203 lines
23 KiB
XML
203 lines
23 KiB
XML
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<mods:title id="805DDEA446B56543C2720395900F021D">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="C2280C9BD5FAC94CD9D9613CF7446F96">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="67D6857DB1E6FDEABBBACF2CE11B9BB3">Karapunar, Baran</mods:namePart>
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<mods:namePart id="EAEB607DB1570644686019F599DE3678">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="7C24A337778A62C2EB29794344789D9B">Nützel, Al- Exander</mods:namePart>
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<mods:title id="708B11C837C61C6F3F410F17EEBBFC00">Zootaxa</mods:title>
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<mods:date id="641976C27BB149F7FBFFEC3CDDAF5870">2020</mods:date>
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<mods:number id="43996FDA96052959195FE7F492AE5241">2020-04-17</mods:number>
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<mods:number id="9D66B24B2399781229C267460C18352B">4766</mods:number>
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<treatment id="03B587AB4F121559FF517EE1FF25FD9E" ID-DOI="http://doi.org/10.5281/zenodo.3803805" ID-GBIF-Taxon="163737190" ID-Zenodo-Dep="3803805" LSID="urn:lsid:plazi:treatment:03B587AB4F121559FF517EE1FF25FD9E" httpUri="http://treatment.plazi.org/id/03B587AB4F121559FF517EE1FF25FD9E" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
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<subSubSection id="C30665364F12155BFF517EE1FE82F93F" box="[151,329,1695,1722]" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="8BA336BD4F12155BFF517EE1FE82F93F" blockId="37.[151,329,1695,1758]" box="[151,329,1695,1722]" pageId="37" pageNumber="38">
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<heading id="D0EB81D14F12155BFF517EE1FE82F93F" bold="true" box="[151,329,1695,1722]" fontSize="10" level="2" pageId="37" pageNumber="38" reason="2">
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<emphasis id="B968EAAF4F12155BFF517EE1FE82F93F" bold="true" box="[151,329,1695,1722]" pageId="37" pageNumber="38">
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<taxonomicName id="4C1C4D3E4F12155BFF517EE1FED5F93F" ID-CoL="7PDM8" authorityName="Mansuy" authorityYear="1914" box="[151,286,1695,1722]" class="Gastropoda" family="Orthonematidae" genus="Protostylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F12155BFF517EE1FED5F93F" bold="true" box="[151,286,1695,1722]" italics="true" pageId="37" pageNumber="38">Protostylus</emphasis>
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</taxonomicName>
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sp.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F12155BFF517EBAFEE0F95B" box="[151,299,1731,1758]" pageId="37" pageNumber="38" type="description">
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<paragraph id="8BA336BD4F12155BFF517EBAFEE0F95B" blockId="37.[151,329,1695,1758]" box="[151,299,1731,1758]" pageId="37" pageNumber="38">
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(
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<figureCitation id="13272A384F12155BFF667EBAFF24F95B" box="[160,239,1731,1758]" captionStart="FIGURE 25" captionStartId="38.[151,250,1830,1855]" captionTargetBox="[197,1390,181,1805]" captionTargetId="figure@38.[197,1390,181,1805]" captionTargetPageId="38" captionText="FIGURE 25. A, Donaldospira sp., ESKU-19-LP 85. B–E, Knightella irregularis (Longstaff, 1933), B–C, ESKU-19-LP 41. D–E, ESKU-19-LP 84. F–H, Protostylus sp., F, ESKU-19-LP 39. G–H, ESKU-19-LP 64. I–N, Trypanocochlea lopburiensis sp. nov. I, M, ESKU-19-LP 210, holotype; oblique view of early whorls to show protoconch. J, ESKU-19-LP 201, paratype. K, ESKU-19-LP 203, paratype. L, ESKU-19-LP 206, paratype; lateral view of early whorls to show protoconch. N, ESKU-19-LP 221, paratype. O–R, Streptacis? khaokhadensis sp. nov., O–P, ESKU-19-LP 214, holotype. Q–R, ESKU-19-LP 213, paratype. Scale bars represent: 2 mm (A–H); 200 μm (I–R)." figureDoi="http://doi.org/10.5281/zenodo.3763957" httpUri="https://zenodo.org/record/3763957/files/figure.png" pageId="37" pageNumber="38">Fig. 25</figureCitation>
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F–H)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F12155BFF517F75FF09F8EB" pageId="37" pageNumber="38" type="materials_examined">
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<paragraph id="8BA336BD4F12155BFF517F75FD7EF8A0" blockId="37.[151,1436,1803,2010]" box="[151,693,1803,1830]" pageId="37" pageNumber="38">
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<emphasis id="B968EAAF4F12155BFF517F75FECDF8A3" bold="true" box="[151,262,1803,1830]" pageId="37" pageNumber="38">Material.</emphasis>
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Two specimens: ESKU-19-
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<collectionCode id="ED0DAE784F12155BFDF97F72FDAAF8A3" box="[575,609,1804,1830]" country="Russia" name="Laboratory of Palaeontology" pageId="37" pageNumber="38">LP</collectionCode>
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39, 64.
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</paragraph>
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<paragraph id="8BA336BD4F12155BFF017F4EFF09F8EB" blockId="37.[151,1436,1803,2010]" pageId="37" pageNumber="38">
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Dimensions (mm): ESKU-19-
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<collectionCode id="ED0DAE784F12155BFDDF7F4EFDF0F8CF" box="[537,571,1840,1866]" country="Russia" name="Laboratory of Palaeontology" pageId="37" pageNumber="38">LP</collectionCode>
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39: height =
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<emphasis id="B968EAAF4F12155BFD137F4EFD2AF8CC" box="[725,737,1840,1865]" italics="true" pageId="37" pageNumber="38">c</emphasis>
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. 16.8; width = 7.8. ESKU-19-
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<collectionCode id="ED0DAE784F12155BFBFF7F4EFB90F8CF" box="[1081,1115,1840,1866]" country="Russia" name="Laboratory of Palaeontology" pageId="37" pageNumber="38">LP</collectionCode>
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64: height =
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<emphasis id="B968EAAF4F12155BFB307F4EFAC9F8CC" box="[1270,1282,1840,1865]" italics="true" pageId="37" pageNumber="38">c</emphasis>
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. 6.9; width = 2.9.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F12155BFF017F09FC85F85F" pageId="37" pageNumber="38" type="description">
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<paragraph id="8BA336BD4F12155BFF017F09FC85F85F" blockId="37.[151,1436,1803,2010]" pageId="37" pageNumber="38">
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<emphasis id="B968EAAF4F12155BFF017F09FE92F817" bold="true" box="[199,345,1911,1938]" pageId="37" pageNumber="38">Description.</emphasis>
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Shell high-spired, slender, consisting of approximately 7 whorls (apex missing); whorls smooth; whorls slightly convex to almost straight, periphery somewhat below mid-whorl; base evenly rounded, convex; anomphalous; sutures shallow but distinct; aperture unknown.
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</paragraph>
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</subSubSection>
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<caption id="DF6366354F111558FF517F58FD5AF876" ID-DOI="http://doi.org/10.5281/zenodo.3763957" ID-Zenodo-Dep="3763957" httpUri="https://zenodo.org/record/3763957/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,1830,1855]" targetBox="[197,1390,181,1805]" targetPageId="38">
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<paragraph id="8BA336BD4F111558FF517F58FD5AF876" blockId="38.[151,1437,1830,2036]" pageId="38" pageNumber="39">
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<emphasis id="B968EAAF4F111558FF517F58FEE9F8BA" bold="true" box="[151,290,1830,1855]" pageId="38" pageNumber="39">FIGURE 25.</emphasis>
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A,
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<taxonomicName id="4C1C4D3E4F111558FE8D7F59FE13F8BA" authorityName="Batten" authorityYear="1966" box="[331,472,1831,1855]" class="Gastropoda" family="Palaeostylidae" genus="Donaldospira" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F111558FE8D7F59FE13F8BA" box="[331,472,1831,1855]" italics="true" pageId="38" pageNumber="39">Donaldospira</emphasis>
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</taxonomicName>
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sp., ESKU-19-LP 85. B–E,
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<taxonomicName id="4C1C4D3E4F111558FCC47F59FB69F8BA" authority="(Longstaff, 1933)" baseAuthorityName="Longstaff" baseAuthorityYear="1933" box="[770,1186,1830,1855]" class="Gastropoda" family="Palaeostylidae" genus="Knightella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="irregularis">
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<emphasis id="B968EAAF4F111558FCC47F59FC29F8BA" box="[770,994,1831,1855]" italics="true" pageId="38" pageNumber="39">Knightella irregularis</emphasis>
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(
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<bibRefCitation id="EF8D4B4C4F111558FC377F58FB51F8BA" author="Longstaff" box="[1009,1178,1830,1855]" firstAuthor="Longstaff" pageId="38" pageNumber="39" pagination="87 - 124" refId="ref25915" refString="Longstaff, J. D. (1933) A revision of the British Carboniferous Loxonematidae, with descriptions of new forms. Quarterly Journal of the Geological Society of London, 89 (2), 87 - 124. https: // doi. org / 10.1144 / GSL. JGS. 1933.089.01 - 04.05" type="journal article" year="1933">Longstaff, 1933</bibRefCitation>
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)
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</taxonomicName>
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, B–C, ESKU-19-LP 41. D–E, ESKU-19-LP 84. F–H,
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<taxonomicName id="4C1C4D3E4F111558FE0F7F35FDF0F8E6" authorityName="Mansuy" authorityYear="1914" box="[457,571,1867,1891]" class="Gastropoda" family="Orthonematidae" genus="Protostylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F111558FE0F7F35FDF0F8E6" box="[457,571,1867,1891]" italics="true" pageId="38" pageNumber="39">Protostylus</emphasis>
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</taxonomicName>
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sp., F, ESKU-19-LP 39. G–H, ESKU-19-LP 64. I–N,
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<taxonomicName id="4C1C4D3E4F111558FBB57F35FA57F8E6" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" box="[1139,1436,1867,1891]" class="Gastropoda" family="Murchisoniidae" genus="Trypanocochlea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="lopburiensis" status="sp. nov.">
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<emphasis id="B968EAAF4F111558FBB57F35FA57F8E6" box="[1139,1436,1867,1891]" italics="true" pageId="38" pageNumber="39">Trypanocochlea lopburiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57D44F111558FF517F0EFF2DF802" box="[151,230,1904,1927]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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I, M, ESKU-19-LP 210, holotype; oblique view of early whorls to show protoconch. J, ESKU-19-LP 201, paratype. K, ESKU-19-LP 203, paratype. L, ESKU-19-LP 206, paratype; lateral view of early whorls to show protoconch. N, ESKU-19-LP 221, paratype. O–R,
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<taxonomicName id="4C1C4D3E4F111558FEAB7FC6FE1AF84A" authorityName="Meek" authorityYear="1871" box="[365,465,1976,1999]" class="Gastropoda" family="Streptacididae" genus="Streptacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F111558FEAB7FC6FE1AF84A" box="[365,465,1976,1999]" italics="true" pageId="38" pageNumber="39">Streptacis</emphasis>
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</taxonomicName>
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?
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<emphasis id="B968EAAF4F111558FE227FC9FDB2F84A" box="[484,633,1975,1999]" italics="true" pageId="38" pageNumber="39">khaokhadensis</emphasis>
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sp. nov., O–P, ESKU-19-LP 214, holotype. Q–R, ESKU-19-LP 213, paratype. Scale bars represent: 2 mm (A–H); 200 μm (I–R).
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</paragraph>
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</caption>
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<subSubSection id="C30665364F101559FF0178E7FF25FD9E" pageId="39" pageNumber="40" type="discussion">
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<paragraph id="8BA336BD4F101559FF0178E7FF25FD9E" blockId="39.[151,1437,153,539]" pageId="39" pageNumber="40">
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<emphasis id="B968EAAF4F101559FF0178E7FEF0FF36" bold="true" box="[199,315,153,179]" pageId="39" pageNumber="40">Remarks.</emphasis>
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These unornamented high-spired shells resemble several Permian
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<taxonomicName id="4C1C4D3E4F101559FBEB78E7FB67FF36" authorityName="Mansuy" authorityYear="1914" box="[1069,1196,153,179]" class="Gastropoda" family="Orthonematidae" genus="Protostylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F101559FBEB78E7FB67FF36" box="[1069,1196,153,179]" italics="true" pageId="39" pageNumber="40">Protostylus</emphasis>
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</taxonomicName>
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species e.g. from the Middle Permian
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<collectingRegion id="49D8F85F4F101559FE9778C3FEB0FF52" box="[337,379,189,215]" country="Thailand" name="Tak" pageId="39" pageNumber="40">Tak</collectingRegion>
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Fa Limestone from
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<collectingCountry id="F30B762D4F101559FD9C78C3FD75FF52" box="[602,702,189,215]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
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(
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<bibRefCitation id="EF8D4B4C4F101559FD0C78C3FC15FF5D" author="Ketwetsuriya" box="[714,990,189,216]" etAl="et al." firstAuthor="Ketwetsuriya" pageId="39" pageNumber="40" pagination="481 - 513" refId="ref24896" refString="Ketwetsuriya, C., Nu ¨ tzel, A. & Kanjanapayont, P. (2016) Permian gastropods from the Tak Fa Limestone, Nakhon Sawan, Northern Thailand. Bulletin of Geosciences, 91, 481 - 513. https: // doi. org / 10.3140 / bull. geosci. 1617" type="journal article" year="2016">
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Ketwetsuriya
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<emphasis id="B968EAAF4F101559FCA178C0FC5CFF52" box="[871,919,189,215]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
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. 2016
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</bibRefCitation>
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) but the body-whorl of the present specimens seems to be more inflated. The Japanese
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<taxonomicName id="4C1C4D3E4F101559FD70789FFCFEFF7E" authorityName="Mansuy" authorityYear="1914" box="[694,821,225,251]" class="Gastropoda" family="Orthonematidae" genus="Protostylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F101559FD70789FFCFEFF7E" box="[694,821,225,251]" italics="true" pageId="39" pageNumber="40">Protostylus</emphasis>
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</taxonomicName>
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species from the Middle Permian Akasaka Limestone (Nützel &
|
||
<bibRefCitation id="EF8D4B4C4F101559FED2797BFE07FE9A" author="Nakazawa" box="[276,460,261,287]" firstAuthor="Nakazawa" pageId="39" pageNumber="40" pagination="103 - 169" refId="ref27024" refString="Nu ¨ tzel, A. & Nakazawa, K. (2012) Permian (Capitanian) gastropods from the Akasaka Limestone (Gifu Prefecture, Japan). Journal of Systematic Palaeontology, 10, 103 - 169. https: // doi. org / 10.1080 / 14772019.2010.549659" type="journal article" year="2012">Nakazawa 2012</bibRefCitation>
|
||
) are much smaller.
|
||
<taxonomicName id="4C1C4D3E4F101559FD77797BFCFBFE9A" authorityName="Mansuy" authorityYear="1914" box="[689,816,261,287]" class="Gastropoda" family="Orthonematidae" genus="Protostylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F101559FD77797BFCFBFE9A" box="[689,816,261,287]" italics="true" pageId="39" pageNumber="40">Protostylus</emphasis>
|
||
</taxonomicName>
|
||
sp. from the Permian of Ratburi Limestone,
|
||
<collectingCountry id="F30B762D4F101559FAFF797BFA56FE9A" box="[1337,1437,261,287]" name="Thailand" pageId="39" pageNumber="40">Thailand</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F101559FF597957FE70FEC6" author="Ketwetsuriya" box="[159,443,297,323]" etAl="et al." firstAuthor="Ketwetsuriya" pageId="39" pageNumber="40" pagination="53 - 77" refId="ref24953" refString="Ketwetsuriya, C., Cook, A. G. & Nu ¨ tzel, A. (2020) Permian gastropods from the Ratburi Limestone, Khao Phrik, Central Thai- land. Pala ¨ ontologische Zeitschrift, 94, 53 - 77. https: // doi. org / 10.1007 / s 12542 - 019 - 00463 - 0" type="journal article" year="2020">
|
||
Ketwetsuriya
|
||
<emphasis id="B968EAAF4F101559FEF97954FEB9FEC6" box="[319,370,297,323]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
) is more slender in shape. The
|
||
<typeStatus id="54A7881F4F101559FCE37957FC9EFEC6" box="[805,853,297,323]" pageId="39" pageNumber="40">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C1C4D3E4F101559FC787957FA96FEC6" authority="Mansuy, 1914" authorityName="Mansuy" authorityYear="1914" box="[958,1373,297,323]" class="Gastropoda" family="Orthonematidae" genus="Protostylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="lantenoisi">
|
||
<emphasis id="B968EAAF4F101559FC787957FB7FFEC6" box="[958,1204,297,323]" italics="true" pageId="39" pageNumber="40">Protostylus lantenoisi</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F101559FB7B7957FA96FEC6" author="Mansuy" box="[1213,1373,297,323]" firstAuthor="Mansuy" pageId="39" pageNumber="40" pagination="1 - 190" refId="ref26085" refString="Mansuy, H. (1914) Nouvelle contribution a la paleontologie du Yunnan. Memoires du Service Geologique de l'Indochine, 3 (2), 1 - 190." type="journal article" year="1914">Mansuy, 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Carboniferous of South
|
||
<collectingCountry id="F30B762D4F101559FE117933FDD1FEE2" box="[471,538,333,359]" name="China" pageId="39" pageNumber="40">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49D8F85F4F101559FDEC7933FD48FEE2" box="[554,643,333,359]" country="China" name="Yunnan" pageId="39" pageNumber="40">Yunnan</collectingRegion>
|
||
) and the specimens illustrated by
|
||
<bibRefCitation id="EF8D4B4C4F101559FBCB7933FB61FEE2" author="Batten" box="[1037,1194,332,359]" firstAuthor="Batten" pageId="39" pageNumber="40" pagination="1 - 40" refId="ref23123" refString="Batten, R. L. (1985) Permian gastropods from Perak, Malaysia. Part 3. The murchisoniids, cerithiids, loxonematids, and subulitids. American Museum Novitates, 2829, 1 - 40." type="journal article" year="1985">Batten (1985)</bibRefCitation>
|
||
from the Permian of
|
||
<collectingCountry id="F30B762D4F101559FF51790FFECBFE0E" box="[151,256,369,395]" name="Malaysia" pageId="39" pageNumber="40">Malaysia</collectingCountry>
|
||
, differ from the present specimens in being more slender smaller and having lower whorls. The Permian Malaysian species
|
||
<taxonomicName id="4C1C4D3E4F101559FEA879EBFC88FE2A" authority="Batten, 1985" authorityName="Batten" authorityYear="1985" box="[366,835,404,431]" class="Gastropoda" family="Coelostylinidae" genus="Omphaloptychia" higherTaxonomySource="CoL" kingdom="Animalia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="paleozoica">
|
||
<emphasis id="B968EAAF4F101559FEA879EBFD61FE2A" box="[366,682,405,431]" italics="true" pageId="39" pageNumber="40">Omphaloptychia paleozoica</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F101559FD7479EBFC88FE2A" author="Batten" box="[690,835,404,431]" firstAuthor="Batten" pageId="39" pageNumber="40" pagination="1 - 40" refId="ref23123" refString="Batten, R. L. (1985) Permian gastropods from Perak, Malaysia. Part 3. The murchisoniids, cerithiids, loxonematids, and subulitids. American Museum Novitates, 2829, 1 - 40." type="journal article" year="1985">Batten, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
is similar in having high whorls, but it has a moderately high-spired turbiniform shape and a more step-like whorl profile. The studied specimens are also similar to
|
||
<taxonomicName id="4C1C4D3E4F101559FF5179A3FD96FE72" authority="(Longstaff, 1933)" baseAuthorityName="Longstaff" baseAuthorityYear="1933" box="[151,605,477,503]" class="Gastropoda" family="Palaeostylidae" genus="Knightella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="irregularis">
|
||
<emphasis id="B968EAAF4F101559FF5179A3FE45FE72" box="[151,398,477,503]" italics="true" pageId="39" pageNumber="40">Knightella irregularis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F101559FE5879A3FD9EFE72" author="Longstaff" box="[414,597,477,503]" firstAuthor="Longstaff" pageId="39" pageNumber="40" pagination="87 - 124" refId="ref25915" refString="Longstaff, J. D. (1933) A revision of the British Carboniferous Loxonematidae, with descriptions of new forms. Quarterly Journal of the Geological Society of London, 89 (2), 87 - 124. https: // doi. org / 10.1144 / GSL. JGS. 1933.089.01 - 04.05" type="journal article" year="1933">Longstaff, 1933</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but its whorls are somewhat lower and less convex, and it has shallower sutures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |