468 lines
65 KiB
XML
468 lines
65 KiB
XML
<document id="D281C691581A457794DBBAE5745C80DC" ID-DOI="10.11646/zootaxa.4803.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3907900" ID-ZooBank="2D391AAF-F105-4252-A090-34E725A05315" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593082538606" checkinUser="plazi" docAuthor="Chirivi-Joya, Daniel, Moreno-González, Jairo A. & Fagua, Giovanny" docDate="2020" docId="03B58796B640FFD2FF7EFC24FF5D75E2" docLanguage="en" docName="zootaxa.4803.1.1.pdf" docOrigin="Zootaxa 4803 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterophrynus elaphus Pocock 1903" docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="FF8CFFEEB652FFC4FFE9FFA9FFE77661" masterDocTitle="Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species" masterLastPageNumber="41" masterPageNumber="1" pageNumber="19" updateTime="1700277040307" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6465F40D7D2FC50E24F0B57AF4D67CCD">Two new species of the whip-spider genus Heterophrynus (Arachnida: Amblypygi) with complementary information of four species</mods:title>
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<mods:namePart id="7AA2E20EDB142645C9E9AE82BEBCFA07">Chirivi-Joya, Daniel</mods:namePart>
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<mods:affiliation id="01DFD06A8DA8572516EA2C023BDE4169">Laboratório de Entomología, Departamento de Biología, Facultad de Ciencias, Pontificia Universidad Javeriana. Carrera 7 N °. 40 - 62, Bogotá. & dhipnosodin @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1198 - 782 X</mods:affiliation>
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<mods:namePart id="BEAA44CE1F4AB08BAE831BCA2AC3A2E1">Moreno-González, Jairo A.</mods:namePart>
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<mods:affiliation id="C2A4B725E25DB9484280EC51D60FEDA5">Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo. Rua do Matão, trav. 14, n ° 321, Cidade Universitária, São Paulo- SP, CEP: 05508 - 090. hansenochrus @ gmail. com; https: // orcid. org / 0000 - 0001 - 5980 - 1417</mods:affiliation>
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<mods:namePart id="5099B754022B5457E2D4538F9822DDBF">Fagua, Giovanny</mods:namePart>
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<mods:affiliation id="19528FC4C84731B318C5CB029C7B011D">Laboratório de Entomología, Departamento de Biología, Facultad de Ciencias, Pontificia Universidad Javeriana. Carrera 7 N °. 40 - 62, Bogotá. & fagua @ javeriana. edu. co; https: // orcid. org / 0000 - 0003 - 1253 - 4416</mods:affiliation>
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<mods:title id="A1B3D82D5CFBB71BC28D61C8D8FAFCA0">Zootaxa</mods:title>
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<mods:part id="B5F0B071ECF7E69A084121BCBD77E5EC">
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<mods:date id="26E3CB90C65197DF122B84E5E83E053E">2020</mods:date>
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<mods:number id="20A0C456B28000BDDB6CA48984E45963">2020-06-25</mods:number>
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<mods:number id="4FA9D9600B0A954982705F42455D339C">4803</mods:number>
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<subSubSection id="C306650BB640FFD6FF7EFC24FDF275C6" box="[151,533,909,936]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8BA33680B640FFD6FF7EFC24FDF275C6" blockId="18.[151,554,909,971]" box="[151,533,909,936]" pageId="18" pageNumber="19">
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<heading id="D0EB81ECB640FFD6FF7EFC24FDF275C6" bold="true" box="[151,533,909,936]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<taxonomicName id="4C1C4D03B640FFD6FF7EFC24FDF275C6" ID-CoL="3LBZL" authority="Pocock" authorityName="Pocock" authorityYear="1903" box="[151,533,909,936]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elaphus">
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<emphasis id="B968EA92B640FFD6FF7EFC24FDF275C6" bold="true" box="[151,533,909,936]" pageId="18" pageNumber="19">
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<emphasis id="B968EA92B640FFD6FF7EFC24FE5375C9" bold="true" box="[151,436,909,936]" italics="true" pageId="18" pageNumber="19">Heterophrynus elaphus</emphasis>
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Pocock
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C306650BB640FFD6FF7EFC18FDCD75AA" box="[151,554,945,971]" pageId="18" pageNumber="19" type="description">
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<paragraph id="8BA33680B640FFD6FF7EFC18FDCD75AA" blockId="18.[151,554,909,971]" box="[151,554,945,971]" pageId="18" pageNumber="19">
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<figureCitation id="13272A05B640FFD6FF7EFC18FEDD75AA" box="[151,314,945,971]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="19.[151,250,1790,1815]" captionStartId-1="20.[151,250,1907,1932]" captionStartId-2="21.[151,250,1904,1929]" captionTargetBox-0="[161,1426,182,1752]" captionTargetBox-1="[165,1426,465,1872]" captionTargetBox-2="[163,1425,747,1874]" captionTargetId-0="figure@19.[151,1436,181,1765]" captionTargetId-1="figure@20.[151,1436,453,1882]" captionTargetId-2="figure@21.[157,1431,741,1879]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 11. Heterophrynus elaphus. Female (MZUSP 74571): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." captionText-1="FIGURE 12. Heterophrynus elaphus. Female (MZUSP 74571): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." captionText-2="FIGURE 13. Male genital: Heterophrynus elaphus. (MZUSP 74571): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MZUSP 74571), white arrow indicates the sclerotized area of the seminal receptacle,), black arrow indicates an extra sclerite at the base of gonopod." figureDoi-0="http://doi.org/10.5281/zenodo.3907924" figureDoi-1="http://doi.org/10.5281/zenodo.3907926" figureDoi-2="http://doi.org/10.5281/zenodo.3907928" httpUri-0="https://zenodo.org/record/3907924/files/figure.png" httpUri-1="https://zenodo.org/record/3907926/files/figure.png" httpUri-2="https://zenodo.org/record/3907928/files/figure.png" pageId="18" pageNumber="19">Figures 11–13</figureCitation>
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,
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<figureCitation id="13272A05B640FFD6FEACFC18FE8675AA" box="[325,353,945,971]" captionStart="FIGURE 22" captionStartId="33.[151,250,1654,1679]" captionTargetBox="[185,1402,574,1621]" captionTargetId="figure@33.[177,1410,561,1629]" captionTargetPageId="33" captionText="FIGURE 22. Distribution map. Heterophrynus alces: purple circle. Heterophrynus armiger: green circle. Heterophrynus cheiracanthus: blue circle. Heterophrynus elaphus: orange circle. Heterophrynus awa sp. nov.: gray circle. Heterophrynus origamii sp. nov.: red circle." figureDoi="http://doi.org/10.5281/zenodo.3907947" httpUri="https://zenodo.org/record/3907947/files/figure.png" pageId="18" pageNumber="19">22</figureCitation>
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(in part).
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<tableCitation id="C69E033BB640FFD6FE38FC18FDC175AA" box="[465,550,945,971]" captionStart="TABLE 4" captionStartId="22.[151,239,946,970]" captionText="TABLE 4. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus elaphus. Empty cells correspond to missing segment or data." pageId="18" pageNumber="19">Table 4</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306650BB640FFD6FF7EFC5CFCCD72C6" pageId="18" pageNumber="19" type="reference_group">
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<paragraph id="8BA33680B640FFD6FF7EFC5CFCCD72C6" blockId="18.[151,1436,1012,1191]" pageId="18" pageNumber="19">
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<treatmentCitation id="0ABD1091B640FFD6FF7EFC5CFD62726C" author="Pocock, R. I." box="[151,645,1012,1037]" page="222" pageId="18" pageNumber="19" year="1903">
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<taxonomicName id="4C1C4D03B640FFD6FF7EFC5CFD62726C" ID-CoL="3LBZL" authority="Pocock, 1903: 220 - 222" authorityName="Pocock" authorityPageNumber="220 - 222" authorityYear="1903" box="[151,645,1012,1037]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elaphus">
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<emphasis id="B968EA92B640FFD6FF7EFC5CFE61726C" box="[151,390,1013,1037]" italics="true" pageId="18" pageNumber="19">Heterophrynus elaphus</emphasis>
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<bibRefCitation id="EF8D4B71B640FFD6FE67FC5DFD62726C" author="Pocock, R. I." box="[398,645,1012,1037]" pageId="18" pageNumber="19" pagination="220 - 226" refId="ref23478" refString="Pocock, R. I. (1903) Descriptions of four new Arachnida of the orders Pedipalpi, Solifugae, and Araneae. Annals and Magazine of Natural History, Series 7, 11, 220 - 226. https: // doi. org / 10.1080 / 00222930308678753" type="journal article" year="1903">Pocock, 1903: 220–222</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="0ABD1091B640FFD6FD7BFC5DFC99726C" author="Mello-Leitao, C." box="[658,894,1012,1037]" page="49" pageId="18" pageNumber="19" year="1931">
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<bibRefCitation id="EF8D4B71B640FFD6FD7BFC5DFC99726C" author="Mello-Leitao, C." box="[658,894,1012,1037]" pageId="18" pageNumber="19" pagination="7 - 72" refId="ref23079" refString="Mello-Leitao, C. (1931) Pedipalpos do Brasil e algumas notas sobre a ordem. Archivos do Museu Nacional, 33, 7 - 72." type="journal article" year="1931">Mello-Leitão 1931: 49</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0ABD1091B640FFD6FC62FC5DFB31726C" author="Mello-Leitao, C. & Feio, J. de" box="[907,1238,1012,1037]" page="318" pageId="18" pageNumber="19" year="1949">
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<bibRefCitation id="EF8D4B71B640FFD6FC62FC5DFB31726C" author="Mello-Leitao, C. & Feio, J. de" box="[907,1238,1012,1037]" pageId="18" pageNumber="19" pagination="313 - 324" refId="ref23131" refString="Mello-Leitao, C. & Feio, J. de A. (1949) Notas sobre pequena coleccao de arachnidios do Peru. Boletim do Museu Goeldi de Historia Naturel e Ethnographia, 10, 313 - 324." type="journal article" year="1949">Mello-Leitão & Feio 1949: 318</bibRefCitation>
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</treatmentCitation>
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;
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||
<treatmentCitation id="0ABD1091B640FFD6FB0BFC5FFA7F726C" author="Roewer, C. F." box="[1250,1432,1012,1037]" page="37" pageId="18" pageNumber="19" year="1952">
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<bibRefCitation id="EF8D4B71B640FFD6FB0BFC5FFA7F726C" author="Roewer, C. F." box="[1250,1432,1012,1037]" pageId="18" pageNumber="19" pagination="37 - 58" refId="ref23693" refString="Roewer, C. F. (1952) Neotropische Arachnida Arthrogastra, zumeist aus Peru. Senckenbergiana Biologica, 33, 37 - 58." type="journal article" year="1952">Roewer 1952: 37</bibRefCitation>
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</treatmentCitation>
|
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;
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<treatmentCitation id="0ABD1091B640FFD6FF2EFBBDFE61724D" author="Roewer, C. F." box="[199,390,1043,1068]" page="430" pageId="18" pageNumber="19" year="1956">
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<bibRefCitation id="EF8D4B71B640FFD6FF2EFBBDFE61724D" author="Roewer, C. F." box="[199,390,1043,1068]" pageId="18" pageNumber="19" pagination="429 - 445" refId="ref23719" refString="Roewer, C. F. (1956) Arachnida Arthrogastra aus Peru, II. Senckenbergiana Biologica, 37, 429 - 445." type="journal article" year="1956">Roewer 1956: 430</bibRefCitation>
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</treatmentCitation>
|
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;
|
||
<treatmentCitation id="0ABD1091B640FFD6FE79FBBDFDA6724D" author="Roewer, C. F." box="[400,577,1043,1068]" page="68" pageId="18" pageNumber="19" year="1957">
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<bibRefCitation id="EF8D4B71B640FFD6FE79FBBDFDA6724D" author="Roewer, C. F." box="[400,577,1043,1068]" pageId="18" pageNumber="19" pagination="67 - 94" refId="ref23744" refString="Roewer, C. F. (1957) Arachnida Arthrogastra aus Peru, III. Senckenbergiana Biologica, 38, 67 - 94." type="journal article" year="1957">Roewer 1957: 68</bibRefCitation>
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</treatmentCitation>
|
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;
|
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<bibRefCitation id="EF8D4B71B640FFD6FDA5FBBAFD0D724D" author="Weygoldt, P." box="[588,746,1043,1068]" pageId="18" pageNumber="19" pagination="95 - 107" refId="ref24153" refString="Weygoldt, P. (1972) Geisselskorpione und Geisselspinnen (Uropygi und Amblypygi). Zeitschrift des Kolner Zoo, 15, 95 - 107." type="journal article" year="1972">Weygoldt 1972</bibRefCitation>
|
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: fig. 10;
|
||
<bibRefCitation id="EF8D4B71B640FFD6FCAAFBBAFC0F724D" author="Weygoldt, P." box="[835,1000,1043,1068]" pageId="18" pageNumber="19" pagination="175 - 191" refId="ref24180" refString="Weygoldt, P. (1974) Vergleichende Untersuchungen au zwei Heterophrynus (Admetus) - Arten, H. longicornis Butler und H. batesii (Arachnida, Amblypygi, Tarantulidae). Zoologischer Anzeiger, 192, 175 - 191." type="journal article" year="1974">Weygoldt, 1974</bibRefCitation>
|
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: figs 2, 6;
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<treatmentCitation id="0ABD1091B640FFD6FBBBFBBAFA7F724D" author="Igelmund, P. & Wendler, G." box="[1106,1432,1043,1068]" page="287" pageId="18" pageNumber="19" year="1985">
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<bibRefCitation id="EF8D4B71B640FFD6FBBBFBBAFA7F724D" author="Igelmund, P. & Wendler, G." box="[1106,1432,1043,1068]" pageId="18" pageNumber="19" pagination="287" refId="ref22827" refString="Igelmund, P. & Wendler, G. (1985) Periphere Rieseninterneurone in den Tastbeinen von Geisselspinnen (Amblypygi, Arachnida). Verhandlungen der Deutschen Zoologischen Gesellschaft, 78, 287." type="journal article" year="1985">Igelmund & Wendler 1985: 287</bibRefCitation>
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</treatmentCitation>
|
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;
|
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<treatmentCitation id="0ABD1091B640FFD6FF2EFB9BFE57722A" author="Igelmund, P." box="[199,432,1074,1099]" page="75" pageId="18" pageNumber="19" year="1987">
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<bibRefCitation id="EF8D4B71B640FFD6FF2EFB9BFE57722A" author="Igelmund, P." box="[199,432,1074,1099]" pageId="18" pageNumber="19" pagination="75 - 89" refId="ref22773" refString="Igelmund, P. (1987) Morphology, sense organs, and regeneration of the forelegs (whips) of the whip spider Heterophrynus elaphus (Arachnida, Amblypygi). Journal of Morphology, 193, 75 - 89. https: // doi. org / 10.1002 / jmor. 1051930108" type="journal article" year="1987">Igelmund 1987: 75–88</bibRefCitation>
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</treatmentCitation>
|
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, figs 1–6;
|
||
<treatmentCitation id="0ABD1091B640FFD6FDF2FB9BFC62722A" author="Igelmund, P. & Wendler, G." box="[539,901,1074,1099]" page="63" pageId="18" pageNumber="19" year="1991">
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<bibRefCitation id="EF8D4B71B640FFD6FDF2FB9BFC62722A" author="Igelmund, P. & Wendler, G." box="[539,901,1074,1099]" pageId="18" pageNumber="19" pagination="63 - 73" refId="ref22862" refString="Igelmund, P. & Wendler, G. (1991 a) The giant fibre system in the forelegs (whips) of the whip spider Heterophrynus elaphus Pocock (Arachnida: Amblypygi). Journal of Comparative Physiology, A, 168, 63 - 73. https: // doi. org / 10.1007 / BF 00217104" type="journal article" year="1991">Igelmund & Wendler 1991a: 63–73</bibRefCitation>
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</treatmentCitation>
|
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, figs 1–10;
|
||
<treatmentCitation id="0ABD1091B640FFD6FC12FB9BFA8B722A" author="Igelmund, P. & Wendler, G." box="[1019,1388,1074,1099]" page="75" pageId="18" pageNumber="19" year="1991">
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<bibRefCitation id="EF8D4B71B640FFD6FC12FB9BFA8B722A" author="Igelmund, P. & Wendler, G." box="[1019,1388,1074,1099]" pageId="18" pageNumber="19" pagination="75 - 83" refId="ref22922" refString="Igelmund, P. & Wendler, G. (1991 b) Morphology and physiology of peripheral giant interneurons in the forelegs (whips) of the whip spider Heterophrynus elaphus Pocock (Arachnida: Amblypygi). Journal of Comparative Physiology, A, 168, 75 - 83. https: // doi. org / 10.1007 / BF 00217105" type="journal article" year="1991">Igelmund & Wendler, 1991b: 75–82</bibRefCitation>
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</treatmentCitation>
|
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, figs 1–6; Selden
|
||
<emphasis id="B968EA92B640FFD6FEA0FBFBFE637208" box="[329,388,1105,1129]" italics="true" pageId="18" pageNumber="19">et. al.</emphasis>
|
||
1991: figs 2, 6;
|
||
<treatmentCitation id="0ABD1091B640FFD6FDD8FBF9FCE27208" author="Weygoldt, P." box="[561,773,1104,1129]" page="245" pageId="18" pageNumber="19" year="1994">
|
||
<bibRefCitation id="EF8D4B71B640FFD6FDD8FBF9FCE27208" author="Weygoldt, P." box="[561,773,1104,1129]" pageId="18" pageNumber="19" pagination="241 - 247" refId="ref24271" refString="Weygoldt, P. (1994) Amblypygi. In: Juberthie, C. & Decu, V. (Eds.), Encyclopaedia biospeologica. Vol. 1. Societe de Biospeologie, Moulis and Bucarest, pp. 241 - 247." type="book chapter" year="1994">Weygoldt 1994: 245</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B71B640FFD6FCF8FBF9FC577208" author="Weygoldt, P." box="[785,944,1104,1129]" pageId="18" pageNumber="19" pagination="185 - 202" refId="ref24317" refString="Weygoldt, P. (1996) Evolutionary morphology of whip spiders: towards a phylogenetic system (Chelicerata: Arachnida: Amblypygi). Journal of Zoological Systematics and Evolution Research, 34, 185 - 202. https: // doi. org / 10.1111 / j. 1439 - 0469.1996. tb 00825. x" type="journal article" year="1996">Weygoldt 1996</bibRefCitation>
|
||
: figs 9, 29, 33;
|
||
<bibRefCitation id="EF8D4B71B640FFD6FBBDFBF9FB127208" author="Weygoldt, P." box="[1108,1269,1104,1129]" pageId="18" pageNumber="19" refId="ref24375" refString="Weygoldt, P. (2000) Whip Spiders (Chelicerata: Amblypygi): Their biology, morphology and systematics. Apollo Books, Stenstrup, 163 pp." type="book" year="2000">Weygoldt 2000</bibRefCitation>
|
||
: figs 20, 25, 32, 79–87;
|
||
<treatmentCitation id="0ABD1091B640FFD6FEFAFBC6FE0772E9" author="Giupponi, A." box="[275,480,1135,1160]" page="173" pageId="18" pageNumber="19" year="2002">
|
||
<bibRefCitation id="EF8D4B71B640FFD6FEFAFBC6FE0772E9" author="Giupponi, A." box="[275,480,1135,1160]" pageId="18" pageNumber="19" pagination="173 - 175" refId="ref22531" refString="Giupponi, A. (2002) duas novas sinonimias no genero Heterophrynus Pocock, 1894 (Amblypygi, Phrynidae). Revista Iberica de Aracnologia, 6, 173 - 175." type="journal article" year="2002">Giupponi 2002: 173</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0ABD1091B640FFD6FE03FBC6FD1072E9" author="Reveillion, F. & Olivier, P. & Hofer, H." box="[490,759,1135,1160]" page="102" pageId="18" pageNumber="19" year="2014">
|
||
<bibRefCitation id="EF8D4B71B640FFD6FE03FBC6FD1072E9" author="Reveillion, F. & Olivier, P. & Hofer, H." box="[490,759,1135,1160]" pageId="18" pageNumber="19" pagination="102" refId="ref23649" refString="Reveillion, F., Olivier, P. & Hofer, H. (2014) First record of Heterophrynus elaphus (Pocock, 1903) (Amblypygi: Phrynidae) for Bolivia. Revista Iberica de Aracnologia, 25, 102." type="journal article" year="2014">
|
||
Réveillion
|
||
<emphasis id="B968EA92B640FFD6FDB0FBD8FD8C72E9" box="[601,619,1137,1160]" italics="true" pageId="18" pageNumber="19">et</emphasis>
|
||
al. 2014: 102
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0ABD1091B640FFD6FD16FBD9FC3C72E9" author="Armas, L. F. & Palomino-Cardenas, A. & Castillo-Espinoza, M." box="[767,987,1135,1160]" page="45" pageId="18" pageNumber="19" year="2016">
|
||
<bibRefCitation id="EF8D4B71B640FFD6FD16FBD9FC3C72E9" author="Armas, L. F. & Palomino-Cardenas, A. & Castillo-Espinoza, M." box="[767,987,1135,1160]" pageId="18" pageNumber="19" pagination="45 - 50" refId="ref22027" refString="Armas, L. F., Palomino-Cardenas, A. & Castillo-Espinoza, M. (2016) Amblipigios de los departamentos Cusco y Madre De Dios, Peru, con la descripcion de un nuevo Charinus (Amblypygi: Charinidae, Phrynidae). Revista Iberica de Aracnologia, 28, 45 - 50." type="journal article" year="2016">
|
||
Armas
|
||
<emphasis id="B968EA92B640FFD6FCA1FBD8FC9372E9" box="[840,884,1136,1160]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
|
||
. 2016: 45
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 48, figs 1A, 2F-I, 3C;
|
||
<treatmentCitation id="0ABD1091B640FFD6FB28FBC6FEC872C6" author="Lehmann, T. & Friedrich, S." page="171" pageId="18" pageNumber="19" year="2018">
|
||
<bibRefCitation id="EF8D4B71B640FFD6FB28FBC6FEC872C6" author="Lehmann, T. & Friedrich, S." pageId="18" pageNumber="19" pagination="169 - 172" refId="ref23034" refString="Lehmann, T. & Friedrich, S. (2018) DNA barcoding the smaller arachnid orders from ACP Panguana, Amazonian Peru (Amblypygi, Phrynidae and Schizomida, Hubbardiidae). Spixiana, 41 (2), 169 - 172." type="journal article" year="2018">Lehmann & Friedrich 2018: 171</bibRefCitation>
|
||
</treatmentCitation>
|
||
, table 2, 3; Ballón-Estacio & Armas 2019: 15, 19.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306650BB640FFD6FF7EFB7CFE1E733A" pageId="18" pageNumber="19" type="materials_examined">
|
||
<paragraph id="8BA33680B640FFD6FF7EFB7CFD46728E" blockId="18.[151,1437,1236,2019]" box="[151,673,1236,1263]" pageId="18" pageNumber="19">
|
||
<materialsCitation id="3B743CDDB640FFD6FF7EFB7CFD7A728E" box="[151,669,1236,1263]" country="Peru" location="Marcapata Valley" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
|
||
<emphasis id="B968EA92B640FFD6FF7EFB7CFEDE728E" bold="true" box="[151,313,1236,1263]" pageId="18" pageNumber="19">
|
||
<typeStatus id="54A78822B640FFD6FF7EFB7CFF35728E" box="[151,210,1237,1263]" pageId="18" pageNumber="19">Type</typeStatus>
|
||
locality:
|
||
</emphasis>
|
||
<location id="8EC3605BB640FFD6FEA9FB7CFDE1728E" LSID="urn:lsid:plazi:treatment:03B58796B640FFD2FF7EFC24FF5D75E2:8EC3605BB640FFD6FEA9FB7CFDE1728E" box="[320,518,1237,1263]" country="Peru" name="Marcapata Valley" pageId="18" pageNumber="19" stateProvince="Cusco">Marcapata Valley</location>
|
||
,
|
||
<collectingRegion id="49D8F862B640FFD6FDF8FB7CFDBC728E" box="[529,603,1237,1263]" country="Peru" name="Cusco" pageId="18" pageNumber="19">Cuzco</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30B7610B640FFD6FD8FFB7CFD7A728E" box="[614,669,1237,1263]" name="Peru" pageId="18" pageNumber="19">Perú</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B640FFD6FF2EFB51FE1E733A" blockId="18.[151,1437,1236,2019]" pageId="18" pageNumber="19">
|
||
<emphasis id="B968EA92B640FFD6FF2EFB51FCFA7372" bold="true" box="[199,797,1272,1299]" pageId="18" pageNumber="19">
|
||
Examined material (
|
||
<specimenCount id="9D1AFD09B640FFD6FE2DFB51FDCA7372" box="[452,557,1272,1299]" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
|
||
,
|
||
<specimenCount id="9D1AFD09B640FFD6FDD5FB50FD777372" box="[572,656,1273,1299]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D1AFD09B640FFD6FD49FB50FCF77372" box="[672,784,1272,1299]" pageId="18" pageNumber="19" type="juvenile">1 juvenil</specimenCount>
|
||
):
|
||
</emphasis>
|
||
<materialsCitation id="3B743CDDB640FFD6FCCEFB51FE12733A" collectingDate="2015-10-01" collectionCode="MZUSP" collectorName="J. Cabra & B. Portuguez" country="Peru" elevation="360" latitude="-10.322917" location="Paujil" longLatPrecision="1" longitude="-75.26241" municipality="Parque Nacional Yanachaga-Chemillen" pageId="18" pageNumber="19" specimenCode="MZUSP 74571" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Pasco">
|
||
<collectingCountry id="F30B7610B640FFD6FCCEFB51FC947372" box="[807,883,1272,1299]" name="Peru" pageId="18" pageNumber="19">PERÚ</collectingCountry>
|
||
:
|
||
<emphasis id="B968EA92B640FFD6FC68FB53FC227372" box="[897,965,1274,1299]" italics="true" pageId="18" pageNumber="19">
|
||
<collectingRegion id="49D8F862B640FFD6FC68FB53FC227372" box="[897,965,1274,1299]" country="Peru" name="Pasco" pageId="18" pageNumber="19">Pasco</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingMunicipality id="6BC7ACFAB640FFD6FC3EFB50FA7E7372" box="[983,1433,1273,1299]" pageId="18" pageNumber="19">Parque Nacional Yanachaga-Chemillén</collectingMunicipality>
|
||
,
|
||
<location id="8EC3605BB640FFD6FF7EFAB4FF3F7356" LSID="urn:lsid:plazi:treatment:03B58796B640FFD2FF7EFC24FF5D75E2:8EC3605BB640FFD6FF7EFAB4FF3F7356" box="[151,216,1309,1335]" country="Peru" latitude="-10.322917" longLatPrecision="1" longitude="-75.26241" municipality="Parque Nacional Yanachaga-Chemillen" name="Paujil" pageId="18" pageNumber="19" stateProvince="Pasco">Paujil</location>
|
||
(
|
||
<geoCoordinate id="EE285047B640FFD6FF01FAB5FE6C7356" box="[232,395,1308,1335]" degrees="10" direction="south" minutes="19" orientation="latitude" pageId="18" pageNumber="19" precision="1" seconds="22.5" value="-10.322917">10°19´22.5´´S</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE285047B640FFD6FE7EFAB5FDA07356" box="[407,583,1308,1335]" degrees="75" direction="west" minutes="15" orientation="longitude" pageId="18" pageNumber="19" precision="1" seconds="44.7" value="-75.26241">75°15´44.7´´W</geoCoordinate>
|
||
):
|
||
<quantity id="4CE49B65B640FFD6FDB0FAB4FD257356" box="[601,706,1309,1335]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="18" pageNumber="19" unit="m" value="360.0">
|
||
<elevation id="0031D1B3B640FFD6FDB0FAB4FD257356" box="[601,706,1309,1335]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="18" pageNumber="19" unit="m" value="360.0">360 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA21040B640FFD6FD24FAB4FC997356" box="[717,894,1308,1335]" pageId="18" pageNumber="19" value="2015-10-01">
|
||
<collectingDate id="EFE6E9A8B640FFD6FD24FAB4FC997356" box="[717,894,1308,1335]" pageId="18" pageNumber="19" value="2015-10-01">1 October 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26E95356B640FFD6FC61FAB4FC007356" box="[904,999,1309,1335]" pageId="18" pageNumber="19">J. Cabra</collectorName>
|
||
,
|
||
<collectorName id="26E95356B640FFD6FC1BFAB4FB617356" box="[1010,1158,1309,1335]" pageId="18" pageNumber="19">B. Portuguéz</collectorName>
|
||
, three males,
|
||
<specimenCount id="9D1AFD09B640FFD6FAF6FAB4FA647356" box="[1311,1411,1309,1335]" pageId="18" pageNumber="19" type="female">1 female</specimenCount>
|
||
,
|
||
<specimenCount id="9D1AFD09B640FFD6FA67FAB4FF2A733A" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
|
||
juvenile (
|
||
<specimenCode id="DBBA9EFBB640FFD6FED4FAE8FE0A733A" box="[317,493,1344,1371]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="18" pageNumber="19">MZUSP 74571</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306650BB640FFD6FF2EFACDFE08738A" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="8BA33680B640FFD6FF2EFACDFE08738A" blockId="18.[151,1437,1236,2019]" pageId="18" pageNumber="19">
|
||
<emphasis id="B968EA92B640FFD6FF2EFACDFE84731E" bold="true" box="[199,355,1380,1407]" pageId="18" pageNumber="19">Distribution:</emphasis>
|
||
Countries:
|
||
<collectingCountry id="F30B7610B640FFD6FE04FACCFDA5731E" box="[493,578,1381,1407]" name="Bolivia" pageId="18" pageNumber="19">Bolivia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F30B7610B640FFD6FDB8FACCFD6F731E" box="[593,648,1381,1407]" name="Peru" pageId="18" pageNumber="19">Perú</collectingCountry>
|
||
. Other records from:
|
||
<collectingCountry id="F30B7610B640FFD6FC6FFACCFC59731E" box="[902,958,1381,1407]" name="Peru" pageId="18" pageNumber="19">Peru</collectingCountry>
|
||
:
|
||
<bibRefCitation id="EF8D4B71B640FFD6FC25FACCFAC1731E" author="Igelmund, P. & Wendler, G." box="[972,1318,1381,1407]" pageId="18" pageNumber="19" pagination="63 - 73" refId="ref22862" refString="Igelmund, P. & Wendler, G. (1991 a) The giant fibre system in the forelegs (whips) of the whip spider Heterophrynus elaphus Pocock (Arachnida: Amblypygi). Journal of Comparative Physiology, A, 168, 63 - 73. https: // doi. org / 10.1007 / BF 00217104" type="journal article" year="1991">Igelmund & Wendler (1991a)</bibRefCitation>
|
||
: Pucallpa;
|
||
<bibRefCitation id="EF8D4B71B640FFD6FF7EFA20FEA473C2" author="Roewer, C. F." box="[151,323,1416,1443]" pageId="18" pageNumber="19" pagination="37 - 58" refId="ref23693" refString="Roewer, C. F. (1952) Neotropische Arachnida Arthrogastra, zumeist aus Peru. Senckenbergiana Biologica, 33, 37 - 58." type="journal article" year="1952">Roewer (1952)</bibRefCitation>
|
||
: Tingo María;
|
||
<bibRefCitation id="EF8D4B71B640FFD6FE01FA20FD7173C2" author="Roewer, C. F." box="[488,662,1416,1443]" pageId="18" pageNumber="19" pagination="67 - 94" refId="ref23744" refString="Roewer, C. F. (1957) Arachnida Arthrogastra aus Peru, III. Senckenbergiana Biologica, 38, 67 - 94." type="journal article" year="1957">Roewer (1957)</bibRefCitation>
|
||
: Cascas, near to Trujillo; La Merced; Saposoa; Puente Cayumba: Rio Huallaga; Tingo Maria: Cueva de Las Lechuzas; Ballón-Estacio & Armas (2019):
|
||
<collectingRegion id="49D8F862B640FFD6FBDEFA04FB6773A6" box="[1079,1152,1453,1479]" country="Peru" name="Cusco" pageId="18" pageNumber="19">Cusco</collectingRegion>
|
||
,
|
||
<collectingRegion id="49D8F862B640FFD6FB65FA04FAD173A6" box="[1164,1334,1453,1479]" country="Peru" name="Madre de Dios" pageId="18" pageNumber="19">Madre de Dios</collectingRegion>
|
||
.
|
||
<collectingCountry id="F30B7610B640FFD6FAABFA04FA7E73A6" box="[1346,1433,1453,1479]" name="Bolivia" pageId="18" pageNumber="19">Bolivia</collectingCountry>
|
||
:
|
||
<bibRefCitation id="EF8D4B71B640FFD6FF7EFA78FE7A738A" author="Reveillion, F. & Olivier, P. & Hofer, H." box="[151,413,1489,1515]" pageId="18" pageNumber="19" pagination="102" refId="ref23649" refString="Reveillion, F., Olivier, P. & Hofer, H. (2014) First record of Heterophrynus elaphus (Pocock, 1903) (Amblypygi: Phrynidae) for Bolivia. Revista Iberica de Aracnologia, 25, 102." type="journal article" year="2014">
|
||
Réveillion
|
||
<emphasis id="B968EA92B640FFD6FEFAFA7BFEAC738A" box="[275,331,1489,1515]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
:
|
||
<collectingRegion id="49D8F862B640FFD6FE5BFA78FE0D738A" box="[434,490,1489,1515]" country="Bolivia" name="Beni" pageId="18" pageNumber="19">Beni</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306650BB640FFD2FF2EFA5DFAE777EA" lastPageId="22" lastPageNumber="23" pageId="18" pageNumber="19" type="diagnosis">
|
||
<paragraph id="8BA33680B640FFD6FF2EFA5DFD9C70FE" blockId="18.[151,1437,1236,2019]" pageId="18" pageNumber="19">
|
||
<emphasis id="B968EA92B640FFD6FF2EFA5DFEA5706E" bold="true" box="[199,322,1524,1551]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C1C4D03B640FFD6FEA1FA5CFDAA706E" authorityName="Pocock" authorityYear="1903" box="[328,589,1525,1551]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elaphus">
|
||
<emphasis id="B968EA92B640FFD6FEA1FA5CFDAA706E" box="[328,589,1525,1551]" italics="true" pageId="18" pageNumber="19">Heterophrynus elaphus</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from other species by present the following characters together: three teeth in the ectal portion of the basal segment of the chelicerae. Six spines in the dorsal
|
||
<taxonomicName id="4C1C4D03B640FFD6FACDF9B0FA947052" box="[1316,1395,1561,1587]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="18" pageNumber="34" phylum="Mollusca" rank="genus">Patella</taxonomicName>
|
||
, P7 and PIII are absent. Four long spines in the ventral femur, spine FIII is smaller than FII and longer than FV, FIV is absent. Female genitalia with gonopod sclerites with claw-like shape, a little curved at the distal portion. Seminal receptacles with a square shape (
|
||
<figureCitation id="13272A05B640FFD6FDEFF92CFD8970FE" box="[518,622,1669,1695]" captionStart="FIGURE 13" captionStartId="21.[151,250,1904,1929]" captionTargetBox="[163,1425,747,1874]" captionTargetId="figure@21.[157,1431,741,1879]" captionTargetPageId="21" captionText="FIGURE 13. Male genital: Heterophrynus elaphus. (MZUSP 74571): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MZUSP 74571), white arrow indicates the sclerotized area of the seminal receptacle,), black arrow indicates an extra sclerite at the base of gonopod." figureDoi="http://doi.org/10.5281/zenodo.3907928" httpUri="https://zenodo.org/record/3907928/files/figure.png" pageId="18" pageNumber="19">Fig. 13D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B640FFD6FF2EF900FB1F7086" blockId="18.[151,1437,1236,2019]" pageId="18" pageNumber="19">
|
||
Similar species: the more similar species is
|
||
<emphasis id="B968EA92B640FFD6FD5AF903FC9C70A2" box="[691,891,1705,1731]" italics="true" pageId="18" pageNumber="19">
|
||
<taxonomicName id="4C1C4D03B640FFD6FD5AF903FC9070A2" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1931" box="[691,887,1705,1731]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cheiracanthus">H. cheiracanthus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
however, this species present the spines P7, and PIII. The female genital differs by presenting gonopod sclerites shorter and more curved (
|
||
<figureCitation id="13272A05B640FFD6FB6AF964FB0B7086" box="[1155,1260,1741,1767]" captionStart="FIGURE 10" captionStartId="17.[151,250,1541,1566]" captionTargetBox="[157,1429,351,1508]" captionTargetId="figure@17.[151,1436,345,1516]" captionTargetPageId="17" captionText="FIGURE 10. Male genital: Heterophrynus cheiracanthus. (MZUSP 15859):A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MZUSP 9459), black arrow indicates the sclerotized area of the seminal receptacle." figureDoi="http://doi.org/10.5281/zenodo.3907922" httpUri="https://zenodo.org/record/3907922/files/figure.png" pageId="18" pageNumber="19">Fig. 10D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B640FFD6FF2EF958FC97714E" blockId="18.[151,1437,1236,2019]" pageId="18" pageNumber="19">
|
||
Male genital of
|
||
<taxonomicName id="4C1C4D03B640FFD6FE9EF95BFE17716A" authorityName="Pocock" authorityYear="1903" box="[375,496,1777,1803]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elaphus">
|
||
<emphasis id="B968EA92B640FFD6FE9EF95BFE17716A" box="[375,496,1777,1803]" italics="true" pageId="18" pageNumber="19">H. elaphus</emphasis>
|
||
</taxonomicName>
|
||
presents a Pi, LaM, and LoL 1 with a very similar length and an LoD very elongated and wide, which differs from the other species (
|
||
<figureCitation id="13272A05B640FFD6FD44F8BCFCE7714E" box="[685,768,1813,1839]" captionStart="FIGURE 13" captionStartId="21.[151,250,1904,1929]" captionTargetBox="[163,1425,747,1874]" captionTargetId="figure@21.[157,1431,741,1879]" captionTargetPageId="21" captionText="FIGURE 13. Male genital: Heterophrynus elaphus. (MZUSP 74571): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MZUSP 74571), white arrow indicates the sclerotized area of the seminal receptacle,), black arrow indicates an extra sclerite at the base of gonopod." figureDoi="http://doi.org/10.5281/zenodo.3907928" httpUri="https://zenodo.org/record/3907928/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
|
||
, A, B, C).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B640FFD6FF2EF891FA7F7116" blockId="18.[151,1437,1236,2019]" pageId="18" pageNumber="19">
|
||
<emphasis id="B968EA92B640FFD6FF2EF891FCB27132" bold="true" box="[199,853,1848,1875]" pageId="18" pageNumber="19">Complementary description. Female (MZUSP 74571):</emphasis>
|
||
Total length
|
||
<quantity id="4CE49B65B640FFD6FC19F890FBB67132" box="[1008,1105,1849,1875]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="18" pageNumber="19" unit="mm" value="32.4">32.4mm</quantity>
|
||
. Carapace, opisthosoma, and legs have a brown coloration, pedipalps brown with red tonalities. Measures of the female are provided in
|
||
<tableCitation id="C69E033BB640FFD6FAD6F8F4FA737116" box="[1343,1428,1885,1911]" captionStart="TABLE 4" captionStartId="22.[151,239,946,970]" captionText="TABLE 4. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus elaphus. Empty cells correspond to missing segment or data." pageId="18" pageNumber="19">Table 4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B640FFD6FF2EF828FDEF7182" blockId="18.[151,1437,1236,2019]" pageId="18" pageNumber="19">
|
||
<emphasis id="B968EA92B640FFD6FF2EF828FEDB71FA" bold="true" box="[199,316,1921,1947]" italics="true" pageId="18" pageNumber="19">Carapace.</emphasis>
|
||
Anterior margin straight without lobes, carapace surface with a widely spaced granulation, posterior margin of with a concave medial region. Lateral and anteromedial eyes clearly visible, anteromedial ocular tubercle with black coloration (
|
||
<figureCitation id="13272A05B640FFD6FE7DF860FE1B7182" box="[404,508,1993,2019]" captionStart="FIGURE 11" captionStartId="19.[151,250,1790,1815]" captionTargetBox="[161,1426,182,1752]" captionTargetId="figure@19.[151,1436,181,1765]" captionTargetPageId="19" captionText="FIGURE 11. Heterophrynus elaphus. Female (MZUSP 74571): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907924" httpUri="https://zenodo.org/record/3907924/files/figure.png" pageId="18" pageNumber="19">Fig. 11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF636608B641FFD7FF7EF957FD2A715A" ID-DOI="http://doi.org/10.5281/zenodo.3907924" ID-Zenodo-Dep="3907924" httpUri="https://zenodo.org/record/3907924/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1790,1815]" targetBox="[161,1426,182,1752]" targetPageId="19">
|
||
<paragraph id="8BA33680B641FFD7FF7EF957FD2A715A" blockId="19.[151,1437,1790,1851]" pageId="19" pageNumber="20">
|
||
<emphasis id="B968EA92B641FFD7FF7EF957FEF97176" bold="true" box="[151,286,1790,1815]" pageId="19" pageNumber="20">FIGURE 11.</emphasis>
|
||
<taxonomicName id="4C1C4D03B641FFD7FECAF956FDE97176" authorityName="Pocock" authorityYear="1903" box="[291,526,1791,1815]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="elaphus">
|
||
<emphasis id="B968EA92B641FFD7FECAF956FDE97176" box="[291,526,1791,1815]" italics="true" pageId="19" pageNumber="20">Heterophrynus elaphus</emphasis>
|
||
</taxonomicName>
|
||
. Female (MZUSP 74571): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA33680B641FFD7FF2EF8C0FDAA71AD" blockId="19.[151,1436,1897,2032]" pageId="19" pageNumber="20">
|
||
<emphasis id="B968EA92B641FFD7FF2EF8C0FED671E2" bold="true" box="[199,305,1897,1923]" italics="true" pageId="19" pageNumber="20">Sternum.</emphasis>
|
||
Tri-segmented, segments are well sclerotized, the area around segments is also sclerotized. Tritosternum projected anteriorly, elongated, conical, with 30 setae in the basal region, eighth in the medial region,
|
||
<quantity id="4CE49B65B641FFD7FA8AF824FA7B71C6" box="[1379,1436,1933,1959]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="19" pageNumber="20" unit="in" value="11.0">11 in</quantity>
|
||
distal region and two more in the apex.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B641FFD0FF2EF87CFE8A76B6" blockId="19.[151,1436,1897,2032]" lastBlockId="20.[151,1436,153,431]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||
The second segment (tetrasternum) oval and with three setae on medial region. The third segment (penta- sternum) rounded, smaller than the second segment, has two setae on medial region. Metasternum longitudinally divided (
|
||
<figureCitation id="13272A05B646FFD0FF12FF14FE8776B6" box="[251,352,189,215]" captionStart="FIGURE 11" captionStartId="19.[151,250,1790,1815]" captionTargetBox="[161,1426,182,1752]" captionTargetId="figure@19.[151,1436,181,1765]" captionTargetPageId="19" captionText="FIGURE 11. Heterophrynus elaphus. Female (MZUSP 74571): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907924" httpUri="https://zenodo.org/record/3907924/files/figure.png" pageId="20" pageNumber="21">Fig. 11B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B646FFD0FF2EFF49FAA0769A" blockId="20.[151,1436,153,431]" box="[199,1351,224,251]" pageId="20" pageNumber="21">
|
||
<emphasis id="B968EA92B646FFD0FF2EFF49FEDB769A" bold="true" box="[199,316,224,251]" italics="true" pageId="20" pageNumber="21">Abdomen.</emphasis>
|
||
Oblong, color brown, with soft punctuations. Carapace wider than the Abdomen (
|
||
<figureCitation id="13272A05B646FFD0FB3CFF48FADD769A" box="[1237,1338,225,251]" captionStart="FIGURE 11" captionStartId="19.[151,250,1790,1815]" captionTargetBox="[161,1426,182,1752]" captionTargetId="figure@19.[151,1436,181,1765]" captionTargetPageId="19" captionText="FIGURE 11. Heterophrynus elaphus. Female (MZUSP 74571): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907924" httpUri="https://zenodo.org/record/3907924/files/figure.png" pageId="20" pageNumber="21">Fig. 11C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B646FFD0FF2EFEADFAEB77CE" blockId="20.[151,1436,153,431]" pageId="20" pageNumber="21">
|
||
<emphasis id="B968EA92B646FFD0FF2EFEADFED9777E" bold="true" box="[199,318,260,287]" italics="true" pageId="20" pageNumber="21">Chelicera.</emphasis>
|
||
Presents a mesal row of three teeth on the basal segment of the chelicerae with the first in the proximal region, with two cuspids, follow by one tooth shorter in the medial portion, and the third bigger than others placed in the distal region (
|
||
<figureCitation id="13272A05B646FFD0FE2AFEE4FDCF7706" box="[451,552,333,359]" captionStart="FIGURE 11" captionStartId="19.[151,250,1790,1815]" captionTargetBox="[161,1426,182,1752]" captionTargetId="figure@19.[151,1436,181,1765]" captionTargetPageId="19" captionText="FIGURE 11. Heterophrynus elaphus. Female (MZUSP 74571): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907924" httpUri="https://zenodo.org/record/3907924/files/figure.png" pageId="20" pageNumber="21">Fig. 11D</figureCitation>
|
||
). The ectal row of the basal segment of the chelicerae have three teeth, two with acuminated apex placed in the proximal portions, fused at the base (tooth 1 and 2), and one more with acuminate apex placed on the medial portion (tooth 3) (
|
||
<figureCitation id="13272A05B646FFD0FD62FE3CFD0977CE" box="[651,750,405,431]" captionStart="FIGURE 11" captionStartId="19.[151,250,1790,1815]" captionTargetBox="[161,1426,182,1752]" captionTargetId="figure@19.[151,1436,181,1765]" captionTargetPageId="19" captionText="FIGURE 11. Heterophrynus elaphus. Female (MZUSP 74571): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907924" httpUri="https://zenodo.org/record/3907924/files/figure.png" pageId="20" pageNumber="21">Fig. 11F</figureCitation>
|
||
). Mobile finger of the chelicerae with five teeth.
|
||
</paragraph>
|
||
<caption id="DF636608B646FFD0FF7EF8DAFCBA71D1" ID-DOI="http://doi.org/10.5281/zenodo.3907926" ID-Zenodo-Dep="3907926" httpUri="https://zenodo.org/record/3907926/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1907,1932]" targetBox="[165,1426,465,1872]" targetPageId="20">
|
||
<paragraph id="8BA33680B646FFD0FF7EF8DAFCBA71D1" blockId="20.[151,1437,1907,1968]" pageId="20" pageNumber="21">
|
||
<emphasis id="B968EA92B646FFD0FF7EF8DAFEF871ED" bold="true" box="[151,287,1907,1932]" pageId="20" pageNumber="21">FIGURE 12.</emphasis>
|
||
<emphasis id="B968EA92B646FFD0FECCF8DDFDFE71ED" box="[293,537,1908,1932]" italics="true" pageId="20" pageNumber="21">
|
||
<taxonomicName id="4C1C4D03B646FFD0FECCF8DDFDF371ED" authorityName="Pocock" authorityYear="1903" box="[293,532,1908,1932]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="elaphus">Heterophrynus elaphus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Female (MZUSP 74571): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral
|
||
<taxonomicName id="4C1C4D03B646FFD0FAB0F8DAFF2171D1" authority="(Pv)" baseAuthorityName="Pv" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="20" pageNumber="34" phylum="Mollusca" rank="species" species="patella">patella (Pv)</taxonomicName>
|
||
. D. Dorsal
|
||
<taxonomicName id="4C1C4D03B646FFD0FED1F83EFE5671D1" authority="(Pd)" baseAuthorityName="Pd" box="[312,433,1943,1968]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="20" pageNumber="34" phylum="Mollusca" rank="species" species="patella">patella (Pd)</taxonomicName>
|
||
. E. Ventral tibia (Tv). F. Dorsal tibia (Td).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA33680B647FFD1FF2EFF31FC1974AE" blockId="21.[151,1437,152,719]" pageId="21" pageNumber="22">
|
||
<emphasis id="B968EA92B647FFD1FF2EFF31FE5F76D2" box="[199,440,152,179]" italics="true" pageId="21" pageNumber="22">
|
||
<emphasis id="B968EA92B647FFD1FF2EFF31FED476D2" bold="true" box="[199,307,152,179]" italics="true" pageId="21" pageNumber="22">Pedipalp.</emphasis>
|
||
Trochanter
|
||
</emphasis>
|
||
: Prolateral face with four spines; spines Tr1 and Tr3 placed in the medial region, Tr2 placed near to ventral margin. Tr3 and Tr4 with subequal lengths. Spines lengths: Tr2>Tr1>Tr3≥Tr4. Dorsal oblique series of nine setal tubercles. The dorsomedial area without spines but with three tubercles (
|
||
<figureCitation id="13272A05B647FFD1FB7EFF48FB1C769A" box="[1175,1275,225,251]" captionStart="FIGURE 11" captionStartId="19.[151,250,1790,1815]" captionTargetBox="[161,1426,182,1752]" captionTargetId="figure@19.[151,1436,181,1765]" captionTargetPageId="19" captionText="FIGURE 11. Heterophrynus elaphus. Female (MZUSP 74571): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.3907924" httpUri="https://zenodo.org/record/3907924/files/figure.png" pageId="21" pageNumber="22">Fig. 11E</figureCitation>
|
||
).
|
||
<emphasis id="B968EA92B647FFD1FAE6FF4BFABD769A" box="[1295,1370,226,251]" italics="true" pageId="21" pageNumber="22">Femur</emphasis>
|
||
: Ventral face with four major spines, with four tubercles between FII–FIII, seven between FIII–FV, and 11 more distally than FV, FIV is absent. Spine lengths: FI>FII>FIII>FV (
|
||
<figureCitation id="13272A05B647FFD1FCF2FE80FC627722" box="[795,901,297,323]" captionStart="FIGURE 12" captionStartId="20.[151,250,1907,1932]" captionTargetBox="[165,1426,465,1872]" captionTargetId="figure@20.[151,1436,453,1882]" captionTargetPageId="20" captionText="FIGURE 12. Heterophrynus elaphus. Female (MZUSP 74571): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907926" httpUri="https://zenodo.org/record/3907926/files/figure.png" pageId="21" pageNumber="22">Fig. 12A</figureCitation>
|
||
). Dorsal face with five major spines, with four tubercles between F3–F4, five between F4–F5, and 11 more distally than F5. F1 and F2 share the same base and are clearly separated from F3. Spine lengths: F3>F4>F1=F2>F5 (
|
||
<figureCitation id="13272A05B647FFD1FC97FED8FC0177EA" box="[894,998,369,395]" captionStart="FIGURE 12" captionStartId="20.[151,250,1907,1932]" captionTargetBox="[165,1426,465,1872]" captionTargetId="figure@20.[151,1436,453,1882]" captionTargetPageId="20" captionText="FIGURE 12. Heterophrynus elaphus. Female (MZUSP 74571): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907926" httpUri="https://zenodo.org/record/3907926/files/figure.png" pageId="21" pageNumber="22">Fig. 12B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1C4D03B647FFD1FC12FED8FB4B77EA" authority=": Ventral" authorityName="Ventral" box="[1019,1196,369,395]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EA92B647FFD1FC12FED8FBAB77EA" box="[1019,1100,369,395]" italics="true" pageId="21" pageNumber="22">Patella</emphasis>
|
||
: Ventral
|
||
</taxonomicName>
|
||
face with four major spines, PVI and PIII are absent, six tubercles more proximally than PV, four between PV–PIV, six between PIV–PII, five between PII–PI, PI wider than the other spines. Spine lengths: PII>PIV>PV>PI (
|
||
<figureCitation id="13272A05B647FFD1FBAFFE10FB4B77B2" box="[1094,1196,441,467]" captionStart="FIGURE 12" captionStartId="20.[151,250,1907,1932]" captionTargetBox="[165,1426,465,1872]" captionTargetId="figure@20.[151,1436,453,1882]" captionTargetPageId="20" captionText="FIGURE 12. Heterophrynus elaphus. Female (MZUSP 74571): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907926" httpUri="https://zenodo.org/record/3907926/files/figure.png" pageId="21" pageNumber="22">Fig. 12C</figureCitation>
|
||
). Dorsal face with six major spines, P7 is absent, with six tubercles more proximally than P6 and five between P6 and P5. Spine lengths: P5=P4>P3>P6>P1>P2 (
|
||
<figureCitation id="13272A05B647FFD1FE4FFDA8FDEA747A" box="[422,525,513,539]" captionStart="FIGURE 12" captionStartId="20.[151,250,1907,1932]" captionTargetBox="[165,1426,465,1872]" captionTargetId="figure@20.[151,1436,453,1882]" captionTargetPageId="20" captionText="FIGURE 12. Heterophrynus elaphus. Female (MZUSP 74571): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907926" httpUri="https://zenodo.org/record/3907926/files/figure.png" pageId="21" pageNumber="22">Fig. 12D</figureCitation>
|
||
).
|
||
<emphasis id="B968EA92B647FFD1FDC9FDA8FDBD747A" box="[544,602,513,539]" italics="true" pageId="21" pageNumber="22">Tibia</emphasis>
|
||
: Ventral face with two major spines, five very small tubercles more distally than TII. Spine lengths: TII>TI (
|
||
<figureCitation id="13272A05B647FFD1FE17FD8CFD84745E" box="[510,611,549,575]" captionStart="FIGURE 12" captionStartId="20.[151,250,1907,1932]" captionTargetBox="[165,1426,465,1872]" captionTargetId="figure@20.[151,1436,453,1882]" captionTargetPageId="20" captionText="FIGURE 12. Heterophrynus elaphus. Female (MZUSP 74571): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907926" httpUri="https://zenodo.org/record/3907926/files/figure.png" pageId="21" pageNumber="22">Fig. 12E</figureCitation>
|
||
). Dorsal face with two major spines, six very small tubercles more distally than T2. Spine lengths: T2>T1 (
|
||
<figureCitation id="13272A05B647FFD1FE17FDE0FD857402" box="[510,610,585,611]" captionStart="FIGURE 12" captionStartId="20.[151,250,1907,1932]" captionTargetBox="[165,1426,465,1872]" captionTargetId="figure@20.[151,1436,453,1882]" captionTargetPageId="20" captionText="FIGURE 12. Heterophrynus elaphus. Female (MZUSP 74571): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.3907926" httpUri="https://zenodo.org/record/3907926/files/figure.png" pageId="21" pageNumber="22">Fig. 12F</figureCitation>
|
||
).
|
||
<emphasis id="B968EA92B647FFD1FD9FFDE3FCAA7402" box="[630,845,586,611]" italics="true" pageId="21" pageNumber="22">Tarsus-metatarsus:</emphasis>
|
||
Internal face with the secondary line of dorsomedial bristles complete, the suture between the tarsus and the apotele is not visible.
|
||
<emphasis id="B968EA92B647FFD1FC1AFDC4FBCE74E6" bold="true" box="[1011,1065,621,647]" italics="true" pageId="21" pageNumber="22">Legs</emphasis>
|
||
. Femora brown. Femora lengths: I>II>III>IV (
|
||
<tableCitation id="C69E033BB647FFD1FEC2FD38FE6474CA" box="[299,387,657,683]" captionStart="TABLE 4" captionStartId="22.[151,239,946,970]" captionText="TABLE 4. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus elaphus. Empty cells correspond to missing segment or data." pageId="21" pageNumber="22">Table 4</tableCitation>
|
||
). Leg I: Tibia with 55 segments and tarsus with 97 segments in the right leg; in left leg, tibia with 26 segments and tarsus is broken. Leg IV: Basitibia with three segments.
|
||
</paragraph>
|
||
<caption id="DF636608B647FFD1FF7EF8D9FB2A7194" ID-DOI="http://doi.org/10.5281/zenodo.3907928" ID-Zenodo-Dep="3907928" httpUri="https://zenodo.org/record/3907928/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1904,1929]" targetBox="[163,1425,747,1874]" targetPageId="21">
|
||
<paragraph id="8BA33680B647FFD1FF7EF8D9FB2A7194" blockId="21.[151,1437,1904,2037]" pageId="21" pageNumber="22">
|
||
<emphasis id="B968EA92B647FFD1FF7EF8D9FEC571E8" bold="true" box="[151,290,1904,1929]" pageId="21" pageNumber="22">FIGURE 13.</emphasis>
|
||
Male genital:
|
||
<emphasis id="B968EA92B647FFD1FE52F8D9FD5671E9" box="[443,689,1904,1928]" italics="true" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C1C4D03B647FFD1FE52F8D9FD4A71E9" authorityName="Pocock" authorityYear="1903" box="[443,685,1904,1928]" class="Arachnida" family="Phrynidae" genus="Heterophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="elaphus">Heterophrynus elaphus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
(MZUSP 74571): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi =
|
||
<emphasis id="B968EA92B647FFD1FE0FF83DFDC271CD" box="[486,549,1940,1964]" italics="true" pageId="21" pageNumber="22">fistula</emphasis>
|
||
; LaM =
|
||
<emphasis id="B968EA92B647FFD1FD6BF83DFCC271CD" box="[642,805,1940,1964]" italics="true" pageId="21" pageNumber="22">lamina medialis</emphasis>
|
||
; LoD =
|
||
<emphasis id="B968EA92B647FFD1FC97F83DFBE971CD" box="[894,1038,1940,1964]" italics="true" pageId="21" pageNumber="22">lobus dorsalis</emphasis>
|
||
; LoL 1=
|
||
<emphasis id="B968EA92B647FFD1FB99F83DFAA871CD" box="[1136,1359,1940,1964]" italics="true" pageId="21" pageNumber="22">lobus lateralis primus</emphasis>
|
||
; LoL 2 =
|
||
<emphasis id="B968EA92B647FFD1FF45F811FE4571B1" box="[172,418,1976,2000]" italics="true" pageId="21" pageNumber="22">lobus lateralis secundus</emphasis>
|
||
; Pi =
|
||
<emphasis id="B968EA92B647FFD1FE08F810FD7971B1" box="[481,670,1977,2000]" italics="true" pageId="21" pageNumber="22">processus internus</emphasis>
|
||
of gonopod. D. Female genital (MZUSP 74571), white arrow indicates the sclerotized area of the seminal receptacle,), black arrow indicates an extra sclerite at the base of gonopod.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA33680B644FFD2FF2EFF30FA9F76D2" blockId="22.[151,1437,153,899]" box="[199,1400,153,179]" pageId="22" pageNumber="23">Basitibia-distitibia lengths: BT1>DT>BT3>BT2. Basitarsus shorter than the telotarsus. Tarsus tetramerous.</paragraph>
|
||
<paragraph id="8BA33680B644FFD2FF2EFF15FAE777EA" blockId="22.[151,1437,153,899]" pageId="22" pageNumber="23">
|
||
<emphasis id="B968EA92B644FFD2FF2EFF15FE7676B6" bold="true" box="[199,401,188,215]" italics="true" pageId="22" pageNumber="23">Female genitalia.</emphasis>
|
||
Genital operculum pentagonal, reaching the second segment of the opisthosoma, seminal receptacles with a rectangular shape, separated at the medial portion, dorsal surface sclerotized at the proximal region and with an additional sclerotized layer around the base of the sclerite of the gonopod, basal region with orange coloration, medial and distal portion white. Sclerites of the gonopod with claw-like shape, basal portion with some transversal grooves, distal portion a little curved, with rounded apex, sclerites placed parallel, longer than the width of the seminal receptacle, sclerites cover almost the total length of the seminal receptacle (
|
||
<figureCitation id="13272A05B644FFD2FB62FED8FB1477EA" box="[1163,1267,369,395]" captionStart="FIGURE 13" captionStartId="21.[151,250,1904,1929]" captionTargetBox="[163,1425,747,1874]" captionTargetId="figure@21.[157,1431,741,1879]" captionTargetPageId="21" captionText="FIGURE 13. Male genital: Heterophrynus elaphus. (MZUSP 74571): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MZUSP 74571), white arrow indicates the sclerotized area of the seminal receptacle,), black arrow indicates an extra sclerite at the base of gonopod." figureDoi="http://doi.org/10.5281/zenodo.3907928" httpUri="https://zenodo.org/record/3907928/files/figure.png" pageId="22" pageNumber="23">Fig. 13D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306650BB644FFD2FF2EFE3DFF5D75E2" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="8BA33680B644FFD2FF2EFE3DFA7B77CE" blockId="22.[151,1437,153,899]" box="[199,1436,404,431]" pageId="22" pageNumber="23">
|
||
<emphasis id="B968EA92B644FFD2FF2EFE3DFED877CE" bold="true" box="[199,319,404,431]" pageId="22" pageNumber="23">Variation.</emphasis>
|
||
Variation in measures and segments of the basitibia IV, tibia I and tarsus I, are provided in
|
||
<tableCitation id="C69E033BB644FFD2FAABFE3CFA7F77CE" box="[1346,1432,405,431]" captionStart="TABLE 4" captionStartId="22.[151,239,946,970]" captionText="TABLE 4. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus elaphus. Empty cells correspond to missing segment or data." pageId="22" pageNumber="23">Table 4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B644FFD2FF7EFE10FD7B74CA" blockId="22.[151,1437,153,899]" pageId="22" pageNumber="23">
|
||
<emphasis id="B968EA92B644FFD2FF7EFE10FEE577B2" bold="true" box="[151,258,441,467]" pageId="22" pageNumber="23">
|
||
<emphasis id="B968EA92B644FFD2FF7EFE10FF1C77B2" bold="true" box="[151,251,441,467]" italics="true" pageId="22" pageNumber="23">Sternum</emphasis>
|
||
.
|
||
</emphasis>
|
||
The number of setae over all segments of the sternum was variable.
|
||
<emphasis id="B968EA92B644FFD2FBFFFE11FB9C77B2" bold="true" box="[1046,1147,440,467]" italics="true" pageId="22" pageNumber="23">Pedipalp</emphasis>
|
||
.
|
||
<emphasis id="B968EA92B644FFD2FB62FE10FAE177B2" box="[1163,1286,441,467]" italics="true" pageId="22" pageNumber="23">Trochanter</emphasis>
|
||
: line of setal tubercles vary between seven and nine.
|
||
<emphasis id="B968EA92B644FFD2FDBFFE77FD467796" box="[598,673,478,503]" italics="true" pageId="22" pageNumber="23">Femur</emphasis>
|
||
: Ventral and Dorsal face: number of tubercles between the spines is variable; tubercles can be more conspicuous and, in some cases, can look as small spines.
|
||
<taxonomicName id="4C1C4D03B644FFD2FB6AFDA8FB33747A" box="[1155,1236,513,539]" class="Gastropoda" family="Patellidae" genus="Patella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pezizales" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EA92B644FFD2FB6AFDA8FB33747A" box="[1155,1236,513,539]" italics="true" pageId="22" pageNumber="23">Patella</emphasis>
|
||
</taxonomicName>
|
||
. Ventral face: can be present a very small spine PVI, number of tubercles is variable. Dorsal face: number and size of the tubercles are variable.
|
||
<emphasis id="B968EA92B644FFD2FEC3FDE0FE8C7402" box="[298,363,585,611]" italics="true" pageId="22" pageNumber="23">Tibia.</emphasis>
|
||
Ventral and Dorsal face: tubercles between spines can be more conspicuous.
|
||
<emphasis id="B968EA92B644FFD2FB3DFDE0FAF57402" bold="true" box="[1236,1298,585,611]" italics="true" pageId="22" pageNumber="23">Legs.</emphasis>
|
||
In leg I, the number of segments of the tibia vary between 26 and 65 and tarsus vary between 97 and 103. The relative sizes of the femora II, III and IV is variable. (
|
||
<tableCitation id="C69E033BB644FFD2FDDEFD38FD6874CA" box="[567,655,657,683]" captionStart="TABLE 4" captionStartId="22.[151,239,946,970]" captionText="TABLE 4. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus elaphus. Empty cells correspond to missing segment or data." pageId="22" pageNumber="23">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA33680B644FFD2FF2EFD1CFB7F7492" blockId="22.[151,1437,153,899]" pageId="22" pageNumber="23">
|
||
<emphasis id="B968EA92B644FFD2FF2EFD1CFEEC74AE" bold="true" box="[199,267,693,719]" pageId="22" pageNumber="23">Male.</emphasis>
|
||
Observed Males have a similar size than the female (
|
||
<tableCitation id="C69E033BB644FFD2FCB0FD1CFC4874AE" box="[857,943,693,719]" captionStart="TABLE 4" captionStartId="22.[151,239,946,970]" captionText="TABLE 4. Measurements (mm) of the body, pedipalp segments, and femora of the legs in three specimens of Heterophrynus elaphus. Empty cells correspond to missing segment or data." pageId="22" pageNumber="23">Table 4</tableCitation>
|
||
), Spination pattern same as females. Genital operculum has an oval posterior margin, expanded to the third segment of the opisthosoma.
|
||
</paragraph>
|
||
<paragraph id="8BA33680B644FFD2FF2EFD55FF5D75E2" blockId="22.[151,1437,153,899]" pageId="22" pageNumber="23">
|
||
<emphasis id="B968EA92B644FFD2FF2EFD55FE937576" bold="true" box="[199,372,764,791]" pageId="22" pageNumber="23">
|
||
<emphasis id="B968EA92B644FFD2FF2EFD55FE8A7576" bold="true" box="[199,365,764,791]" italics="true" pageId="22" pageNumber="23">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
(
|
||
<emphasis id="B968EA92B644FFD2FE6AFD55FDC87576" bold="true" box="[387,559,764,791]" pageId="22" pageNumber="23">MZUSP 74571</emphasis>
|
||
): LaM with similar size to Pi. LoL 1 longer than LoL 2, LoL 1 is wider, densely covered with minute projections, placed close together with the base not visible; LoL 2 is wrinkled and does not present projections. In dorsal view, the LoD are elongated, wide, rounded and cover LoL1 and LoL 2 (
|
||
<figureCitation id="13272A05B644FFD2FAF9FCECFF4A75E2" captionStart="FIGURE 13" captionStartId="21.[151,250,1904,1929]" captionTargetBox="[163,1425,747,1874]" captionTargetId="figure@21.[157,1431,741,1879]" captionTargetPageId="21" captionText="FIGURE 13. Male genital: Heterophrynus elaphus. (MZUSP 74571): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1= lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. D. Female genital (MZUSP 74571), white arrow indicates the sclerotized area of the seminal receptacle,), black arrow indicates an extra sclerite at the base of gonopod." figureDoi="http://doi.org/10.5281/zenodo.3907928" httpUri="https://zenodo.org/record/3907928/files/figure.png" pageId="22" pageNumber="23">Fig. 13A, B, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |