249 lines
47 KiB
XML
249 lines
47 KiB
XML
<document id="9A23ECD7C8B347EA836D92DF68429D2E" ID-DOI="10.11646/zootaxa.3894.1.11" ID-GBIF-Dataset="6c37cabe-0df5-462b-9f82-b5acfc8c37cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="229191" ID-ZooBank="78A995DA-7D11-48CE-A547-C7653E505720" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460997608442" checkinUser="plazi" docAuthor="Golovatch, Sergei I. & Vandenspiegel, Didier" docDate="2014" docId="0389DE08734F255EFF35FA01FD51C671" docLanguage="en" docName="zt03894p151.pdf" docOrigin="Zootaxa 3894 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Koponenius unicornis Golovatch & Vandenspiegel, 2014, sp. nov." docType="treatment" docUuid="4C790A59-BF16-4562-BA43-8AEF3EE2430D" docUuidSource="ZooBank" docVersion="8" lastPageNumber="147" masterDocId="FFB0A670734C2559FFA2FD62FFEBC76C" masterDocTitle="Koponenius gen. nov., a new genus of the millipede family Haplodesmidae from the Himalayas of India and Nepal (Diplopoda: Polydesmida)" masterLastPageNumber="151" masterPageNumber="141" pageNumber="144" updateTime="1698563275390" updateUser="plazi">
|
||
<mods:mods id="CF26174BD7A3B7A63D3CDC0E009A3517" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8D665AC849EAD152C02E1530D7ABE465">
|
||
<mods:title id="5E4ECAD90DDA1FF9241A5DA855E3DFB3">Koponenius gen. nov., a new genus of the millipede family Haplodesmidae from the Himalayas of India and Nepal (Diplopoda: Polydesmida)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6087C40AEB51BD4CC3B4B952F8D4DA34" type="personal">
|
||
<mods:role id="F2C75B789AA631D4A610483F872D8964">
|
||
<mods:roleTerm id="3562ECAE981638C59550D72135DA1B7A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E94BD8199350C5BFE9CF5ED99FA56FF0">Golovatch, Sergei I.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="9E6FAEC8D8287EE5F05497FB81E7B9B1" type="personal">
|
||
<mods:role id="B68E3A4AF2A9F9F8590EDD352A39D2C0">
|
||
<mods:roleTerm id="7CFF09DD1E016F6F4C6D35F2622AF16E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7712703AC2FD42B99A1F88A76F5D9F66">Vandenspiegel, Didier</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="1BE327B86810B31FF5B5A29C6FACCB2E">text</mods:typeOfResource>
|
||
<mods:relatedItem id="55EB2496FA2544F45BC1F922F2238CE9" type="host">
|
||
<mods:titleInfo id="8B8E779842ADF70A8977C68B2EF2BF63">
|
||
<mods:title id="6F7A8554B4A82498E8156B59A9ADDED3">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="84A440D077A608096CA6CA655A77B6C2">
|
||
<mods:date id="15188A5CBCF02F60B3D8AD0EB3EF4905">2014</mods:date>
|
||
<mods:detail id="56084DBEF85ED726110A1AA1FCFAFFD6" type="volume">
|
||
<mods:number id="96BA7D1A8B910D1A5EC2F4857EC6AD4B">3894</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A3509032CF82B0071E2316EEF89D12DD" type="issue">
|
||
<mods:number id="F782C1EC2DAFDE7FDA29B414A462ABB7">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="F8BDE02E27512A66D2B89E34D7D93CE4" unit="page">
|
||
<mods:start id="324F2480BE66F382AAC6BADCF8053EC0">141</mods:start>
|
||
<mods:end id="79933611B1769F833424E84105498B23">151</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F33328076058022F7ABE33A78016DC28">journal article</mods:classification>
|
||
<mods:identifier id="2DFB0127FD782CA8E0505E15C7478FA6" type="DOI">10.11646/zootaxa.3894.1.11</mods:identifier>
|
||
<mods:identifier id="EAE81C5BE442070570D852E45A88D39C" type="GBIF-Dataset">6c37cabe-0df5-462b-9f82-b5acfc8c37cc</mods:identifier>
|
||
<mods:identifier id="B54A7B938AC18BA0976361D69C5AD834" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="2138F8FE89A937DA01E2C4D1C895E423" type="Zenodo-Dep">229191</mods:identifier>
|
||
<mods:identifier id="61BC1CACA3334A3FD2293766D57E307C" type="ZooBank">78A995DA-7D11-48CE-A547-C7653E505720</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="0389DE08734F255EFF35FA01FD51C671" ID-DOI="http://doi.org/10.5281/zenodo.6126354" ID-GBIF-Taxon="119597640" ID-Zenodo-Dep="6126354" LSID="urn:lsid:zoobank.org:act:4C790A59-BF16-4562-BA43-8AEF3EE2430D" httpUri="http://treatment.plazi.org/id/0389DE08734F255EFF35FA01FD51C671" lastPageId="7" lastPageNumber="147" pageId="3" pageNumber="144">
|
||
<subSubSection id="C33A3C95734F255AFF35FA01FF12C0F2" pageId="3" pageNumber="144" type="nomenclature">
|
||
<paragraph id="8B9F6F1E734F255AFF35FA01FDE3C011" blockId="3.[151,520,1891,1950]" box="[151,520,1891,1917]" pageId="3" pageNumber="144">
|
||
<heading id="D0D7D872734F255AFF35FA01FDE3C011" bold="true" box="[151,520,1891,1917]" fontSize="11" level="1" pageId="3" pageNumber="144" reason="1">
|
||
<emphasis id="B954B30C734F255AFF35FA01FDE3C011" bold="true" box="[151,520,1891,1917]" pageId="3" pageNumber="144">
|
||
<taxonomicName id="4C20149D734F255AFF35FA01FE48C011" ID-CoL="3RCZ7" box="[151,419,1891,1917]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
|
||
<emphasis id="B954B30C734F255AFF35FA01FE48C011" bold="true" box="[151,419,1891,1917]" italics="true" pageId="3" pageNumber="144">Koponenius unicornis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2670E77734F255AFE08FA01FDE3C011" box="[426,520,1891,1917]" pageId="3" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9F6F1E734F255AFF35FAE7FF12C0F2" blockId="3.[151,520,1891,1950]" box="[151,249,1925,1950]" pageId="3" pageNumber="144">
|
||
<figureCitation id="131B739B734F255AFF35FAE7FF12C0F2" box="[151,249,1925,1950]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1030,1052]" captionStartId-1="2.[151,249,1778,1800]" captionStartId-2="3.[151,250,1774,1796]" captionStartId-3="4.[151,249,1769,1791]" captionTargetBox-0="[167,1430,341,988]" captionTargetBox-1="[151,1436,195,1753]" captionTargetBox-2="[151,1436,663,1749]" captionTargetBox-3="[174,1412,462,1722]" captionTargetId-0="figure@1.[151,1436,339,1006]" captionTargetId-1="figure@2.[151,1436,195,1754]" captionTargetId-2="figure@3.[151,1436,663,1749]" captionTargetId-3="figure@4.[151,1436,447,1745]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Habitus of Koponenius unicornis sp. nov., male paratype. A: dorsal view. B: lateral view. Pictures by K. Makarov, not taken to scale." captionText-1="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." captionText-2="FIGURE 3. Koponenius unicornis sp. nov., male paratype. A – D: gonopods, lateral, ventral, lateral and mesal views, respectively. Scale bars: B, 0.5 mm; D, 0.2 mm; A, C, 0.1 mm. Designations explained in text." captionText-3="FIGURE 4. Koponenius unicornis sp. nov., male paratype. A: leg 7, lateral view. B: left gonopod, mesal view. Scale bar: 0.3 mm. Designations explained in text." httpUri-0="https://zenodo.org/record/229192/files/figure.png" httpUri-1="https://zenodo.org/record/229193/files/figure.png" httpUri-2="https://zenodo.org/record/229194/files/figure.png" httpUri-3="https://zenodo.org/record/229195/files/figure.png" pageId="3" pageNumber="144">Figs 1–4</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A3C95734F255DFF35FAACFEEFC671" lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144" type="materials_examined">
|
||
<paragraph id="8B9F6F1E734F255DFF35FAACFB2AC7B9" blockId="3.[151,1436,1998,2023]" lastBlockId="4.[151,1436,151,392]" lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144">
|
||
<emphasis id="B954B30C734F255AFF35FAACFE62C08B" bold="true" box="[151,393,1998,2023]" pageId="3" pageNumber="144">Material examined:</emphasis>
|
||
<typeStatus id="549BD1BC734F255AFE35FAACFE14C08B" box="[407,511,1998,2023]" pageId="3" pageNumber="144" type="holotype">Holotype</typeStatus>
|
||
male (
|
||
<collectionCode id="ED31F7DB734F255AFDFBFAADFD5DC08B" box="[601,694,1999,2023]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="3" pageNumber="144">ZMUM</collectionCode>
|
||
),
|
||
<collectingCountry id="F3372F8E734F255AFD72FAACFCE7C08B" box="[720,780,1998,2023]" name="India" pageId="3" pageNumber="144">India</collectingCountry>
|
||
, West Bengal, Darjeeling District, Kalimpong (Gandhi Ashram),
|
||
<geoCoordinate id="EE1409D97348255DFEA9FDF5FE81C7DC" box="[267,362,151,176]" direction="north" orientation="latitude" pageId="4" pageNumber="145" precision="555" value="27.06">27,06ºN</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1409D97348255DFEDBFDF5FE3EC7DC" box="[377,469,151,176]" direction="east" orientation="longitude" pageId="4" pageNumber="145" precision="555" value="88.45">88,45ºE</geoCoordinate>
|
||
, ca
|
||
<quantity id="4CD8C2FB7348255DFDABFDFAFDBFC7C3" box="[521,596,152,176]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="4" pageNumber="145" unit="m" value="880.0">880 m</quantity>
|
||
a.s.l.,
|
||
<date id="FF9E49DE7348255DFD04FDF5FC4EC7DC" box="[678,933,151,176]" pageId="4" pageNumber="145" value="2013-11-05" valueMax="2013-11-15" valueMin="2013-11-05">5–15 November 2013</date>
|
||
, leg. K. Tomkovich.
|
||
<typeStatus id="549BD1BC7348255DFB03FDFAFAF8C7DC" box="[1185,1299,152,176]" pageId="4" pageNumber="145" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D26A4977348255DFA83FDFAFA94C7DC" box="[1313,1407,151,176]" pageId="4" pageNumber="145" type="male">7 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D26A4977348255DFA2CFDFAFF05C7B9" pageId="4" pageNumber="145" type="female">8 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED31F7DB7348255DFF5FFDDEFEB1C7B8" box="[253,346,188,212]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="4" pageNumber="145">ZMUM</collectionCode>
|
||
),
|
||
<specimenCount id="9D26A4977348255DFECFFDDEFE50C7B9" box="[365,443,188,213]" pageId="4" pageNumber="145" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D26A4977348255DFE64FDDEFDCCC7B9" box="[454,551,188,213]" pageId="4" pageNumber="145" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="ED31F7DB7348255DFD94FDDEFD98C7B8" LSID="urn:lsid:biocol.org:col:34838" box="[566,627,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="145">SMF</collectionCode>
|
||
),
|
||
<specimenCount id="9D26A4977348255DFD25FDDEFD36C7B9" box="[647,733,188,213]" pageId="4" pageNumber="145" type="male">2 males</specimenCount>
|
||
(SEM), same data, together with
|
||
<typeStatus id="549BD1BC7348255DFBF5FDDEFB56C7B9" box="[1111,1213,188,213]" pageId="4" pageNumber="145" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F6F1E7348255DFF65FDBDFEEFC671" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="145">
|
||
<emphasis id="B954B30C7348255DFF65FDBDFEFDC794" bold="true" box="[199,278,223,248]" pageId="4" pageNumber="145">Name:</emphasis>
|
||
To emphasize the conspicuous central projection at the fore margin of the collum in both sexes; adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A3C957348255DFF65FC45FE93C6E4" pageId="4" pageNumber="145" type="diagnosis">
|
||
<paragraph id="8B9F6F1E7348255DFF65FC45FE93C6E4" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="145">
|
||
<emphasis id="B954B30C7348255DFF65FC45FEA8C62C" bold="true" box="[199,323,295,320]" pageId="4" pageNumber="145">Diagnosis:</emphasis>
|
||
Differs clearly from the only other known congener,
|
||
<taxonomicName id="4C20149D7348255DFC1EFC4BFBD6C62C" box="[956,1085,296,320]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="biramus" status="sp. nov.">
|
||
<emphasis id="B954B30C7348255DFC1EFC4BFBD6C62C" box="[956,1085,296,320]" italics="true" pageId="4" pageNumber="145">K. biramus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2670E777348255DFBE5FC4AFB4FC653" box="[1095,1188,296,319]" pageId="4" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, in the presence of a highly conspicuous central process at the fore margin of the collum in both sexes, coupled with a uniramous gonopod telopodite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5F3F967348255DFF35FB8BFDEEC072" httpUri="https://zenodo.org/record/229195/files/figure.png" pageId="4" pageNumber="145" targetBox="[174,1412,462,1722]" targetPageId="4">
|
||
<paragraph id="8B9F6F1E7348255DFF35FB8BFDEEC072" blockId="4.[151,1436,1769,1822]" pageId="4" pageNumber="145">
|
||
<emphasis id="B954B30C7348255DFF35FB8BFEF8C192" bold="true" box="[151,275,1769,1791]" pageId="4" pageNumber="145">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C20149D7348255DFEB9FB88FE10C193" box="[283,507,1770,1791]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
|
||
<emphasis id="B954B30C7348255DFEB9FB88FE10C193" box="[283,507,1770,1791]" italics="true" pageId="4" pageNumber="145">Koponenius unicornis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2670E777348255DFDA0FB88FDB8C193" box="[514,595,1770,1791]" pageId="4" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male paratype. A: leg 7, lateral view. B: left gonopod, mesal view. Scale bar: 0.3 mm. Designations explained in text.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33A3C957348255EFF65FA27FD51C671" lastPageId="7" lastPageNumber="148" pageId="4" pageNumber="145" type="description">
|
||
<paragraph id="8B9F6F1E7348255DFF65FA27FA77C082" blockId="4.[151,1436,1861,2030]" pageId="4" pageNumber="145">
|
||
<emphasis id="B954B30C7348255DFF65FA27FEB0C032" bold="true" box="[199,347,1861,1886]" pageId="4" pageNumber="145">Description:</emphasis>
|
||
Length in both sexes ca
|
||
<quantity id="4CD8C2FB7348255DFD24FA24FCEFC031" box="[646,772,1862,1886]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="4" pageNumber="145" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
|
||
, width ca 1.3–1.5 and 1.8–2.0 mm on midbody pro- and metazonae, respectively.
|
||
<typeStatus id="549BD1BC7348255DFE1EFA0BFDCFC0EE" box="[444,548,1897,1922]" pageId="4" pageNumber="145" type="holotype">Holotype</typeStatus>
|
||
ca
|
||
<quantity id="4CD8C2FB7348255DFDFAFA08FD46C0ED" box="[600,685,1898,1922]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="145" unit="mm" value="13.0">13 mm</quantity>
|
||
long, 1.3 and
|
||
<quantity id="4CD8C2FB7348255DFCC4FA08FC29C0ED" box="[870,962,1898,1922]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="145" unit="mm" value="1.8">1.8 mm</quantity>
|
||
wide on midbody pro- and metazonae, respectively. Coloration in alcohol mostly brown to dark grey-brown, including vertigial region of head, as well as most of pleurosternal and midsternal regions, but pattern evident due to contrasting pale, whitish to light greyish clypeolabral and occipital regions of head, antennae, prozonae, porosteles, legs, epi-, hypo- and paraprocts (
|
||
<figureCitation id="131B739B7348255DFAE8FAB7FA60C082" box="[1354,1419,2005,2030]" captionStart="FIGURE 1" captionStartId="1.[151,250,1030,1052]" captionTargetBox="[167,1430,341,988]" captionTargetId="figure@1.[151,1436,339,1006]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Koponenius unicornis sp. nov., male paratype. A: dorsal view. B: lateral view. Pictures by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229192/files/figure.png" pageId="4" pageNumber="145">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F6F1E7349255CFF35FDF5FF2CC6E4" blockId="5.[151,1436,151,392]" pageId="5" pageNumber="146">
|
||
Body with 19 segments in both sexes, subcylindrical, not capable of volvation. Collum and following metaterga clothed with a dense, dull, microvillose cerategument crust (
|
||
<figureCitation id="131B739B7349255CFC99FDDEFC96C7B8" box="[827,893,188,213]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
M–P); vertigial region down to a well-expressed border with clypeolabral region clearly granular (
|
||
<figureCitation id="131B739B7349255CFD43FDBDFCC1C794" box="[737,810,223,248]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
D). Clypeolabral region densely setose (
|
||
<figureCitation id="131B739B7349255CFAAAFDBDFABAC794" box="[1288,1361,223,248]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
D, Q). Antennae short and clavate,
|
||
<emphasis id="B954B30C7349255CFE42FC67FDC2C670" box="[480,553,261,284]" italics="true" pageId="5" pageNumber="146">in situ</emphasis>
|
||
each placed inside a deep, transverse, nearly C-shaped groove (
|
||
<figureCitation id="131B739B7349255CFAB0FC66FAB3C670" box="[1298,1368,260,285]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
D); in length, antennomere 6> 2> 1> 3=4=5; both antennomeres 5 and, especially, 6 with a tight dorso-apical group of bacilliform sensilla (
|
||
<figureCitation id="131B739B7349255CFE21FC2EFE22C608" box="[387,457,332,357]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
A, D, L). Ventral surface of cardo and gnathochilarium largely finely squamate (
|
||
<figureCitation id="131B739B7349255CFACDFC2EFF4DC6E4" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
L).
|
||
</paragraph>
|
||
<caption id="DF5F3F967349255CFF35F850FA68C224" box="[151,1411,1330,1352]" httpUri="https://zenodo.org/record/229196/files/figure.png" pageId="5" pageNumber="146" targetBox="[179,1392,465,1290]" targetPageId="5">
|
||
<paragraph id="8B9F6F1E7349255CFF35F850FA68C224" blockId="5.[151,1411,1330,1352]" box="[151,1411,1330,1352]" pageId="5" pageNumber="146">
|
||
<emphasis id="B954B30C7349255CFF35F850FEF9C22B" bold="true" box="[151,274,1330,1352]" pageId="5" pageNumber="146">FIGURE 5.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C20149D7349255CFE2BF851FDB0C224" box="[393,603,1330,1352]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="biramus" status="sp. nov.">
|
||
<emphasis id="B954B30C7349255CFE2BF851FDB0C224" box="[393,603,1330,1352]" italics="true" pageId="5" pageNumber="146">Koponenius biramus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2670E777349255CFDC3F851FD59C224" box="[609,690,1331,1352]" pageId="5" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male paratype, lateral view. Picture by K. Makarov, not taken to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F6F1E7349255CFF65F812FBA4C030" blockId="5.[151,1437,1391,2028]" pageId="5" pageNumber="146">
|
||
In width, head <collum = segment 3 = 4 <2 = 5=16, thereafter body gradually tapering towards telson (
|
||
<figureCitation id="131B739B7349255CFACDF80DFF4EC2C0" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
H–J). Collum at fore margin dentate, with a highly conspicuous, central, tuberculate protuberance overhanging the head and concealing it from above (
|
||
<figureCitation id="131B739B7349255CFDF5F8D5FD72C2BC" box="[599,665,1463,1488]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
A, D, H). Paraterga rather well developed, strongly declined, mostly wing-shaped, set low (at about lower ¼ of midbody height), starting from collum, laterally vaguely lobulate (
|
||
<figureCitation id="131B739B7349255CFAF4F8B9FA77C298" box="[1366,1436,1499,1524]" captionStart="FIGURE 1" captionStartId="1.[151,250,1030,1052]" captionTargetBox="[167,1430,341,988]" captionTargetId="figure@1.[151,1436,339,1006]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Koponenius unicornis sp. nov., male paratype. A: dorsal view. B: lateral view. Pictures by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229192/files/figure.png" pageId="5" pageNumber="146">Figs 1</figureCitation>
|
||
&
|
||
<figureCitation id="131B739B7349255CFF17FB62FF28C174" box="[181,195,1536,1560]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">2</figureCitation>
|
||
A–J, M, P). Paraterga 2 clearly enlarged, subtending the head on both sides. Dorsum very convex (
|
||
<figureCitation id="131B739B7349255CFA8DF89DFA9FC174" box="[1327,1396,1535,1560]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
M). Postcollum metaterga usually with 4, rarely 5, regular, transverse rows of rather flat, sometimes rather clearly obliterated, setigerous, isostictic (= regular in axial direction), mostly roundish bosses or tubercles (
|
||
<figureCitation id="131B739B7349255CFB4BFB25FAC4C10C" box="[1257,1327,1607,1632]" captionStart="FIGURE 1" captionStartId="1.[151,250,1030,1052]" captionTargetBox="[167,1430,341,988]" captionTargetId="figure@1.[151,1436,339,1006]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Koponenius unicornis sp. nov., male paratype. A: dorsal view. B: lateral view. Pictures by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229192/files/figure.png" pageId="5" pageNumber="146">Figs 1</figureCitation>
|
||
&
|
||
<figureCitation id="131B739B7349255CFAF0FB2AFA8BC10C" box="[1362,1376,1608,1632]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">2</figureCitation>
|
||
A–C, H–J); caudal row of particularly elongate bosses forming a faint waist (
|
||
<figureCitation id="131B739B7349255CFC1EFB09FBEAC1E8" box="[956,1025,1643,1668]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">
|
||
Fig.
|
||
<date id="FF9E49DE7349255CFC53FB0EFBEAC1E8" box="[1009,1025,1644,1668]" pageId="5" pageNumber="146">2</date>
|
||
</figureCitation>
|
||
I), a series of lobulations and ending up in a distinct crenulate limbus (
|
||
<figureCitation id="131B739B7349255CFDBFFBEDFD88C1C4" box="[541,611,1679,1704]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">
|
||
Fig.
|
||
<date id="FF9E49DE7349255CFDF6FBF2FD88C1C4" box="[596,611,1680,1704]" pageId="5" pageNumber="146">2</date>
|
||
</figureCitation>
|
||
I, N); fore row likewise forming a row of similar lobulations (
|
||
<figureCitation id="131B739B7349255CFA91FBEDFA96C1C4" box="[1331,1405,1679,1704]" captionStart="FIGURE 1" captionStartId="1.[151,250,1030,1052]" captionTargetBox="[167,1430,341,988]" captionTargetId="figure@1.[151,1436,339,1006]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Koponenius unicornis sp. nov., male paratype. A: dorsal view. B: lateral view. Pictures by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229192/files/figure.png" pageId="5" pageNumber="146">Figs 1</figureCitation>
|
||
&
|
||
<figureCitation id="131B739B7349255CFF35FBD6FF4EC1A0" box="[151,165,1716,1740]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">2</figureCitation>
|
||
A–C, H–J, N, P). Caudolateral corner of postcollum paraterga mostly well rounded, nearly pointed only in segments 17 and 18 (
|
||
<figureCitation id="131B739B7349255CFE36FBB5FE37C19C" box="[404,476,1751,1776]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
J, K). Ozopores borne on conspicuous dorsolateral porosteles, these lying close to caudolateral corners of paraterga; pore formula 5, 7–18 (
|
||
<figureCitation id="131B739B7349255CFCB8FB99FC8AC078" box="[794,865,1787,1812]" captionStart="FIGURE 1" captionStartId="1.[151,250,1030,1052]" captionTargetBox="[167,1430,341,988]" captionTargetId="figure@1.[151,1436,339,1006]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Koponenius unicornis sp. nov., male paratype. A: dorsal view. B: lateral view. Pictures by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229192/files/figure.png" pageId="5" pageNumber="146">Figs 1</figureCitation>
|
||
&
|
||
<figureCitation id="131B739B7349255CFC25FB9EFC7EC078" box="[903,917,1788,1812]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">2</figureCitation>
|
||
A–C, I, M,
|
||
<collectingCountry id="F3372F8E7349255CFBB8FB9EFBDBC078" box="[1050,1072,1788,1812]" name="Iceland" pageId="5" pageNumber="146">O</collectingCountry>
|
||
, P). Tergal setae filiform, rather long (
|
||
<figureCitation id="131B739B7349255CFF7FFA7DFECDC054" box="[221,294,1823,1848]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
<collectingCountry id="F3372F8E7349255CFE84FA42FED7C054" box="[294,316,1824,1848]" name="Iceland" pageId="5" pageNumber="146">O</collectingCountry>
|
||
, P). Epiproct strongly flattened dorsoventrally, lobulated laterally, subtruncate caudally, but tip ventral, invisible from above (
|
||
<figureCitation id="131B739B7349255CFE48FA21FDC6C030" box="[490,557,1859,1884]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
C, G, J, K). Hypoproct subtrapeziform (
|
||
<figureCitation id="131B739B7349255CFC4AFA21FBC0C030" box="[1000,1067,1859,1884]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
G).
|
||
</paragraph>
|
||
<paragraph id="8B9F6F1E7349255CFF65FA0AFC93C080" blockId="5.[151,1437,1391,2028]" pageId="5" pageNumber="146">
|
||
Sterna narrow, but evident, mostly slightly elevated due to small subtriangular lobules observed between both coxae (
|
||
<figureCitation id="131B739B7349255CFF4AFAE9FEC0C0C8" box="[232,299,1931,1956]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">Fig. 2</figureCitation>
|
||
D–F). Spiracles seem to be absent. Legs mostly tightly appressed to venter, densely setose, short and stout, about as long as body height and subequally incrassate in both sexes (
|
||
<figureCitation id="131B739B7349255CFC42FACDFBCDC0A4" box="[992,1062,1967,1992]" captionStart="FIGURE 1" captionStartId="1.[151,250,1030,1052]" captionTargetBox="[167,1430,341,988]" captionTargetId="figure@1.[151,1436,339,1006]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Koponenius unicornis sp. nov., male paratype. A: dorsal view. B: lateral view. Pictures by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229192/files/figure.png" pageId="5" pageNumber="146">Figs 1</figureCitation>
|
||
&
|
||
<figureCitation id="131B739B7349255CFBEBFAD2FBBCC0A4" box="[1097,1111,1968,1992]" captionStart="FIGURE 2" captionStartId="2.[151,249,1778,1800]" captionTargetBox="[151,1436,195,1753]" captionTargetId="figure@2.[151,1436,195,1754]" captionTargetPageId="2" captionText="FIGURE 2. Koponenius unicornis sp. nov., male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N – P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J – L, 0.2 mm; P – R, 0.1 mm; N, 0.05 mm; O, 0.02 mm." httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="5" pageNumber="146">2</figureCitation>
|
||
B–F, M, Q, R), but only male coxae and, to a lesser degree, prefemora micropapillate (
|
||
<figureCitation id="131B739B7349255CFCB2FAB1FCBFC080" box="[784,852,2003,2028]" captionStart="FIGURE 4" captionStartId="4.[151,249,1769,1791]" captionTargetBox="[174,1412,462,1722]" captionTargetId="figure@4.[151,1436,447,1745]" captionTargetPageId="4" captionText="FIGURE 4. Koponenius unicornis sp. nov., male paratype. A: leg 7, lateral view. B: left gonopod, mesal view. Scale bar: 0.3 mm. Designations explained in text." httpUri="https://zenodo.org/record/229195/files/figure.png" pageId="5" pageNumber="146">Fig. 4</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<caption id="DF5F3F96734A255FFF35FB81FB39C039" httpUri="https://zenodo.org/record/229197/files/figure.png" pageId="6" pageNumber="147" targetBox="[151,1436,195,1737]" targetPageId="6">
|
||
<paragraph id="8B9F6F1E734A255FFF35FB81FB39C039" blockId="6.[151,1437,1763,1877]" pageId="6" pageNumber="147">
|
||
<emphasis id="B954B30C734A255FFF35FB81FEF2C194" bold="true" box="[151,281,1763,1785]" pageId="6" pageNumber="147">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C20149D734A255FFE84FB86FE14C195" box="[294,511,1763,1785]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="biramus" status="sp. nov.">
|
||
<emphasis id="B954B30C734A255FFE84FB86FE14C195" box="[294,511,1763,1785]" italics="true" pageId="6" pageNumber="147">Koponenius biramus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2670E77734A255FFDAEFB86FD8FC195" box="[524,612,1764,1785]" pageId="6" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male paratype. A, D, G, J: anterior body part, dorsal, lateral, ventral and frontoventral views, respectively. B, E, H: midbody segments, dorsal, lateral and ventral views, respectively. C, F, I: posterior body part, dorsal, lateral and ventral views, respectively. K: cross-section of a midbody segment. L: tergal texture, dorsal view. M: ozopore region, dorsolateral view. Scale bars: H, 0.5 mm; A–G, I–L, 0.2 mm; N, 0.05 mm; L, 0.02 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F6F1E734A255EFF65FA1EFD51C671" blockId="6.[151,1436,1916,2013]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="148" pageId="6" pageNumber="147">
|
||
Gonopods (
|
||
<figureCitation id="131B739B734A255FFEECFA1EFE71C0F9" box="[334,410,1916,1941]" captionStart="FIGURE 3" captionStartId="3.[151,250,1774,1796]" captionTargetBox="[151,1436,663,1749]" captionTargetId="figure@3.[151,1436,663,1749]" captionTargetPageId="3" captionText="FIGURE 3. Koponenius unicornis sp. nov., male paratype. A – D: gonopods, lateral, ventral, lateral and mesal views, respectively. Scale bars: B, 0.5 mm; D, 0.2 mm; A, C, 0.1 mm. Designations explained in text." httpUri="https://zenodo.org/record/229194/files/figure.png" pageId="6" pageNumber="147">Figs 3</figureCitation>
|
||
&
|
||
<figureCitation id="131B739B734A255FFE65FA1FFE3DC0F9" box="[455,470,1917,1941]" captionStart="FIGURE 4" captionStartId="4.[151,249,1769,1791]" captionTargetBox="[174,1412,462,1722]" captionTargetId="figure@4.[151,1436,447,1745]" captionTargetPageId="4" captionText="FIGURE 4. Koponenius unicornis sp. nov., male paratype. A: leg 7, lateral view. B: left gonopod, mesal view. Scale bar: 0.3 mm. Designations explained in text." httpUri="https://zenodo.org/record/229195/files/figure.png" pageId="6" pageNumber="147">4</figureCitation>
|
||
B) with large, subcylindrical, mediobasally fused coxae carrying a few long setae ventrally and a small, but evident, rounded, lateral lobe apically; cannulae long and slender. Telopodite strongly elongated, considerably longer than coxae, slender; prefemoral (= densely setose) portion suberect, about half as long as entire telopodite, set off from a subfalcate, rather gradually attenuating and narrowly truncate acropodite (= solenomere,
|
||
<emphasis id="B954B30C734B255EFE84FDDEFED2C7B9" bold="true" box="[294,313,188,213]" pageId="7" pageNumber="148">sl</emphasis>
|
||
) by a strong sulcus (
|
||
<emphasis id="B954B30C734B255EFD86FDDEFDC4C7B9" bold="true" box="[548,559,188,213]" pageId="7" pageNumber="148">s</emphasis>
|
||
) on mesal face and by a ventral projection (
|
||
<emphasis id="B954B30C734B255EFBBCFDDEFBC6C7B9" bold="true" box="[1054,1069,188,213]" pageId="7" pageNumber="148">p</emphasis>
|
||
); seminal groove rapidly shifted laterad due to a twisted prefemoral portion, thereafter running only on lateral side all over
|
||
<emphasis id="B954B30C734B255EFB11FDBDFB2DC794" bold="true" box="[1203,1222,223,248]" pageId="7" pageNumber="148">sl</emphasis>
|
||
extent. Neither an accessory seminal chamber nor a hairy pulvillus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |