403 lines
59 KiB
XML
403 lines
59 KiB
XML
<document id="BEF3B8CDC58937039DA572B5CC67C998" ID-CLB-Dataset="32021" ID-DOI="10.11646/zootaxa.4323.3.1" ID-GBIF-Dataset="f1be4566-adeb-4995-83c7-2aebab3c7458" ID-ISSN="1175-5326" ID-Zenodo-Dep="919606" ID-ZooBank="3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506057765384" checkinUser="plazi" docAuthor="Schmidt, Joachim" docDate="2017" docId="03956A03A3637F6CFF67FF3689E0D8E0" docLanguage="en" docName="zootaxa.4323.3.1.pdf" docOrigin="Zootaxa 4323 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus (Bottchrus) schawalleri Schmidt 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="318" masterDocId="FFAC127BA36D7F7DFFF0FFAE8B5CD930" masterDocTitle="Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species" masterLastPageNumber="358" masterPageNumber="301" pageNumber="315" updateTime="1698468866275" updateUser="ExternalLinkService">
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<mods:title id="B8DEF1A3986FFFDD22B853EB0F876BBA">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<mods:namePart id="17A6E4528F330BBAA0734CE739563405">Schmidt, Joachim</mods:namePart>
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<mods:title id="22A618973E0FA6F0692656F7AEA74A89">Zootaxa</mods:title>
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<mods:date id="9524F2DB144E6732415C0FDFC01F9812">2017</mods:date>
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<mods:number id="556D83EE1DBDCB2B60D4A3C5E0DF1EF8">2017-09-22</mods:number>
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<treatment id="03956A03A3637F6CFF67FF3689E0D8E0" ID-DOI="http://doi.org/10.5281/zenodo.6015747" ID-GBIF-Taxon="134839424" ID-Zenodo-Dep="6015747" LSID="urn:lsid:plazi:treatment:03956A03A3637F6CFF67FF3689E0D8E0" httpUri="http://treatment.plazi.org/id/03956A03A3637F6CFF67FF3689E0D8E0" lastPageId="17" lastPageNumber="318" pageId="14" pageNumber="315">
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<paragraph id="8B83DB15A3637F73FF67FF3689F2D982" blockId="14.[151,686,152,211]" box="[151,686,152,178]" pageId="14" pageNumber="315">
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<heading id="D0CB6C79A3637F73FF67FF3689F2D982" bold="true" box="[151,686,152,178]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
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<emphasis id="B9480707A3637F73FF67FF3689F2D982" bold="true" box="[151,686,152,178]" pageId="14" pageNumber="315">
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<taxonomicName id="4C3CA096A3637F73FF67FF368939D982" ID-CoL="58LHV" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,613,152,178]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov." subGenus="Bottchrus">
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<emphasis id="B9480707A3637F73FF67FF368A1DD982" bold="true" box="[151,321,152,178]" italics="true" pageId="14" pageNumber="315">Trichotichnus</emphasis>
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(
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<emphasis id="B9480707A3637F73FEA2FF368A95D982" bold="true" box="[338,457,152,178]" italics="true" pageId="14" pageNumber="315">Bottchrus</emphasis>
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)
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<emphasis id="B9480707A3637F73FE2AFF368939D982" bold="true" box="[474,613,152,178]" italics="true" pageId="14" pageNumber="315">schawalleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27BBA7CA3637F73FD9CFF3689F2D982" box="[620,686,152,178]" pageId="14" pageNumber="315" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B83DB15A3637F73FF67FF148AF8D9E3" blockId="14.[151,686,152,211]" box="[151,420,186,211]" pageId="14" pageNumber="315">
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(
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<figureCitation id="1307C790A3637F73FF6FFF148BA9D9E3" box="[159,245,186,211]" captionStart="FIGURES 38 – 41" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[273,1314,196,1918]" captionTargetId="figure@15.[271,1315,193,1921]" captionTargetPageId="15" captionText="FIGURES 38 – 41. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 38, T. schawalleri sp. n., holotype; 39, T. obliquebasalis sp. n., paratype, male; 40, T. siklesensis sp. n., paratype, male; 41, T. panchhaseensis sp. n., paratype, male." httpUri="https://zenodo.org/record/919620/files/figure.png" pageId="14" pageNumber="315">Figs 38</figureCitation>
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,
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<figureCitation id="1307C790A3637F73FEF1FF148A16D9E3" box="[257,330,186,211]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="14" pageNumber="315">42–46</figureCitation>
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,
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<figureCitation id="1307C790A3637F73FEA5FF148ADDD9E3" box="[341,385,186,211]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="14" pageNumber="315">151</figureCitation>
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,
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<figureCitation id="1307C790A3637F73FE7DFF158AC7D9E2" box="[397,411,187,210]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="14" pageNumber="315">
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<emphasis id="B9480707A3637F73FE7DFF158AC7D9E2" box="[397,411,187,210]" italics="true" pageId="14" pageNumber="315">5</emphasis>
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</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C326889EA3637F73FF67FEAC8A33D8B7" pageId="14" pageNumber="315" type="materials_examined">
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<paragraph id="8B83DB15A3637F73FF67FEAC8AC8D80F" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="315">
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<materialsCitation id="3B54D148A3637F73FF67FEAC8AC8D80F" ID-GBIF-Occurrence="1655986840" collectingDate="1997-05-26" collectionCode="SMNS" country="Nepal" elevation="2100" location="Bhojpur Distr." pageId="14" pageNumber="315" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B9480707A3637F73FF67FEAC8A65D82B" bold="true" box="[151,313,258,283]" pageId="14" pageNumber="315">
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<typeStatus id="548765B7A3637F73FF67FEAC8B93D82B" box="[151,207,258,283]" pageId="14" pageNumber="315">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="548765B7A3637F73FEB9FEAC8AEDD82B" box="[329,433,258,283]" pageId="14" pageNumber="315" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D3A109CA3637F73FE4EFEAD8A88D82A" box="[446,468,259,282]" pageId="14" pageNumber="315" type="male">♂</specimenCount>
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, “539
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<collectingCountry id="F32B9B85A3637F73FDD1FEAD8926D82B" box="[545,634,259,283]" name="Nepal" pageId="14" pageNumber="315">NEPAL</collectingCountry>
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:
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<location id="8EE38DCEA3637F73FD76FEAC887FD82B" LSID="urn:lsid:plazi:treatment:03956A03A3637F6CFF67FF3689E0D8E0:8EE38DCEA3637F73FD76FEAC887FD82B" box="[646,803,258,283]" country="Nepal" name="Bhojpur Distr." pageId="14" pageNumber="315">Bhojpur Distr.</location>
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Dilkharka
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||
<elevation id="00113C26A3637F73FC51FEAD88ABD82A" box="[929,1015,259,283]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="315" unit="m" value="2100.0">
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<quantity id="4CC476F0A3637F73FC51FEAD88ABD82A" box="[929,1015,259,283]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="315" unit="m" value="2100.0">2100 m</quantity>
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||
</elevation>
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||
,
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||
<collectingDate id="EFC6043DA3637F73FBF5FEAC8F2FD82B" box="[1029,1139,258,283]" pageId="14" pageNumber="315" value="1997-05-26">
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||
<date id="FF82FDD5A3637F73FBF5FEAC8F2FD82B" box="[1029,1139,258,283]" pageId="14" pageNumber="315" value="1997-05-26">26.V.1997</date>
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||
</collectingDate>
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||
leg. W. SCHAWALLER”, “Mus. Stuttg” (
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||
<collectionCode id="ED2D43D0A3637F73FEC9FE898AD4D80F" box="[313,392,295,319]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="14" pageNumber="315">SMNS</collectionCode>
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||
).
|
||
</materialsCitation>
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</paragraph>
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||
<paragraph id="8B83DB15A3637F73FF37FEE58A33D8B7" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="315">
|
||
<typeStatus id="548765B7A3637F73FF37FEE58A75D853" box="[199,297,331,355]" pageId="14" pageNumber="315" type="paratype">Paratype</typeStatus>
|
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:
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||
<collectingCountry id="F32B9B85A3637F73FEC2FEE48A29D853" box="[306,373,330,355]" name="Nepal" pageId="14" pageNumber="315">
|
||
<emphasis id="B9480707A3637F73FEC2FEE48A29D853" bold="true" box="[306,373,330,355]" pageId="14" pageNumber="315">Nepal</emphasis>
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</collectingCountry>
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||
:
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||
<specimenCount id="9D3A109CA3637F73FE72FEE58AF6D852" box="[386,426,331,355]" pageId="14" pageNumber="315" type="male">1 ♂</specimenCount>
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, “536
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<collectingCountry id="F32B9B85A3637F73FE01FEE58916D853" box="[497,586,331,355]" name="Nepal" pageId="14" pageNumber="315">NEPAL</collectingCountry>
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||
: Bhojpur Distr. Valley NW Phedi
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||
<quantity id="4CC476F0A3637F73FC43FEE58F5AD852" box="[947,1030,331,355]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="315" unit="m" value="1900.0">1900 m</quantity>
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,
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<date id="FF82FDD5A3637F73FBE1FEE48F22D853" box="[1041,1150,330,355]" pageId="14" pageNumber="315" value="1997-05-25">25.V.1997</date>
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||
leg. W. SCHAWALLER”, “Mus. Stuttg” (cJS).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C326889EA3637F6DFF37FE3C88E9DCF2" lastPageId="16" lastPageNumber="317" pageId="14" pageNumber="315" type="description">
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<paragraph id="8B83DB15A3637F73FF37FE3C8A02D8FF" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="315">
|
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<emphasis id="B9480707A3637F73FF37FE3C8AF7D89B" bold="true" box="[199,427,402,427]" pageId="14" pageNumber="315">Description (male).</emphasis>
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||
Habitus as in
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||
<figureCitation id="1307C790A3637F73FDBCFE3C89C2D89B" box="[588,670,402,427]" captionStart="FIGURES 38 – 41" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[273,1314,196,1918]" captionTargetId="figure@15.[271,1315,193,1921]" captionTargetPageId="15" captionText="FIGURES 38 – 41. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 38, T. schawalleri sp. n., holotype; 39, T. obliquebasalis sp. n., paratype, male; 40, T. siklesensis sp. n., paratype, male; 41, T. panchhaseensis sp. n., paratype, male." httpUri="https://zenodo.org/record/919620/files/figure.png" pageId="14" pageNumber="315">Fig. 38</figureCitation>
|
||
. Body length in
|
||
<typeStatus id="548765B7A3637F73FCA7FE3C88E5D89B" box="[855,953,402,427]" pageId="14" pageNumber="315" type="holotype">holotype</typeStatus>
|
||
5.8, in
|
||
<typeStatus id="548765B7A3637F73FBFDFE3D8F32D89B" box="[1037,1134,403,427]" pageId="14" pageNumber="315" type="paratype">paratype</typeStatus>
|
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<quantity id="4CC476F0A3637F73FB85FE3C8F91D89A" box="[1141,1229,402,427]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="14" pageNumber="315" unit="mm" value="6.1">6.1 mm</quantity>
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, width 2.7 and
|
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<quantity id="4CC476F0A3637F73FA89FE3D8B99D8FE" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="14" pageNumber="315" unit="mm" value="2.8">2.8 mm</quantity>
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, respectively.
|
||
</paragraph>
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||
<paragraph id="8B83DB15A3637F73FF37FE748AD0DB0B" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="315">Colour: Body piceous, shiny on dorsum, with light bluish or blue greenish tinge; labrum, very narrow margins of pronotum and apical portion of elytral margins as well as of elytral epipleurae slightly paler; palpi, antennae and legs brownish yellow.</paragraph>
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||
<paragraph id="8B83DB15A3637F73FF37FDE889C3DD03" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="315">
|
||
Head: Comparatively large, slightly smaller than in
|
||
<taxonomicName id="4C3CA096A3637F73FCF8FDE98893DB6F" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[776,975,582,607]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="bubsaensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3637F73FCF8FDE98841DB6E" box="[776,797,583,606]" italics="true" pageId="14" pageNumber="315">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3637F73FCDCFDE98861DB6E" box="[812,829,583,606]" italics="true" pageId="14" pageNumber="315">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3637F73FCA4FDE88893DB6F" box="[852,975,582,607]" italics="true" pageId="14" pageNumber="315">bubsaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3637F73FC25FDE88F48DB6F" box="[981,1044,582,607]" pageId="14" pageNumber="315" rank="species">
|
||
<emphasis id="B9480707A3637F73FC25FDE88F48DB6F" bold="true" box="[981,1044,582,607]" pageId="14" pageNumber="315">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
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||
, with wide neck (HWmax/PWmax = 0.67, HWmin/PWmax = 0.55–0.57). Eyes rather convex (HWmax/HWmin = 1.17–1.22), but slightly less than in
|
||
<taxonomicName id="4C3CA096A3637F73FF67FD218A35DB97" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,361,654,679]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="bubsaensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3637F73FF67FD218BF0DB96" box="[151,172,655,678]" italics="true" pageId="14" pageNumber="315">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3637F73FF31FD218B8EDB96" box="[193,210,655,678]" italics="true" pageId="14" pageNumber="315">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3637F73FF1EFD208A35DB97" box="[238,361,654,679]" italics="true" pageId="14" pageNumber="315">bubsaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3637F73FE86FD208AE6DB97" box="[374,442,654,679]" pageId="14" pageNumber="315" rank="species">
|
||
<emphasis id="B9480707A3637F73FE86FD208AE6DB97" bold="true" box="[374,442,654,679]" pageId="14" pageNumber="315">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, in lateral view elongate oval. Tempora about half as long as eye, slightly convex, somewhat abruptly sloped to neck. Genae almost twice as wide as antennomere 1, with few short setae. Lateral margin of head between antenna and eye, like in all the preceding species, slightly protruded laterally, forming an obtuse angle near eyes, very finely bordered. Area between supraorbital furrow and upper margin of eye wide, approximately as wide as width of antennomere 2 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow approximately by width of antennomere 3 apically. Labrum slightly concave anteriorly. Clypeus very slightly emarginate along anterior margin, moderately convex basally, flatly sloped to apex, with a comparatively large setigerous pore at each anterior corner. Frons and vertex convex.
|
||
<collectingRegion id="49F815F7A3637F73FF07FC018A0ADAF6" box="[247,342,943,966]" country="Myanmar" name="Chin" pageId="14" pageNumber="315">Mentum</collectingRegion>
|
||
and submentum completely fused, submentum with one long seta and a shorter one laterally of the long seta on each side. Left mandible not truncate, slightly blunted at apex. Dorsal microsculpture consisting of obliterate weakly transverse meshes. Antennae (in male) almost extending to the basal edge of pronotum, with antennomeres 4–8 markedly longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3637F73FF37FB918E44DF37" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="315">Pronotum: Comparatively wide (PWmax/PL = 1.41), widest at the end of anterior third, moderately narrowed posteriad (PWmax/PWmin = 1.18–1.19), with one lateral seta inserted slightly before the widest point. Sides rounded almost throughout, slightly sinuate just before basal angles. Apical margin very slightly emarginate, bordered only laterally. Apical angles very slightly prominent anteriad, widely rounded at tip. Basal margin nearly straight, only slightly roundly oblique laterally, vaguely bordered in lateral third, slightly longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles slightly more than 90, each with a sharp, subdenticulate apex slightly protruded laterad. Pronotal disc moderately convex, strongly sloped to apical angles and moderately so to basal ones, not depressed basally although latero-basal areas slightly flattened. Lateral gutter very narrow, very hardly widened in apical third. Median line interrupted, very fine, superficial, ended markedly before apical and basal margins. Lateral depressions absent; basal foveae small, somewhat elongate and very shallow. Surface sparsely, somewhat finely and irregularly punctate mainly in latero-basal areas, with middle basal portion impunctate, punctation at apical margin absent. Microsculpture present throughout, distinct; meshes mostly transverse, between apical margin and anterior transverse impression more or less isodiametric.</paragraph>
|
||
<paragraph id="8B83DB15A3637F73FF37F9BC885FDE7B" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="315">Elytra: In lateral and caudal view convex, in dorsal view oval, relatively wide (in male, EL/EW = 1.30–1.31; EL/PL = 2.32–2.45; EW/PWmax = 1.27–1.32), widest approximately at middle, with somewhat acute apex. Humeri prominent, rather widely angularly rounded at tip, without denticle. Subapical sinuation shallow. Sutural angle acute and somewhat sharp at tip. Basal edge markedly sinuate laterally, forming distinct obtuse angle with lateral margin. Striae fine, almost superficial along entire length. Intervals somewhat flat throughout, weakly narrowed posteriad. Parascutellar setigerous pore present, large. Parascutellar striole rudimentary, almost absent. Interval 3 with a very small discal setigerous pore at stria 2 behind middle. Marginal umbilicate series widely interrupted at middle, with 6–8 pores in anterior group and 8–9 pores in posterior group. Microsculpture fine, poorly recognizable, consisting of thin transverse lines.</paragraph>
|
||
<paragraph id="8B83DB15A3637F73FF37F8F8890FDE5F" blockId="14.[151,1437,258,2011]" box="[199,595,1878,1903]" pageId="14" pageNumber="315">Hindwings: Reduced to tiny scales.</paragraph>
|
||
<paragraph id="8B83DB15A3637F73FF37F8D48A8EDEEB" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="315">
|
||
Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum (
|
||
<figureCitation id="1307C790A3637F73FBA1F8D48FF8DEA3" box="[1105,1188,1914,1939]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="14" pageNumber="315">Fig. 42</figureCitation>
|
||
) markedly wider than long, strongly narrowed posteriad. Apex of last visible (VII) abdominal sternite (in male) subtruncate and slightly concave just at tip (
|
||
<figureCitation id="1307C790A3637F73FE81F86C8A9ADEEB" box="[369,454,1986,2011]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="14" pageNumber="315">Fig. 43</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF438B9DA3627F72FF67F8388A6FDEDA" httpUri="https://zenodo.org/record/919620/files/figure.png" pageId="15" pageNumber="316" targetBox="[273,1314,196,1918]" targetPageId="15">
|
||
<paragraph id="8B83DB15A3627F72FF67F8388A6FDEDA" blockId="15.[151,1435,1942,2026]" pageId="15" pageNumber="316">
|
||
<emphasis id="B9480707A3627F72FF67F8388A09DE9C" bold="true" box="[151,341,1942,1965]" pageId="15" pageNumber="316">FIGURES 38–41.</emphasis>
|
||
Himalayan species of
|
||
<taxonomicName id="4C3CA096A3627F72FDCEF8388995DE9C" authorityName="Morawitz" authorityYear="1863" box="[574,713,1942,1964]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A3627F72FDCEF8388995DE9C" box="[574,713,1942,1964]" italics="true" pageId="15" pageNumber="316">Trichotichnus</emphasis>
|
||
</taxonomicName>
|
||
subgenus
|
||
<taxonomicName id="4C3CA096A3627F72FCC6F83888C4DE9C" authorityName="Jedlicka" authorityYear="1935" box="[822,920,1942,1964]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A3627F72FCC6F83888C4DE9C" box="[822,920,1942,1964]" italics="true" pageId="15" pageNumber="316">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
, photographs of habitus. 38,
|
||
<taxonomicName id="4C3CA096A3627F72FB34F8368E08DE9C" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1220,1364,1942,1965]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov.">
|
||
<emphasis id="B9480707A3627F72FB34F8368E08DE9C" box="[1220,1364,1942,1965]" italics="true" pageId="15" pageNumber="316">T. schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3627F72FAACF8398ECADE9C" box="[1372,1430,1943,1964]" pageId="15" pageNumber="316" rank="species">
|
||
<emphasis id="B9480707A3627F72FAACF8398ECADE9C" bold="true" box="[1372,1430,1943,1964]" pageId="15" pageNumber="316">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype; 39,
|
||
<taxonomicName id="4C3CA096A3627F72FED6F8188A89DEFB" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[294,469,1973,1995]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="obliquebasalis" status="sp. nov.">
|
||
<emphasis id="B9480707A3627F72FED6F8188A89DEFB" box="[294,469,1973,1995]" italics="true" pageId="15" pageNumber="316">T. obliquebasalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3627F72FE2DF8188944DEFB" box="[477,536,1974,1995]" pageId="15" pageNumber="316" rank="species">
|
||
<emphasis id="B9480707A3627F72FE2DF8188944DEFB" bold="true" box="[477,536,1974,1995]" pageId="15" pageNumber="316">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male; 40,
|
||
<taxonomicName id="4C3CA096A3627F72FD04F8188826DEFB" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[756,890,1973,1995]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="siklesensis" status="sp. nov.">
|
||
<emphasis id="B9480707A3627F72FD04F8188826DEFB" box="[756,890,1973,1995]" italics="true" pageId="15" pageNumber="316">T. siklesensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3627F72FC72F81888E1DEFB" box="[898,957,1974,1995]" pageId="15" pageNumber="316" rank="species">
|
||
<emphasis id="B9480707A3627F72FC72F81888E1DEFB" bold="true" box="[898,957,1974,1995]" pageId="15" pageNumber="316">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male; 41,
|
||
<taxonomicName id="4C3CA096A3627F72FB69F8188E0DDEFB" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1177,1361,1973,1995]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov.">
|
||
<emphasis id="B9480707A3627F72FB69F8188E0DDEFB" box="[1177,1361,1973,1995]" italics="true" pageId="15" pageNumber="316">T. panchhaseensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3627F72FAABF8188EC9DEFB" box="[1371,1429,1974,1995]" pageId="15" pageNumber="316" rank="species">
|
||
<emphasis id="B9480707A3627F72FAABF8188EC9DEFB" bold="true" box="[1371,1429,1974,1995]" pageId="15" pageNumber="316">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF438B9DA37D7F6DFF67FC6B8FCFDD29" httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="16" pageNumber="317" targetBox="[218,1372,202,935]" targetPageId="16">
|
||
<paragraph id="8B83DB15A37D7F6DFF67FC6B8FCFDD29" blockId="16.[151,1436,965,1049]" pageId="16" pageNumber="317">
|
||
<emphasis id="B9480707A37D7F6DFF67FC6B8A0FDAEB" bold="true" box="[151,339,965,988]" pageId="16" pageNumber="317">FIGURES 42–46.</emphasis>
|
||
<taxonomicName id="4C3CA096A37D7F6DFEA8FC688984DAEC" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[344,728,966,988]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFEA8FC688ABFDAEC" box="[344,483,966,988]" italics="true" pageId="16" pageNumber="317">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFE02FC688909DAEC" box="[498,597,966,988]" italics="true" pageId="16" pageNumber="317">Bottchrus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A37D7F6DFD92FC688984DAEC" box="[610,728,966,988]" italics="true" pageId="16" pageNumber="317">schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA37D7F6DFD2EFC69884ADAEC" box="[734,790,967,988]" pageId="16" pageNumber="317" rank="species">
|
||
<emphasis id="B9480707A37D7F6DFD2EFC69884ADAEC" bold="true" box="[734,790,967,988]" pageId="16" pageNumber="317">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(42–43, paratype; 44–46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42–43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42–44, 46; B: Fig. 45).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B83DB15A37D7F6DFF37FBEC8BA2DDDA" blockId="16.[151,1437,1089,2014]" pageId="16" pageNumber="317">
|
||
Legs: Protibia without longitudinal sulcus on upper surface. Profemur with four setae on anterio-ventral margin. Tarsi short, similar to those in
|
||
<taxonomicName id="4C3CA096A37D7F6DFDACFBC8884DDD4D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[604,785,1124,1149]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFDACFBC8892DDD4D" box="[604,625,1126,1149]" italics="true" pageId="16" pageNumber="317">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFD74FBC889C9DD4D" box="[644,661,1126,1149]" italics="true" pageId="16" pageNumber="317">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37D7F6DFD5FFBCA884DDD4D" box="[687,785,1124,1149]" italics="true" pageId="16" pageNumber="317">parvulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA37D7F6DFCEBFBCA8801DD4D" box="[795,861,1124,1149]" pageId="16" pageNumber="317" rank="species">
|
||
<emphasis id="B9480707A37D7F6DFCEBFBCA8801DD4D" bold="true" box="[795,861,1124,1149]" pageId="16" pageNumber="317">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, metatarsus noticeably shorter than HWmin, with metatarsomeres (particularly 2–4) markedly widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Tarsomere 5 with two pairs of ventro-lateral setae. In male, pro- and mesotarsi moderately widened.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A37D7F6DFF37FB5A88E9DCF2" blockId="16.[151,1437,1089,2014]" pageId="16" pageNumber="317">
|
||
Male genitalia: Median lobe of aedeagus (
|
||
<figureCitation id="1307C790A37D7F6DFD5DFB5A886CDC3D" box="[685,816,1268,1293]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="16" pageNumber="317">Figs 44–46</figureCitation>
|
||
) in lateral view rather evenly arcuate, C-shaped, with almost straight middle portion of ventral margin and with apical third curved ventrad; in dorsal view almost straight, narrowed to apex in distal third. Terminal lamella moderately short, narrowed to apex. Apical capitulum in lateral view (
|
||
<figureCitation id="1307C790A37D7F6DFED9FACF8A27DC49" box="[297,379,1377,1402]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="16" pageNumber="317">Fig. 44</figureCitation>
|
||
) very oblique, poorly delimited, almost not protruded ventrad and dorsad, narrowly rounded at tip; in caudal view (
|
||
<figureCitation id="1307C790A37D7F6DFE77FA2A8A80DCAD" box="[391,476,1412,1437]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="16" pageNumber="317">Fig. 45</figureCitation>
|
||
) triangular, somewhat horseshoe-shaped, rather acute, only slightly blunted at tip. Internal sac without distinct spiny patches and other sclerotic elements.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA37D7F6CFF37FA628805D870" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317" type="etymology">
|
||
<paragraph id="8B83DB15A37D7F6DFF37FA62894BDF3A" blockId="16.[151,1437,1089,2014]" pageId="16" pageNumber="317">
|
||
<emphasis id="B9480707A37D7F6DFF37FA628A1ADCD5" bold="true" box="[199,326,1484,1509]" pageId="16" pageNumber="317">Etymology</emphasis>
|
||
. This new species is named after the famous coleopterologist Wolfgang Schawaller (Stuttgart), who collected the
|
||
<typeStatus id="548765B7A37D7F6DFE93FA5C8ACFDF3A" box="[355,403,1522,1546]" pageId="16" pageNumber="317">type</typeStatus>
|
||
specimens.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A37D7F6CFF37F9BA8805D870" blockId="16.[151,1437,1089,2014]" lastBlockId="17.[151,1437,151,464]" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317">
|
||
<emphasis id="B9480707A37D7F6DFF37F9BA8A06DF1D" bold="true" box="[199,346,1556,1581]" pageId="16" pageNumber="317">Comparison</emphasis>
|
||
.
|
||
<taxonomicName id="4C3CA096A37D7F6DFE9DF9BA8991DF1D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[365,717,1556,1581]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFE9DF9BA895BDF1D" box="[365,519,1556,1581]" italics="true" pageId="16" pageNumber="317">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFDECF9B88971DF1D" box="[540,557,1558,1581]" italics="true" pageId="16" pageNumber="317">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37D7F6DFDBAF9BA8991DF1D" box="[586,717,1556,1581]" italics="true" pageId="16" pageNumber="317">schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA37D7F6DFD28F9BA8840DF1D" box="[728,796,1556,1581]" pageId="16" pageNumber="317" rank="species">
|
||
<emphasis id="B9480707A37D7F6DFD28F9BA8840DF1D" bold="true" box="[728,796,1556,1581]" pageId="16" pageNumber="317">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is very similar to
|
||
<taxonomicName id="4C3CA096A37D7F6DFC0EF9B88F91DF1D" authorityName="Kirschenhofer" authorityYear="1992" box="[1022,1229,1556,1581]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="holzschuhi" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFC0EF9B88F4EDF1D" box="[1022,1042,1558,1581]" italics="true" pageId="16" pageNumber="317">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFBD8F9B88F65DF1D" box="[1064,1081,1558,1581]" italics="true" pageId="16" pageNumber="317">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37D7F6DFBA5F9BA8F91DF1D" box="[1109,1229,1556,1581]" italics="true" pageId="16" pageNumber="317">holzschuhi</emphasis>
|
||
</taxonomicName>
|
||
and all the other preceding species in habitus and in having metepisternum short, protibia not sulcate dorsally, and aedeagal median lobe arcuate, with inclined apical capitulum, but easily recognized from them in the combination of the following characters: pronotum rather wide, moderately narrowed basad; dorsal microsculpture present throughout on head, pronotum and elytra; parascutellar striole rudimentary, almost absent; apex of last visible (VII) abdominal sternite in male (
|
||
<figureCitation id="1307C790A37D7F6DFF0BF9678A11DFD1" box="[251,333,1737,1762]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="16" pageNumber="317">Fig. 43</figureCitation>
|
||
) subtruncate and slightly concave just at tip [in this character,
|
||
<taxonomicName id="4C3CA096A37D7F6DFBE3F9648FB9DFD2" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1043,1253,1737,1762]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFBE3F9648F74DFD1" box="[1043,1064,1738,1761]" italics="true" pageId="16" pageNumber="317">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFBC8F9648F15DFD1" box="[1080,1097,1738,1761]" italics="true" pageId="16" pageNumber="317">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37D7F6DFB92F9678FB9DFD2" box="[1122,1253,1737,1762]" italics="true" pageId="16" pageNumber="317">schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA37D7F6DFB1CF9678E70DFD2" box="[1260,1324,1737,1762]" pageId="16" pageNumber="317" rank="species">
|
||
<emphasis id="B9480707A37D7F6DFB1CF9678E70DFD2" bold="true" box="[1260,1324,1737,1762]" pageId="16" pageNumber="317">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is similar only to
|
||
<taxonomicName id="4C3CA096A37D7F6DFF02F9408AF0DE36" authorityName="Andrewes" authorityYear="1930" box="[242,428,1773,1798]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFF02F9408A5BDE35" box="[242,263,1774,1797]" italics="true" pageId="16" pageNumber="317">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFEE9F9408A76DE35" box="[281,298,1774,1797]" italics="true" pageId="16" pageNumber="317">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37D7F6DFEB4F9438AF0DE36" box="[324,428,1773,1798]" italics="true" pageId="16" pageNumber="317">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
]; median lobe (
|
||
<figureCitation id="1307C790A37D7F6DFD96F94389B4DE36" box="[614,744,1773,1798]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="16" pageNumber="317">Figs 44–46</figureCitation>
|
||
) with shorter terminal lamella and very oblique, horseshoeshaped apical capitulum, which is poorly separated from the terminal lamella in lateral view and almost triangular [as in
|
||
<taxonomicName id="4C3CA096A37D7F6DFF2CF8988ACFDE7E" authorityName="Andrewes" authorityYear="1930" box="[220,403,1845,1870]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFF2CF8988BAEDE7D" box="[220,242,1846,1869]" italics="true" pageId="16" pageNumber="317">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFEF2F8988A4FDE7D" box="[258,275,1846,1869]" italics="true" pageId="16" pageNumber="317">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37D7F6DFEDBF89B8ACFDE7E" box="[299,403,1845,1870]" italics="true" pageId="16" pageNumber="317">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
], with rather acute, only slightly blunted tip, in caudal view (
|
||
<figureCitation id="1307C790A37D7F6DFBB8F89B8FC1DE7E" box="[1096,1181,1845,1870]" captionStart="FIGURES 42 – 46" captionStartId="16.[151,264,965,988]" captionTargetBox="[218,1372,202,935]" captionTargetId="figure@16.[200,1387,193,945]" captionTargetPageId="16" captionText="FIGURES 42 – 46. Trichotichnus (Bottchrus) schawalleri sp. n. (42 – 43, paratype; 44 – 46, holotype). 42, left metepisternum; 43, apex of last (VII) abdominal sternite, male; 44, 46, median lobe of aedeagus; 45, apical capitulum. 42 – 43, ventral view; 44, lateral view; 45, caudal view; 46, dorsal view. Scale bars = 0.25 mm (A: Figs 42 – 44, 46; B: Fig. 45)." httpUri="https://zenodo.org/record/919624/files/figure.png" pageId="16" pageNumber="317">Fig. 45</figureCitation>
|
||
); and internal sac with characteristic pattern of spiny folders in the middle portion of the median lobe.
|
||
<taxonomicName id="4C3CA096A37D7F6DFBCDF8F78EC9DE41" authorityName="Ito" authorityYear="1998" box="[1085,1429,1881,1906]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFBCDF8F78F8BDE42" box="[1085,1239,1881,1906]" italics="true" pageId="16" pageNumber="317">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFB1BF8F48FA0DE41" box="[1259,1276,1882,1905]" italics="true" pageId="16" pageNumber="317">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37D7F6DFAE7F8F48EC9DE41" box="[1303,1429,1882,1905]" italics="true" pageId="16" pageNumber="317">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
, probably occurring sympatricly with
|
||
<taxonomicName id="4C3CA096A37D7F6DFDCCF8D08852DEA6" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[572,782,1917,1942]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37D7F6DFDCCF8D0890DDEA5" box="[572,593,1918,1941]" italics="true" pageId="16" pageNumber="317">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37D7F6DFD91F8D0892EDEA5" box="[609,626,1918,1941]" italics="true" pageId="16" pageNumber="317">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37D7F6DFD7BF8D38852DEA6" box="[651,782,1917,1942]" italics="true" pageId="16" pageNumber="317">schawalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA37D7F6DFCE5F8D3880ADEA6" box="[789,854,1917,1942]" pageId="16" pageNumber="317" rank="species">
|
||
<emphasis id="B9480707A37D7F6DFCE5F8D3880ADEA6" bold="true" box="[789,854,1917,1942]" pageId="16" pageNumber="317">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, is somewhat similar to the new species in habitus, but easily distinguished by having larger size (
|
||
<quantity id="4CC476F0A37D7F6DFD4BF80F8810DE89" box="[699,844,1953,1978]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="6.9" metricValueMin="6.1" pageId="16" pageNumber="317" unit="mm" value="6.5" valueMax="6.9" valueMin="6.1">6.1–6.9 mm</quantity>
|
||
), metepisternum (
|
||
<figureCitation id="1307C790A37D7F6DFBEDF80F8F23DE8A" box="[1053,1151,1953,1978]" captionStart="FIGURES 115 – 119" captionStartId="40.[151,264,1519,1542]" captionTargetBox="[210,1376,415,1485]" captionTargetId="figure@40.[192,1394,411,1498]" captionTargetPageId="40" captionText="FIGURES 115 – 119. Trichotichnus (Bottchrus) cyanescens Ito, 1998 (specimen from Dhabeli). 115, left metepisternum; 116, apex of last (VII) abdominal sternite, male; 117, 119, median lobe of aedeagus; 118, apical capitulum. 115 – 116, ventral view; 117, lateral view; 118, caudal view; 119, dorsal view. Scale bars = 0.25 mm (A: Figs 115 – 117, 119; B: Fig. 118)." httpUri="https://zenodo.org/record/919668/files/figure.png" pageId="16" pageNumber="317">Fig. 115</figureCitation>
|
||
) slightly more elongate, mentum and submentum fused only laterally, apex of last (VII) abdominal sternite (
|
||
<figureCitation id="1307C790A37D7F6DFBB1F86B8FFCDEEE" box="[1089,1184,1989,2014]" captionStart="FIGURES 115 – 119" captionStartId="40.[151,264,1519,1542]" captionTargetBox="[210,1376,415,1485]" captionTargetId="figure@40.[192,1394,411,1498]" captionTargetPageId="40" captionText="FIGURES 115 – 119. Trichotichnus (Bottchrus) cyanescens Ito, 1998 (specimen from Dhabeli). 115, left metepisternum; 116, apex of last (VII) abdominal sternite, male; 117, 119, median lobe of aedeagus; 118, apical capitulum. 115 – 116, ventral view; 117, lateral view; 118, caudal view; 119, dorsal view. Scale bars = 0.25 mm (A: Figs 115 – 117, 119; B: Fig. 118)." httpUri="https://zenodo.org/record/919668/files/figure.png" pageId="16" pageNumber="317">Fig. 116</figureCitation>
|
||
) in male rounded, and median lobe of aedeagus (
|
||
<figureCitation id="1307C790A37C7F6CFE35FF39893ED980" box="[453,610,151,176]" captionStart="FIGURES 115 – 119" captionStartId="40.[151,264,1519,1542]" captionTargetBox="[210,1376,415,1485]" captionTargetId="figure@40.[192,1394,411,1498]" captionTargetPageId="40" captionText="FIGURES 115 – 119. Trichotichnus (Bottchrus) cyanescens Ito, 1998 (specimen from Dhabeli). 115, left metepisternum; 116, apex of last (VII) abdominal sternite, male; 117, 119, median lobe of aedeagus; 118, apical capitulum. 115 – 116, ventral view; 117, lateral view; 118, caudal view; 119, dorsal view. Scale bars = 0.25 mm (A: Figs 115 – 117, 119; B: Fig. 118)." httpUri="https://zenodo.org/record/919668/files/figure.png" pageId="17" pageNumber="318">Figs 117–119</figureCitation>
|
||
) S-shaped, hooked at apex.
|
||
<taxonomicName id="4C3CA096A37C7F6CFC59FF398FB5D980" authorityName="Andrewes" authorityYear="1930" box="[937,1257,151,176]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37C7F6CFC59FF398F1FD980" box="[937,1091,151,176]" italics="true" pageId="17" pageNumber="318">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37C7F6CFBA6FF378F3BD980" box="[1110,1127,153,176]" italics="true" pageId="17" pageNumber="318">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37C7F6CFB71FF398FB5D980" box="[1153,1257,151,176]" italics="true" pageId="17" pageNumber="318">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
, also occurring sympatricly with this new species, is easily distinguished from in having larger size (
|
||
<quantity id="4CC476F0A37C7F6CFBBDFF128F85D9E4" box="[1101,1241,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.7" metricValueMin="6.8" pageId="17" pageNumber="318" unit="mm" value="7.25" valueMax="7.7" valueMin="6.8">6.8–7.7 mm</quantity>
|
||
), genae glabrous, metepisterna (
|
||
<figureCitation id="1307C790A37C7F6CFEC7FF718AC5D9C8" box="[311,409,223,248]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="17" pageNumber="318">Fig. 137</figureCitation>
|
||
) markedly longer, pronotum never sinuate basally and with narrow, apically not widened lateral gutter, protibia longitudinally sulcate on dorsal side, and apical capitulum of the aedeagal median lobe (
|
||
<figureCitation id="1307C790A37C7F6CFA9AFEAA8BA7D870" captionStart="FIGURES 135 – 136" captionStartId="46.[151,264,1244,1267]" captionTargetBox="[301,1277,456,1212]" captionTargetId="figure@46.[271,1316,445,1223]" captionTargetPageId="46" captionText="FIGURES 135 – 136. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930, median lobe of aedeagus, lateral view (135, lectotype; 136, holotype of T. curvatus Ito, 1996). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/919684/files/figure.png" pageId="17" pageNumber="318">Figs 135–136</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A37C7F6CFEF7FE868A6ED870" box="[263,306,296,320]" captionStart="FIGURES 137 – 145" captionStartId="47.[151,264,1705,1728]" captionTargetBox="[194,1406,318,1664]" captionTargetId="figure@47.[151,1436,303,1684]" captionTargetPageId="47" captionText="FIGURES 137 – 145. Trichotichnus (Bottchrus) hingstoni Andrewes, 1930 (137 – 139, Kechu Pheri Lake, W-Sikkim; 140 – 143, Maney Bhanjyang, Darjeeling, 144, above Yamputhin, Nepal; 145, Ting Sang La, Nepal). 137, left metepisternum; 138, apex of last (VII) abdominal sternite, male; 139, same, female; 140, 142, median lobe of aedeagus; 141, apical capitulum; 143 – 145, middle portion of median lobe. 137 – 139, ventral view; 140, left lateral view; 141, caudal view; 142, dorsal view; 143 – 145, right lateral view. Scale bars = 0.25 mm (A: Figs 137 – 140, 142 – 145; B: Fig. 141)." httpUri="https://zenodo.org/record/919688/files/figure.png" pageId="17" pageNumber="318">140</figureCitation>
|
||
) more strongly prominent dorsally and ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA37C7F6CFF37FEE289E0D8E0" pageId="17" pageNumber="318" type="distribution">
|
||
<paragraph id="8B83DB15A37C7F6CFF37FEE289E0D8E0" blockId="17.[151,1437,151,464]" pageId="17" pageNumber="318">
|
||
<emphasis id="B9480707A37C7F6CFF37FEE28A05D855" bold="true" box="[199,345,332,357]" pageId="17" pageNumber="318">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1307C790A37C7F6CFE9AFEE28A97D855" box="[362,459,332,357]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="17" pageNumber="318">Fig. 151</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A37C7F6CFE29FEE38ABBD854" box="[473,487,333,356]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="17" pageNumber="318">
|
||
<emphasis id="B9480707A37C7F6CFE29FEE38ABBD854" box="[473,487,333,356]" italics="true" pageId="17" pageNumber="318">5</emphasis>
|
||
</figureCitation>
|
||
). Up to today only known from the western slope of the middle Arun River Valley towards Salpa pass, Bhojpur District, East
|
||
<collectingCountry id="F32B9B85A37C7F6CFD8CFEC1899ED8B8" box="[636,706,367,392]" name="Nepal" pageId="17" pageNumber="318">Nepal</collectingCountry>
|
||
, at altitudes of approximately
|
||
<quantity id="4CC476F0A37C7F6CFBEAFEDE8FE5D8B7" box="[1050,1209,368,392]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="17" pageNumber="318" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">1900–2100 m</quantity>
|
||
. This species seems to occur sympatricly with
|
||
<taxonomicName id="4C3CA096A37C7F6CFE4AFE3B8932D89D" authorityName="Andrewes" authorityYear="1930" box="[442,622,404,429]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="hingstoni" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37C7F6CFE4AFE3B8A93D89C" box="[442,463,405,428]" italics="true" pageId="17" pageNumber="318">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37C7F6CFE2EFE3B8AB3D89C" box="[478,495,405,428]" italics="true" pageId="17" pageNumber="318">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37C7F6CFDF6FE3A8932D89D" box="[518,622,404,429]" italics="true" pageId="17" pageNumber="318">hingstoni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1307C790A37C7F6CFD8DFE3A8980D89D" box="[637,732,404,429]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="17" pageNumber="318">Fig. 152</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A37C7F6CFD18FE3A89AAD89D" box="[744,758,404,429]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="17" pageNumber="318">
|
||
<emphasis id="B9480707A37C7F6CFD18FE3A89AAD89D" box="[744,758,404,429]" italics="true" pageId="17" pageNumber="318">6</emphasis>
|
||
</figureCitation>
|
||
). Another species,
|
||
<taxonomicName id="4C3CA096A37C7F6CFC3DFE3B8FCBD89C" authorityName="Ito" authorityYear="1998" box="[973,1175,404,428]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A37C7F6CFC3DFE3B88BED89C" box="[973,994,405,428]" italics="true" pageId="17" pageNumber="318">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A37C7F6CFC01FE3B8F5ED89C" box="[1009,1026,405,428]" italics="true" pageId="17" pageNumber="318">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A37C7F6CFBE9FE3B8FCBD89C" box="[1049,1175,405,428]" italics="true" pageId="17" pageNumber="318">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
, was found in the Arun River Valley slightly further north (
|
||
<figureCitation id="1307C790A37C7F6CFDD3FE1989DFD8E0" box="[547,643,439,464]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="17" pageNumber="318">Fig. 151</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A37C7F6CFD7FFE1689F7D8E0" box="[655,683,440,464]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="17" pageNumber="318">
|
||
<emphasis id="B9480707A37C7F6CFD7FFE1689F7D8E0" box="[655,683,440,464]" italics="true" pageId="17" pageNumber="318">12</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |