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<document id="0F277154357F4A5499E475A0B268BB30" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629583631942" checkinUser="felipe" docAuthor="San Martin, G &amp; Hutchings, PA" docDate="2006" docId="038287B3A250FF99A9D42498FEE2F839" docLanguage="en" docName="RecAustMus.58.3.257-370.pdf" docOrigin="Records of the Australian Museum 58" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Pionosyllis koolalya San Martin &amp; Hutchings 2006, n.sp." docType="treatment" docVersion="2" lastPageNumber="340" masterDocId="FFBBFFCBA201FFCAAA142401FFB2FFED" masterDocTitle="Eusyllinae (Polychaeta: Syllidae) from Australia with the Description of a New Genus and Fifteen New Species" masterLastPageNumber="370" masterPageNumber="257" pageNumber="338" updateTime="1699081178577" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="FED09D6EB1D0A8B2E33B2DA6409A7AE2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="72982E1B0A5FCCFBBB4D2A861AE6E254">
<mods:title id="8D49E55BF9DADA4F50FA1D88D897F593">Eusyllinae (Polychaeta: Syllidae) from Australia with the Description of a New Genus and Fifteen New Species</mods:title>
</mods:titleInfo>
<mods:name id="E39A8824F58A8C70F1399DEE3EC53137" type="personal">
<mods:role id="24E328F66F027AB22ED58ABF02B8F61D">
<mods:roleTerm id="5763550E2F82261F95DC01A38BEA60DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DD1FB598F21FFAE9265F0E1ED25AE421">San Martin, G</mods:namePart>
</mods:name>
<mods:name id="4DA1DC4C2B1F9D76DD767EE3932AF17E" type="personal">
<mods:role id="B027A1409C0FD39EC6AAA1079AC83825">
<mods:roleTerm id="C907ADEFED711AEC6B34FB52D10C85C6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC518FC2082584E9F7BC85F4091F2191">Hutchings, PA</mods:namePart>
</mods:name>
<mods:typeOfResource id="BF05F28751E4C2D51C7F4556540C8DE1">text</mods:typeOfResource>
<mods:relatedItem id="AF5571AA415A62587F02A0DFBA42F93F" type="host">
<mods:titleInfo id="33C4C532F7CE1CF18FC848CFCD814BB1">
<mods:title id="F03CEDB4AF5EC44CB9AC009479E4A984">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="774368D0D63DC5C12BC42D3DDA0D7900">
<mods:date id="5D0065F1A077778DF20799E497A474C6">2006</mods:date>
<mods:detail id="38B76950B32862A5AC147E428EA718B1" type="volume">
<mods:number id="30E1C9894D368426F4C475BAFF43C38D">58</mods:number>
</mods:detail>
<mods:extent id="6517FA2DFAEA11BB7872BF8D97363BF5" unit="page">
<mods:start id="E86AA42C7301A48ACB32D31600E8E322">257</mods:start>
<mods:end id="05DF2308B515CE7D4B410D2673C2C48C">370</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="38301C1AE26E05F3BF78FDF0F676DDA5">journal article</mods:classification>
<mods:identifier id="95068B69939C842DEC9BA91FA1346D00" type="ISSN">2201-4349</mods:identifier>
</mods:mods>
<treatment id="038287B3A250FF99A9D42498FEE2F839" LSID="urn:lsid:plazi:treatment:038287B3A250FF99A9D42498FEE2F839" httpUri="http://treatment.plazi.org/id/038287B3A250FF99A9D42498FEE2F839" lastPageId="83" lastPageNumber="340" pageId="81" pageNumber="338">
<subSubSection id="C331652EA250FF9BA9D42498FB40FF09" pageId="81" pageNumber="338" type="nomenclature">
<paragraph id="8B9436A5A250FF9BA9D42498FB55FF5C" blockId="81.[960,1255,153,177]" box="[960,1255,153,177]" pageId="81" pageNumber="338">
<heading id="D0DC81C9A250FF9BA9D42498FB55FF5C" box="[960,1255,153,177]" centered="true" fontSize="11" level="2" pageId="81" pageNumber="338" reason="2">
<emphasis id="B95FEAB7A250FF9BA9D42498FB55FF5C" bold="true" box="[960,1255,153,177]" pageId="81" pageNumber="338">
<taxonomicName id="4C2B4D26A250FF9BA9D42498FB1AFF5C" ID-CoL="4J2TZ" authority="San Martin &amp; Hutchings, 2006" authorityName="San Martin &amp; Hutchings" authorityYear="2006" box="[960,1192,153,177]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="81" pageNumber="338" phylum="Annelida" rank="species" species="koolalya" status="sp. nov.">
<emphasis id="B95FEAB7A250FF9BA9D42498FB1AFF5C" bold="true" box="[960,1192,153,177]" italics="true" pageId="81" pageNumber="338">Pionosyllis koolalya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57CCA250FF9BAEA42498FB55FF5C" box="[1200,1255,153,177]" pageId="81" pageNumber="338" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9436A5A250FF9BA9A124CCFB40FF09" blockId="81.[949,1266,205,228]" box="[949,1266,205,228]" pageId="81" pageNumber="338">Figs 62A–C, 68A–P, 69A–F</paragraph>
</subSubSection>
<subSubSection id="C331652EA250FF9BA90E2500FBD6FEDC" pageId="81" pageNumber="338" type="reference_group">
<paragraph id="8B9436A5A250FF9BA90E2500FBD6FEDC" blockId="81.[794,1421,257,305]" pageId="81" pageNumber="338">
<treatmentCitationGroup id="AB3B118BA250FF9BA90E2500FBD6FEDC" pageId="81" pageNumber="338">
<taxonomicName id="4C2B4D26A250FF9BA90E2500FC53FEFB" ID-CoL="6VL4R" authorityName="Hartmann-Schroder" authorityYear="1979" box="[794,993,257,278]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="81" pageNumber="338" phylum="Annelida" rank="species" species="augeneri">
<emphasis id="B95FEAB7A250FF9BA90E2500FC53FEFB" box="[794,993,257,278]" italics="true" pageId="81" pageNumber="338">Pionosyllis augeneri</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0A8A10B4A250FF9BA9EA2500FAD3FEFB" author="Hartmann-Schroder, G" box="[1022,1377,257,278]" page="359" pageId="81" pageNumber="338" year="1991">
<bibRefCitation id="EFBA4B54A250FF9BA9EA2500FAD3FEFB" author="Hartmann-Schroder, G" box="[1022,1377,257,278]" pageId="81" pageNumber="338" pagination="17 - 71" refId="ref66137" refString="Hartmann-Schroder, G., 1991. Teil 16. Die Polychaeten der subtropisch-tropischen bis tropischen Ostkuste Australiens zwischen Maclean (New South Wales) und Gladstone (Queensland) sowie von Heron Is (Grosses Barriere-Riff). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 88: 17 - 71." type="journal article" year="1991">Hartmann-Schröder, 1991: 359–125</bibRefCitation>
</treatmentCitation>
. Not
<treatmentCitation id="0A8A10B4A250FF9BA92C251DFBD2FEDC" author="Hartmann-Schroder, G" box="[824,1120,284,305]" page="98" pageId="81" pageNumber="338" year="1979">
<bibRefCitation id="EFBA4B54A250FF9BA92C251DFBD2FEDC" author="Hartmann-Schroder, G" box="[824,1120,284,305]" pageId="81" pageNumber="338" pagination="75 - 218" refId="ref65657" refString="Hartmann-Schroder, G., 1979. Teil 2. Die Polychaeten der tropischen Nordwestkuste Australiens (zwischen Port Samson in Norden und Port Hedland in Suden). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 76: 75 - 218." type="journal article" year="1979">Hartmann-Schröder, 1979: 98</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C331652EA250FF9BA90E254AFBE9FDCF" pageId="81" pageNumber="338" type="materials_examined">
<paragraph id="8B9436A5A250FF9BA90E254AFB49FE2F" blockId="81.[794,1423,331,450]" pageId="81" pageNumber="338">
<emphasis id="B95FEAB7A250FF9BA90E254AFC41FE8E" bold="true" box="[794,1011,331,355]" pageId="81" pageNumber="338">Material examined</emphasis>
.
<materialsCitation id="3B433CF8A250FF9BA9E8254EFC5DFE7F" accessionNumber="W28945" collectingDate="1979-03-14" collectionCode="AM" collectorName="P. A. Hutchings" country="Australia" latitude="-32.8" location="Streaky Bay" longLatPrecision="1209" longitude="134.21666" municipality="Speeds Point" pageId="81" pageNumber="338" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<typeStatus id="54908807A250FF9BA9E8254EFBEBFE8D" box="[1020,1113,335,354]" pageId="81" pageNumber="338" type="holotype">HOLOTYPE</typeStatus>
(
<collectionCode id="ED3AAE60A250FF9BAE70254EFB3AFE8F" box="[1124,1160,335,354]" country="Australia" name="Australian Museum" pageId="81" pageNumber="338" type="Museum">AM</collectionCode>
W28945)
<collectingCountry id="F33C7635A250FF9BAEF6254EFAF4FE8D" box="[1250,1350,335,354]" name="Australia" pageId="81" pageNumber="338">AUSTRALIA</collectingCountry>
:
<collectingRegion id="49EFF847A250FF9BAF45254EFC32FE95" country="Australia" name="South Australia" pageId="81" pageNumber="338">SOUTH AUSTRALIA</collectingRegion>
:
<collectingMunicipality id="6BF0ACDFA250FF9BA99A2566FBB4FE97" box="[910,1030,359,378]" pageId="81" pageNumber="338">Speeds Point</collectingMunicipality>
,
<location id="8EF4607EA250FF9BAE062566FB33FE97" LSID="urn:lsid:plazi:treatment:038287B3A250FF99A9D42498FEE2F839:8EF4607EA250FF9BAE062566FB33FE97" box="[1042,1153,359,378]" country="Australia" latitude="-32.8" longLatPrecision="1209" longitude="134.21666" municipality="Speeds Point" name="Streaky Bay" pageId="81" pageNumber="338" stateProvince="South Australia">Streaky Bay</location>
,
<geoCoordinate id="EE1F5062A250FF9BAE992566FB60FE97" box="[1165,1234,359,378]" degrees="32" direction="south" minutes="48" orientation="latitude" pageId="81" pageNumber="338" precision="925" value="-32.8">32°48'S</geoCoordinate>
<geoCoordinate id="EE1F5062A250FF9BAECE2566FA9FFE97" box="[1242,1325,359,378]" degrees="134" direction="east" minutes="13" orientation="longitude" pageId="81" pageNumber="338" precision="925" value="134.21666">134°13'E</geoCoordinate>
, coll.
<collectorName id="26DE5373A250FF9BAF7D2566FCC4FE7F" pageId="81" pageNumber="338">P.A. Hutchings</collectorName>
,
<date id="FF951065A250FF9BA96A257EFC5DFE7F" box="[894,1007,383,402]" pageId="81" pageNumber="338" value="1979-03-14">
<collectingDate id="EFD1E98DA250FF9BA96A257EFC5DFE7F" box="[894,1007,383,402]" pageId="81" pageNumber="338" value="1979-03-14">14 Mar 1979</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B433CF8A250FF9BA9E2257EFAE9FE7F" accessionNumber="W28409" box="[1014,1371,383,402]" collectionCode="AM" country="Australia" pageId="81" pageNumber="338" specimenCount="1" stateProvince="South Australia" typeStatus="paratype">
<typeStatus id="54908807A250FF9BA9E2257EFBE2FE7D" box="[1014,1104,383,402]" pageId="81" pageNumber="338" type="paratype">PARATYPE</typeStatus>
1 on SEM stub (
<collectionCode id="ED3AAE60A250FF9BAECA257EFAB0FE7F" box="[1246,1282,383,402]" country="Australia" name="Australian Museum" pageId="81" pageNumber="338" type="Museum">AM</collectionCode>
W28409)
</materialsCitation>
.
<materialsCitation id="3B433CF8A250FF9BAF77257EFB45FE2F" collectingDate="1989-12-11" collectorName="Bottle &amp; Glass Rocks &amp; G. Clark" country="Australia" elevation="12" latitude="-33.85" location="Port Jackson" longLatPrecision="1202" longitude="151.26666" pageId="81" pageNumber="338" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
<collectingRegion id="49EFF847A250FF9BAF77257EFC29FE45" country="Australia" name="New South Wales" pageId="81" pageNumber="338">NEW SOUTH WALES</collectingRegion>
:
<collectorName id="26DE5373A250FF9BA9B32596FC6EFE47" box="[935,988,407,426]" pageId="81" pageNumber="338">Bottle</collectorName>
&amp;
<collectorName id="26DE5373A250FF9BA9EE2596FBDAFE47" box="[1018,1128,407,426]" pageId="81" pageNumber="338">Glass Rocks</collectorName>
,
<location id="8EF4607EA250FF9BAE662596FB56FE47" LSID="urn:lsid:plazi:treatment:038287B3A250FF99A9D42498FEE2F839:8EF4607EA250FF9BAE662596FB56FE47" box="[1138,1252,407,426]" country="Australia" latitude="-33.85" longLatPrecision="1202" longitude="151.26666" name="Port Jackson" pageId="81" pageNumber="338" stateProvince="New South Wales">Port Jackson</location>
,
<geoCoordinate id="EE1F5062A250FF9BAEF92596FA80FE47" box="[1261,1330,407,426]" degrees="33" direction="south" minutes="51" orientation="latitude" pageId="81" pageNumber="338" precision="925" value="-33.85">33°51'S</geoCoordinate>
<geoCoordinate id="EE1F5062A250FF9BAF2C2596FA3BFE47" box="[1336,1417,407,426]" degrees="151" direction="east" minutes="16" orientation="longitude" pageId="81" pageNumber="338" precision="925" value="151.26666">151°16'E</geoCoordinate>
, substrate unknown,
<quantity id="4CD39B40A250FF9BA9DF25AEFC41FE2F" box="[971,1011,431,450]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="81" pageNumber="338" unit="m" value="12.0">
<elevation id="0006D196A250FF9BA9DF25AEFC41FE2F" box="[971,1011,431,450]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="81" pageNumber="338" unit="m" value="12.0">12m</elevation>
</quantity>
, coll.
<collectorName id="26DE5373A250FF9BAE3E25AEFBCBFE2F" box="[1066,1145,431,450]" pageId="81" pageNumber="338">G. Clark</collectorName>
,
<date id="FF951065A250FF9BAE9725AEFB45FE2F" box="[1155,1271,431,450]" pageId="81" pageNumber="338" value="1989-12-11">
<collectingDate id="EFD1E98DA250FF9BAE9725AEFB45FE2F" box="[1155,1271,431,450]" pageId="81" pageNumber="338" value="1989-12-11">11 Dec 1989</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9436A5A250FF9BA90E25DAFBE9FDCF" blockId="81.[794,1421,475,546]" pageId="81" pageNumber="338">
<emphasis id="B95FEAB7A250FF9BA90E25DAFBDBFE1E" bold="true" box="[794,1129,475,499]" pageId="81" pageNumber="338">Additional material examined</emphasis>
.
<materialsCitation id="3B433CF8A250FF9BAE6725DEFBE5FDCF" collectionCode="HMZ" collectorName="Heron Is. &amp; G. Hartmann-Schroder" latitude="-23.45" location="QUEENSLAND" longLatPrecision="1256" longitude="151.91667" pageId="81" pageNumber="338" specimenCount="1" stateProvince="Queensland">
<collectingRegion id="49EFF847A250FF9BAE6725DEFB5AFE1C" box="[1139,1256,479,498]" country="Australia" name="Queensland" pageId="81" pageNumber="338">QUEENSLAND</collectingRegion>
:
<collectorName id="26DE5373A250FF9BAEE725DEFAF1FE1F" box="[1267,1347,479,498]" pageId="81" pageNumber="338">Heron Is.</collectorName>
,
<geoCoordinate id="EE1F5062A250FF9BAF5E25DEFA3FFE1F" box="[1354,1421,479,498]" degrees="23" direction="south" minutes="27" orientation="latitude" pageId="81" pageNumber="338" precision="925" value="-23.45">23°27'S</geoCoordinate>
<geoCoordinate id="EE1F5062A250FF9BA90E25F6FCDEFDE7" box="[794,876,503,522]" degrees="151" direction="east" minutes="55" orientation="longitude" pageId="81" pageNumber="338" precision="925" value="151.91667">151°55'E</geoCoordinate>
, sand, intertidal, coll.
<collectorName id="26DE5373A250FF9BAE3E25F6FB40FDE7" box="[1066,1266,503,522]" pageId="81" pageNumber="338">G. Hartmann-Schröder</collectorName>
, id. as
<taxonomicName id="4C2B4D26A250FF9BAF3825F6FCD5FDCF" authorityName="Hartmann-Schroder" authorityYear="1979" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="81" pageNumber="338" phylum="Annelida" rank="species" species="augeneri">
<emphasis id="B95FEAB7A250FF9BAF3825F6FCD5FDCF" italics="true" pageId="81" pageNumber="338">Pionosyllis augeneri</emphasis>
</taxonomicName>
(
<collectionCode id="ED3AAE60A250FF9BA960260EFC17FDCF" box="[884,933,527,546]" pageId="81" pageNumber="338">HMZ</collectionCode>
P-21018, P-16796)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331652EA250FF98A90E263FFDE1F86D" lastPageId="82" lastPageNumber="339" pageId="81" pageNumber="338" type="description">
<paragraph id="8B9436A5A250FF98A90E263FFDE1F86D" blockId="81.[794,1422,574,2006]" lastBlockId="82.[166,795,1839,1920]" lastPageId="82" lastPageNumber="339" pageId="81" pageNumber="338">
<emphasis id="B95FEAB7A250FF9BA90E263FFC2BFDBB" bold="true" box="[794,921,574,598]" pageId="81" pageNumber="338">Description</emphasis>
.
<typeStatus id="54908807A250FF9BA9B2263EFBB5FDBB" box="[934,1031,575,598]" pageId="81" pageNumber="338" type="holotype">Holotype</typeStatus>
<quantity id="4CD39B40A250FF9BAE18263EFBDBFDBB" box="[1036,1129,575,598]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="81" pageNumber="338" unit="mm" value="12.8">12.8 mm</quantity>
long,
<quantity id="4CD39B40A250FF9BAEBE263EFB4BFDBB" box="[1194,1273,575,598]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="81" pageNumber="338" unit="mm" value="0.3">0.3 mm</quantity>
wide, with 71 chaetigers;
<typeStatus id="54908807A250FF9BA985265DFC58FD9E" box="[913,1002,604,627]" pageId="81" pageNumber="338" type="paratype">paratype</typeStatus>
much smaller. Prostomium oval, with 4 eyes arranged in open trapezoidal pattern (
<figureCitation id="13102A20A250FF9BAED9267BFA9FFD7C" box="[1229,1325,634,657]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68A</figureCitation>
). Median antenna long, coiled on
<typeStatus id="54908807A250FF9BAE0F2696FBCAFD43" box="[1051,1144,663,686]" pageId="81" pageNumber="338" type="holotype">holotype</typeStatus>
, several times longer than combined length of prostomium and palps, inserted between posterior eyes; lateral antennae distinctly shorter than median antenna, inserted near anterior margin of prostomium (
<figureCitation id="13102A20A250FF9BAF4B26EEFCFBFCCE" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Figs 68A</figureCitation>
,
<figureCitation id="13102A20A250FF9BA946270DFC10FCCE" box="[850,930,780,803]" captionStart="Fig" captionStartId="83.[136,171,1499,1520]" captionTargetBox="[136,1407,151,1487]" captionTargetId="figure-10@83.[121,1422,151,1487]" captionTargetPageId="83" captionText="Fig. 69. SEM of Pionosyllis koolalya n.sp. (A) incomplete specimen, dorsal view; (B) anterior end, dorsal view; (C) detail of same; (D) detail of dorsum, midbody; (E) compound chaetae, midbody; (F) long-bladed compound chaeta, and one short-bladed, compound chaeta. Paratype, AM W28409." pageId="81" pageNumber="338">69A–C</figureCitation>
). Palps broad, similar in length to prostomium or slightly longer.Tentacular and dorsal cirri long, smooth, coiled on
<typeStatus id="54908807A250FF9BA92E2746FC25FCB3" box="[826,919,839,862]" pageId="81" pageNumber="338" type="holotype">holotype</typeStatus>
, long on anterior segments, becoming shorter on segments beyond proventricle, alternating long cirri (
<figureCitation id="13102A20A250FF9BAF3E2765FA3BFC96" box="[1322,1417,868,891]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Figs 68C</figureCitation>
,
<figureCitation id="13102A20A250FF9BA90E2783FCD0FC74" box="[794,866,898,921]" captionStart="Fig" captionStartId="83.[136,171,1499,1520]" captionTargetBox="[136,1407,151,1487]" captionTargetId="figure-10@83.[121,1422,151,1487]" captionTargetPageId="83" captionText="Fig. 69. SEM of Pionosyllis koolalya n.sp. (A) incomplete specimen, dorsal view; (B) anterior end, dorsal view; (C) detail of same; (D) detail of dorsum, midbody; (E) compound chaetae, midbody; (F) long-bladed compound chaeta, and one short-bladed, compound chaeta. Paratype, AM W28409." pageId="81" pageNumber="338">69A,B</figureCitation>
), 2–3 times as long as body width, and short cirri (
<figureCitation id="13102A20A250FF9BAF702783FCF8FC5B" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68B</figureCitation>
), slightly shorter than body width; dorsal cirri with cirrophores. Ciliary bands on dorsum of each segment and areas of dorsal cirri (
<figureCitation id="13102A20A250FF9BA9E727DBFBC1FC1C" box="[1011,1139,986,1009]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68B–D</figureCitation>
), distinct on
<typeStatus id="54908807A250FF9BAEEE27DBFAE7FC1C" box="[1274,1365,986,1009]" pageId="81" pageNumber="338" type="holotype">holotype</typeStatus>
(
<figureCitation id="13102A20A250FF9BAF7727DBFCF9FBE3" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68A</figureCitation>
); anterior segments with single ciliary band (
<figureCitation id="13102A20A250FF9BAF1927F6FA33FBE3" box="[1293,1409,1015,1038]" captionStart="Fig" captionStartId="83.[136,171,1499,1520]" captionTargetBox="[136,1407,151,1487]" captionTargetId="figure-10@83.[121,1422,151,1487]" captionTargetPageId="83" captionText="Fig. 69. SEM of Pionosyllis koolalya n.sp. (A) incomplete specimen, dorsal view; (B) anterior end, dorsal view; (C) detail of same; (D) detail of dorsum, midbody; (E) compound chaetae, midbody; (F) long-bladed compound chaeta, and one short-bladed, compound chaeta. Paratype, AM W28409." pageId="81" pageNumber="338">Fig. 69B,C</figureCitation>
), double from proventricle segments posteriorly (
<figureCitation id="13102A20A250FF9BAF092015FA30FBC6" box="[1309,1410,1044,1067]" captionStart="Fig" captionStartId="83.[136,171,1499,1520]" captionTargetBox="[136,1407,151,1487]" captionTargetId="figure-10@83.[121,1422,151,1487]" captionTargetPageId="83" captionText="Fig. 69. SEM of Pionosyllis koolalya n.sp. (A) incomplete specimen, dorsal view; (B) anterior end, dorsal view; (C) detail of same; (D) detail of dorsum, midbody; (E) compound chaetae, midbody; (F) long-bladed compound chaeta, and one short-bladed, compound chaeta. Paratype, AM W28409." pageId="81" pageNumber="338">Fig. 69D</figureCitation>
). Parapodial lobes conical, ending with 2 lobes (
<figureCitation id="13102A20A250FF9BAF1C2030FA30FBA5" box="[1288,1410,1073,1096]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68B,D</figureCitation>
). Ventral cirri digitiform, shorter than parapodial lobes anteriorly, becoming longer posteriorly (
<figureCitation id="13102A20A250FF9BAE53206DFB0FFB6E" box="[1095,1213,1132,1155]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68B,D</figureCitation>
).Anterior parapodia with about 10–14 compound chaetae, slightly enlarged, distally truncated shafts, provided with short subdistal spines, and bidentate blades, both teeth similar, with long spines on margin, pointing distally, those of basal part coarse, distal ones thin, reaching or extending beyond proximal tooth, 1–3 slightly longer (
<figureCitation id="13102A20A250FF9BA978211DFC79FADE" box="[876,971,1308,1331]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68E</figureCitation>
), rest of chaetae within fascicle with slight dorsoventral gradation in length (
<figureCitation id="13102A20A250FF9BAE622138FB61FABD" box="[1142,1235,1337,1360]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68F</figureCitation>
), 26 µm in length dorsally, 18 µm in length ventrally. Posteriorly, number of compound chaetae per parapodium decreasing, and blades of 1–3 most dorsal compound chaetae becoming proportionally longer (
<figureCitation id="13102A20A250FF9BA97C21AEFC7BFA2B" box="[872,969,1455,1478]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Figs 68H</figureCitation>
,
<figureCitation id="13102A20A250FF9BA9C621AEFC4EFA2B" box="[978,1020,1455,1478]" captionStart="Fig" captionStartId="83.[136,171,1499,1520]" captionTargetBox="[136,1407,151,1487]" captionTargetId="figure-10@83.[121,1422,151,1487]" captionTargetPageId="83" captionText="Fig. 69. SEM of Pionosyllis koolalya n.sp. (A) incomplete specimen, dorsal view; (B) anterior end, dorsal view; (C) detail of same; (D) detail of dorsum, midbody; (E) compound chaetae, midbody; (F) long-bladed compound chaeta, and one short-bladed, compound chaeta. Paratype, AM W28409." pageId="81" pageNumber="338">69F</figureCitation>
) and rest with dorsoventral gradation in length (
<figureCitation id="13102A20A250FF9BA97021CDFC08FA0E" box="[868,954,1484,1507]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Figs 68I</figureCitation>
,
<figureCitation id="13102A20A250FF9BA9D621CDFC5CFA0E" box="[962,1006,1484,1507]" captionStart="Fig" captionStartId="83.[136,171,1499,1520]" captionTargetBox="[136,1407,151,1487]" captionTargetId="figure-10@83.[121,1422,151,1487]" captionTargetPageId="83" captionText="Fig. 69. SEM of Pionosyllis koolalya n.sp. (A) incomplete specimen, dorsal view; (B) anterior end, dorsal view; (C) detail of same; (D) detail of dorsum, midbody; (E) compound chaetae, midbody; (F) long-bladed compound chaeta, and one short-bladed, compound chaeta. Paratype, AM W28409." pageId="81" pageNumber="338">69E</figureCitation>
); midbody parapodia with 1–2 compound chaetae with relatively long blades, about 31 µm in length, and 4 compound chaetae 15–25 µm in length; proximal tooth proportionally longer than those of anterior chaetae (Fig. 62A). Posterior parapodia with 1 compound chaeta with long blades (
<figureCitation id="13102A20A250FF9BA936225EFC35F99B" box="[802,903,1631,1654]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68K</figureCitation>
), 33 µm long, and 3 compound chaetae, similar to those of midbody, but with distinctly larger proximal tooth (
<figureCitation id="13102A20A250FF9BA9352298FC3AF95D" box="[801,904,1689,1712]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Figs 68L</figureCitation>
, 62C), well separated from distal tooth, 18–22 µm in length. Dorsal simple chaetae from midbody, distally bifid to truncate (
<figureCitation id="13102A20A250FF9BA9C022D5FB8EF906" box="[980,1084,1748,1771]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Figs 68G</figureCitation>
, 62B), thicker posteriorly, with short, coarse spines on margin (
<figureCitation id="13102A20A250FF9BAE6422F0FB64F8E5" box="[1136,1238,1777,1800]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68 O</figureCitation>
). Ventral simple chaetae on posterior parapodia, sigmoid, thick, strongly bidentate, both teeth well separated, almost at 90°, proximal tooth long, broad, slightly hooked, with few, long subdistal spines, surpassing level of proximal tooth (
<figureCitation id="13102A20A250FF9BAEF32366FAF4F893" box="[1255,1350,1895,1918]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Figs 68P</figureCitation>
, 62C). Two aciculae on anterior segments (
<figureCitation id="13102A20A250FF9BAEAD2385FA9BF876" box="[1209,1321,1924,1947]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68M</figureCitation>
), 1 from proventricle segments onwards, distally bilobed (
<figureCitation id="13102A20A250FF9BAF3323A0FA33F855" box="[1319,1409,1953,1976]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68J</figureCitation>
), thicker on posterior parapodia (
<figureCitation id="13102A20A250FF9BAE7223BEFB78F83B" box="[1126,1226,1983,2006]" captionStart="Fig" captionStartId="82.[166,201,1657,1678]" captionTargetBox="[204,1426,139,1640]" captionTargetId="figure-9@82.[204,1426,139,1640]" captionTargetPageId="82" captionText="Fig. 68. Pionosyllis koolalya n.sp. (A) anterior end, dorsal view; (B) midbody parapodium with short dorsal cirri; (C) long dorsal cirri, midbody; (D) posterior parapodium;(E) long-bladed compound chaeta, anterior parapodium; (F) compound chaetae, anterior parapodium; (G) dorsal simple chaeta, midbody; (H) long-bladed compound chaeta, midbody; (I) compound chaetae, midbody; (J) acicula, midbody; (K) long-bladed compound chaeta, posterior parapodium; (L) dorsal and ventral compound chaetae, posterior parapodium; (M) aciculae, anterior parapodium; (N) acicula, posterior parapodium; (O) dorsal simple chaeta, posterior parapodium; (P) ventral simple chaeta. AM W28409. Scales: A 0.18 mm, B–D 97 µm, E–P 20 µm." pageId="81" pageNumber="338">Fig. 68N</figureCitation>
). Pharynx through about 6 segments, pharyngeal tooth located on anterior rim. Proventricle through 4 segments, with about 26 muscle cell rows. Pygidium small, with 2 anal cirri.
</paragraph>
</subSubSection>
<caption id="DF54662DA253FF98AAB22278FD59F8F9" pageId="82" pageNumber="339" startId="82.[166,201,1657,1678]" targetBox="[204,1426,139,1640]" targetPageId="82">
<paragraph id="8B9436A5A253FF98AAB22278FD59F8F9" blockId="82.[166,1468,1657,1812]" pageId="82" pageNumber="339">
Fig. 68.
<taxonomicName id="4C2B4D26A253FF98AAE32278FE0CF963" authority="San Martin &amp; Hutchings, 2006" authorityName="San Martin &amp; Hutchings" authorityYear="2006" box="[247,446,1657,1678]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="82" pageNumber="339" phylum="Annelida" rank="species" species="koolalya" status="sp. nov.">
<emphasis id="B95FEAB7A253FF98AAE32278FE0CF963" box="[247,446,1657,1678]" italics="true" pageId="82" pageNumber="339">Pionosyllis koolalya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57CCA253FF98ABD72278FE40F963" box="[451,498,1657,1678]" pageId="82" pageNumber="339" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B95FEAB7A253FF98ABEB2278FDBCF963" box="[511,526,1657,1678]" italics="true" pageId="82" pageNumber="339">A</emphasis>
) anterior end, dorsal view; (
<emphasis id="B95FEAB7A253FF98A9352278FC82F963" box="[801,816,1657,1678]" italics="true" pageId="82" pageNumber="339">B</emphasis>
) midbody parapodium with short dorsal cirri; (
<emphasis id="B95FEAB7A253FF98AEEF2278FAB9F963" box="[1275,1291,1657,1678]" italics="true" pageId="82" pageNumber="339">C</emphasis>
) long dorsal cirri, midbody; (
<emphasis id="B95FEAB7A253FF98AB1A2295FEADF944" box="[270,287,1684,1705]" italics="true" pageId="82" pageNumber="339">D</emphasis>
) posterior parapodium; (
<emphasis id="B95FEAB7A253FF98A81E2295FDAAF944" box="[522,536,1684,1705]" italics="true" pageId="82" pageNumber="339">E</emphasis>
) long-bladed compound chaeta, anterior parapodium; (
<emphasis id="B95FEAB7A253FF98AE0A2295FB9FF944" box="[1054,1069,1684,1705]" italics="true" pageId="82" pageNumber="339">F</emphasis>
) compound chaetae, anterior parapodium; (
<emphasis id="B95FEAB7A253FF98AABA22AEFF0DF929" box="[174,191,1711,1732]" italics="true" pageId="82" pageNumber="339">G</emphasis>
) dorsal simple chaeta, midbody; (
<emphasis id="B95FEAB7A253FF98A81122AEFDA4F929" box="[517,534,1711,1732]" italics="true" pageId="82" pageNumber="339">H</emphasis>
) long-bladed compound chaeta, midbody; (
<emphasis id="B95FEAB7A253FF98A9AD22AEFC73F929" box="[953,961,1711,1732]" italics="true" pageId="82" pageNumber="339">I</emphasis>
) compound chaetae, midbody; (
<emphasis id="B95FEAB7A253FF98AEE322AEFAB0F929" box="[1271,1282,1711,1732]" italics="true" pageId="82" pageNumber="339">J</emphasis>
) acicula, midbody; (
<emphasis id="B95FEAB7A253FF98AABA22C8FF0CF933" box="[174,190,1737,1758]" italics="true" pageId="82" pageNumber="339">K</emphasis>
) long-bladed compound chaeta, posterior parapodium; (
<emphasis id="B95FEAB7A253FF98A8CE22C8FD55F933" box="[730,743,1737,1758]" italics="true" pageId="82" pageNumber="339">L</emphasis>
) dorsal and ventral compound chaetae, posterior parapodium; (
<emphasis id="B95FEAB7A253FF98AF5022C8FAEAF933" box="[1348,1368,1737,1758]" italics="true" pageId="82" pageNumber="339">M</emphasis>
) aciculae, anterior parapodium; (
<emphasis id="B95FEAB7A253FF98AB9122E5FE27F914" box="[389,405,1764,1785]" italics="true" pageId="82" pageNumber="339">N</emphasis>
) acicula, posterior parapodium; (
<emphasis id="B95FEAB7A253FF98A8F522E5FD40F914" box="[737,754,1764,1785]" italics="true" pageId="82" pageNumber="339">O</emphasis>
) dorsal simple chaeta, posterior parapodium; (
<emphasis id="B95FEAB7A253FF98AED022E5FB61F914" box="[1220,1235,1764,1785]" italics="true" pageId="82" pageNumber="339">P</emphasis>
) ventral simple chaeta. AM W28409. Scales:
<emphasis id="B95FEAB7A253FF98AB6B22FEFE3CF8F9" box="[383,398,1791,1812]" italics="true" pageId="82" pageNumber="339">A</emphasis>
0.18 mm,
<emphasis id="B95FEAB7A253FF98ABEC22FEFD96F8F9" box="[504,548,1791,1812]" italics="true" pageId="82" pageNumber="339">B–D</emphasis>
97 µm,
<emphasis id="B95FEAB7A253FF98A86222FEFD2DF8F9" box="[630,671,1791,1812]" italics="true" pageId="82" pageNumber="339">E–P</emphasis>
20 µm.
</paragraph>
</caption>
<subSubSection id="C331652EA253FF99AAB223A2FD71F8CE" lastPageId="83" lastPageNumber="340" pageId="82" pageNumber="339" type="discussion">
<paragraph id="8B9436A5A253FF99AAB223A2FE2EF91A" blockId="82.[166,794,1955,2007]" lastBlockId="83.[121,749,1642,1783]" lastPageId="83" lastPageNumber="340" pageId="82" pageNumber="339">
<emphasis id="B95FEAB7A253FF98AAB223A2FEA3F856" bold="true" box="[166,273,1955,1979]" pageId="82" pageNumber="339">Remarks</emphasis>
.
<taxonomicName id="4C2B4D26A253FF98AB3723A2FDBBF857" authorityName="San Martin &amp; Hutchings" authorityYear="2006" box="[291,521,1955,1978]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="82" pageNumber="339" phylum="Annelida" rank="species" species="koolalya">
<emphasis id="B95FEAB7A253FF98AB3723A2FDBBF857" box="[291,521,1955,1978]" italics="true" pageId="82" pageNumber="339">Pionosyllis koolalya</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C2B4D26A253FF98A8BF23A2FCA8F857" baseAuthorityName="Hartmann-Schroder" baseAuthorityYear="1984" box="[683,794,1955,1978]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="82" pageNumber="339" phylum="Annelida" rank="species" species="serrata">
<emphasis id="B95FEAB7A253FF98A8BF23A2FCA8F857" box="[683,794,1955,1978]" italics="true" pageId="82" pageNumber="339">P. serrata</emphasis>
</taxonomicName>
described above but differs mainly in lacking enlarged anterior parapodia with thick compound chaetae with markedly short blades; remaining compound chaetae are rather similar, but the dorsal simple chaeta of
<taxonomicName id="4C2B4D26A253FF98AF39236BFA10F86C" authorityName="San Martin &amp; Hutchings" authorityYear="2006" box="[1325,1442,1898,1921]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="82" pageNumber="339" phylum="Annelida" rank="species" species="koolalya">
<emphasis id="B95FEAB7A253FF98AF39236BFA10F86C" box="[1325,1442,1898,1921]" italics="true" pageId="82" pageNumber="339">P. koolalya</emphasis>
</taxonomicName>
is truncated, and slightly bifid in
<taxonomicName id="4C2B4D26A253FF98AE862386FB4BF873" baseAuthorityName="Hartmann-Schroder" baseAuthorityYear="1984" box="[1170,1273,1927,1950]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="82" pageNumber="339" phylum="Annelida" rank="species" species="serrata">
<emphasis id="B95FEAB7A253FF98AE862386FB4BF873" box="[1170,1273,1927,1950]" italics="true" pageId="82" pageNumber="339">P. serrata</emphasis>
</taxonomicName>
. All other species of the genus are provided with long, spiniger-like chaetae. These two species also have fewer compound chaetae with distinctly longer blades than other chaetae in the fascicle, but they are relatively short and less than half the length of the longest bladed chaetae. Some material of
<taxonomicName id="4C2B4D26A252FF99A86522A4FE29F934" authority="(Hartmann-Schroder, 1991)" baseAuthorityName="Hartmann-Schroder" baseAuthorityYear="1991" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="83" pageNumber="340" phylum="Annelida" rank="species" species="augeneri">
<emphasis id="B95FEAB7A252FF99A86522A4FD5FF951" box="[625,749,1701,1724]" italics="true" pageId="83" pageNumber="340">P. augeneri</emphasis>
(
<bibRefCitation id="EFBA4B54A252FF99AA9422C3FE21F934" author="Hartmann-Schroder, G" box="[128,403,1730,1753]" pageId="83" pageNumber="340" pagination="17 - 71" refId="ref66137" refString="Hartmann-Schroder, G., 1991. Teil 16. Die Polychaeten der subtropisch-tropischen bis tropischen Ostkuste Australiens zwischen Maclean (New South Wales) und Gladstone (Queensland) sowie von Heron Is (Grosses Barriere-Riff). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 88: 17 - 71." type="journal article" year="1991">Hartmann-Schröder, 1991</bibRefCitation>
)
</taxonomicName>
from Heron Island,
<collectingRegion id="49EFF847A252FF99A87D22C3FD5AF934" box="[617,744,1730,1753]" country="Australia" name="Queensland" pageId="83" pageNumber="340">Queensland</collectingRegion>
, belong to this new species.
</paragraph>
<caption id="DF54662DA252FF99AA9C21DAFE0EF9C8" pageId="83" pageNumber="340" startId="83.[136,171,1499,1520]" targetBox="[136,1407,151,1487]" targetPageId="83">
<paragraph id="8B9436A5A252FF99AA9C21DAFE0EF9C8" blockId="83.[136,1406,1499,1573]" pageId="83" pageNumber="340">
Fig. 69. SEM of
<taxonomicName id="4C2B4D26A252FF99AB3F21DAFE43FA1D" authority="San Martin &amp; Hutchings, 2006" authorityName="San Martin &amp; Hutchings" authorityYear="2006" box="[299,497,1499,1520]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="83" pageNumber="340" phylum="Annelida" rank="species" species="koolalya" status="sp. nov.">
<emphasis id="B95FEAB7A252FF99AB3F21DAFE43FA1D" box="[299,497,1499,1520]" italics="true" pageId="83" pageNumber="340">Pionosyllis koolalya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57CCA252FF99ABE321DAFD97FA1D" box="[503,549,1499,1520]" pageId="83" pageNumber="340" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B95FEAB7A252FF99A82721DAFDF0FA1D" box="[563,578,1499,1520]" italics="true" pageId="83" pageNumber="340">A</emphasis>
) incomplete specimen, dorsal view; (
<emphasis id="B95FEAB7A252FF99A9BB21DAFC0CFA1D" box="[943,958,1499,1520]" italics="true" pageId="83" pageNumber="340">B</emphasis>
) anterior end, dorsal view; (
<emphasis id="B95FEAB7A252FF99AEC521DAFB53FA1D" box="[1233,1249,1499,1520]" italics="true" pageId="83" pageNumber="340">C</emphasis>
) detail of same; (
<emphasis id="B95FEAB7A252FF99AA8421F4FF13F9E7" box="[144,161,1525,1546]" italics="true" pageId="83" pageNumber="340">D</emphasis>
) detail of dorsum, midbody; (
<emphasis id="B95FEAB7A252FF99ABAA21F4FE7FF9E7" box="[446,461,1525,1546]" italics="true" pageId="83" pageNumber="340">E</emphasis>
) compound chaetae, midbody; (
<emphasis id="B95FEAB7A252FF99A91421F4FCBDF9E7" box="[768,783,1525,1546]" italics="true" pageId="83" pageNumber="340">F</emphasis>
) long-bladed compound chaeta, and one short-bladed, compound chaeta. Paratype, AM W28409.
</paragraph>
</caption>
<paragraph id="8B9436A5A252FF99AA6D230DFD71F8CE" blockId="83.[121,707,1804,1828]" box="[121,707,1804,1828]" pageId="83" pageNumber="340">
<emphasis id="B95FEAB7A252FF99AA6D230DFF63F8C9" bold="true" box="[121,209,1804,1828]" pageId="83" pageNumber="340">Habitat</emphasis>
. Unknown, occurring from intertidal to
<specimenCount id="9D2DFD2CA252FF99A891230DFD71F8CE" box="[645,707,1804,1827]" pageId="83" pageNumber="340" type="generic">
<quantity id="4CD39B40A252FF99A891230DFD0DF8CE" box="[645,703,1804,1827]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="83" pageNumber="340" unit="m" value="12.0">12 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C331652EA252FF99AA6D2339FEBBF880" pageId="83" pageNumber="340" type="distribution">
<paragraph id="8B9436A5A252FF99AA6D2339FEBBF880" blockId="83.[121,748,1848,1901]" pageId="83" pageNumber="340">
<emphasis id="B95FEAB7A252FF99AA6D2339FEB6F8BD" bold="true" box="[121,260,1848,1872]" pageId="83" pageNumber="340">Distribution</emphasis>
.
<collectingCountry id="F33C7635A252FF99AB1A2339FEC2F8A2" box="[270,368,1848,1871]" name="Australia" pageId="83" pageNumber="340">Australia</collectingCountry>
(
<collectingRegion id="49EFF847A252FF99AB682339FD91F8A2" box="[380,547,1848,1871]" country="Australia" name="South Australia" pageId="83" pageNumber="340">South Australia</collectingRegion>
,
<collectingRegion id="49EFF847A252FF99A8382339FD5AF8A2" box="[556,744,1848,1871]" country="Australia" name="New South Wales" pageId="83" pageNumber="340">New South Wales</collectingRegion>
,
<collectingRegion id="49EFF847A252FF99AA6D2357FF4CF880" box="[121,254,1878,1901]" country="Australia" name="Queensland" pageId="83" pageNumber="340">Queensland</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C331652EA252FF99AA6D2383FEE2F839" pageId="83" pageNumber="340" type="etymology">
<paragraph id="8B9436A5A252FF99AA6D2383FEE2F839" blockId="83.[121,749,1922,2004]" pageId="83" pageNumber="340">
<emphasis id="B95FEAB7A252FF99AA6D2383FF47F877" bold="true" box="[121,245,1922,1946]" pageId="83" pageNumber="340">Etymology</emphasis>
. The specific name comes from an aboriginal word,
<taxonomicName id="4C2B4D26A252FF99AAA823A1FEABF85A" authorityName="San Martin &amp; Hutchings" authorityYear="2006" box="[188,281,1952,1975]" class="Polychaeta" family="Syllidae" genus="Pionosyllis" kingdom="Animalia" order="Phyllodocida" pageId="83" pageNumber="340" phylum="Annelida" rank="species" species="koolalya">
<emphasis id="B95FEAB7A252FF99AAA823A1FEABF85A" box="[188,281,1952,1975]" italics="true" pageId="83" pageNumber="340">koolalya</emphasis>
</taxonomicName>
, meaning feather epaulettes, in reference to the ciliated dorsum.
</paragraph>
</subSubSection>
</treatment>
</document>
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