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<document id="DCDD9E52B7D7AF51E899CA147DEF1D05" ID-DOI="10.11646/zootaxa.4044.2.2" ID-GBIF-Dataset="8b09a6a1-35c3-407c-a4b5-076c4af030db" ID-ISSN="1175-5326" ID-Zenodo-Dep="241241" ID-ZooBank="7000893A-A8BC-4196-B76D-385C97B96FBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461338858631" checkinUser="plazi" docAuthor="Kanao, Taisuke &amp; Maruyama, Munetoshi" docDate="2015" docId="0395F739FFAFBF29FF02194BFCF3BDB5" docLanguage="en" docName="zt04044p223.pdf" docOrigin="Zootaxa 4044 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Discoxenus kohkongensis Kanao &amp; Maruyama, 2015, n. sp." docType="treatment" docVersion="11" lastPageNumber="213" masterDocId="FFAC8F41FFA5BF25FF951C5CFFC1BC65" masterDocTitle="Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="223" masterPageNumber="201" pageNumber="211" updateTime="1698619795159" updateUser="plazi">
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<mods:title id="413BEC92768B23E9F8B52F7F612920E0">Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="850F0360F289D5083FEB7E24435DC950">Kanao, Taisuke</mods:namePart>
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<mods:namePart id="6E41D2A64DFB6E320A602E5BEA6CFE5F">Maruyama, Munetoshi</mods:namePart>
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<mods:title id="6EFCFBB8DBFC4D59EABB750EF15BFDCA">Zootaxa</mods:title>
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<mods:date id="98E98D573FF5596CA63973CCD8839740">2015</mods:date>
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<treatment id="0395F739FFAFBF29FF02194BFCF3BDB5" ID-DOI="http://doi.org/10.5281/zenodo.6111783" ID-GBIF-Taxon="119644722" ID-Zenodo-Dep="6111783" LSID="urn:lsid:plazi:treatment:0395F739FFAFBF29FF02194BFCF3BDB5" httpUri="http://treatment.plazi.org/id/0395F739FFAFBF29FF02194BFCF3BDB5" lastPageId="12" lastPageNumber="213" pageId="10" pageNumber="211">
<subSubSection id="C32615A4FFAFBF2FFF02194BFE5CB936" pageId="10" pageNumber="211" type="nomenclature">
<paragraph id="8B83462FFFAFBF2FFF02194BFDDBB957" blockId="10.[151,538,1303,1363]" box="[151,538,1303,1330]" pageId="10" pageNumber="211">
<heading id="D0CBF143FFAFBF2FFF02194BFDDBB957" bold="true" box="[151,538,1303,1330]" fontSize="11" level="1" pageId="10" pageNumber="211" reason="1">
<emphasis id="B9489A3DFFAFBF2FFF02194BFDDBB957" bold="true" box="[151,538,1303,1330]" pageId="10" pageNumber="211">
<taxonomicName id="4C3C3DACFFAFBF2FFF02194BFE11B954" ID-CoL="6CZCB" box="[151,464,1303,1329]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="kohkongensis" status="sp. nov.">
<emphasis id="B9489A3DFFAFBF2FFF02194BFE11B954" bold="true" box="[151,464,1303,1329]" italics="true" pageId="10" pageNumber="211">Discoxenus kohkongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFAFBF2FFE421944FDDBB957" box="[471,538,1304,1330]" pageId="10" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B83462FFFAFBF2FFF021966FE5CB936" blockId="10.[151,538,1303,1363]" box="[151,413,1338,1363]" pageId="10" pageNumber="211">
(
<figureCitation id="13075AAAFFAFBF2FFF0A1966FECBB936" box="[159,266,1338,1363]" captionStart="FIGURES 5 12" captionStartId="3.[151,264,1680,1703]" captionTargetBox="[151,1436,483,1659]" captionTargetId="figure@3.[151,1436,483,1659]" captionTargetPageId="3" captionText="FIGURES 5 12. Habitus. 5 6. Discoxenus phourini n. sp.: 5. Dorsal view, 6. Ventral view. 7 8. D. kohkongensis n. sp.: 7. Dorsal view, 8. Ventral view. 9 10. D. hirsutus n. sp.: 9. Dorsal view, 10. Ventral view. 11 12. D. minutus n. sp.: 11. Dorsal view, 12. Ventral view." httpUri="https://zenodo.org/record/241243/files/figure.png" pageId="10" pageNumber="211">Figs. 7, 8</figureCitation>
,
<figureCitation id="13075AAAFFAFBF2FFE831967FE9EB936" box="[278,351,1339,1363]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">7388</figureCitation>
,
<figureCitation id="13075AAAFFAFBF2FFEFF1966FE54B936" box="[362,405,1338,1363]" captionStart="FIGURES 165 166" captionStartId="21.[151,264,581,604]" captionTargetBox="[260,1324,186,557]" captionTargetId="figure@21.[255,1332,183,560]" captionTargetPageId="21" captionText="FIGURES 165 166. Habitus. 165. Discoxenus kohkongensis n. sp., 166. D. lucidus n. sp." httpUri="https://zenodo.org/record/241254/files/figure.png" pageId="10" pageNumber="211">165</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFAFBF2FFF0219DEFE4EB986" pageId="10" pageNumber="211" type="materials_examined">
<paragraph id="8B83462FFFAFBF2FFF0219DEFACFB9FE" blockId="10.[151,1437,1410,2011]" box="[151,1294,1410,1435]" pageId="10" pageNumber="211">
<emphasis id="B9489A3DFFAFBF2FFF0219DEFE0DB9FE" bold="true" box="[151,460,1410,1435]" pageId="10" pageNumber="211">
<typeStatus id="5487F88DFFAFBF2FFF0219DEFF13B9FE" box="[151,210,1410,1435]" pageId="10" pageNumber="211">Type</typeStatus>
materials.
<typeStatus id="5487F88DFFAFBF2FFEC319DEFE06B9FE" box="[342,455,1410,1435]" pageId="10" pageNumber="211" type="holotype">Holotype</typeStatus>
:
</emphasis>
♂, Koh Kong,
<collectingCountry id="F32B06BFFFAFBF2FFDEF19DEFD31B9FE" box="[634,752,1410,1435]" name="Cambodia" pageId="10" pageNumber="211">Cambodia</collectingCountry>
,
<date id="FF8260EFFFAFBF2FFD6E19DFFCB8B9FE" box="[763,889,1411,1435]" pageId="10" pageNumber="211" value="2012-12-07">7 XII 2012</date>
, Kanao T. leg. (Colony no. KT428).
</paragraph>
<paragraph id="8B83462FFFAFBF2FFF5219FAFE4EB986" blockId="10.[151,1437,1410,2011]" pageId="10" pageNumber="211">
<emphasis id="B9489A3DFFAFBF2FFF5219FAFE89B9DA" bold="true" box="[199,328,1446,1471]" pageId="10" pageNumber="211">
<typeStatus id="5487F88DFFAFBF2FFF5219FAFE82B9DA" box="[199,323,1446,1471]" pageId="10" pageNumber="211" type="paratype">Paratypes</typeStatus>
:
</emphasis>
2♂♂, 25♀♀, same data as
<typeStatus id="5487F88DFFAFBF2FFDEB19FAFD21B9DA" box="[638,736,1446,1471]" pageId="10" pageNumber="211" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D3A8DA6FFAFBF2FFD6419FBFCD4B9DA" box="[753,789,1447,1471]" pageId="10" pageNumber="211" type="generic">1♂</specimenCount>
, completely dissected, 1♀, head and abdominal segments VIIIX dissected off).
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFAFBF2FFF5219B2FDA1BADE" pageId="10" pageNumber="211" type="diagnosis">
<paragraph id="8B83462FFFAFBF2FFF5219B2FDA1BADE" blockId="10.[151,1437,1410,2011]" pageId="10" pageNumber="211">
<emphasis id="B9489A3DFFAFBF2FFF5219B2FE80BA62" bold="true" box="[199,321,1518,1543]" pageId="10" pageNumber="211">Diagnosis.</emphasis>
This species is distinguished from other
<taxonomicName id="4C3C3DACFFAFBF2FFC9E19B3FC4DBA63" box="[779,908,1519,1542]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B9489A3DFFAFBF2FFC9E19B3FC4DBA63" box="[779,908,1519,1542]" italics="true" pageId="10" pageNumber="211">Discoxenus</emphasis>
</taxonomicName>
species by a combination of the following four character states: macrochaetotaxy of tergites IIIVIII (6, 6, 6, 6, 6, 6), sternites IIIVII with macrosetae only at each posterior margin, paramere with paramerite narrow (approximately 3 times as wide as condylite) (
<figureCitation id="13075AAAFFAFBF2FFAAC1A6AFA4EBA2A" box="[1337,1423,1590,1615]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Fig. 87</figureCitation>
), and spermatheca with short basal part (approximately 1.5 times longer than apical part) (
<figureCitation id="13075AAAFFAFBF2FFB1E1A06FB21BA16" box="[1163,1248,1626,1651]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Fig. 88</figureCitation>
). This species is most similar to
<taxonomicName id="4C3C3DACFFAFBF2FFEC11A23FE15BAF3" box="[340,468,1663,1686]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B9489A3DFFAFBF2FFEC11A23FE15BAF3" box="[340,468,1663,1686]" italics="true" pageId="10" pageNumber="211">D. minutus</emphasis>
</taxonomicName>
, but is distinguishable from it by a male aedeagal median lobe with apically and moderately pointed distal crest (
<figureCitation id="13075AAAFFAFBF2FFE6A1AFEFD92BADE" box="[511,595,1698,1723]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Fig. 86</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFAFBF29FF521A9AFBB7BDED" lastPageId="12" lastPageNumber="213" pageId="10" pageNumber="211" type="description">
<paragraph id="8B83462FFFAFBF2EFF521A9AFF31BC92" blockId="10.[151,1437,1410,2011]" lastBlockId="11.[151,1437,151,536]" lastPageId="11" lastPageNumber="212" pageId="10" pageNumber="211">
<emphasis id="B9489A3DFFAFBF2FFF521A9AFE98BABA" bold="true" box="[199,345,1734,1759]" pageId="10" pageNumber="211">Description.</emphasis>
Head (
<figureCitation id="13075AAAFFAFBF2FFE3B1A9AFDC3BABA" box="[430,514,1734,1759]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Fig. 73</figureCitation>
) more than 1.3 times wider than long, with anterior margin of clypeus rounded. Antenna (
<figureCitation id="13075AAAFFAFBF2FFE9D1AB6FEBABB66" box="[264,379,1770,1795]" captionStart="FIGURES 5 12" captionStartId="3.[151,264,1680,1703]" captionTargetBox="[151,1436,483,1659]" captionTargetId="figure@3.[151,1436,483,1659]" captionTargetPageId="3" captionText="FIGURES 5 12. Habitus. 5 6. Discoxenus phourini n. sp.: 5. Dorsal view, 6. Ventral view. 7 8. D. kohkongensis n. sp.: 7. Dorsal view, 8. Ventral view. 9 10. D. hirsutus n. sp.: 9. Dorsal view, 10. Ventral view. 11 12. D. minutus n. sp.: 11. Dorsal view, 12. Ventral view." httpUri="https://zenodo.org/record/241243/files/figure.png" pageId="10" pageNumber="211">Figs. 7, 8</figureCitation>
,
<figureCitation id="13075AAAFFAFBF2FFE1F1AB7FE66BB66" box="[394,423,1771,1795]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">74</figureCitation>
) with segment I longer than other segments; segment II as long as segment IV, with 3 macrosetae; segment III trapezoidal, shorter than other segments; segment IV transverse; segment V wider than other segments; segments VIXI successively narrowed distally; segments VIVIII wider than long; segment IX subquadrate; segment X longer than wide; segment XI approximately 2.2 times longer than wide, narrowed apically. Labrum (
<figureCitation id="13075AAAFFAFBF2FFEF61B26FE74BBF6" box="[355,437,1914,1939]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Fig. 75</figureCitation>
, left side) with anterior margin concave; median projection of apodeme broadly rounded; 4 lateral setae present in ventral view (
<figureCitation id="13075AAAFFAFBF2FFDC51BC2FD65BBD2" box="[592,676,1950,1975]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Fig. 75</figureCitation>
, right side). Mandibles (
<figureCitation id="13075AAAFFAFBF2FFC2F1BC2FB83BBD2" box="[954,1090,1950,1975]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Figs. 76, 77</figureCitation>
) with 2 scrobal setae, covered with approximately 30 pores. Left mandible (
<figureCitation id="13075AAAFFAFBF2FFD051B9EFD20BBBE" box="[656,737,1986,2011]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Fig. 76</figureCitation>
) with adoral margin bisinuate. Right mandible (
<figureCitation id="13075AAAFFAFBF2FFB6D1B9EFA88BBBE" box="[1272,1353,1986,2011]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="10" pageNumber="211">Fig. 77</figureCitation>
) with a distinct tooth. Maxillary palpal segment III approximately 2 times longer than wide. Mentum (
<figureCitation id="13075AAAFFAEBF2EFB611CCBFA88BCD5" box="[1268,1353,151,176]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 78</figureCitation>
) short, approximately 3.2 times wider than long, covered with around 20 pores. Labium with prementum covered with 6 7 pores.
</paragraph>
<paragraph id="8B83462FFFAEBF2EFF521D59FA85BD00" blockId="11.[151,1437,151,536]" pageId="11" pageNumber="212">
Pronotum (
<figureCitation id="13075AAAFFAEBF2EFEDC1D58FE5EBD79" box="[329,415,260,285]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 79</figureCitation>
) sparsely covered with approximately 46 macrosetae, few minute setae present around anterior margin. Elytron (
<figureCitation id="13075AAAFFAEBF2EFE2B1D7BFDD2BD25" box="[446,531,295,320]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 80</figureCitation>
) transverse, sparsely covered with pores, several setae present at anterolateral outer corner, 12 macrosetae present on disc. Metaventrite approximately 1.5 times as long as mesoventrite.
</paragraph>
<paragraph id="8B83462FFFAEBF2EFF521D33FCF1BE7D" blockId="11.[151,1437,151,536]" pageId="11" pageNumber="212">
Tergites IIIIV (
<figureCitation id="13075AAAFFAEBF2EFE131D33FE0CBDED" box="[390,461,367,392]" captionStart="FIGURES 5 12" captionStartId="3.[151,264,1680,1703]" captionTargetBox="[151,1436,483,1659]" captionTargetId="figure@3.[151,1436,483,1659]" captionTargetPageId="3" captionText="FIGURES 5 12. Habitus. 5 6. Discoxenus phourini n. sp.: 5. Dorsal view, 6. Ventral view. 7 8. D. kohkongensis n. sp.: 7. Dorsal view, 8. Ventral view. 9 10. D. hirsutus n. sp.: 9. Dorsal view, 10. Ventral view. 11 12. D. minutus n. sp.: 11. Dorsal view, 12. Ventral view." httpUri="https://zenodo.org/record/241243/files/figure.png" pageId="11" pageNumber="212">Fig. 7</figureCitation>
) covered with few setae laterally. Tergites VVIII with posterior half covered with several setae. Tergite VIII (
<figureCitation id="13075AAAFFAEBF2EFE4E1DC8FDF0BDC9" box="[475,561,404,429]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 81</figureCitation>
) with posterior margin pointed, 3 pairs of macrosetae present at posterior margin, 2 pairs of macrosetae present around middle. Macrochaetotaxy of abdominal tergites IIIVIII = 6, 6, 6, 6, 6, 6. Sternite III (
<figureCitation id="13075AAAFFAEBF2EFEF31D80FE6DBD91" box="[358,428,476,501]" captionStart="FIGURES 5 12" captionStartId="3.[151,264,1680,1703]" captionTargetBox="[151,1436,483,1659]" captionTargetId="figure@3.[151,1436,483,1659]" captionTargetPageId="3" captionText="FIGURES 5 12. Habitus. 5 6. Discoxenus phourini n. sp.: 5. Dorsal view, 6. Ventral view. 7 8. D. kohkongensis n. sp.: 7. Dorsal view, 8. Ventral view. 9 10. D. hirsutus n. sp.: 9. Dorsal view, 10. Ventral view. 11 12. D. minutus n. sp.: 11. Dorsal view, 12. Ventral view." httpUri="https://zenodo.org/record/241243/files/figure.png" pageId="11" pageNumber="212">Fig. 8</figureCitation>
) sparsely covered with setae. Sternites IVVIII with a row of yellow setae at middle. Sternites IIIVII with 610 macrosetae at posterior margin.
</paragraph>
<caption id="DF4316A7FFAEBF2EFF021A88FCD0BB22" httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212" targetBox="[162,1418,603,1714]" targetPageId="11">
<paragraph id="8B83462FFFAEBF2EFF021A88FCD0BB22" blockId="11.[151,1436,1748,1863]" pageId="11" pageNumber="212">
<emphasis id="B9489A3DFFAEBF2EFF021A88FE94BA8F" bold="true" box="[151,341,1748,1771]" pageId="11" pageNumber="212">FIGURES 7388.</emphasis>
<taxonomicName id="4C3C3DACFFAEBF2EFEC91A8AFDA0BA8E" box="[348,609,1749,1771]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="kohkongensis" status="sp. nov.">
<emphasis id="B9489A3DFFAEBF2EFEC91A8AFDA0BA8E" box="[348,609,1749,1771]" italics="true" pageId="11" pageNumber="212">Discoxenus kohkongensis</emphasis>
</taxonomicName>
<emphasis id="B9489A3DFFAEBF2EFDFC1A8AFD65BA8E" bold="true" box="[617,676,1750,1771]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A27B2746FFAEBF2EFDFC1A8AFD65BA8E" box="[617,676,1750,1771]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca.
</paragraph>
</caption>
<paragraph id="8B83462FFFAEBF29FF521B33FF12BCB1" blockId="11.[151,1436,1902,2036]" lastBlockId="12.[151,1436,151,464]" lastPageId="12" lastPageNumber="213" pageId="11" pageNumber="212">
<emphasis id="B9489A3DFFAEBF2EFF521B33FEC9BBE2" box="[199,264,1903,1927]" italics="true" pageId="11" pageNumber="212">Male.</emphasis>
Sternite VIII (
<figureCitation id="13075AAAFFAEBF2EFE2D1B32FDCFBBE2" box="[440,526,1902,1927]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 82</figureCitation>
) with posterior half covered with few setae, 2 pairs of macrosetae present at posterior margin and middle. Median lobe of aedeagus narrowed toward apical lobe in ventral view (
<figureCitation id="13075AAAFFAEBF2EFB6E1BCEFA8FBBCE" box="[1275,1358,1938,1963]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 85</figureCitation>
); basal capsule slightly concave at middle, with distal crest moderately pointed apically in lateral view (
<figureCitation id="13075AAAFFAEBF2EFB781BEAFA83BBAA" box="[1261,1346,1974,1999]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 86</figureCitation>
); apical lobe less than half as wide as basal capsule in lateral view (
<figureCitation id="13075AAAFFAEBF2EFCD61B87FC59BB91" box="[835,920,2011,2036]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 86</figureCitation>
). Paramere (
<figureCitation id="13075AAAFFAEBF2EFBBC1B87FBBDBB96" box="[1065,1148,2011,2036]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="11" pageNumber="212">Fig. 87</figureCitation>
) with paramerite narrow, approximately 3 times as wide as condylite; velar sac sclerite with 45 setae; apical lobe with 2 minute setae at apex.
</paragraph>
<paragraph id="8B83462FFFA9BF29FF521CBCFC49BD79" blockId="12.[151,1436,151,464]" pageId="12" pageNumber="213">
<emphasis id="B9489A3DFFA9BF29FF521CBCFEDABC9D" box="[199,283,224,248]" italics="true" pageId="12" pageNumber="213">Female</emphasis>
. Sternite VIII (
<figureCitation id="13075AAAFFA9BF29FE5C1C83FDDDBC9D" box="[457,540,223,248]" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="12" pageNumber="213">Fig. 83</figureCitation>
) with 2 pairs of macrosetae at posterior margin and middle. Spermatheca (
<figureCitation id="13075AAAFFA9BF29FAFB1C83FF75BD79" captionStart="FIGURES 73 88" captionStartId="11.[151,264,1748,1771]" captionTargetBox="[162,1418,603,1714]" captionTargetId="figure@11.[151,1436,589,1728]" captionTargetPageId="11" captionText="FIGURES 73 88. Discoxenus kohkongensis n. sp.: 73. Head, 74. Antenna, 75. Labrum (left side = dorsal view, right side = ventral view), 76. Mandible, left, 77. Mandible, right, 78. Mentum, 79. Pronotum, 80. Elytron, right, 81. Tergite VIII, 82. Sternite VIII, male, 83. Sternite VIII, female. 84, Tergites IX and X. 85, Median lobe of aedeagus, ventral view, 86. Median lobe of aedeagus, lateral view, 87. Paramere. 88. Spermatheca." httpUri="https://zenodo.org/record/241248/files/figure.png" pageId="12" pageNumber="213">Fig. 88</figureCitation>
) with basal part approximately 1.5 times longer than apical part.
</paragraph>
<paragraph id="8B83462FFFA9BF29FF521D7BFBB7BDED" blockId="12.[151,1436,151,464]" pageId="12" pageNumber="213">
<emphasis id="B9489A3DFFA9BF29FF521D7BFEB3BD25" bold="true" box="[199,370,295,320]" pageId="12" pageNumber="213">Measurement.</emphasis>
Body length = average
<quantity id="4CC4EBCAFFA9BF29FD141D74FD27BD5A" box="[641,742,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="213" unit="mm" value="1.4">1.40 mm</quantity>
(
<quantity id="4CC4EBCAFFA9BF29FD601D7BFC5DBD5A" box="[757,924,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" metricValueMax="1.5" metricValueMin="1.38" pageId="12" pageNumber="213" unit="mm" value="1.44" valueMax="1.5" valueMin="1.38">1.381.50 mm</quantity>
, N = 10), pronotal length = average
<quantity id="4CC4EBCAFFA9BF29FAAD1D74FA5DBD5A" box="[1336,1436,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="12" pageNumber="213" unit="mm" value="0.47">0.47 mm</quantity>
(
<quantity id="4CC4EBCAFFA9BF29FF0B1D10FE84BD01" box="[158,325,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="12" pageNumber="213" unit="mm" value="0.45999999999999996" valueMax="0.51" valueMin="0.41">0.410.51 mm</quantity>
, N = 10), pronotal width = average
<quantity id="4CC4EBCAFFA9BF29FD411D10FCF6BD01" box="[724,823,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="213" unit="mm" value="0.7">0.70 mm</quantity>
(
<quantity id="4CC4EBCAFFA9BF29FCD31D10FC2DBD01" box="[838,1004,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.3" metricValueMin="6.7" pageId="12" pageNumber="213" unit="mm" value="0.7" valueMax="0.73" valueMin="0.67">0.670.73 mm</quantity>
, N = 10), elytral length = average
<quantity id="4CC4EBCAFFA9BF29FAFE1D10FF02BDE2" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="213" unit="mm" value="0.32">0.32 mm</quantity>
(
<quantity id="4CC4EBCAFFA9BF29FF441D33FEB9BDE2" box="[209,376,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.5999999999999996" metricValueMin="2.8" pageId="12" pageNumber="213" unit="mm" value="0.32" valueMax="0.36" valueMin="0.28">0.280.36 mm</quantity>
, N = 10), elytral width = average
<quantity id="4CC4EBCAFFA9BF29FD661D33FC96BDE2" box="[755,855,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="12" pageNumber="213" unit="mm" value="0.36">0.36 mm</quantity>
(
<quantity id="4CC4EBCAFFA9BF29FCF01D2CFBCDBDE2" box="[869,1036,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.1" metricValueMin="3.3" pageId="12" pageNumber="213" unit="mm" value="0.37" valueMax="0.41" valueMin="0.33">0.330.41 mm</quantity>
, N = 10).
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFA9BF29FF521DC8FCF3BDB5" pageId="12" pageNumber="213" type="etymology">
<paragraph id="8B83462FFFA9BF29FF521DC8FAE2BDC8" blockId="12.[151,1436,151,464]" box="[199,1315,404,429]" pageId="12" pageNumber="213">
<emphasis id="B9489A3DFFA9BF29FF521DC8FE8FBDC8" bold="true" box="[199,334,404,429]" pageId="12" pageNumber="213">Etymology.</emphasis>
The specific Lathinized epithet is derived from its
<typeStatus id="5487F88DFFA9BF29FC1F1DC9FC7BBDC8" box="[906,954,405,429]" pageId="12" pageNumber="213">type</typeStatus>
locality of Koh Kong Province.
</paragraph>
<paragraph id="8B83462FFFA9BF29FF521DEBFCF3BDB5" blockId="12.[151,1436,151,464]" box="[199,818,439,464]" pageId="12" pageNumber="213">
<emphasis id="B9489A3DFFA9BF29FF521DEBFE9EBDB5" bold="true" box="[199,351,439,464]" pageId="12" pageNumber="213">Host species.</emphasis>
<taxonomicName id="4C3C3DACFFA9BF29FEF31DE5FCEFBDB5" authority="Ahmad, 1965" authorityName="Ahmad" authorityYear="1965" box="[358,814,439,464]" class="Insecta" family="Termitidae" genus="Hypotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="makhamensis">
<emphasis id="B9489A3DFFA9BF29FEF31DE5FD4DBDB5" box="[358,652,440,464]" italics="true" pageId="12" pageNumber="213">Hypotermes makhamensis</emphasis>
<bibRefCitation id="EFAD3BDEFFA9BF29FD071DEBFCEFBDB5" author="Ahmad" box="[658,814,439,464]" pageId="12" pageNumber="213" refString="Ahmad, M. (1965) Termites (Isoptera) of Thailand. Bulletin of the American Museum of Natural History, 131 (1), 1 - 114." type="journal article" year="1965">Ahmad, 1965</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>