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<document id="6DBF979C2D0EC37F9E13BBB6918589D8" ID-DOI="10.5281/zenodo.5330324" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330324" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630166723614" checkinUser="felipe" docAuthor="Starý, Jaroslav" docDate="2012" docId="038287EDFF93F952C18DFE56FE68FD58" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.273-279.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Geranomyia fuscior Starý 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="278" masterDocId="FFBBFF95FF97F957C06EFFE0FFFBFFAB" masterDocTitle="The identity of Geranomyia bezzii and a description of G. fuscior sp. nov. (Diptera: Limoniidae)" masterLastPageNumber="279" masterPageNumber="273" pageNumber="277" updateTime="1699171381103" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="50DA8123B2C502790BCF674391717DCA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F50DF70A70755E2F28F9DDA44E2F87BD">
<mods:title id="3AB1237708A6FFDDDFD8A06FF9405474">The identity of Geranomyia bezzii and a description of G. fuscior sp. nov. (Diptera: Limoniidae)</mods:title>
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<mods:namePart id="3F2B6AE56C1E217EEA5289404DDF6CD6">Starý, Jaroslav</mods:namePart>
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<mods:title id="9DAFB8299D690AC5C6C4D8A7D46C5466">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="0FD63D6B2E69A5E3DAAA46F832719666">
<mods:date id="504B197145DFD6D77466B93CB3DC4E1D">2012</mods:date>
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<mods:number id="A5A050CB3F57233C5F62F9C5391C88F1">2012-06-30</mods:number>
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<mods:number id="2D59AE2FAA08AE98EDCA8E78A9EC1CEB">52</mods:number>
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<mods:classification id="2B24EED636DF67F1DB98BB5646E6A35F">journal article</mods:classification>
<mods:identifier id="7E6B02CD16886723DEAF839D467638FA" type="DOI">10.5281/zenodo.5330324</mods:identifier>
<mods:identifier id="B49D11E0F7CB9613DCC10F047DF90D93" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="5A87124BB56FE395875AB97845D922FA" type="Zenodo-Dep">5330324</mods:identifier>
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<treatment id="038287EDFF93F952C18DFE56FE68FD58" ID-DOI="http://doi.org/10.5281/zenodo.5398017" ID-Zenodo-Dep="5398017" LSID="urn:lsid:plazi:treatment:038287EDFF93F952C18DFE56FE68FD58" httpUri="http://treatment.plazi.org/id/038287EDFF93F952C18DFE56FE68FD58" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277">
<subSubSection id="C3316570FF93F953C18DFE56FCE0FE65" box="[483,795,437,462]" pageId="4" pageNumber="277" type="nomenclature">
<paragraph id="8B9436FBFF93F953C18DFE56FCE0FE65" blockId="4.[483,795,437,462]" box="[483,795,437,462]" pageId="4" pageNumber="277">
<heading id="D0DC8197FF93F953C18DFE56FCE0FE65" bold="true" box="[483,795,437,462]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="277" reason="3">
<emphasis id="B95FEAE9FF93F953C18DFE56FCE0FE65" bold="true" box="[483,795,437,462]" pageId="4" pageNumber="277">
<taxonomicName id="4C2B4D78FF93F953C18DFE56FD44FE65" ID-CoL="3FVG8" authority="Starý, 2012" authorityName="Starý" authorityYear="2012" box="[483,703,437,462]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="fuscior" status="sp. nov.">
<emphasis id="B95FEAE9FF93F953C18DFE56FD44FE65" bold="true" box="[483,703,437,462]" italics="true" pageId="4" pageNumber="277">Geranomyia fuscior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C5792FF93F953C2A8FE56FCE0FE65" box="[710,795,438,462]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3316570FF93F953C23EFE34FD55FE4C" box="[592,686,468,487]" pageId="4" pageNumber="277" type="description">
<paragraph id="8B9436FBFF93F953C23EFE34FD55FE4C" blockId="4.[151,1127,468,663]" box="[592,686,468,487]" pageId="4" pageNumber="277">
(
<figureCitation id="13102A7EFF93F953C238FE34FD5CFE4C" box="[598,679,468,487]" captionStart="Figs" captionStartId="3.[115,154,897,916]" captionTargetBox="[129,1073,223,872]" captionTargetId="figure-247@3.[125,1078,219,877]" captionTargetPageId="3" captionText="Figs. 1–7.1–3 – Geranomyia bezzii Alexander &amp; Leonard, 1912 (Portugal:Alvor): 1 – male terminalia, general view, dorsally; 2 – male terminalia, aedeagal complex, lateral view; 3 – female terminalia, internal structures, ventral view. 4–6 – G. fuscior sp. nov. (paratypes; Portugal: Castro Marim): 4 – male terminalia, general view, dorsally; 5 – male terminalia, aedeagal complex, lateral view; 6 – female terminalia, internal structures, ventral view. 7 – G. unicolor Haliday, 1833 (England: Devon: Beer, coll. JSOC), male terminalia, gonostyli, dorsal view. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5330328" httpUri="https://zenodo.org/record/5330328/files/figure.png" pageId="4" pageNumber="277">Figs. 4–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3316570FF93F953C0F9FE1DFC83FD3C" pageId="4" pageNumber="277" type="materials_examined">
<paragraph id="8B9436FBFF93F953C0F9FE1DFC9FFD87" blockId="4.[151,1127,468,663]" pageId="4" pageNumber="277">
<emphasis id="B95FEAE9FF93F953C0F9FE1DFEB2FDBA" bold="true" box="[151,329,509,529]" pageId="4" pageNumber="277">Material examined.</emphasis>
<materialsCitation id="3B433CA6FF93F953C120FE1FFEACFD87" collectingDate="2009-04-02" collectionCode="SMOC" collectorName="J. Stary" country="Portugal" elevation="24" location="Castro Marim" pageId="4" pageNumber="277" specimenCount="1" typeStatus="holotype">
<typeStatus id="54908859FF93F953C120FE1FFE59FDBB" box="[334,418,511,530]" pageId="4" pageNumber="277" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED3AAE3EFF93F953C1A6FE1EFDF1FDBA" box="[456,522,510,529]" country="Czech Republic" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="4" pageNumber="277">SMOC</collectionCode>
):
<emphasis id="B95FEAE9FF93F953C276FE1DFD09FDBA" bold="true" box="[536,754,509,529]" pageId="4" pageNumber="277">
<collectingCountry id="F33C766BFF93F953C276FE1DFD6FFDBA" box="[536,660,509,529]" name="Portugal" pageId="4" pageNumber="277">PORTUGAL</collectingCountry>
: ALGARVE:
</emphasis>
<location id="8EF46020FF93F953C299FE1EFC8BFDBA" LSID="urn:lsid:plazi:treatment:038287EDFF93F952C18DFE56FE68FD58:8EF46020FF93F953C299FE1EFC8BFDBA" box="[759,880,510,529]" country="Portugal" name="Castro Marim" pageId="4" pageNumber="277">Castro Marim</location>
, 0.3 km S, saltmarsh (
<quantity id="4CD39B1EFF93F953C440FE1EFBA7FDBA" box="[1070,1116,510,529]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="277" unit="m" value="24.0">
<elevation id="0006D1C8FF93F953C440FE1EFBA7FDBA" box="[1070,1116,510,529]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="277" unit="m" value="24.0">24 m</elevation>
</quantity>
),
<date id="FF95103BFF93F953C0F9FDF9FF13FD87" box="[151,232,537,556]" pageId="4" pageNumber="277" value="2009-04-02">
<collectingDate id="EFD1E9D3FF93F953C0F9FDF9FF13FD87" box="[151,232,537,556]" pageId="4" pageNumber="277" value="2009-04-02">2.iv.2009</collectingDate>
</date>
,
<collectorName id="26DE532DFF93F953C09FFDF9FECAFD87" box="[241,305,537,556]" pageId="4" pageNumber="277">J. Starý</collectorName>
leg.
</materialsCitation>
/
<typeStatus id="54908859FF93F953C108FDF9FE22FD87" box="[358,473,537,556]" pageId="4" pageNumber="277" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C2B4D78FF93F953C18CFDFAFD76FD87" authority="Starý, 2012" authorityName="Starý" authorityYear="2012" box="[482,653,538,556]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="fuscior" status="sp. nov.">
<emphasis id="B95FEAE9FF93F953C18CFDFAFD76FD87" box="[482,653,538,556]" italics="true" pageId="4" pageNumber="277">Geranomyia fuscior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C5792FF93F953C2FCFDF9FD3AFD87" box="[658,705,537,556]" pageId="4" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
[J], J. Starý 2011.
</paragraph>
<paragraph id="8B9436FBFF93F953C0DBFDD4FC83FD3C" blockId="4.[151,1127,468,663]" pageId="4" pageNumber="277">
<typeStatus id="54908859FF93F953C0DBFDD4FEF6FDEE" box="[181,269,564,583]" pageId="4" pageNumber="277" type="paratype">PARATYPES</typeStatus>
(20 JJ
<specimenCount id="9D2DFD72FF93F953C136FDD4FE75FDED" box="[344,398,562,583]" pageId="4" pageNumber="277" type="female">12 ♀♀</specimenCount>
):
<emphasis id="B95FEAE9FF93F953C1CFFDD3FDF0FDEC" bold="true" box="[417,523,563,583]" pageId="4" pageNumber="277">
<collectingCountry id="F33C766BFF93F953C1CFFDD3FDFCFDEC" box="[417,519,563,583]" name="Albania" pageId="4" pageNumber="277">ALBANIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49EFF819FF93F953C27CFDD4FDA1FDEC" box="[530,602,564,583]" country="Albania" name="Durres" pageId="4" pageNumber="277">Durazzo</collectingRegion>
[=
<collectingRegion id="49EFF819FF93F953C212FDD4FD41FDEC" box="[636,698,564,583]" country="Albania" name="Durres" pageId="4" pageNumber="277">Durrës</collectingRegion>
],
<date id="FF95103BFF93F953C2A4FDD4FCDBFDEC" box="[714,800,564,583]" pageId="4" pageNumber="277" value="1917-05-30">30.v.1917</date>
,
<specimenCount id="9D2DFD72FF93F953C344FDD4FCB3FDED" box="[810,840,562,583]" pageId="4" pageNumber="277" type="female">1 ♀</specimenCount>
, Karny leg. (
<collectionCode id="ED3AAE3EFF93F953C3D1FDD4FBF3FDEC" box="[959,1032,564,583]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="277" type="Museum">NHMW</collectionCode>
) [listed by
<bibRefCitation id="EFBA4B0AFF93F953C0F9FDAFFEB4FDCA" author="LACKSCHEWITZ P." box="[151,335,590,610]" pageId="4" pageNumber="277" pagination="195 - 244" refId="ref3727" refString="LACKSCHEWITZ P. 1928: Die palaearktischen Limnobiinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien 42: 195 - 244." type="journal article" year="1928">LACKSCHEWITZ (1928)</bibRefCitation>
as
<taxonomicName id="4C2B4D78FF93F953C100FDAFFE64FDCA" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[366,415,591,609]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C100FDAFFE64FDCA" box="[366,415,591,609]" italics="true" pageId="4" pageNumber="277">bezzii</emphasis>
</taxonomicName>
].
<emphasis id="B95FEAE9FF93F953C1DFFDADFD76FDCA" bold="true" box="[433,653,589,609]" pageId="4" pageNumber="277">
<collectingCountry id="F33C766BFF93F953C1DFFDADFDD6FDCA" box="[433,557,589,609]" name="Portugal" pageId="4" pageNumber="277">PORTUGAL</collectingCountry>
: ALGARVE:
</emphasis>
Castro Marim, 0.3 km S, saltmarsh (
<quantity id="4CD39B1EFF93F953C3BFFDAEFBFAFDCA" box="[977,1025,590,609]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="277" unit="m" value="24.0">24 m</quantity>
),
<date id="FF95103BFF93F953C47CFDAEFB98FDCA" box="[1042,1123,590,609]" pageId="4" pageNumber="277" value="2009-04-02">2.iv.2009</date>
, 10 JJ
<specimenCount id="9D2DFD72FF93F953C0BCFD89FF00FDD0" box="[210,251,615,636]" pageId="4" pageNumber="277" type="female">5 ♀♀</specimenCount>
,
<date id="FF95103BFF93F953C169FD89FEA3FDD7" box="[263,344,617,636]" pageId="4" pageNumber="277" value="2009-04-04">4.iv.2009</date>
, 10 JJ
<specimenCount id="9D2DFD72FF93F953C1F2FD89FE3DFDD0" box="[412,454,615,636]" pageId="4" pageNumber="277" type="female">5 ♀♀</specimenCount>
, J. Starý leg. (
<collectionCode id="ED3AAE3EFF93F953C22DFD89FD83FDD7" box="[579,632,617,636]" pageId="4" pageNumber="277">JKLE</collectionCode>
,
<collectionCode id="ED3AAE3EFF93F953C2ECFD89FD4DFDD7" box="[642,694,617,636]" pageId="4" pageNumber="277">JSOC</collectionCode>
,
<collectionCode id="ED3AAE3EFF93F953C2D1FD89FD05FDD7" box="[703,766,617,636]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="277">NMPC</collectionCode>
,
<collectionCode id="ED3AAE3EFF93F953C369FD89FCBCFDD7" box="[775,839,617,636]" country="Czech Republic" httpUri="http://grbio.org/cool/42jk-3h2c" name="Slezske Muzeum Opava" pageId="4" pageNumber="277">SMOC</collectionCode>
).
<emphasis id="B95FEAE9FF93F953C338FD88FC64FDD7" bold="true" box="[854,927,616,636]" pageId="4" pageNumber="277">
<collectingCountry id="F33C766BFF93F953C338FD88FC60FDD7" box="[854,923,616,636]" name="Libya" pageId="4" pageNumber="277">LIBYA</collectingCountry>
:
</emphasis>
Tripolis [=
<collectingRegion id="49EFF819FF93F953C468FD89FBA5FDD7" box="[1030,1118,617,636]" country="Libya" name="Tarabulus" pageId="4" pageNumber="277">Tarabulus</collectingRegion>
],
<date id="FF95103BFF93F953C0F9FD64FF01FD3C" box="[151,250,644,663]" pageId="4" pageNumber="277" value="1906-07-17">17.vii.1906</date>
,
<specimenCount id="9D2DFD72FF93F953C16DFD64FEE4FD3D" box="[259,287,642,663]" pageId="4" pageNumber="277" type="female">1 ♀</specimenCount>
, A. Klaptocz leg. (
<collectionCode id="ED3AAE3EFF93F953C1AEFD64FDF2FD3C" box="[448,521,644,663]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="277" type="Museum">NHMW</collectionCode>
) [listed by
<bibRefCitation id="EFBA4B0AFF93F953C207FD64FCE4FD3C" author="LACKSCHEWITZ P." box="[617,799,644,663]" pageId="4" pageNumber="277" pagination="195 - 244" refId="ref3727" refString="LACKSCHEWITZ P. 1928: Die palaearktischen Limnobiinen (Diptera) des Wiener Naturhistorischen Museums. Annalen des Naturhistorischen Museums in Wien 42: 195 - 244." type="journal article" year="1928">LACKSCHEWITZ (1928)</bibRefCitation>
as
<taxonomicName id="4C2B4D78FF93F953C352FD65FC96FD3C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[828,877,645,663]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C352FD65FC96FD3C" box="[828,877,645,663]" italics="true" pageId="4" pageNumber="277">bezzii</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C3316570FF93F953C0F9FD4FFC38FCEC" pageId="4" pageNumber="277" type="diagnosis">
<paragraph id="8B9436FBFF93F953C0F9FD4FFC38FCEC" blockId="4.[151,1127,687,1575]" pageId="4" pageNumber="277">
<emphasis id="B95FEAE9FF93F953C0F9FD4FFEF7FD6C" bold="true" box="[151,268,687,711]" pageId="4" pageNumber="277">Diagnosis.</emphasis>
Extremely similar to
<taxonomicName id="4C2B4D78FF93F953C181FD50FDB0FD6C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[495,587,688,711]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C181FD50FDB0FD6C" box="[495,587,688,711]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
in general appearance, but with thoracic patterning darker and more extensive. Stripes on mesothoracic dorsum shiny, very dark. Postscutellum mostly brown throughout.
<collectingRegion id="49EFF819FF93F953C1C1FD0FFE26FCAC" box="[431,477,751,775]" country="Algeria" name="Tipaza" pageId="4" pageNumber="277">Tips</collectingRegion>
of femora and tibiae with more pronounced darkenings. Male terminalia with ventral gonostylus distinctly longer and broader than in
<taxonomicName id="4C2B4D78FF93F953C3E8FCF0FC19FC8C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[902,994,784,807]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C3E8FCF0FC19FC8C" box="[902,994,784,807]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
, with rostral prolongation more rounded at tip in dorsal aspect. Wing length 6.2–7.4 mm.
</paragraph>
</subSubSection>
<subSubSection id="C3316570FF93F953C0F9FCAFFE18FB0C" pageId="4" pageNumber="277" type="description">
<paragraph id="8B9436FBFF93F953C0F9FCAFFC64FBAC" blockId="4.[151,1127,687,1575]" pageId="4" pageNumber="277">
<emphasis id="B95FEAE9FF93F953C0F9FCAFFE9DFCCC" bold="true" box="[151,358,846,871]" pageId="4" pageNumber="277">
Description.
<emphasis id="B95FEAE9FF93F953C149FCAEFEA4FCCC" bold="true" box="[295,351,846,871]" italics="true" pageId="4" pageNumber="277">Male</emphasis>
.
</emphasis>
Terminalia (
<figureCitation id="13102A7EFF93F953C182FCAFFDABFCCC" box="[492,592,847,871]" captionStart="Figs" captionStartId="3.[115,154,897,916]" captionTargetBox="[129,1073,223,872]" captionTargetId="figure-247@3.[125,1078,219,877]" captionTargetPageId="3" captionText="Figs. 1–7.1–3 – Geranomyia bezzii Alexander &amp; Leonard, 1912 (Portugal:Alvor): 1 – male terminalia, general view, dorsally; 2 – male terminalia, aedeagal complex, lateral view; 3 – female terminalia, internal structures, ventral view. 4–6 – G. fuscior sp. nov. (paratypes; Portugal: Castro Marim): 4 – male terminalia, general view, dorsally; 5 – male terminalia, aedeagal complex, lateral view; 6 – female terminalia, internal structures, ventral view. 7 – G. unicolor Haliday, 1833 (England: Devon: Beer, coll. JSOC), male terminalia, gonostyli, dorsal view. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5330328" httpUri="https://zenodo.org/record/5330328/files/figure.png" pageId="4" pageNumber="277">Figs. 4–5</figureCitation>
) darkly pigmented. Tergite 9 roughly semicircular. Gonocoxite distinctly shorter than ventral gonostylus; ratio about 2: 3. Ventral gonostylus distinctly longer and broader than in
<taxonomicName id="4C2B4D78FF93F953C275FC70FD8CFC0C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[539,631,912,935]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C275FC70FD8CFC0C" box="[539,631,912,935]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
, rather ovoid, with rostral prolongation shorter and stouter, more rounded at tip in dorsal aspect, and curved rather dorsoanteriorly. Dorsal gonostylus parallel-sided, tapered to point only shortly before apex. Aedeagal complex as in
<figureCitation id="13102A7EFF93F953C0F9FC0FFF2CFBAC" box="[151,215,1007,1031]" captionStart="Figs" captionStartId="3.[115,154,897,916]" captionTargetBox="[129,1073,223,872]" captionTargetId="figure-247@3.[125,1078,219,877]" captionTargetPageId="3" captionText="Figs. 1–7.1–3 – Geranomyia bezzii Alexander &amp; Leonard, 1912 (Portugal:Alvor): 1 – male terminalia, general view, dorsally; 2 – male terminalia, aedeagal complex, lateral view; 3 – female terminalia, internal structures, ventral view. 4–6 – G. fuscior sp. nov. (paratypes; Portugal: Castro Marim): 4 – male terminalia, general view, dorsally; 5 – male terminalia, aedeagal complex, lateral view; 6 – female terminalia, internal structures, ventral view. 7 – G. unicolor Haliday, 1833 (England: Devon: Beer, coll. JSOC), male terminalia, gonostyli, dorsal view. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5330328" httpUri="https://zenodo.org/record/5330328/files/figure.png" pageId="4" pageNumber="277">Fig. 5</figureCitation>
, with aedeagus not as strongly downcurved at apex as in
<taxonomicName id="4C2B4D78FF93F953C355FC10FC63FBAC" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[827,920,1008,1031]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C355FC10FC63FBAC" box="[827,920,1008,1031]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9436FBFF93F953C0DBFBEEFE18FB0C" blockId="4.[151,1127,687,1575]" pageId="4" pageNumber="277">
<emphasis id="B95FEAE9FF93F953C0DBFBEEFEF3FB8C" bold="true" box="[181,264,1038,1063]" italics="true" pageId="4" pageNumber="277">Female</emphasis>
resembling male in general appearance. Female terminalia (
<figureCitation id="13102A7EFF93F953C3EEFBEFFC3BFB8C" box="[896,960,1039,1063]" captionStart="Figs" captionStartId="3.[115,154,897,916]" captionTargetBox="[129,1073,223,872]" captionTargetId="figure-247@3.[125,1078,219,877]" captionTargetPageId="3" captionText="Figs. 1–7.1–3 – Geranomyia bezzii Alexander &amp; Leonard, 1912 (Portugal:Alvor): 1 – male terminalia, general view, dorsally; 2 – male terminalia, aedeagal complex, lateral view; 3 – female terminalia, internal structures, ventral view. 4–6 – G. fuscior sp. nov. (paratypes; Portugal: Castro Marim): 4 – male terminalia, general view, dorsally; 5 – male terminalia, aedeagal complex, lateral view; 6 – female terminalia, internal structures, ventral view. 7 – G. unicolor Haliday, 1833 (England: Devon: Beer, coll. JSOC), male terminalia, gonostyli, dorsal view. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5330328" httpUri="https://zenodo.org/record/5330328/files/figure.png" pageId="4" pageNumber="277">Fig. 6</figureCitation>
): Cercus moderate in length, slightly upturned, occasionally slightly longer than that of
<taxonomicName id="4C2B4D78FF93F953C3F8FBD0FC09FBEC" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[918,1010,1072,1095]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C3F8FBD0FC09FBEC" box="[918,1010,1072,1095]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
. Two parts of infra-anal plate generally convex at posterior margin and more rounded at outer edge than in
<taxonomicName id="4C2B4D78FF93F953C0DFFB90FEF5FB2C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[177,270,1136,1159]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C0DFFB90FEF5FB2C" box="[177,270,1136,1159]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
. Genital fork with its central, more pigmented part not as long as in
<taxonomicName id="4C2B4D78FF93F953C3B6FB90FBCFFB2C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[984,1076,1136,1159]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C3B6FB90FBCFFB2C" box="[984,1076,1136,1159]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
and, on average, somewhat broader.
</paragraph>
</subSubSection>
<subSubSection id="C3316570FF93F952C0F9FB4FFC5AFDD8" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277" type="diagnosis">
<paragraph id="8B9436FBFF93F952C0F9FB4FFC5AFDD8" blockId="4.[151,1127,687,1575]" lastBlockId="5.[113,1090,219,755]" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277">
<emphasis id="B95FEAE9FF93F953C0F9FB4FFE69FB6C" bold="true" box="[151,402,1199,1223]" pageId="4" pageNumber="277">Differential diagnosis.</emphasis>
The two species,
<taxonomicName id="4C2B4D78FF93F953C23AFB50FD49FB6C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[596,690,1200,1223]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C23AFB50FD49FB6C" box="[596,690,1200,1223]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2B4D78FF93F953C289FB50FCA8FB6C" authority="Starý, 2012" authorityName="Starý" authorityYear="2012" box="[743,851,1200,1223]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="fuscior" status="sp. nov.">
<emphasis id="B95FEAE9FF93F953C289FB50FCA8FB6C" box="[743,851,1200,1223]" italics="true" pageId="4" pageNumber="277">G. fuscior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C5792FF93F953C334FB50FC4BFB6C" box="[858,944,1200,1223]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
, may differ from each other in a few external characters, such as the patterning of the thorax (paler and less extensive in
<taxonomicName id="4C2B4D78FF93F953C173FB10FE80FAAC" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[285,379,1264,1287]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C173FB10FE80FAAC" box="[285,379,1264,1287]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
), the length of legs (slightly shorter in
<taxonomicName id="4C2B4D78FF93F953C377FB10FC8CFAAC" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[793,887,1264,1287]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C377FB10FC8CFAAC" box="[793,887,1264,1287]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
), and the length of the male tarsal claws (slightly longer in
<taxonomicName id="4C2B4D78FF93F953C248FAF0FD7DFA8C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[550,646,1296,1319]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C248FAF0FD7DFA8C" box="[550,646,1296,1319]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
). None of these features, however, can be a reliable criterion for species separation. Especially the pattern on the thorax varies greatly in intensity and extent in both species (except for the two small marks on the upper part of the pleuron, which are always shining dark brown to blackish), even within specimens from the same locality (see Portuguese material), and it can be diagnostic only for very pale specimens of
<taxonomicName id="4C2B4D78FF93F953C0DAFA50FEEAFA6C" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[180,273,1456,1479]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF93F953C0DAFA50FEEAFA6C" box="[180,273,1456,1479]" italics="true" pageId="4" pageNumber="277">G. bezzii</emphasis>
</taxonomicName>
and very dark ones of
<taxonomicName id="4C2B4D78FF93F953C26BFA50FD94FA6C" authority="Starý, 2012" authorityName="Starý" authorityYear="2012" box="[517,623,1456,1479]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="fuscior" status="sp. nov.">
<emphasis id="B95FEAE9FF93F953C26BFA50FD94FA6C" box="[517,623,1456,1479]" italics="true" pageId="4" pageNumber="277">G. fuscior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C5792FF93F953C218FA50FD32FA6C" box="[630,713,1456,1479]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
A predominantly brown postscutellum may be indicative for
<taxonomicName id="4C2B4D78FF93F953C112FA30FE1EFA4C" authority="Starý, 2012" authorityName="Starý" authorityYear="2012" box="[380,485,1488,1511]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="fuscior" status="sp. nov.">
<emphasis id="B95FEAE9FF93F953C112FA30FE1EFA4C" box="[380,485,1488,1511]" italics="true" pageId="4" pageNumber="277">G. fuscior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C5792FF93F953C185FA30FDC7FA4C" box="[491,572,1488,1511]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
The two species can be best differentiated from each other based on the male terminalia. The ventral gonostylus is the most indicative structure. It is rather large in
<taxonomicName id="4C2B4D78FF93F953C122F9F0FE4CF98C" authority="Starý, 2012" authorityName="Starý" authorityYear="2012" box="[332,439,1552,1575]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="fuscior" status="sp. nov.">
<emphasis id="B95FEAE9FF93F953C122F9F0FE4CF98C" box="[332,439,1552,1575]" italics="true" pageId="4" pageNumber="277">G. fuscior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C5792FF93F953C1D0F9F0FDE8F98C" box="[446,531,1552,1575]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
, distinctly exceeding the length of the gonocoxite (ratio about 3: 2), ovoid, with the rostral prolongation of moderate length, somewhat rounded at the tip in dorsal aspect and curved dorsocaudally. In
<taxonomicName id="4C2B4D78FF92F952C211FF1DFD26FEBF" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[639,733,253,276]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF92F952C211FF1DFD26FEBF" box="[639,733,253,276]" italics="true" pageId="5" pageNumber="278">G. bezzii</emphasis>
</taxonomicName>
, the ventral gonostylus is shorter compared to the gonocoxite (ratio about 11: 9), more slender, rather conical, with the rostral prolongation longer, subacute at the tip in dorsal aspect and curved caudally (cf.
<figureCitation id="13102A7EFF92F952C3A5FEDBFF7AFEDF" captionStart="Figs" captionStartId="3.[115,154,897,916]" captionTargetBox="[129,1073,223,872]" captionTargetId="figure-247@3.[125,1078,219,877]" captionTargetPageId="3" captionText="Figs. 1–7.1–3 – Geranomyia bezzii Alexander &amp; Leonard, 1912 (Portugal:Alvor): 1 – male terminalia, general view, dorsally; 2 – male terminalia, aedeagal complex, lateral view; 3 – female terminalia, internal structures, ventral view. 4–6 – G. fuscior sp. nov. (paratypes; Portugal: Castro Marim): 4 – male terminalia, general view, dorsally; 5 – male terminalia, aedeagal complex, lateral view; 6 – female terminalia, internal structures, ventral view. 7 – G. unicolor Haliday, 1833 (England: Devon: Beer, coll. JSOC), male terminalia, gonostyli, dorsal view. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5330328" httpUri="https://zenodo.org/record/5330328/files/figure.png" pageId="5" pageNumber="278">Figs. 1 and 4</figureCitation>
). In
<taxonomicName id="4C2B4D78FF92F952C0DDFEBDFEE5FEDF" authority="Starý, 2012" authorityName="Starý" authorityYear="2012" box="[179,286,349,372]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="fuscior" status="sp. nov.">
<emphasis id="B95FEAE9FF92F952C0DDFEBDFEE5FEDF" box="[179,286,349,372]" italics="true" pageId="5" pageNumber="278">G. fuscior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C5792FF92F952C14BFEBCFE81FED8" box="[293,378,348,371]" pageId="5" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
, the dorsal gonostylus is tapered to pointed tip only shortly before apex and the aedeagus is moderately downcurved at apex, whereas in
<taxonomicName id="4C2B4D78FF92F952C304FE9DFC32FE3F" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[874,969,381,404]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF92F952C304FE9DFC32FE3F" box="[874,969,381,404]" italics="true" pageId="5" pageNumber="278">G. bezzii</emphasis>
</taxonomicName>
, the dorsal gonostylus is gradually tapered to pointed tip and the aedeagus is strongly downcurved at apex. The female terminalia also provide some differences (cf.
<figureCitation id="13102A7EFF92F952C2BAFE5BFCA4FE78" box="[724,863,443,468]" captionStart="Figs" captionStartId="3.[115,154,897,916]" captionTargetBox="[129,1073,223,872]" captionTargetId="figure-247@3.[125,1078,219,877]" captionTargetPageId="3" captionText="Figs. 1–7.1–3 – Geranomyia bezzii Alexander &amp; Leonard, 1912 (Portugal:Alvor): 1 – male terminalia, general view, dorsally; 2 – male terminalia, aedeagal complex, lateral view; 3 – female terminalia, internal structures, ventral view. 4–6 – G. fuscior sp. nov. (paratypes; Portugal: Castro Marim): 4 – male terminalia, general view, dorsally; 5 – male terminalia, aedeagal complex, lateral view; 6 – female terminalia, internal structures, ventral view. 7 – G. unicolor Haliday, 1833 (England: Devon: Beer, coll. JSOC), male terminalia, gonostyli, dorsal view. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5330328" httpUri="https://zenodo.org/record/5330328/files/figure.png" pageId="5" pageNumber="278">Figs. 3 and 6</figureCitation>
), but these should be thoroughly evaluated and combined with all available external characters. In the structure of the male terminalia, especially the outline of the ventral gonostylus,
<taxonomicName id="4C2B4D78FF92F952C32CFE1DFC57FDBF" authority="Starý, 2012" authorityName="Starý" authorityYear="2012" box="[834,940,509,532]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="fuscior" status="sp. nov.">
<emphasis id="B95FEAE9FF92F952C32CFE1DFC57FDBF" box="[834,940,509,532]" italics="true" pageId="5" pageNumber="278">G. fuscior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C5792FF92F952C3DCFE1CFBF8FDB8" box="[946,1027,508,531]" pageId="5" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
bears resemblance to
<taxonomicName id="4C2B4D78FF92F952C176FDFDFDCAFD9F" authority="Haliday, 1833" authorityName="Haliday" authorityYear="1833" box="[280,561,539,564]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B95FEAE9FF92F952C176FDFDFE69FD9F" box="[280,402,541,564]" italics="true" pageId="5" pageNumber="278">G. unicolor</emphasis>
Haliday, 1833
</taxonomicName>
, but in the latter species, the ventral gonostylus is still broader, with a very short rostral prolongation and with rostral spines situated distinctly more apart (
<figureCitation id="13102A7EFF92F952C09DFDBBFECFFDDF" box="[243,308,603,628]" captionStart="Figs" captionStartId="3.[115,154,897,916]" captionTargetBox="[129,1073,223,872]" captionTargetId="figure-247@3.[125,1078,219,877]" captionTargetPageId="3" captionText="Figs. 1–7.1–3 – Geranomyia bezzii Alexander &amp; Leonard, 1912 (Portugal:Alvor): 1 – male terminalia, general view, dorsally; 2 – male terminalia, aedeagal complex, lateral view; 3 – female terminalia, internal structures, ventral view. 4–6 – G. fuscior sp. nov. (paratypes; Portugal: Castro Marim): 4 – male terminalia, general view, dorsally; 5 – male terminalia, aedeagal complex, lateral view; 6 – female terminalia, internal structures, ventral view. 7 – G. unicolor Haliday, 1833 (England: Devon: Beer, coll. JSOC), male terminalia, gonostyli, dorsal view. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5330328" httpUri="https://zenodo.org/record/5330328/files/figure.png" pageId="5" pageNumber="278">Fig. 7</figureCitation>
), not to mention other characters, including external ones.
</paragraph>
</subSubSection>
<subSubSection id="C3316570FF92F952C01FFD9CFC01FD18" pageId="5" pageNumber="278" type="etymology">
<paragraph id="8B9436FBFF92F952C01FFD9CFC01FD18" blockId="5.[113,1090,219,755]" pageId="5" pageNumber="278">
<emphasis id="B95FEAE9FF92F952C01FFD9CFF0BFD3F" bold="true" box="[113,240,636,660]" pageId="5" pageNumber="278">Etymology.</emphasis>
The name of the new species (Latin
<taxonomicName id="4C2B4D78FF92F952C20DFD9DFD57FD3F" authorityName="Starý" authorityYear="2012" box="[611,684,637,660]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="fuscior">
<emphasis id="B95FEAE9FF92F952C20DFD9DFD57FD3F" box="[611,684,637,660]" italics="true" pageId="5" pageNumber="278">fuscior</emphasis>
</taxonomicName>
= darker) refers to its body colouration, which is in general darker than that of
<taxonomicName id="4C2B4D78FF92F952C263FD7DFD90FD1F" authorityName="Alexander &amp; Leonard" authorityYear="1912" box="[525,619,669,692]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B95FEAE9FF92F952C263FD7DFD90FD1F" box="[525,619,669,692]" italics="true" pageId="5" pageNumber="278">G. bezzii</emphasis>
</taxonomicName>
. An adjective in nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="C3316570FF92F952C01FFD5BFE68FD58" pageId="5" pageNumber="278" type="distribution">
<paragraph id="8B9436FBFF92F952C01FFD5BFE68FD58" blockId="5.[113,1090,219,755]" pageId="5" pageNumber="278">
<emphasis id="B95FEAE9FF92F952C01FFD5BFEF8FD78" bold="true" box="[113,259,699,723]" pageId="5" pageNumber="278">Distribution.</emphasis>
<collectingCountry id="F33C766BFF92F952C166FD5BFEA4FD78" box="[264,351,699,723]" name="Albania" pageId="5" pageNumber="278">Albania</collectingCountry>
,
<collectingCountry id="F33C766BFF92F952C106FD5BFE3EFD78" box="[360,453,699,723]" name="Portugal" pageId="5" pageNumber="278">Portugal</collectingCountry>
,
<collectingCountry id="F33C766BFF92F952C1A3FD5BFDF4FD78" box="[461,527,699,723]" name="Libya" pageId="5" pageNumber="278">Libya</collectingCountry>
. From my experience both species are rather common on saltmarshes in
<collectingCountry id="F33C766BFF92F952C15FFD3BFE74FD58" box="[305,399,731,755]" name="Portugal" pageId="5" pageNumber="278">Portugal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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