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<mods:title id="76D12464B5079756DA24618EB979A478">The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="2FE0605777DE5B9BC4F9906FD6B081D4">Harzhauser, Mathias</mods:namePart>
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<subSubSection id="689965748A27387CFF4DFC5CFDDFFC2C" box="[151,541,942,969]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="203C36FF8A27387CFF4DFC5CFDDFFC2C" blockId="49.[151,541,942,969]" box="[151,541,942,969]" pageId="49" pageNumber="50">
<emphasis id="12F7EAED8A27387CFF4DFC5CFDDFFC2C" bold="true" box="[151,541,942,969]" pageId="49" pageNumber="50">
Genus
<taxonomicName id="E7834D7C8A27387CFF34FC5CFDDFFC2C" ID-CoL="7NVVF" authority="Swainson, 1840" authorityName="Swainson" authorityYear="1840" box="[238,541,942,969]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="genus">
<emphasis id="12F7EAED8A27387CFF34FC5CFE97FC2D" bold="true" box="[238,341,942,969]" italics="true" pageId="49" pageNumber="50">Cancilla</emphasis>
<bibRefCitation id="44124B0E8A27387CFE86FC5CFDDFFC2C" author="Swainson, W." box="[348,541,942,968]" pageId="49" pageNumber="50" refId="ref66058" refString="Swainson, W. (1840) A treatise on malacology or shells and shell-fish. Longman, London, viii + 419 pp. [http: // www. biodiversitylibrary. org / item / 33450]" type="book" year="1840">Swainson, 1840</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="689965748A27387CFF4DFC05FEEBFBD1" pageId="49" pageNumber="50" type="type_taxon">
<paragraph id="203C36FF8A27387CFF4DFC05FEEBFBD1" blockId="49.[151,1436,1014,1473]" pageId="49" pageNumber="50">
<emphasis id="12F7EAED8A27387CFF4DFC05FEF6FBF4" bold="true" box="[151,308,1014,1041]" pageId="49" pageNumber="50">
<typeStatus id="FF38885D8A27387CFF4DFC05FF10FBF5" box="[151,210,1015,1041]" pageId="49" pageNumber="50">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="E7834D7C8A27387CFEE7FC05FD5FFBF5" ID-CoL="6RLHN" authority="Swainson, 1831" authorityName="Swainson" authorityYear="1831" box="[317,669,1015,1041]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="isabella">
<emphasis id="12F7EAED8A27387CFEE7FC05FE1CFBF5" box="[317,478,1015,1041]" italics="true" pageId="49" pageNumber="50">Mitra isabella</emphasis>
<bibRefCitation id="44124B0E8A27387CFE3CFC05FD5FFBF5" author="Swainson, W. &amp; Zoological Illustrations" box="[486,669,1015,1041]" pageId="49" pageNumber="50" pagination="97 - 136" refId="ref65934" refString="Swainson, W. (1829 - 1833) Zoological Illustrations, or original figures and descriptions of new, rare, or interesting animals, selected chiefly from the classes of ornithology, entomology, and conchology, and arranged according to their apparent affinities. Series 2. Baldwin &amp; Cradock, London, 136 pls. [pls. 1 - 30 (1829) + 31 - 45 (1830) + 46 - 85 (1831) + 86 - 96 (1832) + 97 - 136 (1833), http: // www. biodiversitylibrary. org / bibliography / 42278] https: // doi. org / 10.5962 / bhl. title. 42278" type="journal article" year="1831">Swainson, 1831</bibRefCitation>
</taxonomicName>
, subsequent designation by
<bibRefCitation id="44124B0E8A27387CFC04FC05FAC5FBF6" author="Herrmannsen, A. N." box="[990,1287,1015,1042]" pageId="49" pageNumber="50" pagination="155" refId="ref61765" refString="Herrmannsen, A. N. (1846) Indicis Generum Malacozoorum primordia. Fischer, Cassel, 1, pp. 155. [https: // www. biodiversitylibrary. org / page / 10695423]" type="journal article" year="1846">Herrmannsen (1846: 166)</bibRefCitation>
. Present-day, Indo-Pacific.
</paragraph>
</subSubSection>
<subSubSection id="689965748A27387CFF1DFBCCFC7DFA79" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="203C36FF8A27387CFF1DFBCCFE02FA9D" blockId="49.[151,1436,1014,1473]" pageId="49" pageNumber="50">
<emphasis id="12F7EAED8A27387CFF1DFBCCFE80FBBC" bold="true" box="[199,322,1086,1112]" pageId="49" pageNumber="50">Diagnosis.</emphasis>
<emphasis id="12F7EAED8A27387CFE8FFBCDFA52FAB1" italics="true" pageId="49" pageNumber="50">
Shell small to rather large (
<quantity id="E77B9B1A8A27387CFD50FBCDFCD2FBBD" box="[650,784,1087,1113]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="11.0" metricValueMin="1.5" pageId="49" pageNumber="50" unit="mm" value="62.5" valueMax="110.0" valueMin="15.0">15110 mm</quantity>
), fusiform, with high aperture and long, tapering siphonal canal. Protoconch conical, of about three very slightly convex smooth whorls. Spire whorls evenly convex to subcylindrical; suture distinct, impressed. Shell sculptured with spiral cords that are slightly gemmate or with finely dentate margins, very wide and flat, separated by narrow grooves, or with strong, narrow and elevated cords separated by broad depressions bearing regular, very fine riblets. Shell base gradually extended into rather long, tapering siphonal canal. Siphonal fasciole not pronounced, notch deep or shallow. Aperture elongate, narrow; outer aperture lip smooth, gently convex adapically, and flattened in its lower portion. Inner lip with four oblique columellar folds, adapicalmost strongest. Shell pale or cream, typically with multiple darker spiral strokes on crests of spiral cords
</emphasis>
” (
<bibRefCitation id="44124B0E8A27387CFF45FAADFE71FA9D" author="Fedosov, A. &amp; Puillandre, N. &amp; Herrmann, M. &amp; Kantor, Yu. &amp; Oliverio, M. &amp; Dgebuadze, P. &amp; Modica, M. V. &amp; Bouchet, P." box="[159,435,1374,1401]" pageId="49" pageNumber="50" pagination="253 - 337" refId="ref60492" refString="Fedosov, A., Puillandre, N., Herrmann, M., Kantor, Yu., Oliverio, M., Dgebuadze, P., Modica, M. V. &amp; Bouchet, P. (2018) The collapse of Mitra: molecular systematics and morphology of the Mitridae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 183, 253 - 337. https: // doi. org / 10.1093 / zoolinnean / zlx 073" type="journal article" year="2018">
Fedosov
<emphasis id="12F7EAED8A27387CFEDCFAADFEFAFA9D" box="[262,312,1375,1401]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
., 2018: 54
</bibRefCitation>
).
</paragraph>
<paragraph id="203C36FF8A27387CFF1DFA71FC7DFA79" blockId="49.[151,1436,1014,1473]" box="[199,959,1410,1437]" pageId="49" pageNumber="50">
<emphasis id="12F7EAED8A27387CFF1DFA71FE30FA78" bold="true" box="[199,498,1410,1437]" pageId="49" pageNumber="50">Present-day distribution.</emphasis>
Indo-West Pacific (
<bibRefCitation id="44124B0E8A27387CFD0AFA71FC71FA79" author="Fedosov, A. &amp; Puillandre, N. &amp; Herrmann, M. &amp; Kantor, Yu. &amp; Oliverio, M. &amp; Dgebuadze, P. &amp; Modica, M. V. &amp; Bouchet, P." box="[720,947,1411,1437]" pageId="49" pageNumber="50" pagination="253 - 337" refId="ref60492" refString="Fedosov, A., Puillandre, N., Herrmann, M., Kantor, Yu., Oliverio, M., Dgebuadze, P., Modica, M. V. &amp; Bouchet, P. (2018) The collapse of Mitra: molecular systematics and morphology of the Mitridae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 183, 253 - 337. https: // doi. org / 10.1093 / zoolinnean / zlx 073" type="journal article" year="2018">
Fedosov
<emphasis id="12F7EAED8A27387CFCEDFA71FCB2FA79" box="[823,880,1411,1437]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="689965748A27387CFF1DFA55FC1DFA25" box="[199,991,1447,1473]" pageId="49" pageNumber="50" type="discussion">
<paragraph id="203C36FF8A27387CFF1DFA55FC1DFA25" blockId="49.[151,1436,1014,1473]" box="[199,991,1447,1473]" pageId="49" pageNumber="50">
See
<tableCitation id="6D0103448A27387CFF2CFA55FE8BFA25" box="[246,329,1447,1473]" captionStart="TABLE 2" captionStartId="50.[151,239,151,175]" captionTargetBox="[166,1409,191,769]" captionTargetPageId="50" captionText="TABLE 2. Cancilla species in the Paratethys: Morphometric data as in Table 1. For further explanation see Figure 2. Species SL (mm) SL/MD AL/AW AH/S small SL &lt;20.0 broad SL/MD &lt;2.5 narrow AL/AW ≥ 5.8 long AH/S &lt;2.1 medium-sized mod. broad mod. narrow &lt;5.85.0 mod. long 2.1&lt;2.5 20-40&gt;2.53.0 mod. wide &lt;5.04.0 mod. short &lt;2.52.9 mod. large 40.160 mod. slender wide &lt;4.0 short ≥ 3.0 large SL&gt;60 3.13.5 slender&gt;3.5" httpUri="http://table.plazi.org/id/74FC66778A24387FFF4DFF65FA5EFF54" pageId="49" pageNumber="50" tableUuid="74FC66778A24387FFF4DFF65FA5EFF54">Table 2</tableCitation>
for morphometric data of all Paratethyan
<taxonomicName id="E7834D7C8A27387CFCC5FA55FC1DFA25" box="[799,991,1447,1473]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="12F7EAED8A27387CFCC5FA55FCBDFA25" box="[799,895,1447,1473]" italics="true" pageId="49" pageNumber="50">Cancilla</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="689965748A27387CFF4DF9E1FA5FF80E" pageId="49" pageNumber="50" type="key">
<paragraph id="203C36FF8A27387CFF4DF9E1FD50F9C9" blockId="49.[151,658,1554,1581]" box="[151,658,1554,1581]" pageId="49" pageNumber="50">
<emphasis id="12F7EAED8A27387CFF4DF9E1FD50F9C9" bold="true" box="[151,658,1554,1581]" pageId="49" pageNumber="50">
Key to
<taxonomicName id="E7834D7C8A27387CFF2BF9E0FE77F9C8" box="[241,437,1554,1581]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="12F7EAED8A27387CFF2BF9E0FE9AF9C9" bold="true" box="[241,344,1554,1581]" italics="true" pageId="49" pageNumber="50">Cancilla</emphasis>
species
</taxonomicName>
in the Paratethys
</emphasis>
</paragraph>
<paragraph id="203C36FF8A27387CFF4DF9AAFA5EF96E" blockId="49.[151,1437,1624,2027]" pageId="49" pageNumber="50">1. Cancellate sculpture spire whorls......................................................................... 2 Spiral sculpture only................................................................................... 3</paragraph>
<paragraph id="203C36FF8A27387CFF4DF960FA5FF921" blockId="49.[151,1437,1624,2027]" pageId="49" pageNumber="50">
2. Cancellate sculpture weak, restricted to earliest whorl.........................................
<taxonomicName id="E7834D7C8A27387CFB07F961FE85F922" ID-CoL="8KSMM" authority="Cancellate" authorityName="Cancellate" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="praescrobiculata">
<emphasis id="12F7EAED8A27387CFB07F961FA5EF943" box="[1245,1436,1682,1704]" italics="true" pageId="49" pageNumber="50">C. praescrobiculata</emphasis>
Cancellate
</taxonomicName>
sculpture strong, persisting to penultimate whorl.........................................
<taxonomicName id="E7834D7C8A27387CFAD0F942FA5FF921" ID-CoL="8KSRT" baseAuthorityName="Bellardi" baseAuthorityYear="1887" box="[1290,1437,1712,1733]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="pulcherrima">
<emphasis id="12F7EAED8A27387CFAD0F942FA5FF921" box="[1290,1437,1712,1733]" italics="true" pageId="49" pageNumber="50">C. pulcherrima</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="203C36FF8A27387CFF4DF93FFA5EF8F9" blockId="49.[151,1437,1624,2027]" pageId="49" pageNumber="50">
3. Spiral sculpture of fine spirals........................................................................... 4 Spiral sculpture of flattened cords........................................................................ 5 Strong elevated cords, relatively broad, cancellate interspaces, shell slender fusiform........................
<taxonomicName id="E7834D7C8A27387CFAF1F8FAFA5EF8F9" ID-CoL="8KSF3" baseAuthorityName="Bellardi" baseAuthorityYear="1887" box="[1323,1436,1800,1821]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="exornata">
<emphasis id="12F7EAED8A27387CFAF1F8FAFA5EF8F9" box="[1323,1436,1800,1821]" italics="true" pageId="49" pageNumber="50">C. exornata</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="203C36FF8A27387CFF4DF8D7FA5FF8BC" blockId="49.[151,1437,1624,2027]" pageId="49" pageNumber="50">
4. Shell broadly fusiform, irregular crowded spirals...................................................
<taxonomicName id="E7834D7C8A27387CFAC5F8D7FED1F8BC" ID-CoL="8KS9M" authority="Shell" authorityName="Shell" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="sismondai">
<emphasis id="12F7EAED8A27387CFAC5F8D7FA5EF8DE" box="[1311,1436,1829,1850]" italics="true" pageId="49" pageNumber="50">C. sismondai</emphasis>
Shell
</taxonomicName>
narrower fusiform, regular fine spirals............................................... ..
<taxonomicName id="E7834D7C8A27387CFB0BF8B1FA8EF8B3" ID-CoL="8KRWS" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[1233,1356,1858,1880]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="wagreichi" status="sp. nov.">
<emphasis id="12F7EAED8A27387CFB0BF8B1FA8EF8B3" box="[1233,1356,1858,1880]" italics="true" pageId="49" pageNumber="50">C. wagreichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="09C457968A27387CFA88F8B1FA5FF8BC" box="[1362,1437,1859,1880]" pageId="49" pageNumber="50" rank="species">nov. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="203C36FF8A27387CFF4DF8ADFA5EF877" blockId="49.[151,1437,1624,2027]" pageId="49" pageNumber="50">
5. Flattened cords obsolete mid-whorl on last whorl.........................................
<taxonomicName id="E7834D7C8A27387CFB75F892FA8EF891" ID-CoL="8KSRD" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[1199,1356,1888,1909]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="nanostriatula" status="sp. nov.">
<emphasis id="12F7EAED8A27387CFB75F892FA8EF891" box="[1199,1356,1888,1909]" italics="true" pageId="49" pageNumber="50">C. nanostriatula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="09C457968A27387CFA88F892FA5FF891" box="[1362,1437,1888,1909]" pageId="49" pageNumber="50" rank="species">nov. sp.</taxonomicNameLabel>
Flattened cords separated by narrow punctate interspaces...................................................... 6
</paragraph>
<paragraph id="203C36FF8A27387CFF4DF868FA5FF80E" blockId="49.[151,1437,1624,2027]" pageId="49" pageNumber="50">
6. Last whorls shouldered.......................................................................
<taxonomicName id="E7834D7C8A27387CFACEF869FA5EF84B" ID-CoL="8KSK8" baseAuthorityName="Bellardi" baseAuthorityYear="1887" box="[1300,1436,1946,1968]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="suballigata">
<emphasis id="12F7EAED8A27387CFACEF869FA5EF84B" box="[1300,1436,1946,1968]" italics="true" pageId="49" pageNumber="50">C. suballigata</emphasis>
</taxonomicName>
Base strongly constricted.................................................................
<taxonomicName id="E7834D7C8A27387CFB30F84AFEE1F80F" ID-CoL="8KTBP" authority="Profile" authorityName="Profile" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="capelliniana">
<emphasis id="12F7EAED8A27387CFB30F84AFB38F829" box="[1258,1274,1976,1997]" italics="true" pageId="49" pageNumber="50">C</emphasis>
. cf.
<emphasis id="12F7EAED8A27387CFAFEF84AFA5EF829" box="[1316,1436,1976,1997]" italics="true" pageId="49" pageNumber="50">capelliniana</emphasis>
Profile
</taxonomicName>
slender fusiform.....................................................................
<taxonomicName id="E7834D7C8A27387CFAD0F827FA5FF80E" ID-CoL="8KS36" baseAuthorityName="Bellardi" baseAuthorityYear="1887" box="[1290,1437,2005,2026]" class="Gastropoda" family="Mitridae" genus="Cancilla" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="planicostata">
<emphasis id="12F7EAED8A27387CFAD0F827FA5FF80E" box="[1290,1437,2005,2026]" italics="true" pageId="49" pageNumber="50">C. planicostata</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>