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<document id="72987DD5ACEBE2CCFC2232DE480344D8" ID-CLB-Dataset="290199" ID-DOI="10.11646/zootaxa.5418.5.6" ID-GBIF-Dataset="40fd70f2-37f9-42be-8ae8-97be12bf3b58" ID-ISSN="1175-5326" ID-Zenodo-Dep="10780063" ID-ZooBank="8681F446-C4A0-466F-A763-57F426B14523" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1709588019354" checkinUser="plazi" docAuthor="Dhiya’Ulhaq, Naufal Urfi, Dupérré, Nadine, Buchori, Damayanti, Scheu, Stefan &amp; Drescher, Jochen" docDate="2024" docId="03C0D450D17A952A41ADFB331C87D7AF" docLanguage="en" docName="zootaxa.5418.5.6.pdf" docOrigin="Zootaxa 5418 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5418.5.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Utivarachna angsoduo Dhiya’ulhaq &amp; Dupérré 2024, sp. nov." docType="treatment" docVersion="6" lastPageNumber="556" masterDocId="FFF9AC28D178952F413AFF961C7CD506" masterDocTitle="Four new species of Utivarachna Kishida, 1940 (Araneae: Trachelidae) from Sumatra" masterLastPageNumber="575" masterPageNumber="551" pageNumber="553" updateTime="1713441516709" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="FF5934C068CF8A727E16336E08136D3B">Four new species of Utivarachna Kishida, 1940 (Araneae: Trachelidae) from Sumatra</mods:title>
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<mods:namePart id="B65FBFCEEE4CD66447BA6256504721C7">Dhiya’Ulhaq, Naufal Urfi</mods:namePart>
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<mods:affiliation id="3581F57F0D5A4D399A7CA60B245D7D27">Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany</mods:affiliation>
<mods:nameIdentifier id="4DA2CCEB003DB8D084D461A3DA02609C" type="email">naufalurfi.du@gmail.com</mods:nameIdentifier>
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<mods:role id="853AD2B9635412748058A81100570A40">
<mods:roleTerm id="72C154D2AEC9EF8CE3D976441B7C5FA4">Author</mods:roleTerm>
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<mods:namePart id="764560A82C30D2A85B31DF576790093A">Dupérré, Nadine</mods:namePart>
<mods:affiliation id="C463A0CF1502FD043C75E9BBEFD2AFE6">Centre for Taxonomy and Morphology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Museum of Nature Hamburg, Hamburg, Germany &amp; American Museum of Natural History, New York, NY, United States of America</mods:affiliation>
</mods:name>
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<mods:namePart id="1D5E89AAE8BF49EA2DF00D7866CC580D">Buchori, Damayanti</mods:namePart>
<mods:affiliation id="C2E5624B7C66BFADAD07647BEABB1732">Department of Plant Protection, Faculty of Agriculture, IPB University, Bogor, Indonesia &amp; Centre for Transdisciplinary and Sustainability Sciences, IPB University, Bogor, Indonesia</mods:affiliation>
</mods:name>
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<mods:namePart id="3C3A65DD29A17E9399059D592C7D7CC8">Scheu, Stefan</mods:namePart>
<mods:affiliation id="FCA21AE8BD95526CA1C0080AE1CA64D3">Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany &amp; Centre of Biodiversity and Sustainable Land Use, Göttingen, Germany</mods:affiliation>
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<mods:namePart id="94DCEE3BBC0B8C0A9CFBF1E1C413B2C8">Drescher, Jochen</mods:namePart>
<mods:nameIdentifier id="3EA42F0F3126EC177BFE4270BBDF0CB2" type="ORCID">0000-0002-5162-9779</mods:nameIdentifier>
<mods:affiliation id="80F802DDBF1DAC042569B03F407413E8">Department of Animal Ecology, J. - F. Blumenbach Institute for Zoology and Anthropology, University of Göttingen, Göttingen, Germany</mods:affiliation>
<mods:nameIdentifier id="CFA4AD6E87FEEF1F7F36357148FF4D24" type="email">jochen.drescher@biologie.uni-goettingen.de</mods:nameIdentifier>
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<mods:title id="F58533A53B41CA4D5F645576F2916AE8">Zootaxa</mods:title>
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<mods:date id="D903E64C2966D306EC9DB648CC6F6A4C">2024</mods:date>
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<mods:number id="D78E869EC4A8DED3169F93F1E4F09401">2024-03-04</mods:number>
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<mods:classification id="89E7E0220BBD193E2869260409D15EED">journal article</mods:classification>
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<treatment id="03C0D450D17A952A41ADFB331C87D7AF" ID-DOI="http://doi.org/10.5281/zenodo.10794100" ID-GBIF-Taxon="222728026" ID-Zenodo-Dep="10794100" LSID="urn:lsid:plazi:treatment:03C0D450D17A952A41ADFB331C87D7AF" httpUri="http://treatment.plazi.org/id/03C0D450D17A952A41ADFB331C87D7AF" lastPageId="5" lastPageNumber="556" pageId="2" pageNumber="553">
<subSubSection id="C37336CDD17A952D41ADFB331F31D1C6" box="[151,845,1189,1216]" pageId="2" pageNumber="553" type="nomenclature">
<paragraph id="8BD66546D17A952D41ADFB331F31D1C6" blockId="2.[151,845,1189,1252]" box="[151,845,1189,1216]" pageId="2" pageNumber="553">
<heading id="D09ED22AD17A952D41ADFB331F31D1C6" bold="true" box="[151,845,1189,1216]" fontSize="11" level="1" pageId="2" pageNumber="553" reason="1">
<emphasis id="B91DB954D17A952D41ADFB331F31D1C6" bold="true" box="[151,845,1189,1216]" pageId="2" pageNumber="553">
<taxonomicName id="4C691EC5D17A952D41ADFB331E9ED1C6" authority="Dhiya’ulhaq &amp; Dupérré, 2024" authorityName="Dhiya’ulhaq &amp; Dupérré" authorityYear="2024" box="[151,738,1189,1216]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="angsoduo" status="sp. nov.">
<emphasis id="B91DB954D17A952D41ADFB331DD7D1C6" bold="true" box="[151,427,1189,1216]" italics="true" pageId="2" pageNumber="553">Utivarachna angsoduo</emphasis>
Dhiya’ulhaq &amp; Dupérré
</taxonomicName>
,
<taxonomicNameLabel id="A22E042FD17A952D43D5FB301F31D1C6" box="[751,845,1190,1216]" pageId="2" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD17A952D41ADFB5C1D60D1E2" box="[151,284,1226,1252]" pageId="2" pageNumber="553" type="description">
<paragraph id="8BD66546D17A952D41ADFB5C1D60D1E2" blockId="2.[151,845,1189,1252]" box="[151,284,1226,1252]" pageId="2" pageNumber="553">
<figureCitation id="135279C3D17A952D41ADFB5C1D7CD1E2" box="[151,256,1226,1252]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1959]" captionTargetBox="[151,1436,181,1911]" captionTargetId="figure-15@3.[151,1436,181,1911]" captionTargetPageId="3" captionText="FIGURE 1. Utivarachna angsoduo sp. nov., male holotype. A, B Habitus, dorsal view (A), ventral view (B); C Right leg I, prolateral view. D Sternum, ventral view. Scale bars A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780065" httpUri="https://zenodo.org/record/10780065/files/figure.png" pageId="2" pageNumber="553">Figures 1</figureCitation>
,
<figureCitation id="135279C3D17A952D4034FB5C1D60D1E2" box="[270,284,1226,1252]" captionStart="FIGURE 2" captionStartId="4.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1609]" captionTargetId="figure-23@4.[151,1436,181,1616]" captionTargetPageId="4" captionText="FIGURE 2. Utivarachna angsoduo sp. nov., male holotype. A–E Left palp: A prolateral view; B, D ventral view; C, E retrolateral view. Scale bars = 0.2 mm. Abbreviations: dl, dorsal lobe of RTA; ds, dorsal spine of dorsal RTA lobe; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." figureDoi="http://doi.org/10.5281/zenodo.10780067" httpUri="https://zenodo.org/record/10780067/files/figure.png" pageId="2" pageNumber="553">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD17A952D41ADFA841E46D072" pageId="2" pageNumber="553" type="materials_examined">
<paragraph id="8BD66546D17A952D41ADFA841E46D072" blockId="2.[151,1437,1297,2008]" pageId="2" pageNumber="553">
<emphasis id="B91DB954D17A952D41ADFA841D34D02D" bold="true" box="[151,328,1297,1324]" pageId="2" pageNumber="553">Type material.</emphasis>
<materialsCitation id="3B016F1BD17A952D406BFA841E46D072" ID-GBIF-Occurrence="4535210321" collectingDate="2013-07-14" collectionCode="MZB" collectorName="J. Drescher" elevation="64" latitude="-0.015777778" location="Dusun Baru" longLatPrecision="1" longitude="0.7535" municipality="Jambi Province" pageId="2" pageNumber="553" specimenCount="1" specimenCount-male="1" stateProvince="SUMATRA" typeStatus="holotype">
<emphasis id="B91DB954D17A952D406BFA841F32D02A" bold="true" box="[337,846,1297,1327]" pageId="2" pageNumber="553">
<typeStatus id="54D2DBE4D17A952D406BFA841DC2D02A" box="[337,446,1298,1324]" pageId="2" pageNumber="553" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6FAECFD17A952D40FDFA851DA2D029" box="[455,478,1299,1327]" count="1" pageId="2" pageNumber="553" type="male">♂</specimenCount>
:
<collectingRegion id="49ADABA4D17A952D40D5FA871E03D02D" box="[495,639,1297,1323]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="553">SUMATRA</collectingRegion>
:
<emphasis id="B91DB954D17A952D43B7FA871F39D02A" bold="true" box="[653,837,1297,1324]" italics="true" pageId="2" pageNumber="553">
<collectingMunicipality id="6BB2FF3CD17A952D43B7FA871F39D02A" box="[653,837,1297,1324]" pageId="2" pageNumber="553">Jambi Province</collectingMunicipality>
</emphasis>
:
</emphasis>
<taxonomicName id="4C691EC5D17A952D4262FA841F98D02A" authority="Baru" authorityName="Baru" box="[856,996,1298,1324]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="dusun">
<location id="8EB6339DD17A952D4262FA841F98D02A" LSID="urn:lsid:plazi:treatment:03C0D450D17A952A41ADFB331C87D7AF:8EB6339DD17A952D4262FA841F98D02A" box="[856,996,1298,1324]" latitude="-0.015777778" longLatPrecision="1" longitude="0.7535" municipality="Jambi Province" name="Dusun Baru" pageId="2" pageNumber="553" stateProvince="SUMATRA">Dusun Baru</location>
</taxonomicName>
,
<collectingMunicipality id="6BB2FF3CD17A952D42CAFA841814D02A" box="[1008,1128,1298,1324]" pageId="2" pageNumber="553">Air Hitam</collectingMunicipality>
,
<location id="8EB6339DD17A952D454FFA841889D02A" LSID="urn:lsid:plazi:treatment:03C0D450D17A952A41ADFB331C87D7AF:8EB6339DD17A952D454FFA841889D02A" box="[1141,1269,1298,1324]" latitude="-0.015777778" longLatPrecision="1" longitude="0.7535" municipality="Jambi Province" name="Sarolangun" pageId="2" pageNumber="553" stateProvince="SUMATRA">Sarolangun</location>
(2013_BJ4.2_ AraTrac005N_001), canopy fogging jungle rubber agroforest, 02⁰00’56.8”S, 102⁰45’12.6”E, altitude
<quantity id="4C91C8A3D17A952D446EFAA019EBD056" box="[1364,1431,1334,1360]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="553" unit="m" value="64.0">
<elevation id="00448275D17A952D446EFAA019EBD056" box="[1364,1431,1334,1360]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="553" unit="m" value="64.0">64 m</elevation>
</quantity>
,
<date id="FFD74386D17A952D41ADFACC1D5ED072" box="[151,290,1370,1396]" pageId="2" pageNumber="553" value="2013-07-14">
<collectingDate id="EF93BA6ED17A952D41ADFACC1D5ED072" box="[151,290,1370,1396]" pageId="2" pageNumber="553" value="2013-07-14">14.VII.2013</collectingDate>
</date>
, leg.
<collectorName id="269C0090D17A952D4064FACC1DA0D072" box="[350,476,1370,1396]" pageId="2" pageNumber="553">J. Drescher</collectorName>
(
<collectionCode id="ED78FD83D17A952D40D1FACC1E51D072" box="[491,557,1370,1396]" collectionName="MZB" pageId="2" pageNumber="553">MZB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD17A952D41FDFAE81EE7D0E6" pageId="2" pageNumber="553" type="etymology">
<paragraph id="8BD66546D17A952D41FDFAE81EE7D0E6" blockId="2.[151,1437,1297,2008]" pageId="2" pageNumber="553">
<emphasis id="B91DB954D17A952D41FDFAE81D32D09E" bold="true" box="[199,334,1406,1432]" pageId="2" pageNumber="553">Etymology.</emphasis>
The specific name is taken from
<collectingRegion id="49ADABA4D17A952D43DCFAE81F55D09E" box="[742,809,1406,1432]" country="Indonesia" name="Jambi" pageId="2" pageNumber="553">Jambi</collectingRegion>
Malay
<emphasis id="B91DB954D17A952D42B2FAE9187DD09E" box="[904,1025,1406,1432]" italics="true" pageId="2" pageNumber="553">angso duo</emphasis>
meaning “two swans”, part of the mythological founding story of
<collectingRegion id="49ADABA4D17A952D4338FA341E39D0BA" box="[514,581,1442,1468]" country="Indonesia" name="Jambi" pageId="2" pageNumber="553">Jambi</collectingRegion>
and is a common symbol in
<collectingRegion id="49ADABA4D17A952D42AEFA341FABD0BA" box="[916,983,1442,1468]" country="Indonesia" name="Jambi" pageId="2" pageNumber="553">Jambi</collectingRegion>
culture. Also alluding to the long, twolobed RTA of the species. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD17A952D41FDFA7F1EAED3DA" pageId="2" pageNumber="553" type="diagnosis">
<paragraph id="8BD66546D17A952D41FDFA7F1EAED3DA" blockId="2.[151,1437,1297,2008]" pageId="2" pageNumber="553">
<emphasis id="B91DB954D17A952D41FDFA7F1D3ED305" bold="true" box="[199,322,1513,1539]" pageId="2" pageNumber="553">Diagnosis.</emphasis>
This species belongs to the
<taxonomicName id="4C691EC5D17A952D434CFA7C1F6CD302" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[630,784,1514,1540]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B91DB954D17A952D434CFA7C1F6CD302" box="[630,784,1514,1540]" italics="true" pageId="2" pageNumber="553">kinabaluensis</emphasis>
</taxonomicName>
-group based on the following characters: carapace wedge-shaped with undulating lateral margins; posterior end of carapace not produced into an elongated stalk (
<figureCitation id="135279C3D17A952D4409F99819F1D32E" box="[1331,1421,1550,1576]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1959]" captionTargetBox="[151,1436,181,1911]" captionTargetId="figure-15@3.[151,1436,181,1911]" captionTargetPageId="3" captionText="FIGURE 1. Utivarachna angsoduo sp. nov., male holotype. A, B Habitus, dorsal view (A), ventral view (B); C Right leg I, prolateral view. D Sternum, ventral view. Scale bars A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780065" httpUri="https://zenodo.org/record/10780065/files/figure.png" pageId="2" pageNumber="553">Fig. 1A</figureCitation>
); PER recurved and much longer than AER (
<figureCitation id="135279C3D17A952D434FF9A41EB1D34A" box="[629,717,1586,1612]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1959]" captionTargetBox="[151,1436,181,1911]" captionTargetId="figure-15@3.[151,1436,181,1911]" captionTargetPageId="3" captionText="FIGURE 1. Utivarachna angsoduo sp. nov., male holotype. A, B Habitus, dorsal view (A), ventral view (B); C Right leg I, prolateral view. D Sternum, ventral view. Scale bars A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780065" httpUri="https://zenodo.org/record/10780065/files/figure.png" pageId="2" pageNumber="553">Fig. 1A</figureCitation>
); palpal tegulum not enlarged and bulbous (
<figureCitation id="135279C3D17A952D458EF9A41884D34A" box="[1204,1272,1586,1612]" captionStart="FIGURE 2" captionStartId="4.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1609]" captionTargetId="figure-23@4.[151,1436,181,1616]" captionTargetPageId="4" captionText="FIGURE 2. Utivarachna angsoduo sp. nov., male holotype. A–E Left palp: A prolateral view; B, D ventral view; C, E retrolateral view. Scale bars = 0.2 mm. Abbreviations: dl, dorsal lobe of RTA; ds, dorsal spine of dorsal RTA lobe; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." figureDoi="http://doi.org/10.5281/zenodo.10780067" httpUri="https://zenodo.org/record/10780067/files/figure.png" pageId="2" pageNumber="553">Fig. 2</figureCitation>
). Males of this species have a similar cephalothorax and abdomen to those of
<taxonomicName id="4C691EC5D17A952D427FF9C11FD3D376" authority="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher, 2024" authorityName="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher" authorityYear="2024" box="[837,943,1622,1648]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="trisula" status="sp. nov.">
<emphasis id="B91DB954D17A952D427FF9C11FD3D376" box="[837,943,1622,1648]" italics="true" pageId="2" pageNumber="553">U. trisula</emphasis>
</taxonomicName>
<emphasis id="B91DB954D17A952D428FF9C01871D376" bold="true" box="[949,1037,1622,1648]" pageId="2" pageNumber="553">
<taxonomicNameLabel id="A22E042FD17A952D428FF9C01871D376" box="[949,1037,1622,1648]" pageId="2" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished from it (as well as from all other congeners) by the long bifid shape of the RTA (
<figureCitation id="135279C3D17A952D42A6F9EC1864D392" box="[924,1048,1658,1684]" captionStart="FIGURE 2" captionStartId="4.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1609]" captionTargetId="figure-23@4.[151,1436,181,1616]" captionTargetPageId="4" captionText="FIGURE 2. Utivarachna angsoduo sp. nov., male holotype. A–E Left palp: A prolateral view; B, D ventral view; C, E retrolateral view. Scale bars = 0.2 mm. Abbreviations: dl, dorsal lobe of RTA; ds, dorsal spine of dorsal RTA lobe; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." figureDoi="http://doi.org/10.5281/zenodo.10780067" httpUri="https://zenodo.org/record/10780067/files/figure.png" pageId="2" pageNumber="553">Figs 2B–E</figureCitation>
), each lobe curving in the opposite direction of one another, the ventral lobe additionally with a small spine-like dorsal projection (
<figureCitation id="135279C3D17A952D45F9F9081940D3BE" box="[1219,1340,1694,1720]" captionStart="FIGURE 2" captionStartId="4.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1609]" captionTargetId="figure-23@4.[151,1436,181,1616]" captionTargetPageId="4" captionText="FIGURE 2. Utivarachna angsoduo sp. nov., male holotype. A–E Left palp: A prolateral view; B, D ventral view; C, E retrolateral view. Scale bars = 0.2 mm. Abbreviations: dl, dorsal lobe of RTA; ds, dorsal spine of dorsal RTA lobe; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." figureDoi="http://doi.org/10.5281/zenodo.10780067" httpUri="https://zenodo.org/record/10780067/files/figure.png" pageId="2" pageNumber="553">Figs 2C, E</figureCitation>
) (versus trifid in
<taxonomicName id="4C691EC5D17A952D41C9F9551D22D3DA" authorityName="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher" authorityYear="2024" box="[243,350,1730,1756]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="553" phylum="Arthropoda" rank="species" species="trisula">
<emphasis id="B91DB954D17A952D41C9F9551D22D3DA" box="[243,350,1730,1756]" italics="true" pageId="2" pageNumber="553">U. trisula</emphasis>
</taxonomicName>
,
<figureCitation id="135279C3D17A952D4056F9541D89D3DB" box="[364,501,1730,1757]" captionStart="FIGURE 15" captionStartId="18.[151,250,1923,1947]" captionTargetBox="[164,1423,453,1893]" captionTargetId="figure-110@18.[164,1423,453,1900]" captionTargetPageId="18" captionText="FIGURE 15. Utivarachna trisula sp. nov., male holotype. A–E Left palp: A, prolateral view; B, D, ventral view; C, E retrolateral view. Scale bars 0.2 mm. Abbreviations: ac, apical claw of RTA; dl, dorsal lobe of RTA; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." figureDoi="http://doi.org/10.5281/zenodo.10780108" httpUri="https://zenodo.org/record/10780108/files/figure.png" pageId="2" pageNumber="553">Figs 15B–E</figureCitation>
). Female: unknown
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD17A952A41FDF9731D4ED73B" lastPageId="5" lastPageNumber="556" pageId="2" pageNumber="553" type="description">
<paragraph id="8BD66546D17A952D41FDF9731934D2DE" blockId="2.[151,1437,1297,2008]" pageId="2" pageNumber="553">
<emphasis id="B91DB954D17A952D41FDF9731E5CD3F9" bold="true" box="[199,544,1765,1792]" pageId="2" pageNumber="553">
Description. Male (
<typeStatus id="54D2DBE4D17A952D4092F9731E64D3F9" box="[424,536,1765,1791]" pageId="2" pageNumber="553" type="holotype">Holotype</typeStatus>
)
</emphasis>
. Total length 4.90. Carapace length 2.36; width 2.00. Carapace maroon, strongly sclerotized, surface granulated, with granules ending in long white seta, lateral margins undulating, posterior half of carapace almost wedge-shaped with narrowly truncate end; fovea short but distinct; PER longer than AER, both slightly recurved (
<figureCitation id="135279C3D17A952D4067F8C41DCED26A" box="[349,434,1874,1900]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1959]" captionTargetBox="[151,1436,181,1911]" captionTargetId="figure-15@3.[151,1436,181,1911]" captionTargetPageId="3" captionText="FIGURE 1. Utivarachna angsoduo sp. nov., male holotype. A, B Habitus, dorsal view (A), ventral view (B); C Right leg I, prolateral view. D Sternum, ventral view. Scale bars A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780065" httpUri="https://zenodo.org/record/10780065/files/figure.png" pageId="2" pageNumber="553">Fig. 1A</figureCitation>
); clypeus short with slight projection between the chelicerae. Eye diameters:AME 0.13; ALE 0.17; PLE 0.14; PME 0.15. Eye interdistances: AME–AME 0.08; AME–ALE 0.10; ALE–ALE 0.49; PME–PME 0.17; PME–PLE 0.18; ALE–PLE 0.17; AME–PME 0.14; PLE–PLE 0.87. MOA: length 0.41; anterior width 0.33; posterior width 0.48. Clypeus height 0.32. Sternum heart-shaped, colored as carapace, surface granulated (
<figureCitation id="135279C3D17A952D4585F8281947D2DE" box="[1215,1339,1982,2008]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1959]" captionTargetBox="[151,1436,181,1911]" captionTargetId="figure-15@3.[151,1436,181,1911]" captionTargetPageId="3" captionText="FIGURE 1. Utivarachna angsoduo sp. nov., male holotype. A, B Habitus, dorsal view (A), ventral view (B); C Right leg I, prolateral view. D Sternum, ventral view. Scale bars A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780065" httpUri="https://zenodo.org/record/10780065/files/figure.png" pageId="2" pageNumber="553">Figs 1B, D</figureCitation>
).
</paragraph>
<caption id="DF1635CED17B952C41ADF8191FA1D2CD" ID-DOI="http://doi.org/10.5281/zenodo.10780065" ID-Zenodo-Dep="10780065" httpUri="https://zenodo.org/record/10780065/files/figure.png" pageId="3" pageNumber="554" startId="3.[151,250,1935,1959]" targetBox="[151,1436,181,1911]" targetPageId="3" targetType="figure">
<paragraph id="8BD66546D17B952C41ADF8191FA1D2CD" blockId="3.[151,1437,1935,1995]" pageId="3" pageNumber="554">
<emphasis id="B91DB954D17B952C41ADF8191D69D2A1" bold="true" box="[151,277,1935,1959]" pageId="3" pageNumber="554">FIGURE 1.</emphasis>
<taxonomicName id="4C691EC5D17B952C4026F8191E7BD2A1" authority="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher, 2024" authorityName="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher" authorityYear="2024" box="[284,519,1935,1959]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="angsoduo" status="sp. nov.">
<emphasis id="B91DB954D17B952C4026F8191E7BD2A1" box="[284,519,1935,1959]" italics="true" pageId="3" pageNumber="554">Utivarachna angsoduo</emphasis>
</taxonomicName>
<emphasis id="B91DB954D17B952C4334F8191E1CD2A1" bold="true" box="[526,608,1935,1959]" pageId="3" pageNumber="554">
<taxonomicNameLabel id="A22E042FD17B952C4334F8191E1CD2A1" box="[526,608,1935,1959]" pageId="3" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype.
<emphasis id="B91DB954D17B952C4235F8191F3CD2A1" bold="true" box="[783,832,1935,1959]" pageId="3" pageNumber="554">A, B</emphasis>
Habitus, dorsal view (
<emphasis id="B91DB954D17B952C4516F8191842D2A1" bold="true" box="[1068,1086,1935,1959]" pageId="3" pageNumber="554">A</emphasis>
), ventral view (
<emphasis id="B91DB954D17B952C45DEF8191889D2A1" bold="true" box="[1252,1269,1935,1959]" pageId="3" pageNumber="554">B</emphasis>
);
<emphasis id="B91DB954D17B952C4437F8191963D2A1" bold="true" box="[1293,1311,1935,1959]" pageId="3" pageNumber="554">C</emphasis>
Right leg I, prolateral view.
<emphasis id="B91DB954D17B952C4001F8251D31D2CD" bold="true" box="[315,333,1971,1995]" pageId="3" pageNumber="554">D</emphasis>
Sternum, ventral view. Scale bars A, B = 1 mm; C, D = 0.5 mm.
</paragraph>
</caption>
<caption id="DF1635CED17C952B41ADF9F11F0FD3CE" ID-DOI="http://doi.org/10.5281/zenodo.10780067" ID-Zenodo-Dep="10780067" httpUri="https://zenodo.org/record/10780067/files/figure.png" pageId="4" pageNumber="555" startId="4.[151,250,1639,1663]" targetBox="[151,1436,181,1609]" targetPageId="4" targetType="figure">
<paragraph id="8BD66546D17C952B41ADF9F11F0FD3CE" blockId="4.[151,1436,1639,1736]" pageId="4" pageNumber="555">
<emphasis id="B91DB954D17C952B41ADF9F11D6BD379" bold="true" box="[151,279,1639,1663]" pageId="4" pageNumber="555">FIGURE 2.</emphasis>
<taxonomicName id="4C691EC5D17C952B401BF9FE1E72D386" authority="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher, 2024" authorityName="Dhiya’Ulhaq &amp; Dupérré &amp; Buchori &amp; Scheu &amp; Drescher" authorityYear="2024" box="[289,526,1640,1664]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="angsoduo" status="sp. nov.">
<emphasis id="B91DB954D17C952B401BF9FE1E72D386" box="[289,526,1640,1664]" italics="true" pageId="4" pageNumber="555">Utivarachna angsoduo</emphasis>
</taxonomicName>
<emphasis id="B91DB954D17C952B4322F9FE1E10D386" bold="true" box="[536,620,1640,1664]" pageId="4" pageNumber="555">
<taxonomicNameLabel id="A22E042FD17C952B4322F9FE1E10D386" box="[536,620,1640,1664]" pageId="4" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype.
<emphasis id="B91DB954D17C952B4218F9F11F2ED379" bold="true" box="[802,850,1639,1663]" pageId="4" pageNumber="555">A–E</emphasis>
Left palp:
<emphasis id="B91DB954D17C952B42F7F9F11FA3D379" bold="true" box="[973,991,1639,1663]" pageId="4" pageNumber="555">A</emphasis>
prolateral view;
<emphasis id="B91DB954D17C952B45A2F9FE18B7D386" bold="true" box="[1176,1227,1640,1664]" pageId="4" pageNumber="555">B, D</emphasis>
ventral view;
<emphasis id="B91DB954D17C952B4452F9F119E0D386" bold="true" box="[1384,1436,1639,1664]" pageId="4" pageNumber="555">C, E</emphasis>
retrolateral view. Scale bars = 0.2 mm. Abbreviations: dl, dorsal lobe of RTA; ds, dorsal spine of dorsal RTA lobe; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA.
</paragraph>
</caption>
<paragraph id="8BD66546D17C952B41FDF9601950D2A6" blockId="4.[151,1437,1782,2024]" pageId="4" pageNumber="555">Abdomen length 2.54; width 1.56. Abdomen oval, pale-colored with three pairs of dark patches on the lateral margin, the anterior two pairs rather faint, as well as a darker-shaded cardiac pattern and a pair of small dark patches directly anterior to the spinnerets; most of the dorsal surface of abdomen covered with an orange scutum; ventral side of abdomen sclerotized on the area anterior to the epigastric furrow, projecting a short distance posterior to it on the sides; four longitudinal rows of sclerotized dots present between epigastric furrow and spinnerets.</paragraph>
<paragraph id="8BD66546D17C952A41FDF83C1F16D41B" blockId="4.[151,1437,1782,2024]" lastBlockId="5.[151,1437,151,681]" lastPageId="5" lastPageNumber="556" pageId="4" pageNumber="555">
Legs yellow, covered with long white seta especially on the ventral surface, striated with black bands distally and proximally on tibiae I–IV and femur I, while femora II–IV and metatarsi III–IV with distal bands only; anterior legs stouter and longer than posterior legs; the ventral side of anterior legs filled with leg cuspules from tarsi to tibiae (
<figureCitation id="135279C3D17D952A41D2FF2D1D3DD5D3" box="[232,321,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1935,1959]" captionTargetBox="[151,1436,181,1911]" captionTargetId="figure-15@3.[151,1436,181,1911]" captionTargetPageId="3" captionText="FIGURE 1. Utivarachna angsoduo sp. nov., male holotype. A, B Habitus, dorsal view (A), ventral view (B); C Right leg I, prolateral view. D Sternum, ventral view. Scale bars A, B = 1 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10780065" httpUri="https://zenodo.org/record/10780065/files/figure.png" pageId="5" pageNumber="556">Fig. 1C</figureCitation>
), metatarsi III and IV distally with comb-like structure distally followed by brush of setae. Leg measurements: leg I 7.46 (2.13, 0.73, 2.11, 1.72, 0.77); leg II 7.55 (2.17, 0.72, 2.05, 1.76, 0.85); leg III 5.19 (1.45, 0.48, 1.26, 1.44, 0.56); leg IV 6.48 (1.72, 0.54, 1.50, 2.09, 0.63).
</paragraph>
<paragraph id="8BD66546D17D952A41FDFEB11D4ED73B" blockId="5.[151,1437,151,681]" pageId="5" pageNumber="556">
Male palp (
<figureCitation id="135279C3D17D952A4076FEB11DE8D447" box="[332,404,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1609]" captionTargetId="figure-23@4.[151,1436,181,1616]" captionTargetPageId="4" captionText="FIGURE 2. Utivarachna angsoduo sp. nov., male holotype. A–E Left palp: A prolateral view; B, D ventral view; C, E retrolateral view. Scale bars = 0.2 mm. Abbreviations: dl, dorsal lobe of RTA; ds, dorsal spine of dorsal RTA lobe; e, embolus; rta, retrolateral tibial apophysis; st, subtegulum; vl, ventral lobe of RTA." figureDoi="http://doi.org/10.5281/zenodo.10780067" httpUri="https://zenodo.org/record/10780067/files/figure.png" pageId="5" pageNumber="556">Fig. 2</figureCitation>
): Cymbium and bulb oval. Sperm duct starting out diagonally then turning at 120° angle, before making narrow U-shaped turn. RTA bifid, ventral lobe canoe-shaped with excavated dorsal surface, long and slightly curved, ending in sharp bifid tip; curved spine-like projection grows dorsally in middle of lobe; dorsal lobe canoe-shaped, with excavated ventral surface, curved more prominently than ventral lobe and in opposite direction, separated from ventral lobe by membranous area, tip rounded, longer and slightly wider than the ventral lobe. Prolateral margins of both lobes narrower than retrolateral margin, giving lobes rebordered appearance when viewed retrolaterally. Embolus long, coiled horizontally. as wide as bulb, looping twice, visible dorsally through the cymbium.
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD17D952A41FDFDD018DBD767" box="[199,1191,582,609]" pageId="5" pageNumber="556" type="distribution">
<paragraph id="8BD66546D17D952A41FDFDD018DBD767" blockId="5.[151,1437,151,681]" box="[199,1191,582,609]" pageId="5" pageNumber="556">
<emphasis id="B91DB954D17D952A41FDFDD01D1DD766" bold="true" box="[199,353,582,608]" pageId="5" pageNumber="556">Distribution.</emphasis>
<collectingRegion id="49ADABA4D17D952A405DFDD11E6BD767" box="[359,535,583,609]" country="Indonesia" name="Jambi" pageId="5" pageNumber="556">Jambi Province</collectingRegion>
,
<collectingRegion id="49ADABA4D17D952A4319FDD11EFCD767" box="[547,640,583,609]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="556">Sumatra</collectingRegion>
(only known from
<typeStatus id="54D2DBE4D17D952A4260FDD11FF6D767" box="[858,906,583,609]" pageId="5" pageNumber="556">type</typeStatus>
locality) (
<figureCitation id="135279C3D17D952A42C7FDD1182ED767" box="[1021,1106,583,609]" captionStart="MAP 1" captionStartId="21.[152,212,1172,1196]" captionTargetBox="[151,1435,309,1148]" captionTargetId="figure-78@21.[151,1435,309,1148]" captionText="MAP 1. Distribution map of Utivarachna species collected in the framework of the 2013 EFForTS project in Jambi Province, Sumatra, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.10794102" httpUri="https://zenodo.org/record/10794102/files/figure.png" pageId="5" pageNumber="556">Maps 1</figureCitation>
and
<figureCitation id="135279C3D17D952A45B2FDD118E5D767" box="[1160,1177,583,609]" captionStart="MAP 2" captionStartId="22.[152,211,1994,2018]" captionTargetBox="[151,1436,1065,1971]" captionTargetId="figure-394@22.[151,1436,1065,1971]" captionText="MAP 2. Global distribution map of the Utivarachna species treated in this paper." figureDoi="http://doi.org/10.5281/zenodo.10794104" httpUri="https://zenodo.org/record/10794104/files/figure.png" pageId="5" pageNumber="556">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37336CDD17D952A41FDFDFD1C87D7AF" pageId="5" pageNumber="556" type="discussion">
<paragraph id="8BD66546D17D952A41FDFDFD1C87D7AF" blockId="5.[151,1437,151,681]" pageId="5" pageNumber="556">
<emphasis id="B91DB954D17D952A41FDFDFD1D47D783" bold="true" box="[199,315,619,645]" pageId="5" pageNumber="556">Remarks.</emphasis>
The
<typeStatus id="54D2DBE4D17D952A404CFDFD1DDAD783" box="[374,422,619,645]" pageId="5" pageNumber="556">type</typeStatus>
specimen was collected by canopy fogging in jungle rubber agroforest, and is considered arboreal.
</paragraph>
</subSubSection>
</treatment>
</document>
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