treatments-xml/data/03/B5/87/03B5878EFF8EFFB6FE979B95FE72F998.xml
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<document id="368C6FF16FDBE60B54EE3E960B45D031" ID-DOI="10.5281/zenodo.175293" ID-GBIF-Dataset="d3a3d52e-430c-4ec2-be58-3e60bd9435e1" ID-ISSN="1175­5326" ID-Zenodo-Dep="175293" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459804061328" checkinUser="plazi" docAuthor="Freire, Eliza Maria Xavier, Caramaschi, Ulisses &amp; Argôlo, Antônio Jorge Suzart" docDate="2007" docId="03B5878EFF8EFFB6FE979B95FE72F998" docLanguage="en" docName="zt01393p026.pdf" docOrigin="Zootaxa 1393" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Liotyphlops trefauti Freire, Caramaschi &amp; Argôlo, 2007, sp. nov." docType="treatment" docVersion="9" lastPageNumber="23" masterDocId="FF8CFFF6FF8FFFB2FE00984FFFB3FFEA" masterDocTitle="A new species of Liotyphlops (Serpentes: Anomalepididae) from the Atlantic Rain Forest of Northeastern Brazil" masterLastPageNumber="26" masterPageNumber="19" pageNumber="20" updateTime="1698220391044" updateUser="plazi">
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<mods:title id="4145156002392FDC09F24DFA4F0AB9BB">A new species of Liotyphlops (Serpentes: Anomalepididae) from the Atlantic Rain Forest of Northeastern Brazil</mods:title>
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<mods:name id="0D42D4A9C4754D4BE575B18DFC74134E" type="personal">
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<mods:namePart id="F261E1A868E41FFDF686F6CAA790DDDC">Freire, Eliza Maria Xavier</mods:namePart>
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<mods:namePart id="85B23EA6747F3C4C236A18EB9868FFE9">Caramaschi, Ulisses</mods:namePart>
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<mods:role id="CDF38F69AD2D326A92D407ABF8329292">
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<mods:namePart id="A464F1481D043C13429ABC12344293B0">Argôlo, Antônio Jorge Suzart</mods:namePart>
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<mods:title id="2D1BEE2C6F0F971C33646854F0AFF1F1">Zootaxa</mods:title>
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<mods:date id="8E651D037FB82195AC58CDD798F3AFFD">2007</mods:date>
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<treatment id="03B5878EFF8EFFB6FE979B95FE72F998" ID-DOI="http://doi.org/10.5281/zenodo.5679903" ID-GBIF-Taxon="119352625" ID-Zenodo-Dep="5679903" LSID="urn:lsid:plazi:treatment:03B5878EFF8EFFB6FE979B95FE72F998" httpUri="http://treatment.plazi.org/id/03B5878EFF8EFFB6FE979B95FE72F998" lastPageId="4" lastPageNumber="23" pageId="1" pageNumber="20">
<subSubSection id="C3066513FF8EFFB3FE979B95FE9FFBFD" pageId="1" pageNumber="20" type="nomenclature">
<paragraph id="8BA33698FF8EFFB3FE979B95FE59FC1E" blockId="1.[151,490,986,1047]" box="[151,490,986,1012]" pageId="1" pageNumber="20">
<heading id="D0EB81F4FF8EFFB3FE979B95FE59FC1E" bold="true" box="[151,490,986,1012]" fontSize="11" level="1" pageId="1" pageNumber="20" reason="1">
<emphasis id="B968EA8AFF8EFFB3FE979B95FE59FC1E" bold="true" box="[151,490,986,1012]" pageId="1" pageNumber="20">
<taxonomicName id="4C1C4D1BFF8EFFB3FE979B95FE37FC1E" box="[151,388,986,1012]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8EFFB3FE979B95FE37FC1E" bold="true" box="[151,388,986,1012]" italics="true" pageId="1" pageNumber="20">Liotyphlops trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8EFFB3FF8C9B95FE59FC1E" box="[396,490,986,1012]" pageId="1" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA33698FF8EFFB3FE979BB2FE9FFBFD" blockId="1.[151,490,986,1047]" box="[151,300,1021,1047]" pageId="1" pageNumber="20">
(
<figureCitation id="13272A1DFF8EFFB3FE9F9BB2FF57FBFD" box="[159,228,1021,1047]" captionStart="FIGURE 1" captionStartId="3.[151,255,1521,1545]" captionTargetBox="[186,1382,395,1469]" captionTargetId="figure@3.[179,1408,369,1516]" captionTargetPageId="3" captionText="FIGURE 1. Liotyphlops trefauti sp. nov. (MZUSP 12178, holotype). (A) Dorsal view of head. (B) Lateral view of head. (C) Ventral view of head. (D) Ventral view of tail. (E) Lateral view of tail. Bar = 2 mm." httpUri="https://zenodo.org/record/175294/files/figure.png" pageId="1" pageNumber="20">Fig. 1</figureCitation>
AE)
</paragraph>
</subSubSection>
<subSubSection id="C3066513FF8EFFB3FE979C02FC55FAED" pageId="1" pageNumber="20" type="materials_examined">
<paragraph id="8BA33698FF8EFFB3FE979C02FB33FB65" blockId="1.[151,1436,1101,2010]" pageId="1" pageNumber="20">
<emphasis id="B968EA8AFF8EFFB3FE979C02FEA3FB8D" bold="true" box="[151,272,1101,1127]" pageId="1" pageNumber="20">
<typeStatus id="54A7883AFF8EFFB3FE979C02FEB8FB8D" box="[151,267,1101,1127]" pageId="1" pageNumber="20" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED0DAE5DFF8EFFB3FF199C02FEC8FB8D" box="[281,379,1101,1127]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="20">MZUSP</collectionCode>
12178, male, collected at Fazenda Bananeira, Municipality of Murici (
<geoCoordinate id="EE28505FFF8EFFB3FAC89C02FAADFB8D" box="[1224,1310,1101,1127]" direction="south" orientation="latitude" pageId="1" pageNumber="20" precision="925" value="-9.233334">9 14S</geoCoordinate>
,
<geoCoordinate id="EE28505FFF8EFFB3FB2C9C02FA24FB8D" box="[1324,1431,1101,1127]" direction="west" orientation="longitude" pageId="1" pageNumber="20" precision="925" value="-35.8">35 48W</geoCoordinate>
,
<quantity id="4CE49B7DFF8EFFB3FE979C3AFF52FB65" box="[151,225,1141,1167]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="20" unit="m" value="640.0">640 m</quantity>
elevation), State of Alagoas,
<collectingCountry id="F30B7608FF8EFFB3FC3C9C3AFD35FB65" box="[572,646,1141,1167]" name="Brazil" pageId="1" pageNumber="20">Brazil</collectingCountry>
, on 0
<date id="FFA21058FF8EFFB3FCC59C3AFC2AFB65" box="[709,921,1141,1167]" pageId="1" pageNumber="20" value="1994-11-04">4 November 1994</date>
by E. M. X. Freire.
</paragraph>
<paragraph id="8BA33698FF8EFFB3FEC69CD2FC55FAED" blockId="1.[151,1436,1101,2010]" pageId="1" pageNumber="20">
<emphasis id="B968EA8AFF8EFFB3FEC69CD2FE8CFB5D" bold="true" box="[198,319,1181,1207]" pageId="1" pageNumber="20">
<typeStatus id="54A7883AFF8EFFB3FEC69CD2FE89FB5D" box="[198,314,1181,1207]" pageId="1" pageNumber="20" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectionCode id="ED0DAE5DFF8EFFB3FF479CD2FE1AFB5D" box="[327,425,1181,1207]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="20">MZUSP</collectionCode>
12179, female, collected at Regional Center of the CEPLAC (Comissão Executiva do Plano da Lavoura Cacaueira), Municipality of Ilhéus (
<geoCoordinate id="EE28505FFF8EFFB3FD269C8AFC38FB35" box="[806,907,1221,1247]" direction="south" orientation="latitude" pageId="1" pageNumber="20" precision="925" value="-14.766666">14 46S</geoCoordinate>
,
<geoCoordinate id="EE28505FFF8EFFB3FD9B9C8AFBBBFB35" box="[923,1032,1221,1247]" direction="west" orientation="longitude" pageId="1" pageNumber="20" precision="925" value="-39.216667">39 13W</geoCoordinate>
;
<quantity id="4CE49B7DFF8EFFB3FA189C8AFBE4FB35" box="[1048,1111,1221,1247]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="20" unit="m" value="50.0">50 m</quantity>
elevation), State of Bahia,
<collectingCountry id="F30B7608FF8EFFB3FE979CA2FF51FAED" box="[151,226,1261,1287]" name="Brazil" pageId="1" pageNumber="20">Brazil</collectingCountry>
, on
<date id="FFA21058FF8EFFB3FF129CA2FE64FAED" box="[274,471,1261,1287]" pageId="1" pageNumber="20" value="1992-01-29">29 January 1992</date>
, by C. Jared, M. Antoniazzi, and L. Ferreira.
</paragraph>
</subSubSection>
<subSubSection id="C3066513FF8EFFB3FEC69D57FEC5FA68" pageId="1" pageNumber="20" type="description">
<paragraph id="8BA33698FF8EFFB3FEC69D57FEC5FA68" blockId="1.[151,1436,1101,2010]" pageId="1" pageNumber="20">
<emphasis id="B968EA8AFF8EFFB3FEC69D57FE0AFAD8" bold="true" box="[198,441,1304,1330]" pageId="1" pageNumber="20">Referred specimen.</emphasis>
CHBEZ 649, young, collected at
<collectingCountry id="F30B7608FF8EFFB3FD4C9D57FC3BFAD8" box="[844,904,1304,1330]" name="Malta" pageId="1" pageNumber="20">Mata</collectingCountry>
da Cachoeira (
<geoCoordinate id="EE28505FFF8EFFB3FA3B9D57FB29FAD8" box="[1083,1178,1301,1330]" direction="south" orientation="latitude" pageId="1" pageNumber="20" precision="925" value="-9.0">09o00S</geoCoordinate>
,
<geoCoordinate id="EE28505FFF8EFFB3FAA69D57FAA3FAD8" box="[1190,1296,1301,1330]" direction="west" orientation="longitude" pageId="1" pageNumber="20" precision="925" value="-36.05">36o03W</geoCoordinate>
;
<quantity id="4CE49B7DFF8EFFB3FB1D9D57FADBFAD8" box="[1309,1384,1304,1330]" metricMagnitude="2" metricUnit="m" metricValue="4.37" pageId="1" pageNumber="20" unit="m" value="437.0">437 m</quantity>
elevation), Usina Serra Grande, Municipality of São José da Lage, State of Alagoas,
<collectingCountry id="F30B7608FF8EFFB3FA539D0FFB2EFAB0" box="[1107,1181,1344,1370]" name="Brazil" pageId="1" pageNumber="20">Brazil</collectingCountry>
, on
<date id="FFA21058FF8EFFB3FACE9D0FFA26FAB0" box="[1230,1429,1344,1370]" pageId="1" pageNumber="20" value="2003-10-25">25 October 2003</date>
, by E. M. X. Freire.
</paragraph>
</subSubSection>
<subSubSection id="C3066513FF8EFFB0FEC69DDFFE55FEB8" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="diagnosis">
<paragraph id="8BA33698FF8EFFB3FEC69DDFFAC2F9A0" blockId="1.[151,1436,1101,2010]" pageId="1" pageNumber="20">
<emphasis id="B968EA8AFF8EFFB3FEC69DDFFEF5FA40" bold="true" box="[198,326,1424,1450]" pageId="1" pageNumber="20">Diagnosis.</emphasis>
<taxonomicName id="4C1C4D1BFF8EFFB3FF4D9DDFFD80FA43" box="[333,563,1424,1449]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8EFFB3FF4D9DDFFD80FA43" box="[333,563,1424,1449]" italics="true" pageId="1" pageNumber="20">Liotyphlops trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8EFFB3FC3B9DDFFD2BFA40" box="[571,664,1424,1450]" pageId="1" pageNumber="20" rank="species">
<emphasis id="B968EA8AFF8EFFB3FC3B9DDFFD2BFA40" bold="true" box="[571,664,1424,1450]" pageId="1" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from all other species of the genus by the following combinations of characters: (1) four scales contacting the posterior edge of prefrontal; (2) one scale contacting the posterior edge of nasal, between the second supralabial and the prefrontal; (3) five scales in the first vertical row of lateral head scales; (4) 22­22­22 scales around body; (5) 520543 dorsal scales; (6) 499 532 ventral scales; (7) absence of eye spot; (8) dorsal and ventral color bright yellowish to reddish brown.
</paragraph>
<paragraph id="8BA33698FF8EFFB0FEC69E17FE55FEB8" blockId="1.[151,1436,1101,2010]" lastBlockId="2.[151,1437,151,338]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<emphasis id="B968EA8AFF8EFFB3FEC69E17FDE8F998" bold="true" box="[198,603,1624,1650]" pageId="1" pageNumber="20">Comparisons with other species.</emphasis>
The presence of four scales contacting the posterior border of prefrontal approximates
<taxonomicName id="4C1C4D1BFF8EFFB3FF639ECFFE6BF973" box="[355,472,1664,1689]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8EFFB3FF639ECFFE6BF973" box="[355,472,1664,1689]" italics="true" pageId="1" pageNumber="20">L.trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8EFFB3FFE19ECFFD8CF970" box="[481,575,1664,1690]" pageId="1" pageNumber="20" rank="species">
<emphasis id="B968EA8AFF8EFFB3FFE19ECFFD8CF970" bold="true" box="[481,575,1664,1690]" pageId="1" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
to the Colombian species
<taxonomicName id="4C1C4D1BFF8EFFB3FD7C9ECFFC51F973" box="[892,994,1664,1689]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="anops">
<emphasis id="B968EA8AFF8EFFB3FD7C9ECFFC51F973" box="[892,994,1664,1689]" italics="true" pageId="1" pageNumber="20">L. anops</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D1BFF8EFFB3FA1D9ECFFB11F973" box="[1053,1186,1664,1689]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="argaleus">
<emphasis id="B968EA8AFF8EFFB3FA1D9ECFFB11F973" box="[1053,1186,1664,1689]" italics="true" pageId="1" pageNumber="20">L. argaleus</emphasis>
</taxonomicName>
; however,
<taxonomicName id="4C1C4D1BFF8EFFB3FB269ECFFA28F973" box="[1318,1435,1664,1689]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8EFFB3FB269ECFFA28F973" box="[1318,1435,1664,1689]" italics="true" pageId="1" pageNumber="20">L. trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8EFFB3FE979EE7FF45F928" box="[151,246,1704,1730]" pageId="1" pageNumber="20" rank="species">
<emphasis id="B968EA8AFF8EFFB3FE979EE7FF45F928" bold="true" box="[151,246,1704,1730]" pageId="1" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
presents one scale contacting the posterior border of nasal, between the second supralabial and the prefrontal (two scales in
<taxonomicName id="4C1C4D1BFF8EFFB3FFBD9E9FFD90F903" box="[445,547,1744,1769]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="anops">
<emphasis id="B968EA8AFF8EFFB3FFBD9E9FFD90F903" box="[445,547,1744,1769]" italics="true" pageId="1" pageNumber="20">L. anops</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D1BFF8EFFB3FC5E9E9FFD50F903" box="[606,739,1744,1769]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="argaleus">
<emphasis id="B968EA8AFF8EFFB3FC5E9E9FFD50F903" box="[606,739,1744,1769]" italics="true" pageId="1" pageNumber="20">L. argaleus</emphasis>
</taxonomicName>
) and 22­22­22 scales around the body (27­24­23 to 28­24­
<quantity id="4CE49B7DFF8EFFB3FE979EB7FF60F8F8" box="[151,211,1784,1810]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="1" pageNumber="20" unit="in" value="24.0">24 in</quantity>
<taxonomicName id="4C1C4D1BFF8EFFB3FEDC9EB7FEF1F8FB" box="[220,322,1784,1809]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="anops">
<emphasis id="B968EA8AFF8EFFB3FEDC9EB7FEF1F8FB" box="[220,322,1784,1809]" italics="true" pageId="1" pageNumber="20">L. anops</emphasis>
</taxonomicName>
, 26­22­22 to 28­24­
<quantity id="4CE49B7DFF8EFFB3FC319EB7FDDCF8F8" box="[561,623,1784,1810]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="1" pageNumber="20" unit="in" value="22.0">22 in</quantity>
<taxonomicName id="4C1C4D1BFF8EFFB3FC779EB7FD48F8FB" box="[631,763,1784,1809]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="argaleus">
<emphasis id="B968EA8AFF8EFFB3FC779EB7FD48F8FB" box="[631,763,1784,1809]" italics="true" pageId="1" pageNumber="20">L. argaleus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF8D4B69FF8EFFB3FD0C9EB7FB80F8F8" author="Dixon" box="[780,1075,1784,1810]" pageId="1" pageNumber="24" refString="Dixon, J. R. &amp; Kofron, C. P. (1984 [1983]) The Central and South American anomalepid snakes of the genus Liotyphlops. Amphibia-Reptilia, 4, 241 - 264." type="journal article" year="1984">Dixon and Kofron, 1984</bibRefCitation>
;
<tableCitation id="C69E0323FF8EFFB3FA409EB7FB24F8F8" box="[1088,1175,1784,1810]" captionStart="TABLE 1" captionStartId="2.[151,239,386,410]" captionTargetBox="[151,1422,442,1028]" captionTargetPageId="2" captionText="TABLE 1. Comparative diagnostic characters for the species of Liotyphlops (Dixon and Kofron 1984; our data)." httpUri="http://table.plazi.org/id/DF636610FF8DFFB0FE9799CDFAF4FE73" pageId="1" pageNumber="20" tableUuid="DF636610FF8DFFB0FE9799CDFAF4FE73">Table 1</tableCitation>
). Additionally,
<taxonomicName id="4C1C4D1BFF8EFFB3FB539EB7FF7EF8D3" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8EFFB3FB539EB7FF7EF8D3" italics="true" pageId="1" pageNumber="20">L. trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8EFFB3FED79F6FFE8BF8D0" box="[215,312,1824,1850]" pageId="1" pageNumber="20" rank="species">
<emphasis id="B968EA8AFF8EFFB3FED79F6FFE8BF8D0" bold="true" box="[215,312,1824,1850]" pageId="1" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
has dorsal and ventral color bright yellowish to reddish brown, while
<taxonomicName id="4C1C4D1BFF8EFFB3FA959F6FFB4EF8D3" box="[1173,1277,1824,1849]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="anops">
<emphasis id="B968EA8AFF8EFFB3FA959F6FFB4EF8D3" box="[1173,1277,1824,1849]" italics="true" pageId="1" pageNumber="20">L. anops</emphasis>
</taxonomicName>
has all body scales pigmented although the brown pigment of each scale is confined to the posterior one third to one fourth of each scale and the rostral, nasal, and prefrontal areas, and the anal and caudal tip areas are whitish (
<bibRefCitation id="EF8D4B69FF8EFFB3FB4E9F3FFEDDF858" author="Dixon" pageId="1" pageNumber="24" refString="Dixon, J. R. &amp; Kofron, C. P. (1984 [1983]) The Central and South American anomalepid snakes of the genus Liotyphlops. Amphibia-Reptilia, 4, 241 - 264." type="journal article" year="1984">Dixon and Kofron, 1984</bibRefCitation>
), and
<taxonomicName id="4C1C4D1BFF8EFFB3FFB89FD7FD8DF85B" box="[440,574,1944,1969]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="argaleus">
<emphasis id="B968EA8AFF8EFFB3FFB89FD7FD8DF85B" box="[440,574,1944,1969]" italics="true" pageId="1" pageNumber="20">L. argaleus</emphasis>
</taxonomicName>
is uniform dark brown with the posterior one third of each scale with a transverse blackish brown spot, anterior two thirds of each scale dark gray, anal area and caudal spine whitish, and rostral and nasal areas pinkish white (
<bibRefCitation id="EF8D4B69FF8DFFB0FC8198D7FC1BFF58" author="Dixon" box="[641,936,152,178]" pageId="2" pageNumber="24" refString="Dixon, J. R. &amp; Kofron, C. P. (1984 [1983]) The Central and South American anomalepid snakes of the genus Liotyphlops. Amphibia-Reptilia, 4, 241 - 264." type="journal article" year="1984">Dixon and Kofron, 1984</bibRefCitation>
).
<taxonomicName id="4C1C4D1BFF8DFFB0FDBC98D7FB10FF5B" box="[956,1187,152,177]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8DFFB0FDBC98D7FB10FF5B" box="[956,1187,152,177]" italics="true" pageId="2" pageNumber="21">Liotyphlops trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8DFFB0FAAB98D8FABBFF5B" box="[1195,1288,151,177]" pageId="2" pageNumber="21" rank="species">
<emphasis id="B968EA8AFF8DFFB0FAAB98D8FABBFF5B" bold="true" box="[1195,1288,151,177]" pageId="2" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C1C4D1BFF8DFFB0FE97988FFE83FF33" box="[151,304,192,217]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="albirostris">
<emphasis id="B968EA8AFF8DFFB0FE97988FFE83FF33" box="[151,304,192,217]" italics="true" pageId="2" pageNumber="21">L. albirostris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D1BFF8DFFB0FF40988FFE21FF33" box="[320,402,192,217]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="beui">
<emphasis id="B968EA8AFF8DFFB0FF40988FFE21FF33" box="[320,402,192,217]" italics="true" pageId="2" pageNumber="21">L. beui</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D1BFF8DFFB0FFA2988FFD82FF33" box="[418,561,192,217]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="schubarti">
<emphasis id="B968EA8AFF8DFFB0FFA2988FFD82FF33" box="[418,561,192,217]" italics="true" pageId="2" pageNumber="21">L. schubarti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D1BFF8DFFB0FC41988FFD72FF33" box="[577,705,192,217]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="ternetzii">
<emphasis id="B968EA8AFF8DFFB0FC41988FFD72FF33" box="[577,705,192,217]" italics="true" pageId="2" pageNumber="21">L. ternetzii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1C4D1BFF8DFFB0FD04988FFCC4FF33" box="[772,887,192,217]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="wilderi">
<emphasis id="B968EA8AFF8DFFB0FD04988FFCC4FF33" box="[772,887,192,217]" italics="true" pageId="2" pageNumber="21">L. wilderi</emphasis>
</taxonomicName>
by having four scales contacting the posterior border of prefrontal (three scales in those species) and higher number of dorsals scales, 520543 (
<taxonomicName id="4C1C4D1BFF8DFFB0FB2098A7FF0DFEC3" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="albirostris">
<emphasis id="B968EA8AFF8DFFB0FB2098A7FF0DFEC3" italics="true" pageId="2" pageNumber="21">L. albirostris</emphasis>
</taxonomicName>
, 370455;
<taxonomicName id="4C1C4D1BFF8DFFB0FF42995FFE20FEC3" box="[322,403,272,297]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="beui">
<emphasis id="B968EA8AFF8DFFB0FF42995FFE20FEC3" box="[322,403,272,297]" italics="true" pageId="2" pageNumber="21">L. beui</emphasis>
</taxonomicName>
, 375464;
<taxonomicName id="4C1C4D1BFF8DFFB0FC17995FFD16FEC3" box="[535,677,272,297]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="schubarti">
<emphasis id="B968EA8AFF8DFFB0FC17995FFD16FEC3" box="[535,677,272,297]" italics="true" pageId="2" pageNumber="21">L. schubarti</emphasis>
</taxonomicName>
, 372387;
<taxonomicName id="4C1C4D1BFF8DFFB0FD29995FFC1BFEC3" box="[809,936,272,297]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="ternetzii">
<emphasis id="B968EA8AFF8DFFB0FD29995FFC1BFEC3" box="[809,936,272,297]" italics="true" pageId="2" pageNumber="21">L. ternetzii</emphasis>
</taxonomicName>
, 463510, and
<taxonomicName id="4C1C4D1BFF8DFFB0FA5C995FFB7DFEC3" box="[1116,1230,272,297]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="wilderi">
<emphasis id="B968EA8AFF8DFFB0FA5C995FFB7DFEC3" box="[1116,1230,272,297]" italics="true" pageId="2" pageNumber="21">L. wilderi</emphasis>
</taxonomicName>
, 304358;
<bibRefCitation id="EF8D4B69FF8DFFB0FB53995FFED9FEB8" author="Dixon" pageId="2" pageNumber="24" refString="Dixon, J. R. &amp; Kofron, C. P. (1984 [1983]) The Central and South American anomalepid snakes of the genus Liotyphlops. Amphibia-Reptilia, 4, 241 - 264." type="journal article" year="1984">Dixon and Kofron, 1984</bibRefCitation>
; our data).
</paragraph>
</subSubSection>
<caption id="DF636610FF8DFFB0FE9799CDFAF4FE73" ID-Table-UUID="DF636610FF8DFFB0FE9799CDFAF4FE73" box="[151,1351,386,410]" httpUri="http://table.plazi.org/id/DF636610FF8DFFB0FE9799CDFAF4FE73" pageId="2" pageNumber="21" targetBox="[151,1422,442,1028]" targetIsTable="true" targetPageId="2">
<paragraph id="8BA33698FF8DFFB0FE9799CDFAF4FE73" blockId="2.[151,1351,386,410]" box="[151,1351,386,410]" pageId="2" pageNumber="21">
<emphasis id="B968EA8AFF8DFFB0FE9799CDFEB9FE70" bold="true" box="[151,266,386,410]" pageId="2" pageNumber="21">TABLE 1.</emphasis>
Comparative diagnostic characters for the species of
<taxonomicName id="4C1C4D1BFF8DFFB0FD4599CDFC70FE73" box="[837,963,386,409]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B968EA8AFF8DFFB0FD4599CDFC70FE73" box="[837,963,386,409]" italics="true" pageId="2" pageNumber="21">Liotyphlops</emphasis>
</taxonomicName>
(Dixon and Kofron 1984; our data).
</paragraph>
</caption>
<paragraph id="8BA33698FF8DFFB0FC9699F5FEBBFBEE" pageId="2" pageNumber="21">
<table id="F91CC438FF8D004DFE9799F5FA3DFBEE" box="[151,1422,442,1028]" gridcols="5" gridrows="9" pageId="2" pageNumber="21">
<tr id="352C34DAFF8D004DFE9799F5FA3DFE38" box="[151,1422,442,466]" gridrow="0" pageId="2" pageNumber="21" rowspan-0="1">
<th id="76FD5DA6FF8D004DFC9699F5FC82FE38" box="[662,817,442,466]" gridcol="1" gridrow="0" pageId="2" pageNumber="21">
<taxonomicName id="4C1C4D1BFF8DFFB0FC9699F5FCBDFE3B" box="[662,782,442,465]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="argaleus">
<emphasis id="B968EA8AFF8DFFB0FC9699F5FCBDFE3B" box="[662,782,442,465]" italics="true" pageId="2" pageNumber="21">L. argaleus</emphasis>
</taxonomicName>
</th>
<th id="76FD5DA6FF8D004DFD4D99F5FBBAFE38" box="[845,1033,442,466]" gridcol="2" gridrow="0" pageId="2" pageNumber="21">
<taxonomicName id="4C1C4D1BFF8DFFB0FD4D99F5FC1AFE3B" box="[845,937,442,465]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="anops">
<emphasis id="B968EA8AFF8DFFB0FD4D99F5FC1AFE3B" box="[845,937,442,465]" italics="true" pageId="2" pageNumber="21">L. anops</emphasis>
</taxonomicName>
</th>
<th id="76FD5DA6FF8D004DFA2E99F5FB47FE38" box="[1070,1268,442,466]" gridcol="3" gridrow="0" pageId="2" pageNumber="21">
<taxonomicName id="4C1C4D1BFF8DFFB0FA2E99F5FB2AFE3B" box="[1070,1177,442,465]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8DFFB0FA2E99F5FB2AFE3B" box="[1070,1177,442,465]" italics="true" pageId="2" pageNumber="21">L. trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8DFFB0FA9E99F5FB47FE38" box="[1182,1268,442,466]" pageId="2" pageNumber="21" rank="species">
<emphasis id="B968EA8AFF8DFFB0FA9E99F5FB47FE38" bold="true" box="[1182,1268,442,466]" pageId="2" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<th id="76FD5DA6FF8D004DFB0499F5FA3DFE38" box="[1284,1422,442,466]" gridcol="4" gridrow="0" pageId="2" pageNumber="21">
<taxonomicName id="4C1C4D1BFF8DFFB0FB0499F5FA3DFE3B" box="[1284,1422,442,465]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="albirostris">
<emphasis id="B968EA8AFF8DFFB0FB0499F5FA3DFE3B" box="[1284,1422,442,465]" italics="true" pageId="2" pageNumber="21">L. albirostris</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="352C34DAFF8D004DFE9799AAFA3DFDF6" box="[151,1422,485,540]" gridrow="1" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE9799AAFD36FDF6" box="[151,645,485,540]" gridcol="0" gridrow="1" pageId="2" pageNumber="21">Scales number contacting posterior edge of prefrontal</th>
<td id="76FD5DA6FF8D004DFC9699AAFC82FDF6" box="[662,817,485,540]" gridcol="1" gridrow="1" pageId="2" pageNumber="21">4</td>
<td id="76FD5DA6FF8D004DFD4D99AAFBBAFDF6" box="[845,1033,485,540]" gridcol="2" gridrow="1" pageId="2" pageNumber="21">4</td>
<td id="76FD5DA6FF8D004DFA2E99AAFB47FDF6" box="[1070,1268,485,540]" gridcol="3" gridrow="1" pageId="2" pageNumber="21">4</td>
<td id="76FD5DA6FF8D004DFB0499AAFA3DFDF6" box="[1284,1422,485,540]" gridcol="4" gridrow="1" pageId="2" pageNumber="21">3</td>
</tr>
<tr id="352C34DAFF8D004DFE979A7FFA3DFD6D" box="[151,1422,560,647]" gridrow="2" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE979A7FFD36FD6D" box="[151,645,560,647]" gridcol="0" gridrow="2" pageId="2" pageNumber="21">Scales number contacting posterior edge of nasal between second supralabial and pre­ frontal</th>
<td id="76FD5DA6FF8D004DFC969A7FFC82FD6D" box="[662,817,560,647]" gridcol="1" gridrow="2" pageId="2" pageNumber="21">2</td>
<td id="76FD5DA6FF8D004DFD4D9A7FFBBAFD6D" box="[845,1033,560,647]" gridcol="2" gridrow="2" pageId="2" pageNumber="21">2</td>
<td id="76FD5DA6FF8D004DFA2E9A7FFB47FD6D" box="[1070,1268,560,647]" gridcol="3" gridrow="2" pageId="2" pageNumber="21">1</td>
<td id="76FD5DA6FF8D004DFB049A7FFA3DFD6D" box="[1284,1422,560,647]" gridcol="4" gridrow="2" pageId="2" pageNumber="21">1</td>
</tr>
<tr id="352C34DAFF8D004DFE979AD5FA3DFD3B" box="[151,1422,666,721]" gridrow="3" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE979AD5FD36FD3B" box="[151,645,666,721]" gridcol="0" gridrow="3" pageId="2" pageNumber="21">Eye spot</th>
<td id="76FD5DA6FF8D004DFC969AD5FC82FD3B" box="[662,817,666,721]" gridcol="1" gridrow="3" pageId="2" pageNumber="21">little visible or ausent</td>
<td id="76FD5DA6FF8D004DFD4D9AD5FBBAFD3B" box="[845,1033,666,721]" gridcol="2" gridrow="3" pageId="2" pageNumber="21">absent</td>
<td id="76FD5DA6FF8D004DFA2E9AD5FB47FD3B" box="[1070,1268,666,721]" gridcol="3" gridrow="3" pageId="2" pageNumber="21">absent</td>
<td id="76FD5DA6FF8D004DFB049AD5FA3DFD3B" box="[1284,1422,666,721]" gridcol="4" gridrow="3" pageId="2" pageNumber="21">little visible</td>
</tr>
<tr id="352C34DAFF8D004DFE979AAAFA3DFCF6" box="[151,1422,741,796]" gridrow="4" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE979AAAFD36FCF6" box="[151,645,741,796]" gridcol="0" gridrow="4" pageId="2" pageNumber="21">Scales number in the first vertical row of dor­ sals</th>
<td id="76FD5DA6FF8D004DFC969AAAFC82FCF6" box="[662,817,741,796]" gridcol="1" gridrow="4" pageId="2" pageNumber="21">5</td>
<td id="76FD5DA6FF8D004DFD4D9AAAFBBAFCF6" box="[845,1033,741,796]" gridcol="2" gridrow="4" pageId="2" pageNumber="21">6</td>
<td id="76FD5DA6FF8D004DFA2E9AAAFB47FCF6" box="[1070,1268,741,796]" gridcol="3" gridrow="4" pageId="2" pageNumber="21">5</td>
<td id="76FD5DA6FF8D004DFB049AAAFA3DFCF6" box="[1284,1422,741,796]" gridcol="4" gridrow="4" pageId="2" pageNumber="21">5</td>
</tr>
<tr id="352C34DAFF8D004DFE979B7FFA3DFC8D" box="[151,1422,816,871]" gridrow="5" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE979B7FFD36FC8D" box="[151,645,816,871]" gridcol="0" gridrow="5" pageId="2" pageNumber="21">Scales rows around body</th>
<td id="76FD5DA6FF8D004DFC969B7FFC82FC8D" box="[662,817,816,871]" gridcol="1" gridrow="5" pageId="2" pageNumber="21">2628/2224/ 22</td>
<td id="76FD5DA6FF8D004DFD4D9B7FFBBAFC8D" box="[845,1033,816,871]" gridcol="2" gridrow="5" pageId="2" pageNumber="21">2728/2224/23 24</td>
<td id="76FD5DA6FF8D004DFA2E9B7FFB47FC8D" box="[1070,1268,816,871]" gridcol="3" gridrow="5" pageId="2" pageNumber="21">22/22/22</td>
<td id="76FD5DA6FF8D004DFB049B7FFA3DFC8D" box="[1284,1422,816,871]" gridcol="4" gridrow="5" pageId="2" pageNumber="21">2426/22/22</td>
</tr>
<tr id="352C34DAFF8D004DFE979B35FA3DFC7B" box="[151,1422,890,913]" gridrow="6" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE979B35FD36FC7B" box="[151,645,890,913]" gridcol="0" gridrow="6" pageId="2" pageNumber="21">Scales rows dorsals</th>
<td id="76FD5DA6FF8D004DFC969B35FC82FC7B" box="[662,817,890,913]" gridcol="1" gridrow="6" pageId="2" pageNumber="21">351533</td>
<td id="76FD5DA6FF8D004DFD4D9B35FBBAFC7B" box="[845,1033,890,913]" gridcol="2" gridrow="6" pageId="2" pageNumber="21">548 or more</td>
<td id="76FD5DA6FF8D004DFA2E9B35FB47FC7B" box="[1070,1268,890,913]" gridcol="3" gridrow="6" pageId="2" pageNumber="21">520543</td>
<td id="76FD5DA6FF8D004DFB049B35FA3DFC7B" box="[1284,1422,890,913]" gridcol="4" gridrow="6" pageId="2" pageNumber="21">370455</td>
</tr>
<tr id="352C34DAFF8D004DFE979BEAFA3DFC56" box="[151,1422,933,956]" gridrow="7" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE979BEAFD36FC56" box="[151,645,933,956]" gridcol="0" gridrow="7" pageId="2" pageNumber="21">Total length (mm)</th>
<td id="76FD5DA6FF8D004DFC969BEAFC82FC56" box="[662,817,933,956]" gridcol="1" gridrow="7" pageId="2" pageNumber="21">101261</td>
<td id="76FD5DA6FF8D004DFD4D9BEAFBBAFC56" box="[845,1033,933,956]" gridcol="2" gridrow="7" pageId="2" pageNumber="21">125395</td>
<td id="76FD5DA6FF8D004DFA2E9BEAFB47FC56" box="[1070,1268,933,956]" gridcol="3" gridrow="7" pageId="2" pageNumber="21">366389</td>
<td id="76FD5DA6FF8D004DFB049BEAFA3DFC56" box="[1284,1422,933,956]" gridcol="4" gridrow="7" pageId="2" pageNumber="21">70233</td>
</tr>
<tr id="352C34DAFF8D004DFE979BA2FA3DFBEE" box="[151,1422,1005,1028]" gridrow="8" pageId="2" pageNumber="21" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76FD5DA6FF8D004DFE979BA2FD36FBEE" box="[151,645,1005,1028]" gridcol="0" gridrow="8" pageId="2" pageNumber="21">continued.</th>
</tr>
</table>
</paragraph>
<subSubSection id="C3066513FF8DFFB6FC969C6AFF5BFA10" lastPageId="4" lastPageNumber="23" pageId="2" pageNumber="21" type="description">
<paragraph id="8BA33698FF8DFFB0FC969C6AFAE5FBD6" blockId="2.[159,1366,1061,1382]" box="[662,1366,1061,1084]" pageId="2" pageNumber="21">
<emphasis id="B968EA8AFF8DFFB0FC969C6AFAE5FBD6" box="[662,1366,1061,1084]" italics="true" pageId="2" pageNumber="21">
<taxonomicName id="4C1C4D1BFF8DFFB0FC969C6AFD53FBD6" box="[662,736,1061,1084]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="beui">L. beui</taxonomicName>
<taxonomicName id="4C1C4D1BFF8DFFB0FD3C9C6AFC0EFBD6" box="[828,957,1061,1084]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="schubarti">L. schubarti</taxonomicName>
<taxonomicName id="4C1C4D1BFF8DFFB0FA1E9C6AFB21FBD6" box="[1054,1170,1061,1084]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="ternetzii">L. ternetzii</taxonomicName>
<taxonomicName id="4C1C4D1BFF8DFFB0FAEE9C6AFAE5FBD6" box="[1262,1366,1061,1084]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="wilderi">L. wilderi</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BA33698FF8DFFB0FE9F9C00FEBBFB6C" blockId="2.[159,1366,1061,1382]" pageId="2" pageNumber="21">Scales number contacting posterior edge of 3 3 3 3 prefrontal</paragraph>
<paragraph id="8BA33698FF8DFFB0FE9F9CD5FDD2FB3B" blockId="2.[159,1366,1061,1382]" pageId="2" pageNumber="21">Scales number contacting posterior edge of 2 2 2 1 nasal between second supralabial and pre­</paragraph>
<paragraph id="8BA33698FF8DFFB0FE9F9C95FF55FB1B" blockId="2.[159,1366,1061,1382]" box="[159,230,1242,1265]" pageId="2" pageNumber="21">frontal</paragraph>
<paragraph id="8BA33698FF8DFFB0FE9F9D4AFF74FA8C" blockId="2.[159,1366,1061,1382]" pageId="2" pageNumber="21">Eye spot little visible little visible evident evident Scales number in the first vertical row of dor­ 5 5 5 5 sals</paragraph>
<paragraph id="8BA33698FF8DFFB0FE9F9D35FA8DFA0C" pageId="2" pageNumber="21">
<table id="F91CC438FF8D004DFE9F9D35FAE5FA0C" box="[159,1366,1402,1510]" gridcols="4" gridrows="3" pageId="2" pageNumber="21">
<tr id="352C34DAFF8D004DFE9F9D35FAE5FA7B" box="[159,1366,1402,1425]" gridrow="0" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE9F9D35FE19FA7B" box="[159,426,1402,1425]" gridcol="0" gridrow="0" pageId="2" pageNumber="21">Scales rows around body</th>
<th id="76FD5DA6FF8D004DFC969D35FC28FA7B" box="[662,923,1402,1425]" gridcol="1" gridrow="0" pageId="2" pageNumber="21">22/20/20 22/20/20</th>
<th id="76FD5DA6FF8D004DFA1D9D35FBCEFA7B" box="[1053,1149,1402,1425]" gridcol="2" gridrow="0" pageId="2" pageNumber="21">22/24/22</th>
<th id="76FD5DA6FF8D004DFAED9D35FAE5FA7B" box="[1261,1366,1402,1425]" gridcol="3" gridrow="0" pageId="2" pageNumber="21">22/20/20</th>
</tr>
<tr id="352C34DAFF8D004DFE9F9DEAFAE5FA56" box="[159,1366,1445,1468]" gridrow="1" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE9F9DEAFE19FA56" box="[159,426,1445,1468]" gridcol="0" gridrow="1" pageId="2" pageNumber="21">Scales rows dorsals</th>
<td id="76FD5DA6FF8D004DFC969DEAFC28FA56" box="[662,923,1445,1468]" gridcol="1" gridrow="1" pageId="2" pageNumber="21">375464 372387</td>
<td id="76FD5DA6FF8D004DFA1D9DEAFBCEFA56" box="[1053,1149,1445,1468]" gridcol="2" gridrow="1" pageId="2" pageNumber="21">463510</td>
<td id="76FD5DA6FF8D004DFAED9DEAFAE5FA56" box="[1261,1366,1445,1468]" gridcol="3" gridrow="1" pageId="2" pageNumber="21">304358</td>
</tr>
<tr id="352C34DAFF8D004DFE9F9D80FAE5FA0C" box="[159,1366,1487,1510]" gridrow="2" pageId="2" pageNumber="21">
<th id="76FD5DA6FF8D004DFE9F9D80FE19FA0C" box="[159,426,1487,1510]" gridcol="0" gridrow="2" pageId="2" pageNumber="21">Total length (mm)</th>
<td id="76FD5DA6FF8D004DFC969D80FC28FA0C" box="[662,923,1487,1510]" gridcol="1" gridrow="2" pageId="2" pageNumber="21">106381 80245</td>
<td id="76FD5DA6FF8D004DFA1D9D80FBCEFA0C" box="[1053,1149,1487,1510]" gridcol="2" gridrow="2" pageId="2" pageNumber="21">88413</td>
<td id="76FD5DA6FF8D004DFAED9D80FAE5FA0C" box="[1261,1366,1487,1510]" gridcol="3" gridrow="2" pageId="2" pageNumber="21">68290</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA33698FF8DFFB1FEC69E75FBDCFEE8" blockId="2.[151,1437,1594,2020]" lastBlockId="3.[151,1437,152,338]" lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21">
<emphasis id="B968EA8AFF8DFFB0FEC69E75FD97F9BE" bold="true" box="[198,548,1594,1620]" pageId="2" pageNumber="21">
Description of the
<typeStatus id="54A7883AFF8DFFB0FFB19E75FDACF9BE" box="[433,543,1594,1620]" pageId="2" pageNumber="21" type="holotype">holotype</typeStatus>
.
</emphasis>
Rostral large, higher than wide, contacting anterolaterally nasals, laterally prefrontals, and posteriorly the single frontal, which prevents the prefrontals from contacting each other. Prefrontals pairs triangulars, bordered anteriorly and laterally by the rostral, ventrally by the large divided nasal, and dorsoposteriorly partially by the single frontal. Posterior edge of prefrontal passing in front of frontal. In lateral view, four scales contact posterior edge of prefrontal, including the frontal and three scales that lie between the frontal and the second supralabial. One scale bordering the posterior edge of nasal between second supralabial and prefrontal. Divided nasal scale bordered anteriorly by the rostral, dorsally by the prefrontal, ventrally by the first and second supralabials, and posteriorly by a scale that lie between the second supralabial and prefrontal. Five scales in the first vertical row of lateral head scales, consisting of the second supralabial, the frontal, and three scales one above the other in between, namely one supraocular, one ocular, and one subocular, respectively. Eye spot absent. Supralabials 44; infralabials 44. Dorsals, laterals, and ventrals scales of body and tail smooth; no enlarged ventral plates. Scales around body 22­22­22; dorsals 543; ventrals 531; subcaudals 8. In life, dorsal and ventral color uniform bright yellowish brown; in preservative, the color faded to uniform yellowish brown. Total length
<quantity id="4CE49B7DFF8CFFB1FD3698A7FC28FEE8" box="[822,923,232,258]" metricMagnitude="-1" metricUnit="m" metricValue="3.62" pageId="3" pageNumber="22" unit="mm" value="362.0">362 mm</quantity>
; tail length
<quantity id="4CE49B7DFF8CFFB1FA2498A7FBD8FEE8" box="[1060,1131,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="22" unit="mm" value="4.0">4 mm</quantity>
.
</paragraph>
<paragraph id="8BA33698FF8CFFB1FEC69940FD98FEB8" blockId="3.[151,1437,152,338]" pageId="3" pageNumber="22">
<emphasis id="B968EA8AFF8CFFB1FEC69940FEF2FEC3" bold="true" box="[198,321,271,297]" pageId="3" pageNumber="22">Variation.</emphasis>
The female
<typeStatus id="54A7883AFF8CFFB1FFD5995FFD8EFEC0" box="[469,573,272,298]" pageId="3" pageNumber="22" type="paratype">paratype</typeStatus>
, in life, had dorsum and venter bright reddish brown (
<figureCitation id="13272A1DFF8CFFB1FAB1995FFB4BFEC0" box="[1201,1272,272,298]" captionStart="FIGURE 2" captionStartId="4.[151,255,1074,1098]" captionTargetBox="[188,1398,194,1049]" captionTargetId="figure@4.[188,1398,194,1050]" captionTargetPageId="4" captionText="FIGURE 2. Liotyphlops trefauti sp. nov. (MZUSP 12179, paratype), in life." httpUri="https://zenodo.org/record/175295/files/figure.png" pageId="3" pageNumber="22">Fig. 2</figureCitation>
); dorsals 520; ventrals 499; total length
<quantity id="4CE49B7DFF8CFFB1FFC29977FD9BFEB8" box="[450,552,312,338]" metricMagnitude="-1" metricUnit="m" metricValue="3.85" pageId="3" pageNumber="22" unit="mm" value="385.0">385 mm</quantity>
.
</paragraph>
<caption id="DF636610FF8CFFB1FE979DBEFB88F9C1" httpUri="https://zenodo.org/record/175294/files/figure.png" pageId="3" pageNumber="22" targetBox="[186,1382,395,1469]" targetPageId="3">
<paragraph id="8BA33698FF8CFFB1FE979DBEFB88F9C1" blockId="3.[151,1436,1521,1579]" pageId="3" pageNumber="22">
<emphasis id="B968EA8AFF8CFFB1FE979DBEFEAAF9E3" bold="true" box="[151,281,1521,1545]" pageId="3" pageNumber="22">FIGURE 1.</emphasis>
<taxonomicName id="4C1C4D1BFF8CFFB1FF259DBDFE45F9E3" box="[293,502,1522,1545]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8CFFB1FF259DBDFE45F9E3" box="[293,502,1522,1545]" italics="true" pageId="3" pageNumber="22">Liotyphlops trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8CFFB1FFFD9DBEFDE1F9E3" box="[509,594,1521,1545]" pageId="3" pageNumber="22" rank="species">
<emphasis id="B968EA8AFF8CFFB1FFFD9DBEFDE1F9E3" bold="true" box="[509,594,1521,1545]" pageId="3" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
(MZUSP 12178, holotype). (A) Dorsal view of head. (B) Lateral view of head. (C) Ventral view of head. (D) Ventral view of tail. (E) Lateral view of tail. Bar = 2 mm.
</paragraph>
</caption>
<paragraph id="8BA33698FF8CFFB1FEC69E10FD68F853" blockId="3.[151,1438,1631,1977]" pageId="3" pageNumber="22">
<emphasis id="B968EA8AFF8CFFB1FEC69E10FE09F993" bold="true" box="[198,442,1631,1657]" pageId="3" pageNumber="22">Habitat and habits.</emphasis>
The
<typeStatus id="54A7883AFF8CFFB1FFF99E10FDECF993" box="[505,607,1631,1657]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1C4D1BFF8CFFB1FC899E10FD4DF992" box="[649,766,1631,1656]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8CFFB1FC899E10FD4DF992" box="[649,766,1631,1656]" italics="true" pageId="3" pageNumber="22">L. trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8CFFB1FD079E10FCD5F993" box="[775,870,1631,1657]" pageId="3" pageNumber="22" rank="species">
<emphasis id="B968EA8AFF8CFFB1FD079E10FCD5F993" bold="true" box="[775,870,1631,1657]" pageId="3" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
was collected at Fazenda Bananeira, in Murici, Alagoas, where is located one of the biggest and currently the more significative remaining spot of the Atlantic Rain Forest in Northern and Northeastern
<collectingCountry id="F30B7608FF8CFFB1FCAA9EE0FD45F923" box="[682,758,1711,1737]" name="Brazil" pageId="3" pageNumber="22">Brazil</collectingCountry>
. With about 1,200 hectares and 640 meters in elevation, it is covered by a dense forest with a canopy up to 40 meters high. The closed canopy maintains the shadow and humidity inside the forest. The low­growing layer is composed by plantules of the arboreal regeneration, some palms, and large number of lianes and epiphyte plants. The litter is thick and moist. Surrounding the Murici Forest are extensive sugar cane plantations and these isolate completely the forest island from all other similar forest spots. The specimen was collected at 12:30h. through active search inside the thick and moist litter around the trunk of a large tree in the forest.
</paragraph>
<caption id="DF636610FF8BFFB6FE979C7DFC78FBA0" box="[151,971,1074,1098]" httpUri="https://zenodo.org/record/175295/files/figure.png" pageId="4" pageNumber="23" targetBox="[188,1398,194,1049]" targetPageId="4">
<paragraph id="8BA33698FF8BFFB6FE979C7DFC78FBA0" blockId="4.[151,971,1074,1098]" box="[151,971,1074,1098]" pageId="4" pageNumber="23">
<emphasis id="B968EA8AFF8BFFB6FE979C7DFEAAFBA0" bold="true" box="[151,281,1074,1098]" pageId="4" pageNumber="23">FIGURE 2.</emphasis>
<taxonomicName id="4C1C4D1BFF8BFFB6FF269C7CFE4AFBA0" box="[294,505,1075,1098]" class="Reptilia" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="trefauti" status="sp. nov.">
<emphasis id="B968EA8AFF8BFFB6FF269C7CFE4AFBA0" box="[294,505,1075,1098]" italics="true" pageId="4" pageNumber="23">Liotyphlops trefauti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57F1FF8BFFB6FFFF9C7DFDE6FBA0" box="[511,597,1074,1098]" pageId="4" pageNumber="23" rank="species">
<emphasis id="B968EA8AFF8BFFB6FFFF9C7DFDE6FBA0" bold="true" box="[511,597,1074,1098]" pageId="4" pageNumber="23">sp. nov.</emphasis>
</taxonomicNameLabel>
(MZUSP 12179, paratype), in life.
</paragraph>
</caption>
<paragraph id="8BA33698FF8BFFB6FEC69C37FB66FAE0" blockId="4.[151,1437,1144,1650]" pageId="4" pageNumber="23">
The
<typeStatus id="54A7883AFF8BFFB6FEFB9C37FED3FB78" box="[251,352,1144,1170]" pageId="4" pageNumber="23" type="paratype">paratype</typeStatus>
was collected at the CEPLAC Regional Center in Ilhéus, Bahia, in a cocoa plantation shadowed by the former high Atlantic Rain Forest. The specimen was obtained also by active search in the thick and moist litter, at 10:00h. Another specimen was seen on the same day and in the same area, inside the nest of ants of the genus
<taxonomicName id="4C1C4D1BFF8BFFB6FF649CBFFE4AFAE3" box="[356,505,1264,1289]" class="Insecta" family="Formicidae" genus="Acromyrmex" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B968EA8AFF8BFFB6FF649CBFFE4AFAE3" box="[356,505,1264,1289]" italics="true" pageId="4" pageNumber="23">Acromyrmex</emphasis>
</taxonomicName>
(
<taxonomicName id="4C1C4D1BFF8BFFB6FC079CBFFD18FAE0" box="[519,683,1264,1290]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
,
<taxonomicName id="4C1C4D1BFF8BFFB6FCB79CBFFCF2FAE0" box="[695,833,1264,1290]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
), but the efforts to collect it failed.
</paragraph>
<paragraph id="8BA33698FF8BFFB6FEC69D57FDB8FA68" blockId="4.[151,1437,1144,1650]" pageId="4" pageNumber="23">
The very young and poorly preserved referred specimen was collected in the thick and moist litter of the
<collectingCountry id="F30B7608FF8BFFB6FE979D0FFF60FAB0" box="[151,211,1344,1370]" name="Malta" pageId="4" pageNumber="23">Mata</collectingCountry>
da Cachoeira, a secondary forest island of about 100 hectares and 437 meters in elevation inside a large matrix of sugar cane plantation.
</paragraph>
<paragraph id="8BA33698FF8BFFB6FEC69DDFFF5BFA10" blockId="4.[151,1437,1144,1650]" pageId="4" pageNumber="23">
<emphasis id="B968EA8AFF8BFFB6FEC69DDFFDA5FA40" bold="true" box="[198,534,1424,1450]" pageId="4" pageNumber="23">Geographical distribution.</emphasis>
The species is known from three localities in the Atlantic Rain Forest of Northeastern
<collectingCountry id="F30B7608FF8BFFB6FF359DF7FE33FA38" box="[309,384,1464,1490]" name="Brazil" pageId="4" pageNumber="23">Brazil</collectingCountry>
, in the states of Alagoas (Murici and São José da Lage) and Bahia (Ilhéus),
<quantity id="4CE49B7DFF8BFFB6FAF29DF7FADAFA38" box="[1266,1385,1464,1490]" metricMagnitude="2" metricUnit="m" metricValue="3.45" metricValueMax="6.4" metricValueMin="0.5" pageId="4" pageNumber="23" unit="m" value="345.0" valueMax="640.0" valueMin="50.0">50 640 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="C3066513FF8BFFB6FEC69E47FE72F998" pageId="4" pageNumber="23" type="etymology">
<paragraph id="8BA33698FF8BFFB6FEC69E47FE72F998" blockId="4.[151,1437,1144,1650]" pageId="4" pageNumber="23">
<emphasis id="B968EA8AFF8BFFB6FEC69E47FEE0F9C8" bold="true" box="[198,339,1544,1570]" pageId="4" pageNumber="23">Etymology.</emphasis>
The specific name, a noun in the genitive case, honors Miguel Trefaut Rodrigues, by his extensive contribution to the knowledge of the Brazilian herpetology. We are very glad in to prive his pleasant and profitable friendship.
</paragraph>
</subSubSection>
</treatment>
</document>