC41C4BA513FDA90F4DFCC5AD65" box="[598,833,1555,1581]" italics="true" pageId="5" pageNumber="263">Soyedina nevadensis</emphasis>
(Claassen, 1923), 
<emphasis id="B968EAC41C4BA513FBE60F4DFB18AD65" box="[1049,1180,1555,1581]" italics="true" pageId="5" pageNumber="263">S. producta</emphasis>
(Claassen, 1923), and 
<emphasis id="B968EAC41C4BA513FF680F69FEE1AD19" box="[151,357,1591,1617]" italics="true" pageId="5" pageNumber="263">Zapada cordillera</emphasis>
(Baumann &amp; Gaufin, 1971). Adult Trichoptera collected were 
<emphasis id="B968EAC41C4BA513FBC20F69FA85AD18" box="[1085,1281,1591,1617]" italics="true" pageId="5" pageNumber="263">Goeracea genota</emphasis>
(Ross, 1941) (Goeridae) and 
<emphasis id="B968EAC41C4BA513FEB40F05FDA1AD3C" box="[331,549,1627,1653]" italics="true" pageId="5" pageNumber="263">Moselyana comosa</emphasis>
Denning, 1949 (Apataniidae). 
<taxonomicName id="4C1C4D551C4BA513FC760F05FC7BAD3D" box="[905,1023,1627,1653]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
larvae collected were 
<taxonomicName id="4C1C4D551C4BA513FAFC0F05FEF8ADD1" authority=", Yoraperla Ricker, 1952" authorityName="Yoraperla Ricker" authorityYear="1952" class="Insecta" family="Leuctridae" genus="Moselia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C4BA513FAFC0F05FAD9AD3D" box="[1283,1373,1627,1653]" italics="true" pageId="5" pageNumber="263">Moselia</emphasis>
, 
<emphasis id="B968EAC41C4BA513FA910F05FF64ADD1" italics="true" pageId="5" pageNumber="263">Yoraperla</emphasis>
Ricker, 1952
</taxonomicName>
, 
<emphasis id="B968EAC41C4BA513FE770F21FDE6ADD1" box="[392,610,1663,1689]" italics="true" pageId="5" pageNumber="263">Sasquaperla hoopa</emphasis>
Stark &amp; Baumann, 2001, 
<emphasis id="B968EAC41C4BA513FC740F21FC65ADD1" box="[907,993,1663,1689]" italics="true" pageId="5" pageNumber="263">Sweltsa</emphasis>
pr. 
<emphasis id="B968EAC41C4BA513FBF10F21FBD6ADD1" box="[1038,1106,1663,1689]" italics="true" pageId="5" pageNumber="263">fidelis</emphasis>
(Banks, 1920), 
<emphasis id="B968EAC41C4BA513FAF70F21FE83ADF4" italics="true" pageId="5" pageNumber="263">Chernokrilus misnomus</emphasis>
(Claassen, 1936), and 
<emphasis id="B968EAC41C4BA513FDF90FFDFD2AADF5" box="[518,686,1699,1725]" italics="true" pageId="5" pageNumber="263">Isoperla laucki</emphasis>
Baumann &amp; Lee, 2009. We have not visited the spring in late summer but presume it is perennial.
</paragraph>
<paragraph id="8BA336D61C4BA513FF380FB5FD42ACF1" blockId="5.[151,1437,150,1977]" pageId="5" pageNumber="263">
Initially, only females were caught by beating low vegetation next to running water. Some males were caught in low vegetation near the springhead, but it was later discovered that males were more easily caught by beating the vertical branches of Sitka Alder&gt; 
<quantity id="4CE49B331C4BA513FDEC0E6DFDB9AC05" box="[531,573,1843,1869]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="263" unit="m" value="2.0">2 m</quantity>
high in the upper section of the spring creek. Anecdotally, a small female was found on the beating sheet with a male. We are guessing that the females higher up in the vegetation were virgins while many of the females caught nearer the ground were larger and probably gravid. We hypothesize that the males climb up the branches in search of virgin females.
</paragraph>
<caption id="DF63665E1C48A510FF680EDEFAA7ACF5" ID-DOI="http://doi.org/10.5281/zenodo.6352412" ID-Zenodo-Dep="6352412" httpUri="https://zenodo.org/record/6352412/files/figure.png" pageId="6" pageNumber="264" startId="6.[151,264,1920,1944]" targetBox="[165,1423,182,1896]" targetPageId="6">
<paragraph id="8BA336D61C48A510FF680EDEFAA7ACF5" blockId="6.[151,1437,1920,1981]" pageId="6" pageNumber="264">
<emphasis id="B968EAC41C48A510FF680EDEFED7ACD0" bold="true" box="[151,339,1920,1944]" pageId="6" pageNumber="264">FIGURES 15–20.</emphasis>
Siskiyou Spring: 15. Spring head. 16. Spring creek/seep. 17. Spring creek. 18. Spring creek and Sitka Alders. 19. Sitka Alder stand covering wetted area, 28 April 2020. 20. Sitka Alder stand covering wetted area, 2 July 2020.
</paragraph>
</caption>
<paragraph id="8BA336D61C49A511FF3809C8FE2BAF11" blockId="7.[151,1437,150,1257]" pageId="7" pageNumber="265">
<emphasis id="B968EAC41C49A511FF3809C8FD52ABF8" bold="true" box="[199,726,150,177]" pageId="7" pageNumber="265">Apterism in the Klamath Mountains Region.</emphasis>
Relatively few apterous stoneflies are known from North America compared to other parts of the world, such as regions in Europe and 
<collectingCountry id="F30B76461C49A511FC5409E5FC6AAB9D" box="[939,1006,187,213]" name="Japan" pageId="7" pageNumber="265">Japan</collectingCountry>
. Outside of the Klamath Mountains region, fully apterous North American 
<taxonomicName id="4C1C4D551C49A511FDCA0981FD2FABB1" box="[565,683,223,249]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
include 
<emphasis id="B968EAC41C49A511FCF30981FC66ABB1" box="[780,994,223,249]" italics="true" pageId="7" pageNumber="265">Allocapnia harperi</emphasis>
Kirchner, 1980 and 
<emphasis id="B968EAC41C49A511FB370981FAC0ABB0" box="[1224,1348,223,249]" italics="true" pageId="7" pageNumber="265">A. vivipara</emphasis>
(Claassen, 1924) in the eastern 
<collectingCountry id="F30B76461C49A511FE47085DFDD6AA55" box="[440,594,259,285]" name="United States of America" pageId="7" pageNumber="265">United States</collectingCountry>
; 
<emphasis id="B968EAC41C49A511FD9F085DFC93AA55" box="[608,791,259,285]" italics="true" pageId="7" pageNumber="265">Capnia lacustra</emphasis>
Jewett, 1963, from Lake Tahoe, 
<collectingRegion id="49D8F8341C49A511FB6E085DFA82AA55" box="[1169,1286,259,285]" country="United States of America" name="California" pageId="7" pageNumber="265">California</collectingRegion>
; 
<emphasis id="B968EAC41C49A511FAEC085DFF55AA09" italics="true" pageId="7" pageNumber="265">Mesocapnia bergi</emphasis>
(Ricker, 1965), known from the lower Yukon River, 
<collectingRegion id="49D8F8341C49A511FCD40879FCF8AA09" box="[811,892,295,321]" country="United States of America" name="Alaska" pageId="7" pageNumber="265">Alaska</collectingRegion>
; 
<taxonomicName id="4C1C4D551C49A511FC760879FA81AA0A" authority="(Jewett, 1962)" baseAuthorityName="Jewett" baseAuthorityYear="1962" box="[905,1285,295,322]" class="Insecta" family="Capniidae" genus="Paracapnia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="species" species="disala">
<emphasis id="B968EAC41C49A511FC760879FBDDAA09" box="[905,1113,295,321]" italics="true" pageId="7" pageNumber="265">Paracapnia disala</emphasis>
(Jewett, 1962)
</taxonomicName>
, known from 
<collectingRegion id="49D8F8341C49A511FF680815FE8DAA2D" box="[151,265,331,357]" country="United States of America" name="California" pageId="7" pageNumber="265">California</collectingRegion>
and 
<collectingRegion id="49D8F8341C49A511FEBF0815FE13AA2D" box="[320,407,331,357]" country="United States of America" name="Oregon" pageId="7" pageNumber="265">Oregon</collectingRegion>
, 
<taxonomicName id="4C1C4D551C49A511FE5C0815FD43AA2E" authority="(Jewett, 1962)" baseAuthorityName="Jewett" baseAuthorityYear="1962" box="[419,711,331,358]" class="Insecta" family="Capniidae" genus="Paracapnia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="species" species="ensicala">
<emphasis id="B968EAC41C49A511FE5C0815FD98AA2D" box="[419,540,331,357]" italics="true" pageId="7" pageNumber="265">P. ensicala</emphasis>
(Jewett, 1962)
</taxonomicName>
, known from 
<collectingRegion id="49D8F8341C49A511FC9A0815FC3DAA2D" box="[869,953,331,357]" country="United States of America" name="Oregon" pageId="7" pageNumber="265">Oregon</collectingRegion>
and 
<collectingRegion id="49D8F8341C49A511FC0F0815FBFDAA2D" box="[1008,1145,331,357]" country="United States of America" name="Washington" pageId="7" pageNumber="265">Washington</collectingRegion>
; and 
<emphasis id="B968EAC41C49A511FB4A0815FA18AA2D" box="[1205,1436,331,357]" italics="true" pageId="7" pageNumber="265">Utacapnia tahoensis</emphasis>
(Nebeker &amp; Gaufin, 1965), Lake Tahoe, 
<collectingRegion id="49D8F8341C49A511FD9C0831FD5CAAC1" box="[611,728,367,393]" country="United States of America" name="California" pageId="7" pageNumber="265">California</collectingRegion>
. Although the environmental processes leading to apterism are not well established in the literature, isolation in a stable environment over time has been suggested. 
<bibRefCitation id="EF8D4B271C49A511FB0108CDFA18AAE5" author="Brinck, P." box="[1278,1436,403,429]" pageId="7" pageNumber="265" pagination="1 - 250" refId="ref3695" refString="Brinck, P. (1949) Studies on Swedish stoneflies. Opuscula Entomologica Supplementum, XI, 1 - 250." type="journal article" year="1949">Brinck (1949)</bibRefCitation>
states that “species with both sexes micropterous or apterous exist only in regions that have changed little during a long period of time.” 
<bibRefCitation id="EF8D4B271C49A511FE760885FD96AABE" author="Illies, J." box="[393,530,475,502]" pageId="7" pageNumber="265" pagination="328 - 333" refId="ref3791" refString="Illies, J. (1968) The first wingless stonefly from Australia. Psyche. A Journal of Entomology, 75, 328 - 333. https: // doi. org / 10.1155 / 1968 / 905824" type="journal article" year="1968">Illies (1968)</bibRefCitation>
echoed Brinck. 
<bibRefCitation id="EF8D4B271C49A511FD350885FBA4AABE" author="Loskutova, O. A. &amp; Zhiltzova, L. A." box="[714,1056,475,502]" pageId="7" pageNumber="265" pagination="26596" refId="ref3862" refString="Loskutova, O. A. &amp; Zhiltzova, L. A. (2016) Wing and body size polymorphism in populations of the stonefly Arcynopteryx dichroa McL. (Plecoptera: Perlodidae) in the Ural Mountains, Russia. Polar Research, 35, 26596. https: // doi. org / 10.3402 / polar. v 35.26596" type="journal article" year="2016">Loskutova &amp; Zhiltzova (2016)</bibRefCitation>
also suggested that short wingedness is associated with populations being isolated over a long period of time. Stability of a region over a long period of time may explain the degree of apterism in the Klamath Mountains, however, it might not be a predictive cause in other regions. The Klamath Mountains region is known for its great biological diversity containing more than 3,500 plant species, more than 200 of which are endemic (
<bibRefCitation id="EF8D4B271C49A511FD220B35FCF2A9CD" author="Sawyer, J. O." box="[733,886,619,645]" pageId="7" pageNumber="265" pagination="3 - 11" refId="ref4051" refString="Sawyer, J. O. (2007) Why are the Klamath Mountains and adjacent north coast floristically diverse? Fremontia, 35, 3 - 11." type="journal article" year="2007">Sawyer 2007</bibRefCitation>
). 
<bibRefCitation id="EF8D4B271C49A511FC770B35FBAAA9CD" author="Sawyer, J. O." box="[904,1070,619,645]" pageId="7" pageNumber="265" pagination="3 - 11" refId="ref4051" refString="Sawyer, J. O. (2007) Why are the Klamath Mountains and adjacent north coast floristically diverse? Fremontia, 35, 3 - 11." type="journal article" year="2007">Sawyer (2007)</bibRefCitation>
also notes that a large portion of the Klamath Mountains have been above sea level for 65 million years, with no records of volcanic deposits, and no glaciers were regionally extensive. This region is also isolated with poor access, particularly during winter months. Several species of apterous capniids are known from the Klamath Mountains of southern 
<collectingRegion id="49D8F8341C49A511FB8A0B89FB4DA9B9" box="[1141,1225,727,753]" country="United States of America" name="Oregon" pageId="7" pageNumber="265">Oregon</collectingRegion>
(OR) and northern 
<collectingRegion id="49D8F8341C49A511FF680BA5FE8DA85D" box="[151,265,763,789]" country="United States of America" name="California" pageId="7" pageNumber="265">California</collectingRegion>
(CA), including 
<emphasis id="B968EAC41C49A511FE360BA5FDD9A85D" box="[457,605,763,789]" italics="true" pageId="7" pageNumber="265">Capnia fialai</emphasis>
Nelson &amp; Baumann, 1990, known from one perennial headwater creek in Humboldt Co., CA; 
<taxonomicName id="4C1C4D551C49A511FE830A41FDCFA871" baseAuthorityName="Jewett" baseAuthorityYear="1962" box="[380,587,799,825]" class="Insecta" family="Capniidae" genus="Paracapnia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="species" species="disala">
<emphasis id="B968EAC41C49A511FE830A41FDCFA871" box="[380,587,799,825]" italics="true" pageId="7" pageNumber="265">Paracapnia disala</emphasis>
</taxonomicName>
, known from several headwater sites in southwestern OR and northwestern CA; the “ 
<collectingRegion id="49D8F8341C49A511FF020A1DFEF6A815" box="[253,370,835,861]" country="United States of America" name="California" pageId="7" pageNumber="265">California</collectingRegion>
<taxonomicName id="4C1C4D551C49A511FE870A1DFE78A814" authorityName="Hanson" authorityYear="1946" box="[376,508,835,860]" class="Insecta" family="Capniidae" genus="Paracapnia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C49A511FE870A1DFE78A814" box="[376,508,835,860]" italics="true" pageId="7" pageNumber="265">Paracapnia</emphasis>
</taxonomicName>
” (very probably a true phylogenetic lineage, David Murányi, unpublished), 
<taxonomicName id="4C1C4D551C49A511FAB70A1DFE2CA8C9" authority="Stark &amp; Baumann, 2004" authorityName="Stark &amp; Baumann" authorityYear="2004" class="Insecta" family="Capniidae" genus="Paracapnia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="species" species="boris">
<emphasis id="B968EAC41C49A511FAB70A1DFA19A815" box="[1352,1437,835,861]" italics="true" pageId="7" pageNumber="265">P. boris</emphasis>
Stark &amp; Baumann, 2004
</taxonomicName>
, known from perennial and perhaps seasonal creeks in Colusa, Shasta and Trinity counties, CA, 
<taxonomicName id="4C1C4D551C49A511FF340AD5FDE4A8ED" authority="Baumann &amp; Lee, 2007" authorityName="Baumann &amp; Lee" authorityYear="2007" box="[203,608,907,933]" class="Insecta" family="Capniidae" genus="Paracapnia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="species" species="humboldta">
<emphasis id="B968EAC41C49A511FF340AD5FED9A8ED" box="[203,349,907,933]" italics="true" pageId="7" pageNumber="265">P. humboldta</emphasis>
Baumann &amp; Lee, 2007
</taxonomicName>
, known from two seasonal creeks in Humboldt Co., CA, 
<taxonomicName id="4C1C4D551C49A511FB2C0AD5FE04A881" authority="Kondratieff &amp; Lee, 2010" authorityName="Kondratieff &amp; Lee" authorityYear="2010" class="Insecta" family="Capniidae" genus="Paracapnia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="species" species="baumanni">
<emphasis id="B968EAC41C49A511FB2C0AD5FAD9A8ED" box="[1235,1373,907,933]" italics="true" pageId="7" pageNumber="265">P. baumanni</emphasis>
Kondratieff &amp; Lee, 2010
</taxonomicName>
, known from seasonal creeks in Shasta and Trinity counties, CA; and 
<taxonomicName id="4C1C4D551C49A511FB6D0AF1FEEEA8A5" authority="Lee &amp; Baumann, 2011" authorityName="Lee &amp; Baumann" authorityYear="2011" class="Insecta" family="Capniidae" genus="Mesocapnia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="species" species="aptera">
<emphasis id="B968EAC41C49A511FB6D0AF1FAEFA880" box="[1170,1387,943,968]" italics="true" pageId="7" pageNumber="265">Mesocapnia aptera</emphasis>
Lee &amp; Baumann, 2011
</taxonomicName>
, known from one seasonal creek in Mendocino Co., CA. The addition of 
<taxonomicName id="4C1C4D551C49A511FB4D0A8DFACDA8A5" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[1202,1353,979,1005]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C49A511FB4D0A8DFACDA8A5" box="[1202,1353,979,1005]" italics="true" pageId="7" pageNumber="265">Dracoleuctra</emphasis>
</taxonomicName>
and an undescribed apterous capniid (known only from females and early instar larvae that DNA sequence analysis places near 
<emphasis id="B968EAC41C49A511FF310D45FEADAF7D" box="[206,297,1051,1077]" italics="true" pageId="7" pageNumber="265">C. fialai</emphasis>
) suggests that the region is a relative hotbed for apterous stoneflies. Future collecting may continue to produce surprising finds.
</paragraph>
<paragraph id="8BA336D61C49A511FF380D3CFD75AFA1" blockId="7.[151,1437,150,1257]" pageId="7" pageNumber="265">
<emphasis id="B968EAC41C49A511FF380D3CFEBCAF34" bold="true" box="[199,312,1122,1148]" pageId="7" pageNumber="265">Epilogue.</emphasis>
The 2020 Slater Fire in northern 
<collectingRegion id="49D8F8341C49A511FD390D3DFCBCAF35" box="[710,824,1123,1149]" country="United States of America" name="California" pageId="7" pageNumber="265">California</collectingRegion>
and southern 
<collectingRegion id="49D8F8341C49A511FC1F0D3DFBB0AF35" box="[992,1076,1123,1149]" country="United States of America" name="Oregon" pageId="7" pageNumber="265">Oregon</collectingRegion>
burned through the entire area where Siskiyou Spring occurs. Hazardous tree removal has kept Happy Camp Road (the access road) closed at least into 
<date id="FFA210161C49A511FEFA0DF5FE2AAF8D" box="[261,430,1195,1221]" pageId="7" pageNumber="265" value="2022-02">February 2022</date>
. Fire has probably burned through the area many times in the past and we expect that the 
<taxonomicName id="4C1C4D551C49A511FF680D91FEAAAFA1" authorityName="Lee &amp; Baumann" authorityYear="2022" box="[151,302,1231,1257]" class="Insecta" family="Leuctridae" genus="Dracoleuctra" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAC41C49A511FF680D91FEAAAFA1" box="[151,302,1231,1257]" italics="true" pageId="7" pageNumber="265">Dracoleuctra</emphasis>
</taxonomicName>
population has survived this recent fire.
</paragraph>
</subSubSection>
</treatment>
</document>