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<document id="63D85F8F6A84382CBA7E57700B710B9A" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629573285391" checkinUser="felipe" docAuthor="Rasmussen, Claus" docDate="2008" docId="03B08272CB39FF94FCF3FB17FB56F953" docLanguage="en" docName="zt01935p080.pdf" docOrigin="Zootaxa 1935" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Tetragonula sarawakensis" docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="FF89FA0ACB09FFA4FC64FFD8FFF7FF99" masterDocTitle="Catalog of the Indo-Malayan / Australasian stingless bees (Hymenoptera: Apidae: Meliponini)" masterLastPageNumber="80" masterPageNumber="1" pageNumber="49" updateTime="1699077896071" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4F384566FA81E432469E8A788167D240">Catalog of the Indo-Malayan / Australasian stingless bees (Hymenoptera: Apidae: Meliponini)</mods:title>
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<mods:namePart id="F829F406A1011DEEB02D1E024E673819">Rasmussen, Claus</mods:namePart>
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<mods:date id="0286353936ECA30F1D98ABCF97D86B18">2008</mods:date>
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<mods:number id="7A6C3E5B41EC234EE045AE2BBACE82F3">1935</mods:number>
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<treatment id="03B08272CB39FF94FCF3FB17FB56F953" ID-DOI="http://doi.org/10.5281/zenodo.6319601" ID-Zenodo-Dep="6319601" LSID="urn:lsid:plazi:treatment:03B08272CB39FF94FCF3FB17FB56F953" httpUri="http://treatment.plazi.org/id/03B08272CB39FF94FCF3FB17FB56F953" lastPageNumber="49" pageId="48" pageNumber="49">
<subSubSection id="C30360EFCB39FF94FCF3FB17FD56FB70" box="[151,673,1231,1257]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="8BA63364CB39FF94FCF3FB17FD56FB70" blockId="48.[151,673,1231,1257]" box="[151,673,1231,1257]" pageId="48" pageNumber="49">
<heading id="D0EE8408CB39FF94FCF3FB17FD56FB70" bold="true" box="[151,673,1231,1257]" fontSize="11" level="1" pageId="48" pageNumber="49" reason="1">
<taxonomicName id="4C1948E7CB39FF94FCF3FB17FD56FB70" ID-CoL="55ND9" authority="(Schwarz, 1937)" baseAuthorityName="Schwarz" baseAuthorityYear="1937" box="[151,673,1231,1257]" class="Insecta" family="Apidae" genus="Tetragonula" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="B96DEF76CB39FF94FCF3FB17FD56FB70" bold="true" box="[151,673,1231,1257]" pageId="48" pageNumber="49">
<emphasis id="B96DEF76CB39FF94FCF3FB17FE25FB70" bold="true" box="[151,466,1231,1257]" italics="true" pageId="48" pageNumber="49">Tetragonula sarawakensis</emphasis>
(
<bibRefCitation id="EF884E95CB39FF94FD86FB17FD6EFB70" author="Schwarz, H. F." box="[482,665,1231,1257]" pageId="48" pageNumber="49" pagination="281 - 328" refId="ref70708" refString="Schwarz, H. F. (1937) Results of the Oxford University Sarawak (Borneo) expedition: Bornean stingless bees of the genus Trigona. Bulletin of the American Museum of Natural History, 73, 281 - 328." type="journal article" year="1937">Schwarz, 1937</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30360EFCB39FF94FCF3FAF8FB56F953" pageId="48" pageNumber="49" type="reference_group">
<paragraph id="8BA63364CB39FF94FCF3FAF8FB56F953" blockId="48.[151,1436,1311,1738]" pageId="48" pageNumber="49">
<treatmentCitationGroup id="AB09144ACB39FF94FCF3FAF8FB56F953" pageId="48" pageNumber="49">
<treatmentCitation id="0AB81575CB39FF94FCF3FAF8FD7EFAA3" author="Schwarz, H. F." box="[151,649,1311,1338]" page="283" pageId="48" pageNumber="49" year="1937">
<taxonomicName id="4C1948E7CB39FF94FCF3FAF8FD7EFAA3" ID-CoL="58QYV" authority="Schwarz 1937: 283" authorityName="Schwarz" authorityPageNumber="283" authorityYear="1937" box="[151,649,1311,1338]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="B96DEF76CB39FF94FCF3FAF8FE6EFAA0" box="[151,409,1312,1337]" italics="true" pageId="48" pageNumber="49">Trigona sarawakensis</emphasis>
<bibRefCitation id="EF884E95CB39FF94FDFBFAC7FD7EFAA3" author="Schwarz, H. F." box="[415,649,1311,1338]" pageId="48" pageNumber="49" pagination="281 - 328" refId="ref70708" refString="Schwarz, H. F. (1937) Results of the Oxford University Sarawak (Borneo) expedition: Bornean stingless bees of the genus Trigona. Bulletin of the American Museum of Natural History, 73, 281 - 328." type="journal article" year="1937">
<emphasis id="B96DEF76CB39FF94FDFBFAC7FDFDFAA0" bold="true" box="[415,522,1311,1337]" pageId="48" pageNumber="49">Schwarz</emphasis>
1937: 283
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 290, 313-315*, 316, 328, plate 2, 4:
<emphasis id="B96DEF76CB39FF94F859FAC7FB58FAA0" bold="true" box="[1085,1199,1311,1337]" pageId="48" pageNumber="49">
<typeStatus id="54A28DC6CB39FF94F859FAC7FB58FAA0" box="[1085,1199,1311,1337]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
</emphasis>
(BMNH 17b.1110);
<typeStatus id="54A28DC6CB39FF94FCA1FA90FEC2FAFB" box="[197,309,1352,1378]" pageId="48" pageNumber="49" type="paratype">paratypes</typeStatus>
(AMNH, BMNH?) (key to species, taxonomy);
<emphasis id="B96DEF76CB39FF94FFE8FA9FFB15FAF8" bold="true" box="[908,1250,1351,1377]" pageId="48" pageNumber="49">
Type locality:
<collectingCountry id="F30E73F4CB39FF94F821FA9FFB15FAF8" box="[1093,1250,1351,1377]" name="Malaysia" pageId="48" pageNumber="49">MALAYSIA</collectingCountry>
</emphasis>
&quot;
<collectingRegion id="49DDFD86CB39FF94F892FA90FA94FAFB" box="[1270,1379,1352,1378]" country="Malaysia" name="Sarawak" pageId="48" pageNumber="49">Sarawak</collectingRegion>
: Mt. Dulit,
<quantity id="4CE19E81CB39FF94FD76FAA8FE85FA13" box="[274,370,1392,1418]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="48" pageNumber="49" unit="ft" value="4000.0">4000 ft.</quantity>
&quot; (
<specimenCount id="9D1FF8EDCB39FF94FDEAFAA8FDF2FA13" box="[398,517,1392,1418]" pageId="48" pageNumber="49" type="worker">2 workers</specimenCount>
<date id="FFA715A4CB39FF94FE6BFAA8FD40FA13" box="[527,695,1392,1418]" pageId="48" pageNumber="49" value="1932-10-18">Oct. 18, 1932</date>
;
<specimenCount id="9D1FF8EDCB39FF94FEA2FAA8FCCBFA13" box="[710,828,1392,1418]" pageId="48" pageNumber="49" type="worker">2 workers</specimenCount>
<date id="FFA715A4CB39FF94FF22FAA8FC19FA13" box="[838,1006,1392,1418]" pageId="48" pageNumber="49" value="1932-10-19">Oct. 19, 1932</date>
;
<specimenCount id="9D1FF8EDCB39FF94FF98FAA8FB90FA13" box="[1020,1127,1392,1418]" pageId="48" pageNumber="49" type="worker">1 worker</specimenCount>
<date id="FFA715A4CB39FF94F815FAA8FAEDFA13" box="[1137,1306,1392,1418]" pageId="48" pageNumber="49" value="1932-10-22">Oct. 22, 1932</date>
);
<treatmentCitation id="0AB81575CB39FF94F954FAB7FECBFA2B" author="Schwarz, H. F." page="85" pageId="48" pageNumber="49" year="1939">
<bibRefCitation id="EF884E95CB39FF94F954FAB7FECBFA2B" author="Schwarz, H. F." pageId="48" pageNumber="49" pagination="83 - 141" refId="ref70751" refString="Schwarz, H. F. (1939 a) The Indo-Malayan species of Trigona. Bulletin of the American Museum of Natural History, 76, 83 - 141." type="journal article" year="1939">
<emphasis id="B96DEF76CB39FF94F954FAB7FA6CFA10" bold="true" box="[1328,1435,1391,1417]" pageId="48" pageNumber="49">Schwarz</emphasis>
1939a: 85
</bibRefCitation>
</treatmentCitation>
, 92, 106* (distribution, key to species, taxonomy);
<treatmentCitation id="0AB81575CB39FF94FFFAFA4FFB7CFA2B" author="Schwarz, H. F." box="[926,1163,1431,1458]" page="118" pageId="48" pageNumber="49" year="1948">
<bibRefCitation id="EF884E95CB39FF94FFFAFA4FFB7CFA2B" author="Schwarz, H. F." box="[926,1163,1431,1458]" pageId="48" pageNumber="49" pagination="1 - 546" refId="ref70858" refString="Schwarz, H. F. (1948) Stingless bees (Meliponidae) of the western hemisphere. Lestrimelitta and the following subgenera of Trigona: Trigona, Paratrigona, Schwarziana, Parapartamona, Cephalotrigona, Oxytrigona, Scaura, and Mourella. Bulletin of the American Museum of Natural History, 90, 1 - 546." type="journal article" year="1948">
<emphasis id="B96DEF76CB39FF94FFFAFA4FFBFEFA28" bold="true" box="[926,1033,1431,1457]" pageId="48" pageNumber="49">Schwarz</emphasis>
1948: 118
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB81575CB39FF94F8FCFA4FFF06FA43" author="Sakagami, S. F." page="119" pageId="48" pageNumber="49" year="1959">
<bibRefCitation id="EF884E95CB39FF94F8FCFA4FFF06FA43" author="Sakagami, S. F." pageId="48" pageNumber="49" pagination="119 - 121" refId="ref68708" refString="Sakagami, S. F. (1959) Stingless bees collected by Prof. S. Matsumura from Singapore (Hymenoptera, Apidae). Insecta Matsumurana, 22, 119 - 121." type="journal article" year="1959">
<emphasis id="B96DEF76CB39FF94F8FCFA4FFAE3FA28" bold="true" box="[1176,1300,1431,1457]" pageId="48" pageNumber="49">Sakagami</emphasis>
1959: 119- 120
</bibRefCitation>
</treatmentCitation>
(distribution, morphology, variation);
<collectingCountry id="F30E73F4CB39FF94FEAAFA18FC80FA43" box="[718,887,1472,1498]" name="Singapore" pageId="48" pageNumber="49">SINGAPORE</collectingCountry>
;
<treatmentCitation id="0AB81575CB39FF94FFECFA67FB90FA43" author="Moure, J. S." box="[904,1127,1471,1498]" page="210" pageId="48" pageNumber="49" year="1961">
<bibRefCitation id="EF884E95CB39FF94FFECFA67FB90FA43" author="Moure, J. S." box="[904,1127,1471,1498]" pageId="48" pageNumber="49" pagination="181 - 242" refId="ref65305" refString="Moure, J. S. (1961) A preliminary supra-specific classification of the old world meliponine bees (Hymenoptera, Apoidea). Studia Entomologica, 4, 181 - 242." type="journal article" year="1961">
<emphasis id="B96DEF76CB39FF94FFECFA67FC2BFA40" bold="true" box="[904,988,1471,1497]" pageId="48" pageNumber="49">Moure</emphasis>
1961: 210
</bibRefCitation>
</treatmentCitation>
(morphology, systematic position);
<treatmentCitation id="0AB81575CB39FF94FD59FA3FFD70F99B" author="Yoshikawa, K. &amp; Ohgushi, R. &amp; Sakagami, S. F." box="[317,647,1511,1538]" page="168" pageId="48" pageNumber="49" year="1969">
<bibRefCitation id="EF884E95CB39FF94FD59FA3FFD70F99B" author="Yoshikawa, K. &amp; Ohgushi, R. &amp; Sakagami, S. F." box="[317,647,1511,1538]" pageId="48" pageNumber="49" pagination="153 - 182" refId="ref75107" refString="Yoshikawa, K., Ohgushi, R., &amp; Sakagami, S. F. (1969) Preliminary report on entomology of the Osaka city University 5 th scientific expedition to Southeast Asia 1966. With descriptions of two new genera of stenogastrine waps by J. van der Vecht. Nature and Life in Southeast Asia, 6, 153 - 182." type="journal article" year="1969">
<emphasis id="B96DEF76CB39FF94FD59FA3FFDF0F998" bold="true" box="[317,519,1511,1537]" pageId="48" pageNumber="49">
Yoshikawa
<emphasis id="B96DEF76CB39FF94FDA8FA3FFDF0F998" bold="true" box="[460,519,1511,1537]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
</emphasis>
1969: 168
</bibRefCitation>
</treatmentCitation>
, 169 (nest);
<treatmentCitation id="0AB81575CB39FF94FF7FFA3FFB98F99B" author="Wille, A. &amp; Michener, C. D." box="[795,1135,1511,1538]" page="14" pageId="48" pageNumber="49" year="1973">
<bibRefCitation id="EF884E95CB39FF94FF7FFA3FFB98F99B" author="Wille, A. &amp; Michener, C. D." box="[795,1135,1511,1538]" pageId="48" pageNumber="49" pagination="9 - 278" refId="ref74408" refString="Wille, A. &amp; Michener, C. D. (1973) The nest architecture of stingless bees with special reference to those of Costa Rica. Revista de Biologia Tropical, 21 (supl.), 9 - 278." type="journal article" year="1973">
<emphasis id="B96DEF76CB39FF94FF7FFA3FFC0AF998" bold="true" box="[795,1021,1511,1537]" pageId="48" pageNumber="49">Wille &amp; Michener</emphasis>
1973: 14
</bibRefCitation>
</treatmentCitation>
, 25, 59 (nest);
<treatmentCitation id="0AB81575CB39FF94F944FA3FFE84F9B3" author="Sakagami, S. F." page="166" pageId="48" pageNumber="49" year="1978">
<bibRefCitation id="EF884E95CB39FF94F944FA3FFE84F9B3" author="Sakagami, S. F." pageId="48" pageNumber="49" pagination="165 - 247" refId="ref68895" refString="Sakagami, S. F. (1978) Tetragonula stingless bees of the continental Asia and Sri Lanka (Hymenoptera, Apidae). Journal of the Faculty of Science, Hokkaido University, Series VI, Zoology, 21, 165 - 247." type="journal article" year="1978">
<emphasis id="B96DEF76CB39FF94F944FA3FFA6BF998" bold="true" box="[1312,1436,1511,1537]" pageId="48" pageNumber="49">Sakagami</emphasis>
1978: 166-194
</bibRefCitation>
</treatmentCitation>
, 210-211*, 228, 229, 230, 236, 237, 238 (taxonomy);
<treatmentCitation id="0AB81575CB39FF94FF94F9D7FAE8F9B3" author="Sakagami, S. F. &amp; Yamane, S. &amp; Hambali, G. G." box="[1008,1311,1551,1578]" page="2" pageId="48" pageNumber="49" year="1983">
<bibRefCitation id="EF884E95CB39FF94FF94F9D7FAE8F9B3" author="Sakagami, S. F. &amp; Yamane, S. &amp; Hambali, G. G." box="[1008,1311,1551,1578]" pageId="48" pageNumber="49" pagination="1 - 21" refId="ref69768" refString="Sakagami, S. F., Yamane, S., &amp; Hambali, G. G. (1983 b) Nests of some southeast Asian stingless bees. Bulletin of the Faculty of Education, Ibaraki University (Natural Sciences), 32, 1 - 21." type="journal article" year="1983">
<emphasis id="B96DEF76CB39FF94FF94F9D7FB59F9B0" bold="true" box="[1008,1198,1551,1577]" pageId="48" pageNumber="49">
Sakagami
<emphasis id="B96DEF76CB39FF94F817F9D7FB59F9B0" bold="true" box="[1139,1198,1551,1577]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
</emphasis>
1983b: 2
</bibRefCitation>
</treatmentCitation>
, 10 (nest);
<treatmentCitation id="0AB81575CB39FF94FCA1F9EFFDF9F9CB" author="Sakagami, S. F. &amp; Inoue, T. &amp; Yamane, S. &amp; Salmah, S." box="[197,526,1591,1618]" page="106" pageId="48" pageNumber="49" year="1983">
<bibRefCitation id="EF884E95CB39FF94FCA1F9EFFDF9F9CB" author="Sakagami, S. F. &amp; Inoue, T. &amp; Yamane, S. &amp; Salmah, S." box="[197,526,1591,1618]" pageId="48" pageNumber="49" pagination="100 - 111" refId="ref69479" refString="Sakagami, S. F., Inoue, T., Yamane, S., &amp; Salmah, S. (1983 a) Nest architecture and colony composition of the Sumatran stingless bee Trigona (Tetragonula) laeviceps. Kontyu, 51, 100 - 111." type="journal article" year="1983">
<emphasis id="B96DEF76CB39FF94FCA1F9EFFE74F9C8" bold="true" box="[197,387,1591,1617]" pageId="48" pageNumber="49">
Sakagami
<emphasis id="B96DEF76CB39FF94FD2CF9EFFE74F9C8" bold="true" box="[328,387,1591,1617]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
</emphasis>
1983a: 106
</bibRefCitation>
</treatmentCitation>
(comparative note);
<emphasis id="B96DEF76CB39FF94FF60F9EFFC18F9C8" bold="true" box="[772,1007,1591,1617]" pageId="48" pageNumber="49">Sakagami &amp; Inoue</emphasis>
1985: 174, 176, 177, 178, 179, 180, 181, 182*, 184, 186 (key to species, taxonomy);
<emphasis id="B96DEF76CB39FF94FF64F987FC49F9E0" bold="true" box="[768,958,1631,1657]" pageId="48" pageNumber="49">
Sakagami
<emphasis id="B96DEF76CB39FF94FFE7F987FC49F9E0" bold="true" box="[899,958,1631,1657]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
</emphasis>
1985: (distribution);
<treatmentCitation id="0AB81575CB39FF94F8DCF987FED9F93B" author="Starr, C. K. &amp; Sakagami, S. F." page="97" pageId="48" pageNumber="49" year="1987">
<bibRefCitation id="EF884E95CB39FF94F8DCF987FED9F93B" author="Starr, C. K. &amp; Sakagami, S. F." pageId="48" pageNumber="49" pagination="96 - 107" refId="ref71777" refString="Starr, C. K. &amp; Sakagami, S. F. (1987) An extraordinary concentration of stingless bee colonies in the Philippines, with notes on nest structure (Hymenoptera: Apidae: Trigona spp.). Insectes Sociaux, 34, 96 - 107." type="journal article" year="1987">
<emphasis id="B96DEF76CB39FF94F8DCF987FA6BF9E0" bold="true" box="[1208,1436,1631,1657]" pageId="48" pageNumber="49">Starr &amp; Sakagami</emphasis>
1987: 97
</bibRefCitation>
</treatmentCitation>
(citation);
<emphasis id="B96DEF76CB39FF94FDD2F95FFD8FF938" bold="true" box="[438,632,1671,1697]" pageId="48" pageNumber="49">
Sakagami
<emphasis id="B96DEF76CB39FF94FE5FF95FFD8FF938" bold="true" box="[571,632,1671,1697]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
</emphasis>
1990: 131;
<treatmentCitation id="0AB81575CB39FF94FF68F95FFB8CF93B" author="Nagamitsu, T. &amp; Inoue, T." box="[780,1147,1671,1698]" page="77" pageId="48" pageNumber="49" year="2005">
<bibRefCitation id="EF884E95CB39FF94FF68F95FFB8CF93B" author="Nagamitsu, T. &amp; Inoue, T." box="[780,1147,1671,1698]" pageId="48" pageNumber="49" pagination="73 - 88" refId="ref65553" refString="Nagamitsu, T. &amp; Inoue, T. (2005). Floral resource utilization by stingless bees (Apidae, Meliponini). In: D. W. Roubik, S. Sakai &amp; A. A. Hamid (Eds.), Pollination ecology and the rain forest, Sarawak studies. Vol. 174, Springer, pp. 73 - 88." type="book chapter" year="2005">
<emphasis id="B96DEF76CB39FF94FF68F95FFBF0F938" bold="true" box="[780,1031,1671,1697]" pageId="48" pageNumber="49">Nagamitsu &amp; Inoue</emphasis>
2005: 77
</bibRefCitation>
</treatmentCitation>
(distribution, foraging);
<treatmentCitation id="0AB81575CB39FF94FCA1F977FDF3F953" author="Klakasikorn, A. &amp; Wongsiri, S. &amp; Deowanish, S. &amp; Duangphakdee, O." box="[197,516,1711,1738]" page="6" pageId="48" pageNumber="49" year="2005">
<bibRefCitation id="EF884E95CB39FF94FCA1F977FDF3F953" author="Klakasikorn, A. &amp; Wongsiri, S. &amp; Deowanish, S. &amp; Duangphakdee, O." box="[197,516,1711,1738]" pageId="48" pageNumber="49" pagination="1 - 7" refId="ref63285" refString="Klakasikorn, A., Wongsiri, S., Deowanish, S., &amp; Duangphakdee, O. (2005) New record of stingless bees (Meliponini: Trigona) in Thailand. Natural History Journal of Chulalongkorn University, 5, 1 - 7." type="journal article" year="2005">
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