528 lines
85 KiB
XML
528 lines
85 KiB
XML
<document id="3C40A3FE39194D10906D3C50FA7DD4C8" ID-CLB-Dataset="29256" ID-DOI="10.11646/zootaxa.4496.1.28" ID-GBIF-Dataset="059d48b1-7ed5-4d58-9183-f851e0389a94" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446907" ID-ZooBank="BDCE08F3-0743-471A-A667-2D6A9D913EAC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538747702184" checkinUser="plazi" docAuthor="Zhang, Junqian, Lu, Yajing & Xie, Zhicai" docDate="2018" docId="03A9E95F2724007A60EFCABF980BFE76" docLanguage="en" docName="zootaxa.4496.1.28.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesenchytraeus spermatoglomeratus Zhang & Lu & Xie 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="386" masterDocId="FF9091272725007F6078CF3F991DFFFE" masterDocTitle="Two new Mesenchytraeus species (Annelida: Clitellata: Enchytraeidae) from Changbai Mountain, China" masterLastPageNumber="394" masterPageNumber="382" pageNumber="383" updateTime="1698566824926" updateUser="ExternalLinkService">
|
||
<mods:mods id="65D8714C5EEFB5D29EFF230010205292" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="12C36BC77407DCE896C31271516297E6">
|
||
<mods:title id="510A49BEF2BFE8A39E84D50AFC59F6FC">Two new Mesenchytraeus species (Annelida: Clitellata: Enchytraeidae) from Changbai Mountain, China</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6129D421125590FB31A8F2D8693BD66F" type="personal">
|
||
<mods:role id="FFE5A62A7E43725B5B3986B1AA10E57B">
|
||
<mods:roleTerm id="520CE19483D2749B778DD39A2D8E86B9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B94EE0518A16BED0CC3B0F6F02FA831F">Zhang, Junqian</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D48566CC35D654E5B590B6037E22BF5F" type="personal">
|
||
<mods:role id="11305C2D9014682E3F9F1E4603D51DF0">
|
||
<mods:roleTerm id="7EF19837E3329871C0A0F937B864F05A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="094FE8A0C7F2DF05714BD88EA3155A8B">Lu, Yajing</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D7180A0E08BBA2090F4CD97C9A99D35C" type="personal">
|
||
<mods:role id="520FF30E96F579DE4D0915EFF7E17D7E">
|
||
<mods:roleTerm id="44D779BBCB61875931528D229DEA6C75">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="90360ED98FB4D55690C2CA7F5E65096C">Xie, Zhicai</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="343303A8DCE624B279D3B010243EB3F3">text</mods:typeOfResource>
|
||
<mods:relatedItem id="CC9A91BA2EF6ADBEC3FA7001967B6AF5" type="host">
|
||
<mods:titleInfo id="2BE8A66F2C1AD6C6F14BEEF916ECC28B">
|
||
<mods:title id="72C5460BC2999B3639797F4719A4AB5D">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="DD6434706DB9636CE432232DD33E98C9">
|
||
<mods:date id="839273ED279A595910284ABD255762FE">2018</mods:date>
|
||
<mods:detail id="733E14C7CAD4DA69A352501A81CEDABC" type="pubDate">
|
||
<mods:number id="E0BAC24D4C2B83646824CB1336A76477">2018-10-05</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A31B74AE41CE77394AD9336335AF18F0" type="volume">
|
||
<mods:number id="D652B0DCA2D6EDE341062F0FF434D9A7">4496</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4809CA3A86F9CCE0A51458A9706F7FA1" type="issue">
|
||
<mods:number id="4743ABDBF3C206E3B6593F7EC37ACB64">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="8A52C98371B1C4BE7DE5E1E9E7EB053A" unit="page">
|
||
<mods:start id="21B2FAD43903E5A98F4DE850866F4CFC">382</mods:start>
|
||
<mods:end id="D9BC2D2B43243966FBEF758FAAFB7116">394</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="005E0B3772FDF2E9569777D13E76C474">journal article</mods:classification>
|
||
<mods:identifier id="3F3FF34A539A3679D46B031D01EBD385" type="CLB-Dataset">29256</mods:identifier>
|
||
<mods:identifier id="1DF3D7D9A1E152856F87650B91A40893" type="DOI">10.11646/zootaxa.4496.1.28</mods:identifier>
|
||
<mods:identifier id="CB36EFC4F35A26886EBE4F7FA7CC4813" type="GBIF-Dataset">059d48b1-7ed5-4d58-9183-f851e0389a94</mods:identifier>
|
||
<mods:identifier id="FF4B6E52513977CFDD55AE486F5E4DAA" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="24F7D4F78C2777F34CBD7A04399C1B15" type="Zenodo-Dep">1446907</mods:identifier>
|
||
<mods:identifier id="79ACD34C628B8D4EB19B20BB51451A23" type="ZooBank">BDCE08F3-0743-471A-A667-2D6A9D913EAC</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03A9E95F2724007A60EFCABF980BFE76" ID-DOI="http://doi.org/10.5281/zenodo.5978310" ID-GBIF-Taxon="148639925" ID-Zenodo-Dep="5978310" LSID="urn:lsid:plazi:treatment:03A9E95F2724007A60EFCABF980BFE76" httpUri="http://treatment.plazi.org/id/03A9E95F2724007A60EFCABF980BFE76" lastPageId="5" lastPageNumber="386" pageId="1" pageNumber="383">
|
||
<subSubSection id="C31A0BC22724007E60EFCABF98ABFA42" pageId="1" pageNumber="383" type="nomenclature">
|
||
<paragraph id="8BBF58492724007E60EFCABF9BA0FA65" blockId="1.[151,701,1408,1469]" box="[151,701,1408,1435]" pageId="1" pageNumber="383">
|
||
<heading id="D0F7EF252724007E60EFCABF9BA0FA65" bold="true" box="[151,701,1408,1435]" fontSize="11" level="1" pageId="1" pageNumber="383" reason="1">
|
||
<emphasis id="B974845B2724007E60EFCABF9BA0FA65" bold="true" box="[151,701,1408,1435]" pageId="1" pageNumber="383">
|
||
<taxonomicName id="4C0023CA2724007E60EFCABF9B44FA64" authority="Zhang & Lu & Xie, 2018" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[151,601,1408,1434]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="383" phylum="Annelida" rank="species" species="spermatoglomeratus" status="sp. nov.">
|
||
<emphasis id="B974845B2724007E60EFCABF9B44FA64" bold="true" box="[151,601,1408,1434]" italics="true" pageId="1" pageNumber="383">Mesenchytraeus spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24739202724007E6218CABE9BA0FA65" box="[608,701,1409,1435]" pageId="1" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBF58492724007E60EFCA9B98ABFA42" blockId="1.[151,701,1408,1469]" box="[151,438,1444,1469]" pageId="1" pageNumber="383">
|
||
(
|
||
<figureCitation id="133B44CC2724007E60E6CA9B9814FA42" box="[158,265,1444,1469]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="1" pageNumber="383">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="133B44CC2724007E616FCA9B983AFA42" box="[279,295,1444,1468]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="1" pageNumber="383">2</figureCitation>
|
||
,
|
||
<tableCitation id="C6826DF22724007E614BCA9B988CFA42" box="[307,401,1444,1469]" captionStart="TABLE 1" captionStartId="2.[151,238,868,891]" captionText="TABLE 1. Chaetal sizes and numbers in different segments of Mesenchytraeus spermatoglomeratus sp. nov." pageId="1" pageNumber="383">Tables 1</tableCitation>
|
||
, 3)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A0BC22724007E60EFCAD3989BF8DB" pageId="1" pageNumber="383" type="materials_examined">
|
||
<paragraph id="8BBF58492724007E60EFCAD39AEFF9FB" blockId="1.[151,1437,1516,2009]" box="[151,1010,1516,1541]" pageId="1" pageNumber="383">
|
||
<emphasis id="B974845B2724007E60EFCAD39816F9FB" bold="true" box="[151,267,1516,1541]" pageId="1" pageNumber="383">Holotype.</emphasis>
|
||
Fully mature, whole-mounted specimen, stained, JLO201412005.
|
||
</paragraph>
|
||
<paragraph id="8BBF58492724007E60BFC9309AABF9B2" blockId="1.[151,1437,1516,2009]" pageId="1" pageNumber="383">
|
||
<materialsCitation id="3B6852142724007E60BFC9309AABF9B2" ID-GBIF-Occurrence="1914265511" collectingDate="2014-12" collectorName="Z. J. Piao" elevation="1034" location="Mt. Changbai" longitude="127.84088" pageId="1" pageNumber="383" specimenCount="1" stateProvince="Jilin Province" typeStatus="holotype">
|
||
<emphasis id="B974845B2724007E60BFC930987BF9D6" bold="true" box="[199,358,1551,1576]" pageId="1" pageNumber="383">
|
||
<typeStatus id="54BBE6EB2724007E60BFC930981FF9D6" box="[199,258,1551,1576]" pageId="1" pageNumber="383">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EDF0E922724007E6115C92F9B17F9D6" LSID="urn:lsid:plazi:treatment:03A9E95F2724007A60EFCABF980BFE76:8EDF0E922724007E6115C92F9B17F9D6" box="[365,522,1551,1576]" longitude="127.84088" name="Mt. Changbai" pageId="1" pageNumber="383" stateProvince="Jilin Province">Mt. Changbai</location>
|
||
,
|
||
<collectingRegion id="49C496AB2724007E626EC9309BA8F9D6" box="[534,693,1551,1576]" country="China" name="Jilin" pageId="1" pageNumber="383">Jilin Province</collectingRegion>
|
||
, hardwood forest, 42°180.607''N,
|
||
<geoCoordinate id="EE343E8E2724007E644CC9309DC9F9D6" box="[1076,1236,1551,1576]" direction="east" orientation="longitude" pageId="1" pageNumber="383" precision="1" value="127.84088">127°50.453''E</geoCoordinate>
|
||
,
|
||
<elevation id="002DBF7A2724007E6498C92F9C42F9D6" box="[1248,1375,1551,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.034" pageId="1" pageNumber="383" unit="m" value="1034.0">
|
||
<quantity id="4CF8F5AC2724007E6498C92F9C2BF9D9" box="[1248,1334,1552,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.034" pageId="1" pageNumber="383" unit="m" value="1034.0">1034 m</quantity>
|
||
asl
|
||
</elevation>
|
||
, dark brown forest soil under
|
||
<taxonomicName id="4C0023CA2724007E61DAC90B9B60F9B2" box="[418,637,1588,1612]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="383" phylum="Tracheophyta" rank="species" species="mongolica">
|
||
<emphasis id="B974845B2724007E61DAC90B9B60F9B2" box="[418,637,1588,1612]" italics="true" pageId="1" pageNumber="383">Quercus mongolica</emphasis>
|
||
</taxonomicName>
|
||
, coll.
|
||
<collectorName id="26F53D9F2724007E62BBC90A9A2DF9B3" box="[707,816,1588,1613]" pageId="1" pageNumber="383">Z. J. Piao</collectorName>
|
||
,
|
||
<collectingDate id="EFFA87612724007E6343C90A9AAFF9B2" box="[827,946,1588,1612]" pageId="1" pageNumber="383" value="2014-12">
|
||
<date id="FFBE7E892724007E6343C90A9AAFF9B2" box="[827,946,1588,1612]" pageId="1" pageNumber="383" value="2014-12">Dec. 2014</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BBF58492724007E60BFC968989BF8DB" blockId="1.[151,1437,1516,2009]" pageId="1" pageNumber="383">
|
||
<emphasis id="B974845B2724007E60BFC968985BF98E" bold="true" box="[199,326,1623,1648]" pageId="1" pageNumber="383">Paratypes.</emphasis>
|
||
JLO201412003–JLO201412004, two whole mounted adult specimens from Korean pine broadleaved mixed forest, 42°20.150''N, 127°53.056''E,
|
||
<quantity id="4CF8F5AC2724007E62AFC9439A33F96A" box="[727,814,1660,1684]" metricMagnitude="3" metricUnit="m" metricValue="1.084" pageId="1" pageNumber="383" unit="m" value="1084.0">1084 m</quantity>
|
||
asl, dark brown forest soil under
|
||
<taxonomicName id="4C0023CA2724007E64CCC9429C6DF96A" box="[1204,1392,1660,1684]" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="383" phylum="Tracheophyta" rank="species" species="koraiensis">
|
||
<emphasis id="B974845B2724007E64CCC9429C6DF96A" box="[1204,1392,1660,1684]" italics="true" pageId="1" pageNumber="383">Pinus koraiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="FFBE7E892724007E65F8C9439851F946" pageId="1" pageNumber="383" value="2014-12-10">10 December 2014</date>
|
||
, coll. Z. J. Piao; JLO201412006, one whole mounted adult specimen from type locality, same data as holotype. JLO201503001–JLO201503002, two whole mounted specimens, fully mature, from secondary
|
||
<taxonomicName id="4C0023CA2724007E652BC9FB9808F8FE" authorityName="Sukaczev" authorityYear="1911" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="383" phylum="Tracheophyta" rank="species" species="platyphylla">
|
||
<emphasis id="B974845B2724007E652BC9FB9808F8FE" italics="true" pageId="1" pageNumber="383">Betula platyphylla</emphasis>
|
||
</taxonomicName>
|
||
forest, 42°22.719''N, 128°01.020''E,
|
||
<quantity id="4CF8F5AC2724007E62CCC9D89BE6F901" box="[692,763,1767,1792]" metricMagnitude="2" metricUnit="m" metricValue="7.859999999999999" pageId="1" pageNumber="383" unit="m" value="786.0">786 m</quantity>
|
||
asl, dark brown forest soil under
|
||
<taxonomicName id="4C0023CA2724007E6409C9D79C23F8FE" authorityName="Sukaczev" authorityYear="1911" box="[1137,1342,1768,1792]" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="383" phylum="Tracheophyta" rank="species" species="platyphylla">
|
||
<emphasis id="B974845B2724007E6409C9D79C23F8FE" box="[1137,1342,1768,1792]" italics="true" pageId="1" pageNumber="383">Betula platyphylla</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="FFBE7E892724007E6534C9D799CFF8DB" pageId="1" pageNumber="383" value="2015-04-02">2 April 2015</date>
|
||
, coll. Z. J. Piao.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A0BC22724007E60BFC8109DE4F893" pageId="1" pageNumber="383" type="etymology">
|
||
<paragraph id="8BBF58492724007E60BFC8109DE4F893" blockId="1.[151,1437,1516,2009]" pageId="1" pageNumber="383">
|
||
<emphasis id="B974845B2724007E60BFC8109853F8B6" bold="true" box="[199,334,1839,1864]" pageId="1" pageNumber="383">Etymology.</emphasis>
|
||
Named after the shape of sperm bundles: "glomeratus" refers to "condensed into a ball". "
|
||
<taxonomicName id="4C0023CA2724007E60E7C86B9895F893" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[159,392,1876,1901]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="383" phylum="Annelida" rank="species" species="spermatoglomeratus">spermatoglomeratus</taxonomicName>
|
||
" refers to the sperm bundles of the new species, rolled into a round ball shape.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A0BC22724007D60BFC84899C7FF09" lastPageId="2" lastPageNumber="384" pageId="1" pageNumber="383" type="diagnosis">
|
||
<paragraph id="8BBF58492724007D60BFC84899C7FF09" blockId="1.[151,1437,1516,2009]" lastBlockId="2.[151,1437,151,824]" lastPageId="2" lastPageNumber="384" pageId="1" pageNumber="383">
|
||
<emphasis id="B974845B2724007E60BFC848985CF86E" bold="true" box="[199,321,1911,1936]" pageId="1" pageNumber="383">Diagnosis.</emphasis>
|
||
This new species is recognized by the following characters, diagnostic in combination: 1) enlarged ventral chaetae in segments V–VI and XI, 2 per bundle; 2) five pairs of preclitellar nephridia in VI/VII to X/XI; 3) sperm sacs asymmetrically paired from XI–XIV to XV–XXIV, containing curly ball-shaped sperm bundles; 4) spermathecae without diverticula, ental ducts connected with oesophagus in VI–VII; 5) atrium tubular, with 4–6 developed prostate glands, distinct from vas deferens; 6) vas deferens very long, extending backwards to XV– XXII.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A0BC22727007B60BFCE3B9D87FA08" lastPageId="4" lastPageNumber="386" pageId="2" pageNumber="384" type="description">
|
||
<paragraph id="8BBF58492727007D60BFCE3B9A40FCC6" blockId="2.[151,1437,151,824]" pageId="2" pageNumber="384">
|
||
<emphasis id="B974845B2727007D60BFCE3B9844FEE3" bold="true" box="[199,345,260,285]" pageId="2" pageNumber="384">Description.</emphasis>
|
||
Small worms, active, twisting when submerged in water, pale
|
||
<emphasis id="B974845B2727007D646ACE3A9D41FEE2" box="[1042,1116,261,284]" italics="true" pageId="2" pageNumber="384">in vivo</emphasis>
|
||
, body wall thin, transparent. Body length
|
||
<quantity id="4CF8F5AC2727007D615FCE1898A6FEC1" box="[295,443,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="11.5" metricValueMin="6.5" pageId="2" pageNumber="384" unit="mm" value="9.0" valueMax="11.5" valueMin="6.5">6.5–11.5 mm</quantity>
|
||
(
|
||
<typeStatus id="54BBE6EB2727007D61B2CE189B33FEBE" box="[458,558,295,320]" pageId="2" pageNumber="384" type="holotype">holotype</typeStatus>
|
||
8.0 mm), 7.0–
|
||
<quantity id="4CF8F5AC2727007D62ACCE189A2BFEC1" box="[724,822,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="384" unit="mm" value="12.5">12.5 mm</quantity>
|
||
<emphasis id="B974845B2727007D6345CE179A95FEC1" box="[829,904,296,319]" italics="true" pageId="2" pageNumber="384">in vivo</emphasis>
|
||
; body width 270–355 µm at V (330–450 µm
|
||
<emphasis id="B974845B2727007D65FECE1799DBFE9A" italics="true" pageId="2" pageNumber="384">in vivo</emphasis>
|
||
), 470–575 µm at clitellum (320–625 µm
|
||
<emphasis id="B974845B2727007D62DFCE729BE9FE9A" box="[679,756,333,356]" italics="true" pageId="2" pageNumber="384">in vivo</emphasis>
|
||
). Segments 28–46 (
|
||
<typeStatus id="54BBE6EB2727007D63A3CE739D22FE9B" box="[987,1087,332,357]" pageId="2" pageNumber="384" type="holotype">holotype</typeStatus>
|
||
34). Head pore transverse slit near the apex of prostomium (
|
||
<figureCitation id="133B44CC2727007D619FCE509B5FFE76" box="[487,578,367,392]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="2" pageNumber="384">Fig. 2A</figureCitation>
|
||
). Epidermal gland cells inconspicuous, almost absent. Chaetal formula: 2,3 - 3,4: 2–5(6) - 2–5(6), up to 6 per bundle in the last segments. Chaetae sigmoid, nodulated, distally thinner and single-pointed. Lateral chaetae 44–100 µm in length and 3.5–5 µm in maximal width, shortest in II (44–48 µm), gradually increasing in size from III backwards, maximal size in IX (80–100 µm) (
|
||
<tableCitation id="C6826DF22727007D6457CEE39D9FFE0A" box="[1071,1154,476,501]" captionStart="TABLE 1" captionStartId="2.[151,238,868,891]" captionText="TABLE 1. Chaetal sizes and numbers in different segments of Mesenchytraeus spermatoglomeratus sp. nov." pageId="2" pageNumber="384">Table 1</tableCitation>
|
||
). Ventral chaetae, except those in V, VI and XI, 63–120 µm in length and 4–7.5 µm in maximal width (
|
||
<figureCitation id="133B44CC2727007D6478CEC09D60FDE6" box="[1024,1149,511,536]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="2" pageNumber="384">Fig. 1A, D</figureCitation>
|
||
). Ventral chaetae in V, VI and XI reduced to 2 per bundle (occasionally 3) (
|
||
<figureCitation id="133B44CC2727007D62C0CD1B9A53FDC2" box="[696,846,548,573]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="2" pageNumber="384">Fig. 1B, C, E</figureCitation>
|
||
). Ventral chaetae in V and VI longer and stouter than the others even in immature specimens, 126–158 µm in length and 7.5–11 µm in maximal thickness (
|
||
<figureCitation id="133B44CC2727007D656CCD789C92FD9E" box="[1300,1423,583,608]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="2" pageNumber="384">Fig. 1B, C</figureCitation>
|
||
). Ventral chaetae in XI shorter and thinner than in V and VI, but thicker than the ordinary ventral ones, 6–8.5 µm in maximal thickness (
|
||
<figureCitation id="133B44CC2727007D610FCDB098D6FD56" box="[375,459,655,680]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="2" pageNumber="384">Fig. 1E</figureCitation>
|
||
). Chaetae of XII lacking in fully mature specimens. Clitellum extending from 1/
|
||
<date id="FFBE7E892727007D6534CDAF9C62FD59" box="[1356,1407,656,680]" pageId="2" pageNumber="384">2 XI</date>
|
||
to XIII, elevated conspicuously. Hyalocytes (12–24 µm in diameter) more abundant than granulocytes, irregularly arranged and in contact with each other. Granulocytes (13–22 µm in diameter) irregular in shape, having some connection with each other and forming a loose net (
|
||
<figureCitation id="133B44CC2727007D6295CDC39A78FCEA" box="[749,869,764,789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="2" pageNumber="384">Fig. 2B, F</figureCitation>
|
||
). Male pores distinct in the ventral middle of XII. Spermathecal pores paired, in the lateral line at IV/V (
|
||
<figureCitation id="133B44CC2727007D628ECC209A4DFCC6" box="[758,848,799,824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="2" pageNumber="384">Fig. 2K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7F08C12727007D60EFCC5B9C3FFC85" box="[151,1314,867,891]" pageId="2" pageNumber="384">
|
||
<paragraph id="8BBF58492727007D60EFCC5B9C3FFC85" blockId="2.[151,1314,867,891]" box="[151,1314,867,891]" pageId="2" pageNumber="384">
|
||
<emphasis id="B974845B2727007D60EFCC5B9817FC85" bold="true" box="[151,266,868,891]" pageId="2" pageNumber="384">TABLE 1.</emphasis>
|
||
Chaetal sizes and numbers in different segments of
|
||
<taxonomicName id="4C0023CA2727007D634FCC5C9DDBFC85" authority="Zhang & Lu & Xie, 2018" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[823,1222,867,891]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="384" phylum="Annelida" rank="species" species="spermatoglomeratus" status="sp. nov.">
|
||
<emphasis id="B974845B2727007D634FCC5C9DDBFC85" box="[823,1222,867,891]" italics="true" pageId="2" pageNumber="384">Mesenchytraeus spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24739202727007D64B4CC5B9C3FFC85" box="[1228,1314,868,891]" pageId="2" pageNumber="384" rank="species">
|
||
<emphasis id="B974845B2727007D64B4CC5B9DF8FC85" bold="true" box="[1228,1253,868,891]" pageId="2" pageNumber="384">sp</emphasis>
|
||
.
|
||
<emphasis id="B974845B2727007D648BCC5B9C06FC85" bold="true" box="[1267,1307,868,891]" pageId="2" pageNumber="384">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBF58492727007D60E7CCA99C28FAEF" pageId="2" pageNumber="384">
|
||
<table id="F900AAE92727FF8060EFCCA99C75FAEF" box="[151,1384,918,1297]" gridcols="7" gridrows="9" pageId="2" pageNumber="384">
|
||
<tr id="35305A0B2727FF8060EFCCA99C75FC52" box="[151,1384,918,940]" gridrow="0" pageId="2" pageNumber="384">
|
||
<th id="76E133772727FF8060EFCCA998C4FC52" box="[151,473,918,940]" gridcol="0" gridrow="0" pageId="2" pageNumber="384">Segments</th>
|
||
<th id="76E133772727FF80621FCCA99BB2FC52" box="[615,687,918,940]" gridcol="1" gridrow="0" pageId="2" pageNumber="384">II</th>
|
||
<th id="76E133772727FF806298CCA99A31FC52" box="[736,812,918,940]" gridcol="2" gridrow="0" pageId="2" pageNumber="384">III</th>
|
||
<th id="76E133772727FF80631CCCA99AA0FC52" box="[868,957,918,940]" gridcol="3" gridrow="0" pageId="2" pageNumber="384">V</th>
|
||
<th id="76E133772727FF8063A3CCA99D4DFC52" box="[987,1104,918,940]" gridcol="4" gridrow="0" pageId="2" pageNumber="384">VI</th>
|
||
<th id="76E133772727FF806416CCA99DCBFC52" box="[1134,1238,918,940]" gridcol="5" gridrow="0" pageId="2" pageNumber="384">VII</th>
|
||
<th id="76E133772727FF806486CCA99C75FC52" box="[1278,1384,918,940]" gridcol="6" gridrow="0" pageId="2" pageNumber="384">IX</th>
|
||
</tr>
|
||
<table id="F900AAE92727FF8060EFCCFF9C75F9F7" box="[151,1384,960,1545]" gridrows="14" pageId="2" pageNumber="384">
|
||
<tr id="35305A0B2727FF8060EFCCFF9C75FC28" box="[151,1384,960,982]" gridrow="1" pageId="2" pageNumber="384">
|
||
<th id="76E133772727FF8060EFCCFF98C4FC28" box="[151,473,960,982]" gridcol="0" gridrow="1" pageId="2" pageNumber="384">Length of lateral chaetae (µm)</th>
|
||
<td id="76E133772727FF80621FCCFF9BB2FC28" box="[615,687,960,982]" gridcol="1" gridrow="1" pageId="2" pageNumber="384">44–48</td>
|
||
<td id="76E133772727FF806298CCFF9A31FC28" box="[736,812,960,982]" gridcol="2" gridrow="1" pageId="2" pageNumber="384">56–61</td>
|
||
<td id="76E133772727FF80631CCCFF9AA0FC28" box="[868,957,960,982]" gridcol="3" gridrow="1" pageId="2" pageNumber="384">61–66</td>
|
||
<td id="76E133772727FF8063A3CCFF9D4DFC28" box="[987,1104,960,982]" gridcol="4" gridrow="1" pageId="2" pageNumber="384">66–80</td>
|
||
<td id="76E133772727FF806416CCFF9DCBFC28" box="[1134,1238,960,982]" gridcol="5" gridrow="1" pageId="2" pageNumber="384">75–85</td>
|
||
<td id="76E133772727FF806486CCFF9C75FC28" box="[1278,1384,960,982]" gridcol="6" gridrow="1" pageId="2" pageNumber="384">80–100</td>
|
||
</tr>
|
||
<tr id="35305A0B2727FF8060EFCCD69C75FC01" box="[151,1384,1001,1023]" gridrow="2" pageId="2" pageNumber="384">
|
||
<th id="76E133772727FF8060EFCCD698C4FC01" box="[151,473,1001,1023]" gridcol="0" gridrow="2" pageId="2" pageNumber="384">Width of lateral chaetae (µm)</th>
|
||
<td id="76E133772727FF80621FCCD69BB2FC01" box="[615,687,1001,1023]" gridcol="1" gridrow="2" pageId="2" pageNumber="384">3.5–4</td>
|
||
<td id="76E133772727FF806298CCD69A31FC01" box="[736,812,1001,1023]" gridcol="2" gridrow="2" pageId="2" pageNumber="384">3.5–4.5</td>
|
||
<td id="76E133772727FF80631CCCD69AA0FC01" box="[868,957,1001,1023]" gridcol="3" gridrow="2" pageId="2" pageNumber="384">4.5–5</td>
|
||
<td id="76E133772727FF8063A3CCD69D4DFC01" box="[987,1104,1001,1023]" gridcol="4" gridrow="2" pageId="2" pageNumber="384">4.5–5</td>
|
||
<td id="76E133772727FF806416CCD69DCBFC01" box="[1134,1238,1001,1023]" gridcol="5" gridrow="2" pageId="2" pageNumber="384">4.5–5</td>
|
||
<td id="76E133772727FF806486CCD69C75FC01" box="[1278,1384,1001,1023]" gridcol="6" gridrow="2" pageId="2" pageNumber="384">3.5–5</td>
|
||
</tr>
|
||
<tr id="35305A0B2727FF8060EFCB2D9C75FBD6" box="[151,1384,1042,1064]" gridrow="3" pageId="2" pageNumber="384">
|
||
<th id="76E133772727FF8060EFCB2D98C4FBD6" box="[151,473,1042,1064]" gridcol="0" gridrow="3" pageId="2" pageNumber="384">Number per bundle</th>
|
||
<td id="76E133772727FF80621FCB2D9BB2FBD6" box="[615,687,1042,1064]" gridcol="1" gridrow="3" pageId="2" pageNumber="384">2</td>
|
||
<td id="76E133772727FF806298CB2D9A31FBD6" box="[736,812,1042,1064]" gridcol="2" gridrow="3" pageId="2" pageNumber="384">2</td>
|
||
<td id="76E133772727FF80631CCB2D9AA0FBD6" box="[868,957,1042,1064]" gridcol="3" gridrow="3" pageId="2" pageNumber="384">2,3</td>
|
||
<td id="76E133772727FF8063A3CB2D9D4DFBD6" box="[987,1104,1042,1064]" gridcol="4" gridrow="3" pageId="2" pageNumber="384">2,3</td>
|
||
<td id="76E133772727FF806416CB2D9DCBFBD6" box="[1134,1238,1042,1064]" gridcol="5" gridrow="3" pageId="2" pageNumber="384">2,3</td>
|
||
<td id="76E133772727FF806486CB2D9C75FBD6" box="[1278,1384,1042,1064]" gridcol="6" gridrow="3" pageId="2" pageNumber="384">3</td>
|
||
</tr>
|
||
<tr id="35305A0B2727FF8060EFCB039C75FBAC" box="[151,1384,1084,1106]" gridrow="4" pageId="2" pageNumber="384">
|
||
<th id="76E133772727FF8060EFCB0398C4FBAC" box="[151,473,1084,1106]" gridcol="0" gridrow="4" pageId="2" pageNumber="384">Length of ventral chaetae (µm)</th>
|
||
<td id="76E133772727FF80621FCB039BB2FBAC" box="[615,687,1084,1106]" gridcol="1" gridrow="4" pageId="2" pageNumber="384">63–75</td>
|
||
<td id="76E133772727FF806298CB039A31FBAC" box="[736,812,1084,1106]" gridcol="2" gridrow="4" pageId="2" pageNumber="384">70–82</td>
|
||
<td id="76E133772727FF80631CCB039AA0FBAC" box="[868,957,1084,1106]" gridcol="3" gridrow="4" pageId="2" pageNumber="384">126–158</td>
|
||
<td id="76E133772727FF8063A3CB039D4DFBAC" box="[987,1104,1084,1106]" gridcol="4" gridrow="4" pageId="2" pageNumber="384">126–158</td>
|
||
<td id="76E133772727FF806416CB039DCBFBAC" box="[1134,1238,1084,1106]" gridcol="5" gridrow="4" pageId="2" pageNumber="384">95–120</td>
|
||
<td id="76E133772727FF806486CB039C75FBAC" box="[1278,1384,1084,1106]" gridcol="6" gridrow="4" pageId="2" pageNumber="384">100–120</td>
|
||
</tr>
|
||
<tr id="35305A0B2727FF8060EFCB5A9C75FB85" box="[151,1384,1125,1147]" gridrow="5" pageId="2" pageNumber="384">
|
||
<th id="76E133772727FF8060EFCB5A98C4FB85" box="[151,473,1125,1147]" gridcol="0" gridrow="5" pageId="2" pageNumber="384">Width of ventral chaetae (µm)</th>
|
||
<td id="76E133772727FF80621FCB5A9BB2FB85" box="[615,687,1125,1147]" gridcol="1" gridrow="5" pageId="2" pageNumber="384">4–4.5</td>
|
||
<td id="76E133772727FF806298CB5A9A31FB85" box="[736,812,1125,1147]" gridcol="2" gridrow="5" pageId="2" pageNumber="384">4–5</td>
|
||
<td id="76E133772727FF80631CCB5A9AA0FB85" box="[868,957,1125,1147]" gridcol="3" gridrow="5" pageId="2" pageNumber="384">7.5–9.5</td>
|
||
<td id="76E133772727FF8063A3CB5A9D4DFB85" box="[987,1104,1125,1147]" gridcol="4" gridrow="5" pageId="2" pageNumber="384">8–11</td>
|
||
<td id="76E133772727FF806416CB5A9DCBFB85" box="[1134,1238,1125,1147]" gridcol="5" gridrow="5" pageId="2" pageNumber="384">5.5–7.5</td>
|
||
<td id="76E133772727FF806486CB5A9C75FB85" box="[1278,1384,1125,1147]" gridcol="6" gridrow="5" pageId="2" pageNumber="384">5.5–7</td>
|
||
</tr>
|
||
<tr id="35305A0B2727FF8060EFCBB19C75FB5A" box="[151,1384,1166,1188]" gridrow="6" pageId="2" pageNumber="384">
|
||
<th id="76E133772727FF8060EFCBB198C4FB5A" box="[151,473,1166,1188]" gridcol="0" gridrow="6" pageId="2" pageNumber="384">Number per bundle</th>
|
||
<td id="76E133772727FF80621FCBB19BB2FB5A" box="[615,687,1166,1188]" gridcol="1" gridrow="6" pageId="2" pageNumber="384">3–5</td>
|
||
<td id="76E133772727FF806298CBB19A31FB5A" box="[736,812,1166,1188]" gridcol="2" gridrow="6" pageId="2" pageNumber="384">4,5</td>
|
||
<td id="76E133772727FF80631CCBB19AA0FB5A" box="[868,957,1166,1188]" gridcol="3" gridrow="6" pageId="2" pageNumber="384">2</td>
|
||
<td id="76E133772727FF8063A3CBB19D4DFB5A" box="[987,1104,1166,1188]" gridcol="4" gridrow="6" pageId="2" pageNumber="384">2</td>
|
||
<td id="76E133772727FF806416CBB19DCBFB5A" box="[1134,1238,1166,1188]" gridcol="5" gridrow="6" pageId="2" pageNumber="384">3,4</td>
|
||
<td id="76E133772727FF806486CBB19C75FB5A" box="[1278,1384,1166,1188]" gridcol="6" gridrow="6" pageId="2" pageNumber="384">4</td>
|
||
</tr>
|
||
<tr id="35305A0B2727FF8060EFCBF79C75FB21" box="[151,1384,1224,1247]" gridrow="7" pageId="2" pageNumber="384" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
||
<th id="76E133772727FF8060EFCBF798C4FB21" box="[151,473,1224,1247]" gridcol="0" gridrow="7" pageId="2" pageNumber="384">continued.</th>
|
||
</tr>
|
||
<tr id="35305A0B2727FF8060EFCBC49C75FAEF" box="[151,1384,1275,1297]" gridrow="8" pageId="2" pageNumber="384" rowspan-2="1" rowspan-3="1">
|
||
<th id="76E133772727FF8060EFCBC498C4FAEF" box="[151,473,1275,1297]" gridcol="0" gridrow="8" pageId="2" pageNumber="384">Segments</th>
|
||
<td id="76E133772727FF80621FCBC49BB2FAEF" box="[615,687,1275,1297]" gridcol="1" gridrow="8" pageId="2" pageNumber="384">XI</td>
|
||
XIII
|
||
<td id="76E133772727FF8063A3CBC49D4DFAEF" box="[987,1104,1275,1297]" gridcol="4" gridrow="8" pageId="2" pageNumber="384">XVII</td>
|
||
<td id="76E133772727FF806416CBC49DCBFAEF" box="[1134,1238,1275,1297]" gridcol="5" gridrow="8" pageId="2" pageNumber="384">XX</td>
|
||
<td id="76E133772727FF806486CBC49C75FAEF" box="[1278,1384,1275,1297]" gridcol="6" gridrow="8" pageId="2" pageNumber="384">XXX</td>
|
||
</tr>
|
||
<paragraph id="8BBF58492727007D60E7CA1B9C16F9F7" pageId="2" pageNumber="384">Length of lateral chaetae (µm) 71–90 71–85 71–80 71–80 65–75 Width of lateral chaetae (µm) 3.5–5 3.5–5 3.5–5 3.5–5 3.5–4.5 Number per bundle 3 3 3 3,4 3 Length of ventral chaetae (µm) 110–120 105–120 90–105 85–102 75–85 Width of ventral chaetae (µm) 6–8.5 6–7.5 5–6 5–6 5–6 Number per bundle 2 2,3 4 4 5</paragraph>
|
||
</table>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BBF58492727007D60BFC9759BD8F87C" blockId="2.[151,1437,1610,2031]" pageId="2" pageNumber="384">
|
||
Brain in I-II (
|
||
<figureCitation id="133B44CC2727007D6124C97598ABF99C" box="[348,438,1610,1635]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="2" pageNumber="384">Figs. 1J</figureCitation>
|
||
,
|
||
<figureCitation id="133B44CC2727007D61BAC97598F9F99C" box="[450,484,1610,1634]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="2" pageNumber="384">2E</figureCitation>
|
||
), trapezoidal, deeply concave anteriorly and slightly incised posteriorly, 137–170 µm long and 105–125 µm in maximal width. Blood colourless or slightly yellowish. Dorsal vessel arising from XIV– XVI. Two pairs of primary pharyngeal glands in IV and V, attached to septa of 4/5 and 5/6 respectively, and both not connected dorsally. Three pairs of secondary lobes ventrally in V–VII, largest in VI. No oesophageal appendages. Intestinal diverticula absent. Chloragogen cells developed, originating from VIII, 12–45 µm high, absent in clitellum. Coelomocytes abundant, disc-shaped with conspicuous nuclei, 7.5–15.0 µm in diameter, with uneven refractile granules (
|
||
<figureCitation id="133B44CC2727007D61A6C81D9B20F8C4" box="[478,573,1826,1851]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="2" pageNumber="384">Fig. 2C</figureCitation>
|
||
). Five pairs of nephridia in preclitellar segments from VI/VII to X/XI. Anteseptale stalked, containing nephrostome only; postseptale bilobed and with fewer interstitial tissue. Efferent duct arising between the two lobes (
|
||
<figureCitation id="133B44CC2727007D6252C8559B9AF87C" box="[554,647,1898,1923]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="2" pageNumber="384">Figs. 1F</figureCitation>
|
||
,
|
||
<figureCitation id="133B44CC2727007D62EBC8559BA4F87C" box="[659,697,1898,1922]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="2" pageNumber="384">2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF58492727007C60BFC8B09B54FDC3" blockId="2.[151,1437,1610,2031]" lastBlockId="3.[151,1437,151,573]" lastPageId="3" lastPageNumber="385" pageId="2" pageNumber="384">
|
||
Sperm funnels cylindrical, occupying 1/3–1/2 body cavity in XI, 390–440 µm in length and 122–200 µm in maximal width. Their ental openings much narrower than funnel body, 43–73 µm wide. The heads of spermatozoa 17–20 µm in length (
|
||
<figureCitation id="133B44CC2727007D61FBC8E998C1F811" box="[387,476,2006,2031]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="2" pageNumber="384">
|
||
Figs.
|
||
<date id="FFBE7E892727007D61BBC8E898C1F811" box="[451,476,2007,2031]" pageId="2" pageNumber="384">1I</date>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="133B44CC2727007D6190C8E89B1EF811" box="[488,515,2007,2031]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="2" pageNumber="384">
|
||
<date id="FFBE7E892727007D6190C8E89B1EF811" box="[488,515,2007,2031]" pageId="2" pageNumber="384">2I</date>
|
||
</figureCitation>
|
||
). Vasa deferentia with conspicuous cilia in the canal, wound in irregular spirals in coelom from XII and posteriorly into XV–XXII, 18–30 µm in width, lumen of vasa deferentia 14–24 µm wide (
|
||
<figureCitation id="133B44CC2726007C6517CFA899A7FF2A" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="3" pageNumber="385">Fig. 2L</figureCitation>
|
||
). The transition between vas deferens and atrium gradual. Atria in XII–XIII, cylindrical, remarkably wider than vasa deferentia, 415–650 µm long and 65–85 µm in maximal width. No cilia observed in atrial canal. Each atrium possessing 4–6 tubular prostate-like glands (each 117–195 µm in length and 50–68 µm in width) (
|
||
<figureCitation id="133B44CC2726007C6489CE3B9C4BFEE2" box="[1265,1366,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="3" pageNumber="385">Figs. 1H</figureCitation>
|
||
,
|
||
<figureCitation id="133B44CC2726007C651BCE3B9C92FEE2" box="[1379,1423,260,284]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="3" pageNumber="385">
|
||
<date id="FFBE7E892726007C651BCE3B9C63FEE2" box="[1379,1406,260,284]" pageId="3" pageNumber="385">2I</date>
|
||
,J
|
||
</figureCitation>
|
||
). Each atrium divided into two distinct parts, an ental part (length: 217–400 µm; width: 27.5–58 µm) with thin muscle layer and thick epithelium, and an ectal part (length: 200–350 µm; width: 25–44 µm) with a muscular duct connecting with the penial bulb. Penial bulb large, compact in ventral of XII, consisting of muscle strands and surrounded by masses of accessory glands of different size (
|
||
<figureCitation id="133B44CC2726007C634FCEAB9A87FE52" box="[823,922,404,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="3" pageNumber="385">Figs. 1H</figureCitation>
|
||
,
|
||
<figureCitation id="133B44CC2726007C63DECEAB9ADEFE52" box="[934,963,404,428]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="3" pageNumber="385">2J</figureCitation>
|
||
). A pair of sperm sacs large from XI–XIV in different size, and backwards extending to XV–XXIV, each containing numerous ball-shaped sperm bundles (diameter: 50–110 µm), a large number of spermatozoa coiled inside in vivo but much more loose in fixed condition (
|
||
<figureCitation id="133B44CC2726007C616ECEC0988FFDE6" box="[278,402,511,536]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="3" pageNumber="385">Fig. 2D,H</figureCitation>
|
||
). Egg sacs paired, large from XIV–XXII, continuing caudad to XVII–XXXIII, usually containing 2–11 mature eggs at a time.
|
||
</paragraph>
|
||
<caption id="DF7F08C12726007C60EFC9F99D3FF88B" httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="3" pageNumber="385" startId="3.[151,250,1734,1756]" targetBox="[202,1365,658,1684]" targetPageId="3">
|
||
<paragraph id="8BBF58492726007C60EFC9F99D3FF88B" blockId="3.[151,1436,1734,1909]" pageId="3" pageNumber="385">
|
||
<emphasis id="B974845B2726007C60EFC9F9980EF925" bold="true" box="[151,275,1734,1756]" pageId="3" pageNumber="385">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C0023CA2726007C6161C9F99B89F922" authority="Zhang & Lu & Xie, 2018" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[281,660,1734,1756]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="385" phylum="Annelida" rank="species" species="spermatoglomeratus" status="sp. nov.">
|
||
<emphasis id="B974845B2726007C6161C9F99B89F922" box="[281,660,1734,1756]" italics="true" pageId="3" pageNumber="385">Mesenchytraeus spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B974845B2726007C62E3C9F99A18F925" bold="true" box="[667,773,1734,1755]" pageId="3" pageNumber="385">
|
||
<taxonomicNameLabel id="A24739202726007C62E3C9F99BF6F925" box="[667,747,1734,1755]" pageId="3" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A
|
||
</emphasis>
|
||
. Ventral chaetae in II.
|
||
<emphasis id="B974845B2726007C6394C9F99AE0F925" bold="true" box="[1004,1021,1734,1755]" pageId="3" pageNumber="385">B</emphasis>
|
||
. Ventral chaetae in V.
|
||
<emphasis id="B974845B2726007C6499C9F99DEEF922" bold="true" box="[1249,1267,1734,1756]" pageId="3" pageNumber="385">C</emphasis>
|
||
. Ventral chaetae in VI.
|
||
<emphasis id="B974845B2726007C60AFC9DA99F4F904" bold="true" box="[215,233,1765,1786]" pageId="3" pageNumber="385">D</emphasis>
|
||
. Ventral chaetae in VIII.
|
||
<emphasis id="B974845B2726007C619FC9DA98E5F904" bold="true" box="[487,504,1765,1786]" pageId="3" pageNumber="385">E</emphasis>
|
||
. Ventral chaetae in XI.
|
||
<emphasis id="B974845B2726007C629CC9DA9BECF904" bold="true" box="[740,753,1765,1786]" pageId="3" pageNumber="385">F</emphasis>
|
||
. Nephridium in VIII/IX.
|
||
<emphasis id="B974845B2726007C6389C9DA9D1FF905" bold="true" box="[1009,1026,1765,1787]" pageId="3" pageNumber="385">G</emphasis>
|
||
. Spermatheca.
|
||
<emphasis id="B974845B2726007C64E4C9DA9DADF904" bold="true" box="[1180,1200,1765,1786]" pageId="3" pageNumber="385">H</emphasis>
|
||
. Male organ in clitellar region.
|
||
<emphasis id="B974845B2726007C609FC83B99ECF8E7" bold="true" box="[231,241,1796,1817]" pageId="3" pageNumber="385">I</emphasis>
|
||
. Sperm funnel.
|
||
<emphasis id="B974845B2726007C61E3C83B98B5F8E7" bold="true" box="[411,424,1796,1817]" pageId="3" pageNumber="385">J</emphasis>
|
||
. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7F08C12721007B60EFCB6299D6FAB4" httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="4" pageNumber="386" startId="4.[151,250,1117,1139]" targetBox="[191,1395,193,1096]" targetPageId="4">
|
||
<paragraph id="8BBF58492721007B60EFCB6299D6FAB4" blockId="4.[151,1436,1117,1354]" pageId="4" pageNumber="386">
|
||
<emphasis id="B974845B2721007B60EFCB629809FB8D" bold="true" box="[151,276,1117,1139]" pageId="4" pageNumber="386">FIGURE 2.</emphasis>
|
||
Micrographs of
|
||
<taxonomicName id="4C0023CA2721007B61BCCB629A5CFB8D" authority="Zhang & Lu & Xie, 2018" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[452,833,1117,1139]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="spermatoglomeratus" status="sp. nov.">
|
||
<emphasis id="B974845B2721007B61BCCB629A5CFB8D" box="[452,833,1117,1139]" italics="true" pageId="4" pageNumber="386">Mesenchytraeus spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24739202721007B6331CB619A86FB8D" box="[841,923,1118,1139]" pageId="4" pageNumber="386" rank="species">
|
||
<emphasis id="B974845B2721007B6331CB619A86FB8D" bold="true" box="[841,923,1118,1139]" pageId="4" pageNumber="386">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A–D in
|
||
<emphasis id="B974845B2721007B6382CB609D39FB8A" box="[1018,1060,1119,1140]" italics="true" pageId="4" pageNumber="386">vivo</emphasis>
|
||
, E–L fixed, stained.
|
||
<emphasis id="B974845B2721007B6487CB619C0CFB8D" bold="true" box="[1279,1297,1118,1139]" pageId="4" pageNumber="386">A</emphasis>
|
||
. Prostomium and head pore;
|
||
<emphasis id="B974845B2721007B614DCB42985BFB6C" bold="true" box="[309,326,1149,1170]" pageId="4" pageNumber="386">B</emphasis>
|
||
. Pattern of clitellar glands.
|
||
<emphasis id="B974845B2721007B621DCB439B6AFB6C" bold="true" box="[613,631,1148,1170]" pageId="4" pageNumber="386">C</emphasis>
|
||
. Coelomocytes.
|
||
<emphasis id="B974845B2721007B635CCB429A2BFB6C" bold="true" box="[804,822,1149,1170]" pageId="4" pageNumber="386">D</emphasis>
|
||
. Sperm bundles just coming out from broken sperm sac.
|
||
<emphasis id="B974845B2721007B65FCCB429C88FB6C" bold="true" box="[1412,1429,1149,1170]" pageId="4" pageNumber="386">E</emphasis>
|
||
. Brain.
|
||
<emphasis id="B974845B2721007B60A3CBA499F5FB4E" bold="true" box="[219,232,1179,1200]" pageId="4" pageNumber="386">F</emphasis>
|
||
. Pattern of clitellar gland cells.
|
||
<emphasis id="B974845B2721007B6251CBA49B27FB4F" bold="true" box="[553,570,1179,1201]" pageId="4" pageNumber="386">G</emphasis>
|
||
. Nephridium in VIII/IX.
|
||
<emphasis id="B974845B2721007B6342CBA49A53FB4E" bold="true" box="[826,846,1179,1200]" pageId="4" pageNumber="386">H</emphasis>
|
||
. Sperm bundle.
|
||
<emphasis id="B974845B2721007B638BCBA49AE0FB4E" bold="true" box="[1011,1021,1179,1200]" pageId="4" pageNumber="386">I</emphasis>
|
||
. Sperm funnel and atrium.
|
||
<emphasis id="B974845B2721007B6568CBA49C01FB4E" bold="true" box="[1296,1308,1179,1200]" pageId="4" pageNumber="386">J</emphasis>
|
||
. Male organ in clitellar region.
|
||
<emphasis id="B974845B2721007B612CCB859873FB31" bold="true" box="[340,366,1210,1231]" pageId="4" pageNumber="386">K.</emphasis>
|
||
Spermathecae.
|
||
<emphasis id="B974845B2721007B6268CB859B35FB31" bold="true" box="[528,552,1210,1231]" pageId="4" pageNumber="386">L.</emphasis>
|
||
Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBF58492721007B60BFCA4D9D87FA08" blockId="4.[151,1437,1393,2030]" pageId="4" pageNumber="386">
|
||
One pair of spermathecae in V–VII, without ectal glands. Ectal duct 215–385 µm long and 37–50 µm wide, projecting into ampulla. Ampulla spherical, 83–102 µm in diameter. No diverticula visible. Ampulla filled with spermatozoa in fully mature specimens. Ental ducts stout, 195–244 µm long and 62–67 µm wide, connecting to the dorsal side of oesophagus separately in the posterior of VI or anterior of VII (
|
||
<figureCitation id="133B44CC2721007B6383CAE29D46FA08" box="[1019,1115,1501,1526]" captionStart="FIGURE 1" captionStartId="3.[151,250,1734,1756]" captionTargetBox="[202,1365,658,1684]" captionTargetId="figure@3.[188,1398,628,1710]" captionTargetPageId="3" captionText="FIGURE 1. Mesenchytraeus spermatoglomeratus sp. nov. A. Ventral chaetae in II. B. Ventral chaetae in V. C. Ventral chaetae in VI. D. Ventral chaetae in VIII. E. Ventral chaetae in XI. F. Nephridium in VIII/IX. G. Spermatheca. H. Male organ in clitellar region. I. Sperm funnel. J. Brain. Scale bars: A–E 25 µm; F–G 50 µm; H 100 µm; I 50 µm; J 25 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; as, nephridial anteseptale; at, atrium; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; hs, head of spermatozoa; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore; vd, vas deferens." httpUri="https://zenodo.org/record/1446909/files/figure.png" pageId="4" pageNumber="386">Figs. 1G</figureCitation>
|
||
,
|
||
<figureCitation id="133B44CC2721007B641FCAE19D90FA08" box="[1127,1165,1502,1526]" captionStart="FIGURE 2" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[191,1395,193,1096]" captionTargetId="figure@4.[191,1395,193,1096]" captionTargetPageId="4" captionText="FIGURE 2. Micrographs of Mesenchytraeus spermatoglomeratus sp. nov. A–D in vivo, E–L fixed, stained. A. Prostomium and head pore; B. Pattern of clitellar glands. C. Coelomocytes. D. Sperm bundles just coming out from broken sperm sac. E. Brain. F. Pattern of clitellar gland cells. G. Nephridium in VIII/IX. H. Sperm bundle. I. Sperm funnel and atrium. J. Male organ in clitellar region. K. Spermathecae. L. Vasa deferentia. Scale bars: A 50 µm; B 25 µm; C 15 µm; D 25 µm; E–H 50 µm; I–K 100 µm; L 50 µm. Abbreviations: acg, accessory copulatory glands; ag, atrial glands; at, atrium; br, brain; bsf, body of sperm funnel; ecd, spermathecal ectal duct; ed, nephridial efferent duct; end, spermathecal ental duct; gr, granulocyte; hp, head pore; hs, head of spermatozoa; hy, hyalocyte; pb, penial bulb; ps, nephridial postseptale; sa, spermathecal ampulla; sp, spermathecal pore." httpUri="https://zenodo.org/record/1446911/files/figure.png" pageId="4" pageNumber="386">2K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A0BC22721007A60BFC93E980BFE76" lastPageId="5" lastPageNumber="387" pageId="4" pageNumber="386" type="diagnosis">
|
||
<paragraph id="8BBF58492721007B60BFC93E9C58F858" blockId="4.[151,1437,1393,2030]" pageId="4" pageNumber="386">
|
||
<emphasis id="B974845B2721007B60BFC93E98D8F9E4" bold="true" box="[199,453,1537,1562]" pageId="4" pageNumber="386">Differential diagnosis</emphasis>
|
||
(Table 3). This new species is morphologically most similar to the co-occurring species
|
||
<taxonomicName id="4C0023CA2721007B60EFC91998A6F9C0" authority="Xie, 2012" authorityName="Xie" authorityYear="2012" box="[151,443,1573,1598]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="gigachaetus">
|
||
<emphasis id="B974845B2721007B60EFC919985EF9C3" box="[151,323,1573,1597]" italics="true" pageId="4" pageNumber="386">M. gigachaetus</emphasis>
|
||
Xie, 2012
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0023CA2721007B61BEC9199A72F9C0" authority="Shen et al., 2012" authorityName="Shen et al." authorityYear="2012" box="[454,879,1573,1598]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="anisodiverticulus">
|
||
<emphasis id="B974845B2721007B61BEC9199BB1F9C3" box="[454,684,1573,1597]" italics="true" pageId="4" pageNumber="386">M. anisodiverticulus</emphasis>
|
||
Shen
|
||
<emphasis id="B974845B2721007B628AC9199A37F9C3" box="[754,810,1573,1597]" italics="true" pageId="4" pageNumber="386">et al.</emphasis>
|
||
, 2012
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0023CA2721007B63DDC9199C4DF9C0" authority="Shen et al., 2012" authorityName="Shen et al." authorityYear="2012" box="[933,1360,1573,1598]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="monodiverticulus">
|
||
<emphasis id="B974845B2721007B63DDC9199D91F9C3" box="[933,1164,1573,1597]" italics="true" pageId="4" pageNumber="386">M. monodiverticulus</emphasis>
|
||
Shen
|
||
<emphasis id="B974845B2721007B64AAC9199C17F9C3" box="[1234,1290,1573,1597]" italics="true" pageId="4" pageNumber="386">et al.</emphasis>
|
||
, 2012
|
||
</taxonomicName>
|
||
and to a further terrestrial species,
|
||
<taxonomicName id="4C0023CA2721007B61B4C9759D19F99C" authority="Christensen & Dozsa-Farkas, 2012" authorityName="Christensen & Dozsa-Farkas" authorityYear="2012" box="[460,1028,1609,1634]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="longiductus">
|
||
<emphasis id="B974845B2721007B61B4C9759B68F99C" box="[460,629,1610,1634]" italics="true" pageId="4" pageNumber="386">M. longiductus</emphasis>
|
||
<bibRefCitation id="EF9125B82721007B6203C9769D19F99C" box="[635,1028,1609,1634]" pageId="4" pageNumber="386" refId="ref6257" refString="Christensen, B. & Dozsa-Farkas, K. (2012) A new genus Globulidrilus and three new enchytraeid species (Oligochaeta: Enchytraeidae) from Seoraksan National Park (Korea). Journal of Natural History, 46, 2769 - 2785. https: // doi. org / 10.1080 / 00222933.2012.737038" type="journal article">Christensen & Dózsa-Farkas, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
, collected in
|
||
<collectingCountry id="F31718D92721007B64EFC9759DC3F99F" box="[1175,1246,1610,1633]" name="South Korea" pageId="4" pageNumber="386">Korea</collectingCountry>
|
||
, which also have enlarged ventral chaetae in V–VI, and atria, egg sacs, and spermathecae attached to oesophagus. However,
|
||
<taxonomicName id="4C0023CA2721007B60EFC9AD9B23F954" authority="Zhang & Lu & Xie, 2018" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[151,574,1682,1706]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="spermatoglomeratus" status="sp. nov.">
|
||
<emphasis id="B974845B2721007B60EFC9AD9B23F954" box="[151,574,1682,1706]" italics="true" pageId="4" pageNumber="386">Mesenchytraeus spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B974845B2721007B6232C9AE9BBAF954" bold="true" box="[586,679,1681,1706]" pageId="4" pageNumber="386">
|
||
<taxonomicNameLabel id="A24739202721007B6232C9AE9BBAF954" box="[586,679,1681,1706]" pageId="4" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
possesses enlarged ventral chaetae in XI and ball-shaped sperm bundles in the sperm sacs, two characteristics absent in the latter four species. Besides,
|
||
<taxonomicName id="4C0023CA2721007B641CC9899C12F933" authorityName="Xie" authorityYear="2012" box="[1124,1295,1717,1741]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="gigachaetus">
|
||
<emphasis id="B974845B2721007B641CC9899C12F933" box="[1124,1295,1717,1741]" italics="true" pageId="4" pageNumber="386">M. gigachaetus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0023CA2721007B60EFC9E598BFF90C" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[151,418,1754,1778]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="spermatoglomeratus">
|
||
<emphasis id="B974845B2721007B60EFC9E598BFF90C" box="[151,418,1754,1778]" italics="true" pageId="4" pageNumber="386">M. spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
by a higher number per bundle of enlarged ventral chaetae (3–4), no sperm sac visible and no atrial prostate gland.
|
||
<taxonomicName id="4C0023CA2721007B61D0C9C29A3BF8EB" box="[424,806,1789,1813]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="anisodiverticulus">
|
||
<emphasis id="B974845B2721007B61D0C9C29A3BF8EB" box="[424,806,1789,1813]" italics="true" pageId="4" pageNumber="386">Mesenchytraeus anisodiverticulus</emphasis>
|
||
</taxonomicName>
|
||
is distinct from this new species by its absence of atrial prostate glands and the presence of two asymmetrical spermathecal diverticula.
|
||
<taxonomicName id="4C0023CA2721007B6464C81D9C81F8C4" box="[1052,1436,1826,1850]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="monodiverticulus">
|
||
<emphasis id="B974845B2721007B6464C81D9C81F8C4" box="[1052,1436,1826,1850]" italics="true" pageId="4" pageNumber="386">Mesenchytraeus monodiverticulus</emphasis>
|
||
</taxonomicName>
|
||
is different from
|
||
<taxonomicName id="4C0023CA2721007B6126C8799B70F8A3" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[350,621,1861,1885]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="spermatoglomeratus">
|
||
<emphasis id="B974845B2721007B6126C8799B70F8A3" box="[350,621,1861,1885]" italics="true" pageId="4" pageNumber="386">M. spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
by its higher number per bundle of enlarged ventral chaetae (4 or 5), absence of atrial prostate glands, and spermatheca with one diverticulum. In
|
||
<emphasis id="B974845B2721007B6477C8559C45F87C" box="[1039,1368,1898,1922]" italics="true" pageId="4" pageNumber="386">
|
||
<taxonomicName id="4C0023CA2721007B6477C8559C49F87C" authorityName="Christensen & Dozsa-Farkas" authorityYear="2012" box="[1039,1364,1898,1922]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="longiductus">Mesenchytraeus longiductus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
atrial prostate glands are less numerous (only 2) and fewer accessory glands surround the penial bulb (only 4–5).
|
||
</paragraph>
|
||
<paragraph id="8BBF58492721007A60BFC88E980BFE76" blockId="4.[151,1437,1393,2030]" lastBlockId="5.[151,1436,151,392]" lastPageId="5" lastPageNumber="387" pageId="4" pageNumber="386">
|
||
Five other species (
|
||
<taxonomicName id="4C0023CA2721007B61D2C88D9BCBF834" authority="Timm, 1994" authorityName="Timm" authorityYear="1994" box="[426,726,1969,1994]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="crenobius">
|
||
<emphasis id="B974845B2721007B61D2C88D9B22F834" box="[426,575,1970,1994]" italics="true" pageId="4" pageNumber="386">M. crenobius</emphasis>
|
||
<bibRefCitation id="EF9125B82721007B6232C88E9BCBF834" box="[586,726,1969,1994]" pageId="4" pageNumber="386" refId="ref7247" refString="Timm, T. (1994) Propappidae and aquatic Enchytraeidae (Oligochaeta) from the farthest southeast of Russia. Hydrobiologia, 278, 67 - 78. https: // doi. org / 10.1007 / BF 00142312" type="journal article">Timm, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0023CA2721007B629FC88D9D84F834" authority="Piper et al., 1982" authorityName="Piper et al." authorityYear="1982" box="[743,1177,1969,1994]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="kontrimavichusi">
|
||
<emphasis id="B974845B2721007B629FC88D9ADEF834" box="[743,963,1970,1994]" italics="true" pageId="4" pageNumber="386">M. kontrimavichusi</emphasis>
|
||
<bibRefCitation id="EF9125B82721007B63B6C88E9D84F834" box="[974,1177,1969,1994]" pageId="4" pageNumber="386" refId="ref6775" refString="Piper, S. R., MacLean, S. F. & Christensen, B. (1982) Enchytraeidae (Oligochaeta) from taiga and tundra habitats of northeastern U. S. S. R. Canadian Journal of Zoology, 60 (11), 2594 - 2609 https: // doi. org / 10.1139 / z 82 - 334" type="journal article">
|
||
Piper
|
||
<emphasis id="B974845B2721007B646BC88D9D53F834" box="[1043,1102,1970,1994]" italics="true" pageId="4" pageNumber="386">et al.</emphasis>
|
||
, 1982
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0023CA2721007B64DFC88D99CFF810" authority="Timm, 1978" authorityName="Timm" authorityYear="1978" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="tetrapodus">
|
||
<emphasis id="B974845B2721007B64DFC88D9C55F834" box="[1191,1352,1970,1994]" italics="true" pageId="4" pageNumber="386">M. tetrapodus</emphasis>
|
||
<bibRefCitation id="EF9125B82721007B6529C88E99CFF810" pageId="4" pageNumber="386" refId="ref7208" refString="Timm, T. & Popchenko, V. (1978) The aquatic Oligochaeta of the Murmansk Region. Hydrobiological Researches (Tartu), 7, 71 - 132. [in Russian with English abstract]" type="journal article">Timm, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0023CA2721007B6095C8E99B90F810" authority="Bretscher, 1900" authorityName="Bretscher" authorityYear="1900" box="[237,653,2005,2030]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="monochaetus">
|
||
<emphasis id="B974845B2721007B6095C8E998ABF813" box="[237,438,2005,2029]" italics="true" pageId="4" pageNumber="386">M. monochaetus</emphasis>
|
||
<bibRefCitation id="EF9125B82721007B61B5C8EA9B90F810" box="[461,653,2005,2030]" pageId="4" pageNumber="386" refId="ref6011" refString="Bretscher, K. (1900) Mitteilungen uber die Oligochaeten fauna der Schweiz. Revue Suisse de Zoologie, 8, 1 - 44." type="journal article">Bretscher, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0023CA2721007B629CC8E99DA0F810" authority="Dozsa-Farkas et al., 2015" authorityName="Dozsa-Farkas et al." authorityYear="2015" box="[740,1213,2005,2030]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="calyx">
|
||
<emphasis id="B974845B2721007B629CC8E99A49F813" box="[740,852,2005,2029]" italics="true" pageId="4" pageNumber="386">M. calyx</emphasis>
|
||
<bibRefCitation id="EF9125B82721007B6314C8EA9DA0F810" box="[876,1213,2005,2030]" pageId="4" pageNumber="386" refId="ref6312" refString="Dozsa-Farkas, K., Felfoldi, T. & Hong, Y. (2015) New enchytraeid species (Enchytraeidae, Oligochaeta) from Korea. Zootaxa, 4006 (1), 171 - 197. https: // doi. org / 10.11646 / zootaxa. 4006.1.9" type="journal article">
|
||
Dózsa-Farkas
|
||
<emphasis id="B974845B2721007B6464C8E99D79F813" box="[1052,1124,2005,2029]" italics="true" pageId="4" pageNumber="386">et al.</emphasis>
|
||
, 2015
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
) differ from
|
||
<taxonomicName id="4C0023CA2721007A6505C8E99860FF4E" authorityName="Zhang & Lu & Xie" authorityYear="2018" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="5" lastPageNumber="387" order="Haplotaxida" pageId="4" pageNumber="386" phylum="Annelida" rank="species" species="spermatoglomeratus">
|
||
<emphasis id="B974845B2721007A6505C8E99860FF4E" italics="true" lastPageId="5" lastPageNumber="387" pageId="4" pageNumber="386">M. spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
in terms of location and number of enlarged chaetae, chaetae formula, spermatheca, origin of dorsal vessel and habitat requirement. Unlike the new species,
|
||
<emphasis id="B974845B2720007A631CCF829C81FF2A" box="[868,1436,188,212]" italics="true" pageId="5" pageNumber="387">
|
||
<taxonomicName id="4C0023CA2720007A631CCF829AE6FF2A" authorityName="Timm" authorityYear="1994" box="[868,1019,188,212]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="387" phylum="Annelida" rank="species" species="crenobius">M. crenobius</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0023CA2720007A647FCF829DBBFF2A" authorityName="Timm" authorityYear="1978" box="[1031,1190,188,212]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="387" phylum="Annelida" rank="species" species="tetrapodus">M. tetrapodus</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0023CA2720007A6499CF829C81FF2A" authorityName="Bretscher" authorityYear="1900" box="[1249,1436,188,212]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="387" phylum="Annelida" rank="species" species="monochaetus">M. monochaetus</taxonomicName>
|
||
</emphasis>
|
||
are aquatic enchytraids. They also have 1–2 spermathecal diverticula. The terrestrial
|
||
<taxonomicName id="4C0023CA2720007A6414CFDE9C54FF06" box="[1132,1353,224,248]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="387" phylum="Annelida" rank="species" species="kontrimavichusi">
|
||
<emphasis id="B974845B2720007A6414CFDE9C54FF06" box="[1132,1353,224,248]" italics="true" pageId="5" pageNumber="387">M. kontrimavichusi</emphasis>
|
||
</taxonomicName>
|
||
differs from this new species in: spermatheca with one diverticulum, not attached to oesophagus, atria absent, dorsal vessel arising from XV–XVIII. The other terrestrial member,
|
||
<taxonomicName id="4C0023CA2720007A633DCE169AB8FEBE" box="[837,933,296,320]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="387" phylum="Annelida" rank="species" species="calyx">
|
||
<emphasis id="B974845B2720007A633DCE169AB8FEBE" box="[837,933,296,320]" italics="true" pageId="5" pageNumber="387">M. calyx</emphasis>
|
||
</taxonomicName>
|
||
, is different from
|
||
<taxonomicName id="4C0023CA2720007A6416CE169C64FEBE" authorityName="Zhang & Lu & Xie" authorityYear="2018" box="[1134,1401,296,320]" class="Oligochaeta" family="Enchytraeidae" genus="Mesenchytraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="387" phylum="Annelida" rank="species" species="spermatoglomeratus">
|
||
<emphasis id="B974845B2720007A6416CE169C64FEBE" box="[1134,1401,296,320]" italics="true" pageId="5" pageNumber="387">M. spermatoglomeratus</emphasis>
|
||
</taxonomicName>
|
||
by its larger body size (
|
||
<quantity id="4CF8F5AC2720007A61F0CE739B17FE9A" box="[392,522,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.9" metricValueMin="2.0" pageId="5" pageNumber="387" unit="mm" value="29.5" valueMax="39.0" valueMin="20.0">20–39 mm</quantity>
|
||
), the location of enlarged chaetae (VII–IX) and spermatheca with 7–9 pointed diverticula.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |