364 lines
52 KiB
XML
364 lines
52 KiB
XML
<document id="1D984AC9DF51964A87778F198A944D3E" ID-CLB-Dataset="32021" ID-DOI="10.11646/zootaxa.4323.3.1" ID-GBIF-Dataset="f1be4566-adeb-4995-83c7-2aebab3c7458" ID-ISSN="1175-5326" ID-Zenodo-Dep="919606" ID-ZooBank="3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506057765384" checkinUser="plazi" docAuthor="Schmidt, Joachim" docDate="2017" docId="03956A03A3497F5BFF67FAB68908DA24" docLanguage="en" docName="zootaxa.4323.3.1.pdf" docOrigin="Zootaxa 4323 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus (Bottchrus) manasluensis Schmidt 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="339" masterDocId="FFAC127BA36D7F7DFFF0FFAE8B5CD930" masterDocTitle="Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species" masterLastPageNumber="358" masterPageNumber="301" pageNumber="337" updateTime="1698468866275" updateUser="ExternalLinkService">
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<mods:title id="D4E5B8AB54BF46AF63702D328B3A69F1">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<mods:namePart id="B1231D3411D22A860E6B078B50007424">Schmidt, Joachim</mods:namePart>
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<mods:title id="83C79D94FA360A8B5C99F5F5BCCB5C6C">Zootaxa</mods:title>
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<mods:part id="760EE1E7F4C7D14B095260E5743494ED">
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<mods:date id="C19EDA290982048B653C576E01E6351E">2017</mods:date>
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<mods:number id="726D9AC7751D705A7B48F8C834D08D55">2017-09-22</mods:number>
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<mods:number id="94EA684A0B5D81CAAC6FD4446635D72B">4323</mods:number>
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<mods:identifier id="5066B3ECE32F81C498FA5384620EF3FB" type="DOI">10.11646/zootaxa.4323.3.1</mods:identifier>
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<treatment id="03956A03A3497F5BFF67FAB68908DA24" ID-DOI="http://doi.org/10.5281/zenodo.6015765" ID-GBIF-Taxon="134839428" ID-Zenodo-Dep="6015765" LSID="urn:lsid:plazi:treatment:03956A03A3497F5BFF67FAB68908DA24" httpUri="http://treatment.plazi.org/id/03956A03A3497F5BFF67FAB68908DA24" lastPageId="38" lastPageNumber="339" pageId="36" pageNumber="337">
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<subSubSection id="C326889EA3497F59FF67FAB68AE3DC63" pageId="36" pageNumber="337" type="nomenclature">
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<paragraph id="8B83DB15A3497F59FF67FAB68994DC02" blockId="36.[151,712,1304,1363]" box="[151,712,1304,1330]" pageId="36" pageNumber="337">
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<heading id="D0CB6C79A3497F59FF67FAB68994DC02" bold="true" box="[151,712,1304,1330]" fontSize="11" level="1" pageId="36" pageNumber="337" reason="1">
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<emphasis id="B9480707A3497F59FF67FAB68994DC02" bold="true" box="[151,712,1304,1330]" pageId="36" pageNumber="337">
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<taxonomicName id="4C3CA096A3497F59FF67FAB68923DC02" ID-CoL="58LH8" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,639,1304,1330]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov." subGenus="Bottchrus">
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<emphasis id="B9480707A3497F59FF67FAB68A1DDC02" bold="true" box="[151,321,1304,1330]" italics="true" pageId="36" pageNumber="337">Trichotichnus</emphasis>
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(
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<emphasis id="B9480707A3497F59FEA2FAB68A95DC02" bold="true" box="[338,457,1304,1330]" italics="true" pageId="36" pageNumber="337">Bottchrus</emphasis>
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)
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<emphasis id="B9480707A3497F59FE2AFAB68923DC02" bold="true" box="[474,639,1304,1330]" italics="true" pageId="36" pageNumber="337">manasluensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27BBA7CA3497F59FD75FAB68994DC02" box="[645,712,1304,1330]" pageId="36" pageNumber="337" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B83DB15A3497F59FF67FA948AE3DC63" blockId="36.[151,712,1304,1363]" box="[151,447,1338,1363]" pageId="36" pageNumber="337">
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(
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<figureCitation id="1307C790A3497F59FF6FFA948BA9DC63" box="[159,245,1338,1363]" captionStart="FIGURES 91 – 94" captionStartId="32.[151,264,1940,1963]" captionTargetBox="[265,1303,196,1917]" captionTargetId="figure@32.[261,1305,193,1921]" captionTargetPageId="32" captionText="FIGURES 91 – 94. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 91, T. gupchiensis sp. n., paratype, male. 92, T. brancuccii sp. n., paratype, male. 93, T. manasluensis sp. n., paratype, male. 94, T. cyanescens Ito, 1998, male from Dhabeli." httpUri="https://zenodo.org/record/919654/files/figure.png" pageId="36" pageNumber="337">Figs 93</figureCitation>
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,
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<figureCitation id="1307C790A3497F59FEF1FA958A39DC63" box="[257,357,1339,1363]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="36" pageNumber="337">109–114</figureCitation>
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,
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<figureCitation id="1307C790A3497F59FE80FA948AC0DC63" box="[368,412,1338,1363]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="36" pageNumber="337">152</figureCitation>
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,
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<emphasis id="B9480707A3497F59FE58FA958AEADC63" box="[424,438,1339,1363]" italics="true" pageId="36" pageNumber="337">3</emphasis>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C326889EA3497F59FF67FA2C8872DCD3" pageId="36" pageNumber="337" type="materials_examined">
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<paragraph id="8B83DB15A3497F59FF67FA2C8867DC8F" blockId="36.[151,1437,1410,2011]" pageId="36" pageNumber="337">
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<materialsCitation id="3B54D148A3497F59FF67FA2C8867DC8F" ID-GBIF-Occurrence="1655986729" collectingDate="2005-04-29" collectorName="J. Schmidt" country="Nepal" elevation="1700" latitude="28.241112" location="Manaslu Mts." longLatPrecision="20" longitude="84.40889" pageId="36" pageNumber="337" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B9480707A3497F59FF67FA2C8A62DCAB" bold="true" box="[151,318,1410,1435]" pageId="36" pageNumber="337">
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<typeStatus id="548765B7A3497F59FF67FA2C8B8EDCAB" box="[151,210,1410,1435]" pageId="36" pageNumber="337">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="548765B7A3497F59FEBCFA2C8AE5DCAB" box="[332,441,1410,1435]" pageId="36" pageNumber="337" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D3A109CA3497F59FE33FA2D8A86DCAA" box="[451,474,1411,1434]" pageId="36" pageNumber="337" type="male">♂</specimenCount>
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, “
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<collectingCountry id="F32B9B85A3497F59FE1FFA2D8916DCAB" box="[495,586,1411,1435]" name="Nepal" pageId="36" pageNumber="337">NEPAL</collectingCountry>
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<location id="8EE38DCEA3497F59FDA1FA2C89B0DCAA" LSID="urn:lsid:plazi:treatment:03956A03A3497F5BFF67FAB68908DA24:8EE38DCEA3497F59FDA1FA2C89B0DCAA" box="[593,748,1410,1435]" country="Nepal" latitude="28.241112" longLatPrecision="20" longitude="84.40889" name="Manaslu Mts." pageId="36" pageNumber="337">Manaslu Mts.</location>
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<elevation id="00113C26A3497F59FD03FA2C88CCDCAA" box="[755,912,1410,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="36" pageNumber="337" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
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<quantity id="4CC476F0A3497F59FD03FA2C88CCDCAA" box="[755,912,1410,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="36" pageNumber="337" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">1600–1800 m</quantity>
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</elevation>
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, Bara Pokhari Lekh above Bhachok Gaon vill.
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<collectingDate id="EFC6043DA3497F59FF67FA088A55DC8F" box="[151,265,1446,1471]" pageId="36" pageNumber="337" value="2005-04-29">
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<date id="FF82FDD5A3497F59FF67FA088A55DC8F" box="[151,265,1446,1471]" pageId="36" pageNumber="337" value="2005-04-29">29.4.2005</date>
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</collectingDate>
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, leg
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<collectorName id="26C9BEC3A3497F59FECEFA088AF1DC8F" box="[318,429,1446,1471]" pageId="36" pageNumber="337">J.Schmidt</collectorName>
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ca
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<geoCoordinate id="EE08BDD2A3497F59FE24FA098907DC8F" box="[468,603,1447,1471]" direction="north" orientation="latitude" pageId="36" pageNumber="337" precision="15" value="28.241112">28°14'28"N</geoCoordinate>
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,
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<geoCoordinate id="EE08BDD2A3497F59FD96FA0989B5DC8F" box="[614,745,1447,1471]" direction="east" orientation="longitude" pageId="36" pageNumber="337" precision="15" value="84.40889">84°24'32"E</geoCoordinate>
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” (cJS).
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8B83DB15A3497F59FF37FA658872DCD3" blockId="36.[151,1437,1410,2011]" box="[199,814,1482,1507]" pageId="36" pageNumber="337">
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<materialsCitation id="3B54D148A3497F59FF37FA658872DCD3" ID-GBIF-Occurrence="1655986746" box="[199,814,1482,1507]" collectingDate="2005-04-29" collectionCode="ZIN" collectorName="J. Schmidt" country="Nepal" elevation="1700" latitude="28.241112" location="Manaslu Mts." longLatPrecision="20" longitude="84.40889" pageId="36" pageNumber="337" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="548765B7A3497F59FF37FA658A65DCD3" box="[199,313,1483,1507]" pageId="36" pageNumber="337" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D3A109CA3497F59FEB3FA658A32DCD2" box="[323,366,1483,1507]" pageId="36" pageNumber="337" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="9D3A109CA3497F59FE8BFA658AF8DCD2" box="[379,420,1483,1507]" pageId="36" pageNumber="337" type="female">2 ♀</specimenCount>
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, same data as holotype (cJS,
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<collectionCode id="ED2D43D0A3497F59FD1EFA65887DDCD3" box="[750,801,1483,1507]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="36" pageNumber="337">ZIN</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C326889EA3497F58FF37FA408FC9DF60" lastPageId="37" lastPageNumber="338" pageId="36" pageNumber="337" type="description">
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<paragraph id="8B83DB15A3497F59FF37FA408F0CDF36" blockId="36.[151,1437,1410,2011]" box="[199,1104,1518,1543]" pageId="36" pageNumber="337">
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<emphasis id="B9480707A3497F59FF37FA408A0DDF37" bold="true" box="[199,337,1518,1543]" pageId="36" pageNumber="337">Description</emphasis>
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. Habitus as in
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<figureCitation id="1307C790A3497F59FE08FA408917DF37" box="[504,587,1518,1543]" captionStart="FIGURES 91 – 94" captionStartId="32.[151,264,1940,1963]" captionTargetBox="[265,1303,196,1917]" captionTargetId="figure@32.[261,1305,193,1921]" captionTargetPageId="32" captionText="FIGURES 91 – 94. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 91, T. gupchiensis sp. n., paratype, male. 92, T. brancuccii sp. n., paratype, male. 93, T. manasluensis sp. n., paratype, male. 94, T. cyanescens Ito, 1998, male from Dhabeli." httpUri="https://zenodo.org/record/919654/files/figure.png" pageId="36" pageNumber="337">Fig. 93</figureCitation>
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. Body length
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<quantity id="4CC476F0A3497F59FD17FA408833DF36" box="[743,879,1518,1543]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.3" metricValueMin="5.9" pageId="36" pageNumber="337" unit="mm" value="6.1" valueMax="6.3" valueMin="5.9">5.9–6.3 mm</quantity>
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, width
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<quantity id="4CC476F0A3497F59FC32FA408F10DF36" box="[962,1100,1518,1543]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="36" pageNumber="337" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
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.
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</paragraph>
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<paragraph id="8B83DB15A3497F59FF37F9BC8F4CDF43" blockId="36.[151,1437,1410,2011]" pageId="36" pageNumber="337">Colour: Body piceous, shiny on dorsum, with more or less distinct greenish tinge on pronotum and elytra; labrum, in most specimens also base of mandibles, very narrow lateral margins of pronotum and apical portion of elytral margins paler, reddish brown; palpi, antennae and legs brownish yellow.</paragraph>
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<paragraph id="8B83DB15A3497F58FF37F9D08A90D82D" blockId="36.[151,1437,1410,2011]" lastBlockId="37.[151,1437,151,2013]" lastPageId="37" lastPageNumber="338" pageId="36" pageNumber="337">
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Head: Comparatively large for subgenus, with wide neck (in males, HWmax/PWmax = 0.66–0.67, HWmin/ PWmax = 0.52–0.54; in females, these indices 0.67–0.68 and 0.55–0.56, respectively). Eyes convex (in males, HWmax/HWmin = 1.24–1.25, and in females, 1.21–1.22), oval in lateral view. Tempora at most third as long as eye, slightly convex, somewhat abrupt to neck. Genae wide, about 1.5–1.7 as wide as width of antennomere 1, with few short setae. Lateral margin of head between antenna and eye slightly protruded laterally, forming an obtuse angle near eyes, distinctly bordered. Area between supraorbital furrow and upper margin of eye slightly narrower than width of antennomere 2 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow by distance slightly shorter than width of antennomere 3 apically. Labrum slightly concave anteriorly. Clypeus slightly emarginate along anterior margin, moderately convex basally, flatly sloped to apex, with one setigerous pore at each apical angle. Frons and vertex somewhat convex.
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<collectingRegion id="49F815F7A3497F59FAFCF86D8E37DEEA" box="[1292,1387,1987,2010]" country="Myanmar" name="Chin" pageId="36" pageNumber="337">Mentum</collectingRegion>
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and submentum completely fused, submentum on each side with one long seta and one short seta laterally of long seta. Left mandible not truncate, slightly blunted at apex. Dorsal microsculpture not evident even at high magnifications (100x). Antennae in both sexes extending approximately to the basal edge of pronotum, with antennomeres 4–8 markedly longer than wide.
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</paragraph>
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<paragraph id="8B83DB15A3487F58FF37FE86891ADBC0" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="338">Pronotum: Moderately transverse (PWmax/PL = 1.40–1.43), widest slightly before the end of anterior third, moderately narrowed posteriad (PWmax/PWmin = 1.18–1.20), with one lateral seta inserted slightly before widest point. Sides rounded (more widely in basal third) and slightly sinuate near basal angles. Apical margin slightly emarginate, bordered throughout. Apical angles more or less widely rounded, not or only slightly protruded anteriad. Basal margin nearly straight, slightly oblique laterally, vaguely bordered almost throughout, somewhat longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles obtuse, each with sharp denticulate or subdenticulate apex. Pronotal disc convex, strongly sloped to apical angles and moderately so to basal ones, not depressed basally. Lateral gutter very narrow almost throughout, only hardly widened apically. Median line fine, superficial, ending before apical and basal margins. Lateral depressions absent; basal foveae very small and shallow, poorly delimited, or absent. Surface comparatively coarsely and irregularly punctate along base and in latero-basal area; at anterior margin either impunctate or with few vague punctures. Microsculpture not visible even at high magnification (100x), occasionally present at apical angles, consisting of very fine obliterate transverse meshes.</paragraph>
|
||
<paragraph id="8B83DB15A3487F58FF37FD5289B4DD05" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="338">Elytra: In lateral and caudal view convex, in dorsal view oval, elongate, moderately wide (in males, EL/EW = 1.40–1.41; EL/PL = 2.40–2.44; EW/PWmax = 1.22–1.25; in females, these indices 1.35–1.36, 2.37–2.40, 1.23– 1.25, respectively), widest slightly behind middle, with somewhat acute apex. Humeri prominent, angularly rounded at tip, with a tiny acute denticle visible from behind. Subapical sinuation distinct, but not deep. Sutural angle acute, sharp or slightly blunted at tip. Basal edge slightly sinuate, forming an obtuse angle with lateral margin. Striae smooth, slightly impressed along entire length. Parascutellar setigerous pore present, large. Parascutellar striole rudimentary. Intervals slightly convex, interval 3 with a discal setigerous pore at stria 2 at or behind middle. Marginal umbilicate series widely interrupted at middle, with 6 pores in anterior group and 8–9 pores in posterior group. Microsculpture not evident.</paragraph>
|
||
<paragraph id="8B83DB15A3487F58FF37FB91890FDD68" blockId="37.[151,1437,151,2013]" box="[199,595,1087,1112]" pageId="37" pageNumber="338">Hindwings: Reduced to tiny scales.</paragraph>
|
||
<paragraph id="8B83DB15A3487F58FF37FBCA8962DDF4" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="338">
|
||
Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum (
|
||
<figureCitation id="1307C790A3487F58FBA7FBCA8FEBDD4C" box="[1111,1207,1124,1149]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="37" pageNumber="338">Fig. 109</figureCitation>
|
||
) slightly wider than long, moderately narrowed posteriad. Apex of last visible (VII) abdominal sternite in male rounded (
|
||
<figureCitation id="1307C790A3487F58FAFFFB298E2DDD90" box="[1295,1393,1159,1184]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="37" pageNumber="338">Fig. 110</figureCitation>
|
||
), in female somewhat angulate (
|
||
<figureCitation id="1307C790A3487F58FE21FB02896DDDF4" box="[465,561,1196,1221]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="37" pageNumber="338">Fig. 111</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3487F58FF37FB7E88D4DC65" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="338">Legs: Protibia sulcate on upper surface. Profemur with three to five setae on anterio-ventral margin. Tarsi short, metatarsus noticeably shorter than HWmin, with metatarsomeres (particularly 2–4) moderately widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Tarsomere 5 with two pairs of ventrolateral setae. In male, pro- and mesotarsi very moderately widened.</paragraph>
|
||
<paragraph id="8B83DB15A3487F58FF37FAF1893FDCF0" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="338">Female genitalia: Laterotergite (hemisternite) symmetrical, with two thick setae apically. Basal stylomere with one thick preapical spine on external margin. Apical stylomere moderately curved, with a peg-like spine at both ventral and dorsal edges of outer margin.</paragraph>
|
||
<paragraph id="8B83DB15A3487F58FF37FA628FC9DF60" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="338">
|
||
Male genitalia: Median lobe of aedeagus (
|
||
<figureCitation id="1307C790A3487F58FD6AFA628868DCD4" box="[666,820,1484,1509]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="37" pageNumber="338">Figs 112–114</figureCitation>
|
||
) relatively large, in lateral view very slightly S-shaped, strongly curved at apex ventrad forming a hook with acute tip; ventral margin convex; in dorsal view median lobe almost straight, evenly narrowed to apex in distal third. Terminal lamella in dorsal and caudal views short and narrow, not sinuate at sides (
|
||
<figureCitation id="1307C790A3487F58FE25F999892FDF60" box="[469,627,1591,1616]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="37" pageNumber="338">Figs 113–114</figureCitation>
|
||
). Internal sac without distinct sclerotic elements.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA3487F5BFF37F9F28BB1DB98" lastPageId="38" lastPageNumber="339" pageId="37" pageNumber="338" type="etymology">
|
||
<paragraph id="8B83DB15A3487F58FF37F9F28F84DF45" blockId="37.[151,1437,151,2013]" box="[199,1240,1628,1653]" pageId="37" pageNumber="338">
|
||
<emphasis id="B9480707A3487F58FF37F9F28A1ADF45" bold="true" box="[199,326,1628,1653]" pageId="37" pageNumber="338">Etymology</emphasis>
|
||
. The specific name refers to the Manaslu Himal where the
|
||
<typeStatus id="548765B7A3487F58FC2EF9F38F52DF45" box="[990,1038,1629,1653]" pageId="37" pageNumber="338">type</typeStatus>
|
||
series was found.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3487F5BFF37F9D18BB1DB98" blockId="37.[151,1437,151,2013]" lastBlockId="38.[151,1436,151,789]" lastPageId="38" lastPageNumber="339" pageId="37" pageNumber="338">
|
||
<emphasis id="B9480707A3487F58FF37F9D18A06DFA8" bold="true" box="[199,346,1663,1688]" pageId="37" pageNumber="338">Comparison</emphasis>
|
||
. This new species is very similar to
|
||
<taxonomicName id="4C3CA096A3487F58FD1DF92F88EBDFA8" authorityName="Ito" authorityYear="1998" box="[749,951,1664,1688]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3487F58FD1DF92F885EDFA8" box="[749,770,1665,1688]" italics="true" pageId="37" pageNumber="338">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3487F58FCE1F92F887EDFA8" box="[785,802,1665,1688]" italics="true" pageId="37" pageNumber="338">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3487F58FCC9F92F88EBDFA8" box="[825,951,1665,1688]" italics="true" pageId="37" pageNumber="338">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
from East
|
||
<collectingCountry id="F32B9B85A3487F58FBC1F9D18F28DFA8" box="[1073,1140,1663,1688]" name="Nepal" pageId="37" pageNumber="338">Nepal</collectingCountry>
|
||
in general habitus, femora not infuscate (similar in colour to tibiae), protibia finely sulcate, and pronotal lateral gutter scarcely widened anteriad, but readily distinguished from the latter species by slightly smaller size (
|
||
<quantity id="4CC476F0A3487F58FBC5F9698FE2DFEF" box="[1077,1214,1735,1760]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.3" metricValueMin="5.9" pageId="37" pageNumber="338" unit="mm" value="6.1" valueMax="6.3" valueMin="5.9">5.9–6.3 mm</quantity>
|
||
versus
|
||
<quantity id="4CC476F0A3487F58FAE5F9698EC1DFEF" box="[1301,1437,1735,1760]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.2" metricValueMin="6.6" pageId="37" pageNumber="338" unit="mm" value="6.9" valueMax="7.2" valueMin="6.6">6.6–7.2 mm</quantity>
|
||
in the examined specimens and
|
||
<quantity id="4CC476F0A3487F58FDE6F94289F8DE34" box="[534,676,1772,1797]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="6.7" metricValueMin="6.1" pageId="37" pageNumber="338" unit="mm" value="6.4" valueMax="6.7" valueMin="6.1">6.1–6.7 mm</quantity>
|
||
, according to the original description), mentum and submentum completely fused [in
|
||
<taxonomicName id="4C3CA096A3487F58FE77F8BF8908DE18" authorityName="Ito" authorityYear="1998" box="[391,596,1808,1832]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3487F58FE77F8BF8AC0DE18" box="[391,412,1809,1832]" italics="true" pageId="37" pageNumber="338">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3487F58FE5DF8BF8AE2DE18" box="[429,446,1809,1832]" italics="true" pageId="37" pageNumber="338">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3487F58FE26F8BF8908DE18" box="[470,596,1809,1832]" italics="true" pageId="37" pageNumber="338">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
, fused laterally and separated by suture medially], metepisternum slightly wider and less markedly narrowed posteriad, humerus with a tiny denticle at tip [in
|
||
<taxonomicName id="4C3CA096A3487F58FB95F89B8E6ADE7C" authorityName="Ito" authorityYear="1998" box="[1125,1334,1844,1868]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3487F58FB95F89B8F26DE7C" box="[1125,1146,1845,1868]" italics="true" pageId="37" pageNumber="338">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3487F58FB7CF89B8FC1DE7C" box="[1164,1181,1845,1868]" italics="true" pageId="37" pageNumber="338">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3487F58FB48F89B8E6ADE7C" box="[1208,1334,1845,1868]" italics="true" pageId="37" pageNumber="338">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
, without denticle], apex of last (VII) abdominal sternite narrowly rounded in male and angularly rounded in female [in
|
||
<taxonomicName id="4C3CA096A3487F58FA77F8F78A1BDEA4" authorityName="Ito" authorityYear="1998" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3487F58FA77F8F78EC1DE40" box="[1415,1437,1881,1904]" italics="true" pageId="37" pageNumber="338">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3487F58FF50F8D38BEDDEA4" box="[160,177,1917,1940]" italics="true" pageId="37" pageNumber="338">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3487F58FF39F8D38A1BDEA4" box="[201,327,1917,1940]" italics="true" pageId="37" pageNumber="338">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
, more widely rounded in both sexes], and microsculpture not evident on pronotal and elytral discs [in
|
||
<taxonomicName id="4C3CA096A3487F58FF4DF80F8AD6DE88" authorityName="Ito" authorityYear="1998" box="[189,394,1952,1976]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3487F58FF4DF80F8B8EDE88" box="[189,210,1953,1976]" italics="true" pageId="37" pageNumber="338">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3487F58FF13F80F8BA8DE88" box="[227,244,1953,1976]" italics="true" pageId="37" pageNumber="338">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3487F58FEFCF80F8AD6DE88" box="[268,394,1953,1976]" italics="true" pageId="37" pageNumber="338">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
, meshes visible throughout]. The median lobe of the aedeagus of
|
||
<taxonomicName id="4C3CA096A3487F58FB80F80F8E09DE89" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1136,1365,1952,1977]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3487F58FB80F80F8FD9DE88" box="[1136,1157,1953,1976]" italics="true" pageId="37" pageNumber="338">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3487F58FB65F80F8FFADE88" box="[1173,1190,1953,1976]" italics="true" pageId="37" pageNumber="338">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3487F58FB4EF80E8E09DE89" box="[1214,1365,1952,1977]" italics="true" pageId="37" pageNumber="338">manasluensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3487F58FAACF80E8EC0DE89" box="[1372,1436,1952,1977]" pageId="37" pageNumber="338" rank="species">
|
||
<emphasis id="B9480707A3487F58FAACF80E8EC0DE89" bold="true" box="[1372,1436,1952,1977]" pageId="37" pageNumber="338">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="1307C790A3487F58FF6FF86A8A66DEEC" box="[159,314,1988,2013]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="37" pageNumber="338">Figs 112–114</figureCitation>
|
||
) is also somewhat similar to that of
|
||
<taxonomicName id="4C3CA096A3487F58FD27F86B88F9DEEC" authorityName="Ito" authorityYear="1998" box="[727,933,1988,2012]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3487F58FD27F86B89B1DEEC" box="[727,749,1989,2012]" italics="true" pageId="37" pageNumber="338">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3487F58FD0DF86B8852DEEC" box="[765,782,1989,2012]" italics="true" pageId="37" pageNumber="338">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3487F58FCD7F86B88F9DEEC" box="[807,933,1989,2012]" italics="true" pageId="37" pageNumber="338">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1307C790A3487F58FC44F86A8F13DEEC" box="[948,1103,1988,2013]" captionStart="FIGURES 115 – 119" captionStartId="40.[151,264,1519,1542]" captionTargetBox="[210,1376,415,1485]" captionTargetId="figure@40.[192,1394,411,1498]" captionTargetPageId="40" captionText="FIGURES 115 – 119. Trichotichnus (Bottchrus) cyanescens Ito, 1998 (specimen from Dhabeli). 115, left metepisternum; 116, apex of last (VII) abdominal sternite, male; 117, 119, median lobe of aedeagus; 118, apical capitulum. 115 – 116, ventral view; 117, lateral view; 118, caudal view; 119, dorsal view. Scale bars = 0.25 mm (A: Figs 115 – 117, 119; B: Fig. 118)." httpUri="https://zenodo.org/record/919668/files/figure.png" pageId="37" pageNumber="338">Figs 117–119</figureCitation>
|
||
) in general shape and having short terminal lamella hooked at apex, but in lateral view much less sinuate, with more thin terminal lamella and without distinct sclerotic elements in the internal sac. Based on the original description (
|
||
<bibRefCitation id="EFADA6E4A34B7F5BFB6AFF128E61D9E4" author="Ito" box="[1178,1341,188,212]" pageId="38" pageNumber="339" refString="Ito, N. (1998) Some species of the genus Trichotichnus from Nepal and the adjascent regions (Coleoptera, Carabidae, Harpalini). Japanese Journal of Systematic Entomology, 4 (2), 277 - 288." type="journal article" year="1998">Ito 1998: 279</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C3CA096A34B7F5BFAA3FF138B8BD9C8" authorityName="Ito" authorityYear="1998" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="loebli" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFAA3FF138E34D9E4" box="[1363,1384,189,212]" italics="true" pageId="38" pageNumber="339">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFA8AFF138ED7D9E4" box="[1402,1419,189,212]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFF67FF718B8BD9C8" box="[151,215,223,248]" italics="true" pageId="38" pageNumber="339">loebli</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49F815F7A34B7F5BFEEAFF4E8AFBD9C8" box="[282,423,223,248]" country="India" name="West Bengal" pageId="38" pageNumber="339">West Bengal</collectingRegion>
|
||
(Darjeeling District) is also similar to
|
||
<taxonomicName id="4C3CA096A34B7F5BFCA6FF4F8F66D9C8" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[854,1082,223,248]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFCA6FF4F8830D9C8" box="[854,876,225,248]" italics="true" pageId="38" pageNumber="339">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFC8BFF4F88D0D9C8" box="[891,908,225,248]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFC53FF718F66D9C8" box="[931,1082,223,248]" italics="true" pageId="38" pageNumber="339">manasluensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34B7F5BFBB0FF718F23D9C8" box="[1088,1151,223,248]" pageId="38" pageNumber="339" rank="species">
|
||
<emphasis id="B9480707A34B7F5BFBB0FF718F23D9C8" bold="true" box="[1088,1151,223,248]" pageId="38" pageNumber="339">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3CA096A34B7F5BFB45FF4F8EDCD9C8" authorityName="Ito" authorityYear="1998" box="[1205,1408,224,248]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="cyanescens" subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFB45FF4F8F96D9C8" box="[1205,1226,225,248]" italics="true" pageId="38" pageNumber="339">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFB29FF4F8FB6D9C8" box="[1241,1258,225,248]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFAF2FF4F8EDCD9C8" box="[1282,1408,225,248]" italics="true" pageId="38" pageNumber="339">cyanescens</emphasis>
|
||
</taxonomicName>
|
||
in many characters (general habitus, femora not infuscate, pronotal lateral gutter slightly widened anteriad, protibia very finely sulcate on dorsal side, and S-shaped aedeagal median lobe), but distinctly differs from
|
||
<taxonomicName id="4C3CA096A34B7F5BFABEFE878A72D855" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFABEFE878E38D870" box="[1358,1380,297,320]" italics="true" pageId="38" pageNumber="339">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFA8AFE878ED7D870" box="[1402,1419,297,320]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFF67FEE28A72D855" box="[151,302,332,357]" italics="true" pageId="38" pageNumber="339">manasluensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34B7F5BFEC7FEE28A24D855" box="[311,376,332,357]" pageId="38" pageNumber="339" rank="species">
|
||
<emphasis id="B9480707A34B7F5BFEC7FEE28A24D855" bold="true" box="[311,376,332,357]" pageId="38" pageNumber="339">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
at least in having larger size (
|
||
<quantity id="4CC476F0A34B7F5BFD26FEE28839D854" box="[726,869,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.2" metricValueMin="6.3" pageId="38" pageNumber="339" unit="mm" value="6.75" valueMax="7.2" valueMin="6.3">6.3–7.2 mm</quantity>
|
||
), mentum and submentum separated by complete transverse suture, humerus without denticle, and median lobe with a large transverse apical capitulum approximately equally prominent dorsally and ventrally.
|
||
<taxonomicName id="4C3CA096A34B7F5BFCCCFE3A8FC8D89D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[828,1172,404,429]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="brancuccii" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFCCCFE3A888AD89D" box="[828,982,404,429]" italics="true" pageId="38" pageNumber="339">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFC1DFE3B88A2D89C" box="[1005,1022,405,428]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFBECFE3A8FC8D89D" box="[1052,1172,404,429]" italics="true" pageId="38" pageNumber="339">brancuccii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34B7F5BFB52FE3A8FB4D89D" box="[1186,1256,404,429]" pageId="38" pageNumber="339" rank="species">
|
||
<emphasis id="B9480707A34B7F5BFB52FE3A8FB4D89D" bold="true" box="[1186,1256,404,429]" pageId="38" pageNumber="339">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, which is also characterized by somewhat S-shaped median lobe, differs from
|
||
<taxonomicName id="4C3CA096A34B7F5BFC98FE178F12D8E0" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[872,1102,439,464]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFC98FE178820D8E0" box="[872,892,441,464]" italics="true" pageId="38" pageNumber="339">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFC7EFE1788C3D8E0" box="[910,927,441,464]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFC47FE198F12D8E0" box="[951,1102,439,464]" italics="true" pageId="38" pageNumber="339">manasluensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34B7F5BFBA6FE198FCAD8E0" box="[1110,1174,439,464]" pageId="38" pageNumber="339" rank="species">
|
||
<emphasis id="B9480707A34B7F5BFBA6FE198FCAD8E0" bold="true" box="[1110,1174,439,464]" pageId="38" pageNumber="339">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in having mentum and submentum fused laterally and separated by transverse suture medially, protibia not sulcate, femora distinctly infuscate, and aedeagal median lobe (
|
||
<figureCitation id="1307C790A34B7F5BFDBAFE5189B4DB28" box="[586,744,511,536]" captionStart="FIGURES 103 – 108" captionStartId="35.[151,264,1454,1477]" captionTargetBox="[216,1385,597,1417]" captionTargetId="figure@35.[195,1392,589,1433]" captionTargetPageId="35" captionText="FIGURES 103 – 108. Trichotichnus (Bottchrus) brancuccii sp. n. (103, 105, paratype; 104, 106 – 108, holotype). 103, left metepisternum; 104, 105, apex of last (VII) abdominal sternite, males; 106, 108, median lobe of aedeagus; 107, apical capitulum. 103 – 105, ventral view; 106, lateral view; 107, caudal view; 108, dorsal view. Scale bars = 0.25 mm (A: Figs 103 – 106, 108; B: Fig. 107)." httpUri="https://zenodo.org/record/919662/files/figure.png" pageId="38" pageNumber="339">Figs 106–108</figureCitation>
|
||
) more strongly bent basally and with much longer terminal lamella markedly widened at apex.
|
||
<taxonomicName id="4C3CA096A34B7F5BFDA1FD8A88D1DB0D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[593,909,548,573]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFDA1FD8A89B7DB0D" box="[593,747,548,573]" italics="true" pageId="38" pageNumber="339">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFCF4FD8B8849DB0C" box="[772,789,549,572]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFCC6FD8A88D1DB0D" box="[822,909,548,573]" italics="true" pageId="38" pageNumber="339">pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34B7F5BFC6DFD8A88B9DB0D" box="[925,997,548,573]" pageId="38" pageNumber="339" rank="species">
|
||
<emphasis id="B9480707A34B7F5BFC6DFD8A88B9DB0D" bold="true" box="[925,997,548,573]" pageId="38" pageNumber="339">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, occurring sympatricly with
|
||
<taxonomicName id="4C3CA096A34B7F5BFABDFD8B8A72DB50" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFABDFD8B8E3CDB0C" box="[1357,1376,549,572]" italics="true" pageId="38" pageNumber="339">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFA8AFD8B8ED7DB0C" box="[1402,1419,549,572]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFF67FDE98A72DB50" box="[151,302,583,608]" italics="true" pageId="38" pageNumber="339">manasluensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34B7F5BFECBFDE98ADDDB50" box="[315,385,583,608]" pageId="38" pageNumber="339" rank="species">
|
||
<emphasis id="B9480707A34B7F5BFECBFDE98ADDDB50" bold="true" box="[315,385,583,608]" pageId="38" pageNumber="339">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, is easily distinguished from the latter by smaller size (
|
||
<quantity id="4CC476F0A34B7F5BFBDAFDE98FE1DB50" box="[1066,1213,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="38" pageNumber="339" unit="mm" value="5.1" valueMax="5.4" valueMin="4.8">4.8–5.4 mm</quantity>
|
||
), last visible (VII) abdominal sternite in female rounded at apex (
|
||
<figureCitation id="1307C790A34B7F5BFD5DFDC2885CDBB4" box="[685,768,620,645]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="38" pageNumber="339">Fig. 79</figureCitation>
|
||
) and median lobe of the aedeagus in male C-shaped (
|
||
<figureCitation id="1307C790A34B7F5BFA9AFDC28B80DB97" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="38" pageNumber="339">Figs 72–77</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA34B7F5BFF37FD1A8908DA24" pageId="38" pageNumber="339" type="distribution">
|
||
<paragraph id="8B83DB15A34B7F5BFF37FD1A8908DA24" blockId="38.[151,1436,151,789]" pageId="38" pageNumber="339">
|
||
<emphasis id="B9480707A34B7F5BFF37FD1A8A05DBFD" bold="true" box="[199,345,692,717]" pageId="38" pageNumber="339">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1307C790A34B7F5BFE99FD1A8A96DBFD" box="[361,458,692,717]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="38" pageNumber="339">Fig. 152</figureCitation>
|
||
,
|
||
<emphasis id="B9480707A34B7F5BFE27FD1A8AB9DBFC" box="[471,485,692,716]" italics="true" pageId="38" pageNumber="339">3</emphasis>
|
||
). Bara Pokhari Lekh mountain range on the south-western slope of Manaslu Himal, Lamjung District, western
|
||
<collectingRegion id="49F815F7A34B7F5BFE35FD798944DBC0" box="[453,536,727,752]" country="Sri Lanka" name="Central" pageId="38" pageNumber="339">Central</collectingRegion>
|
||
<collectingCountry id="F32B9B85A34B7F5BFDEEFD798938DBC0" box="[542,612,727,752]" name="Nepal" pageId="38" pageNumber="339">Nepal</collectingCountry>
|
||
, at altitudes of approximately
|
||
<quantity id="4CC476F0A34B7F5BFC4AFD798F04DBDF" box="[954,1112,727,752]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="38" pageNumber="339" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">1600–1800 m</quantity>
|
||
, where it occurs sympatricly with
|
||
<taxonomicName id="4C3CA096A34B7F5BFF3FFD538A28DA25" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[207,372,764,789]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34B7F5BFF3FFD538BB9DA24" box="[207,229,765,788]" italics="true" pageId="38" pageNumber="339">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34B7F5BFF04FD538A59DA24" box="[244,261,765,788]" italics="true" pageId="38" pageNumber="339">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34B7F5BFEEDFD528A28DA25" box="[285,372,764,789]" italics="true" pageId="38" pageNumber="339">pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34B7F5BFE8BFD528AE6DA25" box="[379,442,764,789]" pageId="38" pageNumber="339" rank="species">
|
||
<emphasis id="B9480707A34B7F5BFE8BFD528AE6DA25" bold="true" box="[379,442,764,789]" pageId="38" pageNumber="339">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="1307C790A34B7F5BFE39FD528976DA25" box="[457,554,764,789]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="38" pageNumber="339">Fig. 151</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A34B7F5BFDC5FD52891FDA24" box="[565,579,764,788]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="38" pageNumber="339">
|
||
<emphasis id="B9480707A34B7F5BFDC5FD52891FDA24" box="[565,579,764,788]" italics="true" pageId="38" pageNumber="339">9</emphasis>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |