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<document id="0642249C731D783744EF0D9693E7CD30" ID-DOI="10.5852/ejt.2020.626" ID-GBIF-Dataset="c906d9cd-cfcb-43db-a24c-b57b39d9184b" ID-Zenodo-Dep="3754989" ID-ZooBank="D91881D6-55D8-48FC-A383-069BC643A91E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587066773978" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2020" docId="5C11879EFFBDFFD7FD87EFB3FB281065" docLanguage="en" docName="ejt-626_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 626" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor fasciatus" docType="treatment" docVersion="6" lastPageNumber="14" masterDocId="A028FFE6FFB6FFDAFFC1ED40FFC7143C" masterDocTitle="A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="12" updateTime="1698738618004" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A42E7445F6689B83DCF4D5446CDA9862">A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<heading id="8F4F81E4FFBDFFD1FD87EFB3FC2A1732" box="[582,1005,755,782]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<taxonomicName id="13B84D0BFFBDFFD1FD87EFB3FC2A1732" ID-CoL="FZKZ" authority="(Attems, 1896)" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[582,1005,755,782]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFBDFFD1FD87EFB3FCF21731" bold="true" box="[582,821,755,781]" italics="true" pageId="11" pageNumber="12">Aquattuor fasciatus</emphasis>
(
<bibRefCitation id="B0294B79FFBDFFD1FC85EFB4FC221732" author="Attems C. G." box="[836,997,755,782]" pageId="11" pageNumber="12" pagination="23 - 42" refId="ref11215" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13, Suppl.: 23 - 42. Available from: https: // biodiversitylibrary. org / page / 28745426 [accessed 1 Oct. 2019]." type="book chapter" year="1896">Attems, 1896</bibRefCitation>
)
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<paragraph id="D4073688FFBDFFD1FD0AEE56FCAF170D" blockId="11.[582,1005,755,817]" box="[715,872,790,817]" pageId="11" pageNumber="12">
<figureCitation id="4C832A0DFFBDFFD1FD0AEE56FCD3170D" box="[715,788,790,817]" captionStart="Fig" captionStartId="2.[189,232,1948,1974]" captionTargetBox="[192,1385,1023,1872]" captionTargetId="figure@2.[186,1391,1009,1876]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites for Aquattuor spp. in the Udzungwa Mts. Based on Marshall et al. (2010: fig. 1); inset by permission of the Eastern Arc Mountains Conservation Endowment Fund." figureDoi="http://doi.org/10.5281/zenodo.3754991" httpUri="https://zenodo.org/record/3754991/files/figure.png" pageId="11" pageNumber="12">Figs 1</figureCitation>
,
<figureCitation id="4C832A0DFFBDFFD1FCE3EE57FCE9170D" box="[802,814,791,817]" captionStart="Fig" captionStartId="5.[189,232,1849,1875]" captionTargetBox="[205,1381,684,1788]" captionTargetId="figure@5.[189,1399,675,1801]" captionTargetPageId="5" captionText="Fig. 3. Aquattuor spp., first pair of male legs. A–C. A. fasciatus (Attems, 1896), non-type male from Zanzibar (NHMD 621641). D–F. A. claudiahempae Enghoff &amp; Frederiksen, 2015, paratype from Kilimanjaro (NHMD 621638). G–I. A. claudiahempae, specimen from Udzungwa Mts (NHMD 621639). A, D, G. Anterior view. B, E, H. Sublateral view. C, F, I. Ventral view. Abbreviation: pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754995" httpUri="https://zenodo.org/record/3754995/files/figure.png" pageId="11" pageNumber="12">3</figureCitation>
A–C
</paragraph>
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<treatmentCitation id="55191099FFBDFFD1FF7CEE14FD411752" author="Attems C. G." box="[189,646,852,878]" page="40" pageId="11" pageNumber="12" year="1896">
<taxonomicName id="13B84D0BFFBDFFD1FF7CEE14FD411752" ID-CoL="7XK7M" authority="Attems, 1896: 40" authorityName="Attems" authorityPageNumber="40" authorityYear="1896" box="[189,646,852,878]" class="Diplopoda" family="Odontopygidae" genus="Odontopyge" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="E6CCEA9AFFBDFFD1FF7CEE14FE761752" box="[189,433,852,878]" italics="true" pageId="11" pageNumber="12">Odontopyge fasciata</emphasis>
<bibRefCitation id="B0294B79FFBDFFD1FE79EE14FD411752" author="Attems C. G." box="[440,646,852,878]" pageId="11" pageNumber="12" pagination="23 - 42" refId="ref11215" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13, Suppl.: 23 - 42. Available from: https: // biodiversitylibrary. org / page / 28745426 [accessed 1 Oct. 2019]." type="book chapter" year="1896">Attems, 1896: 40</bibRefCitation>
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.
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<paragraph id="D4073688FFBDFFD1FF7CEED2FC2017F3" blockId="11.[189,1398,914,1046]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="F4A811A6FFBDFFD1FF7CEED2FC2017F3" pageId="11" pageNumber="12">
<taxonomicName id="13B84D0BFFBDFFD1FF7CEED2FE2A1790" ID-CoL="4MG36" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[189,493,914,940]" class="Diplopoda" family="Odontopygidae" genus="Prionopetalum" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fasciatum">
<emphasis id="E6CCEA9AFFBDFFD1FF7CEED2FE2A1790" box="[189,493,914,940]" italics="true" pageId="11" pageNumber="12">Prionopetalum fasciatum</emphasis>
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–
<treatmentCitation id="55191099FFBDFFD1FDD7EED2FD3E1790" author="Attems C. G." box="[534,761,914,940]" page="210" pageId="11" pageNumber="12" year="1914">
<bibRefCitation id="B0294B79FFBDFFD1FDD7EED2FD3E1790" author="Attems C. G." box="[534,761,914,940]" pageId="11" pageNumber="12" pagination="1 - 233" refId="ref11344" refString="Attems C. G. 1914. Afrikanisiche Spirostreptiden nebst Uberblick uber die Spirostreptiden orbis terrarum. Zoologica Stuttgart 65 - 66: 1 - 233." type="journal article" year="1914">
Attems
<date id="A0061048FFBDFFD1FDB9EED2FD701790" box="[632,695,914,940]" pageId="11" pageNumber="12" value="1914">1914</date>
: 210
</bibRefCitation>
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. —
<treatmentCitation id="55191099FFBDFFD1FCF4EED2FB821790" author="Brolemann H. W." box="[821,1093,914,940]" page="123" pageId="11" pageNumber="12" year="1920">
<bibRefCitation id="B0294B79FFBDFFD1FCF4EED2FB821790" author="Brolemann H. W." box="[821,1093,914,940]" pageId="11" pageNumber="12" pagination="49 - 298" refId="ref11402" refString="Brolemann H. W. 1920. Myriapodes. In: Allaud Ch. (ed.) Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale 1911 - 1912. Resultats Scientifiques, Myriapoda 3: 49 - 298. A. Schultz, Paris." type="journal article" year="1920">
Brolemann
<date id="A0061048FFBDFFD1FC04EED2FBC31790" box="[965,1028,914,940]" pageId="11" pageNumber="12" value="1920">1920</date>
: 123
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. —
<treatmentCitation id="55191099FFBDFFD1FB43EED2FA8F1790" author="Kraus O." box="[1154,1352,914,940]" page="86" pageId="11" pageNumber="12" year="1960">
<bibRefCitation id="B0294B79FFBDFFD1FB43EED2FA8F1790" author="Kraus O." box="[1154,1352,914,940]" pageId="11" pageNumber="12" pagination="1 - 207" refId="ref12249" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." type="journal article" year="1960">
Kraus
<date id="A0061048FFBDFFD1FB17EED2FAD21790" box="[1238,1301,914,940]" pageId="11" pageNumber="12" value="1960">1960</date>
: 86
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. —
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<bibRefCitation id="B0294B79FFBDFFD1FF2BEEF5FD5817EC" author="VandenSpiegel D. &amp; Pierrard G." box="[234,671,949,976]" pageId="11" pageNumber="12" pagination="149 - 163" refId="ref12440" refString="VandenSpiegel D. &amp; Pierrard G. 2009. Revision du genre Prionopetalum (Odontopygidae, Diplopoda) et descriptions de nouvelles especes d'Afrique de l'Est. Journal of Afrotropical Zoology 5: 149 - 163." type="journal article" year="2009">
VandenSpiegel &amp; Pierrard
<date id="A0061048FFBDFFD1FDE7EEF5FDA217F3" box="[550,613,949,975]" pageId="11" pageNumber="12" value="2009">2009</date>
: 152
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<treatmentCitation id="55191099FFBDFFD1FD0EEEF5FC2417F3" author="Enghoff H. &amp; Hoffman R. L. &amp; Howell K. M." box="[719,995,949,975]" page="69" pageId="11" pageNumber="12" year="2016">
<bibRefCitation id="B0294B79FFBDFFD1FD0EEEF5FC2417F3" author="Enghoff H. &amp; Hoffman R. L. &amp; Howell K. M." box="[719,995,949,975]" pageId="11" pageNumber="12" pagination="51 - 113" refId="ref11986" refString="Enghoff H., Hoffman R. L. &amp; Howell K. M. 2016. Checklist of the millipedes (Diplopoda) of Tanzania. Journal of East African Natural History 105 (1): 51 - 113. https: // doi. org / 10.2982 / 028.105.0103" type="journal article" year="2016">
Enghoff
<emphasis id="E6CCEA9AFFBDFFD1FCF7EEF6FCB617F3" box="[822,881,949,975]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
<date id="A0061048FFBDFFD1FCB9EEF5FC7017F3" box="[888,951,949,975]" pageId="11" pageNumber="12" value="2016">2016</date>
: 69
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.
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<paragraph id="D4073688FFBDFFD1FF7CEE99FC0717CE" blockId="11.[189,1398,914,1046]" box="[189,960,984,1011]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="F4A811A6FFBDFFD1FF7CEE99FC0717CE" box="[189,960,984,1011]" pageId="11" pageNumber="12">
<taxonomicName id="13B84D0BFFBDFFD1FF7CEE99FDDA17CE" ID-CoL="FZKX" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[189,541,984,1011]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBDFFD1FF7CEE99FEF317CF" box="[189,308,985,1011]" italics="true" pageId="11" pageNumber="12">Aquattuor</emphasis>
aff.
<emphasis id="E6CCEA9AFFBDFFD1FEABEE98FDDA17CE" box="[362,541,984,1010]" italics="true" pageId="11" pageNumber="12">claudiahempae</emphasis>
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–
<treatmentCitation id="55191099FFBDFFD1FDFAEE98FC7B17CE" ID-DOI="http://doi.org/10.5281/zenodo.3793775" ID-Zenodo-Dep="3793775" LSID="urn:lsid:zoobank.org:act:66387356-4B9A-467E-A3CA-4BC76EF8FCAE" author="Enghoff H. &amp; Frederiksen S. B." box="[571,956,984,1011]" httpUri="http://treatment.plazi.org/id/03A487F2FFEBFFAC756B6AB29465B6C2" page="19" pageId="11" pageNumber="12" year="2015">
<bibRefCitation id="B0294B79FFBDFFD1FDFAEE98FC7B17CE" author="Enghoff H. &amp; Frederiksen S. B." box="[571,956,984,1011]" pageId="11" pageNumber="12" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen 2015: 19</bibRefCitation>
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<paragraph id="D4073688FFBDFFD1FF7CEEBCFD62102A" blockId="11.[189,1398,914,1046]" box="[189,677,1019,1046]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="F4A811A6FFBDFFD1FF7CEEBCFD62102A" box="[189,677,1019,1046]" pageId="11" pageNumber="12">
<taxonomicName id="13B84D0BFFBDFFD1FF7CEEBCFE64102A" ID-CoL="FZKZ" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[189,419,1020,1046]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFBDFFD1FF7CEEBCFE64102A" box="[189,419,1020,1046]" italics="true" pageId="11" pageNumber="12">Aquattuor fasciatus</emphasis>
</taxonomicName>
–
<treatmentCitation id="55191099FFBDFFD1FE00EEBBFD66102A" ID-DOI="http://doi.org/10.5281/zenodo.5695715" ID-Zenodo-Dep="5695715" author="Enghoff H." box="[449,673,1019,1046]" httpUri="http://treatment.plazi.org/id/03D3D100FFE4C416F028FD757984FC12" page="14" pageId="11" pageNumber="12" year="2016">
<bibRefCitation id="B0294B79FFBDFFD1FE00EEBBFD66102A" author="Enghoff H." box="[449,673,1019,1046]" pageId="11" pageNumber="12" pagination="1 - 23" refId="ref11582" refString="Enghoff H. 2016 b. A mountain of millipedes IV: Species of Prionopetalum Attems, 1909, from the Udzungwa Mountains, Tanzania. With notes on &quot; P. &quot; fasciatum (Attems, 1896) and a revised species key (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 215: 1 - 23. https: // doi. org / 10.5852 / ejt. 2016.215" type="journal article" year="2016">Enghoff 2016b: 14</bibRefCitation>
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.
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<caption id="80C76600FFBDFFD1FF7AEA25FE8813D5" ID-DOI="http://doi.org/10.5281/zenodo.3755007" ID-Zenodo-Dep="3755007" httpUri="https://zenodo.org/record/3755007/files/figure.png" pageId="11" pageNumber="12" startId="11.[187,230,1893,1919]" targetBox="[193,1391,1095,1856]" targetPageId="11">
<paragraph id="D4073688FFBDFFD1FF7AEA25FE8813D5" blockId="11.[187,1397,1893,2025]" pageId="11" pageNumber="12">
<emphasis id="E6CCEA9AFFBDFFD1FF7AEA25FECE1343" bold="true" box="[187,265,1893,1919]" pageId="11" pageNumber="12">Fig. 9.</emphasis>
<taxonomicName id="13B84D0BFFBDFFD1FED3EA26FE4E13BC" authorityName="Frederiksen" authorityYear="2013" box="[274,393,1894,1920]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="E6CCEA9AFFBDFFD1FED3EA26FE4E13BC" box="[274,393,1894,1920]" italics="true" pageId="11" pageNumber="12">Aquattuor</emphasis>
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spp. Body size (number of podous rings and midbody vertical diameter) in males of
<taxonomicName id="13B84D0BFFBDFFD1FF7AEAC9FEF5139F" authorityName="Frederiksen" authorityYear="2013" box="[187,306,1929,1955]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="E6CCEA9AFFBDFFD1FF7AEAC9FEF5139F" box="[187,306,1929,1955]" italics="true" pageId="11" pageNumber="12">Aquattuor</emphasis>
</taxonomicName>
species from the Udzungwa Mts, and of
<taxonomicName id="13B84D0BFFBDFFD1FCF1EAC9FAB2139F" authority="Enghoff &amp; Frederiksen, 2015" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[816,1397,1928,1955]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBDFFD1FCF1EAC9FBC0139E" box="[816,1031,1928,1955]" italics="true" pageId="11" pageNumber="12">A. claudiahempae</emphasis>
<bibRefCitation id="B0294B79FFBDFFD1FBD3EAC8FAB2139F" author="Enghoff H. &amp; Frederiksen S. B." box="[1042,1397,1928,1955]" pageId="11" pageNumber="12" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen, 2015</bibRefCitation>
</taxonomicName>
from Mt Kilimanjaro. In case of (almost) coinciding values, symbols have been slightly displaced horizontally.
</paragraph>
</caption>
<caption id="80C76600FFBAFFD6FF7CEA6FFAAD13EB" ID-DOI="http://doi.org/10.5281/zenodo.3755009" ID-Zenodo-Dep="3755009" httpUri="https://zenodo.org/record/3755009/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1839,1865]" targetBox="[241,1350,283,1781]" targetPageId="12">
<paragraph id="D4073688FFBAFFD6FF7CEA6FFAAD13EB" blockId="12.[189,1399,1839,2007]" pageId="12" pageNumber="13">
<emphasis id="E6CCEA9AFFBAFFD6FF7CEA6FFEDD1376" bold="true" box="[189,282,1839,1866]" pageId="12" pageNumber="13">Fig. 10.</emphasis>
<taxonomicName id="13B84D0BFFBAFFD6FEE2EA70FC7A1376" authority="Enghoff &amp; Frederiksen, 2015" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[291,957,1839,1866]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBAFFD6FEE2EA70FD921376" box="[291,597,1840,1866]" italics="true" pageId="12" pageNumber="13">Aquattuor claudiahempae</emphasis>
<bibRefCitation id="B0294B79FFBAFFD6FD9FEA70FC7A1376" author="Enghoff H. &amp; Frederiksen S. B." box="[606,957,1839,1866]" pageId="12" pageNumber="13" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen, 2015</bibRefCitation>
</taxonomicName>
, male, specimen from the Udzungwa Mts (NHMD 621639), left gonopod.
<emphasis id="E6CCEA9AFFBAFFD6FDB9EA13FD761351" bold="true" box="[632,689,1875,1901]" pageId="12" pageNumber="13">A–C</emphasis>
. Coxa.
<emphasis id="E6CCEA9AFFBAFFD6FCCFEA13FCE41351" bold="true" box="[782,803,1875,1901]" pageId="12" pageNumber="13">A</emphasis>
. Anterior view.
<emphasis id="E6CCEA9AFFBAFFD6FC24EA13FC3E1351" bold="true" box="[997,1017,1875,1901]" pageId="12" pageNumber="13">B</emphasis>
. Mesal view.
<emphasis id="E6CCEA9AFFBAFFD6FB61EA13FB721351" bold="true" box="[1184,1205,1875,1901]" pageId="12" pageNumber="13">C</emphasis>
. Posterior view.
<emphasis id="E6CCEA9AFFBAFFD6FF7CEA36FF3F13AC" bold="true" box="[189,248,1910,1937]" pageId="12" pageNumber="13">D–G</emphasis>
. Telopodite.
<emphasis id="E6CCEA9AFFBAFFD6FE4EEA36FE6313AC" bold="true" box="[399,420,1910,1936]" pageId="12" pageNumber="13">D</emphasis>
. Tip of telomere.
<emphasis id="E6CCEA9AFFBAFFD6FDB3EA36FD4113AC" bold="true" box="[626,646,1910,1936]" pageId="12" pageNumber="13">E</emphasis>
. Anterior view.
<emphasis id="E6CCEA9AFFBAFFD6FCFEEA36FC9613AC" bold="true" box="[831,849,1910,1936]" pageId="12" pageNumber="13">F</emphasis>
. Posterior view.
<emphasis id="E6CCEA9AFFBAFFD6FBD2EA36FBED13AC" bold="true" box="[1043,1066,1910,1936]" pageId="12" pageNumber="13">G</emphasis>
. Mesal view. Abbreviations:
<emphasis id="E6CCEA9AFFBAFFD6FF7CEAD9FF1B138F" box="[189,220,1945,1971]" italics="true" pageId="12" pageNumber="13">btl</emphasis>
= basal telomeral lamella;
<emphasis id="E6CCEA9AFFBAFFD6FDD6EAD9FDE0138F" box="[535,551,1945,1971]" italics="true" pageId="12" pageNumber="13">li</emphasis>
= lateral incision;
<emphasis id="E6CCEA9AFFBAFFD6FCC0EAD9FCEA138F" box="[769,813,1945,1971]" italics="true" pageId="12" pageNumber="13">mbl</emphasis>
= meso-basal lobe of palette;
<emphasis id="E6CCEA9AFFBAFFD6FB4AEADAFB6F138F" box="[1163,1192,1946,1971]" italics="true" pageId="12" pageNumber="13">mi</emphasis>
= mesal incision;
<emphasis id="E6CCEA9AFFBAFFD6FF7CEAFCFF2E13EA" box="[189,233,1980,2006]" italics="true" pageId="12" pageNumber="13">mpl</emphasis>
= mesal-posterior telomeral lamella;
<emphasis id="E6CCEA9AFFBAFFD6FD63EAFDFD7813EA" box="[674,703,1981,2006]" italics="true" pageId="12" pageNumber="13">pa</emphasis>
= palette. Scale bars: A–C, E–G = 0.1 mm; D = 0.02 mm.
</paragraph>
</caption>
<subSubSection id="9CA26503FFBBFFD7FF7CEC4BFDBD179A" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="D4073688FFBBFFD7FF7CEC4BFEF11519" blockId="13.[189,310,267,293]" box="[189,310,267,293]" pageId="13" pageNumber="14">
<heading id="8F4F81E4FFBBFFD7FF7CEC4BFEF11519" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="E6CCEA9AFFBBFFD7FF7CEC4BFEF11519" bold="true" box="[189,310,267,293]" pageId="13" pageNumber="14">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFBBFFD7FF7CEC79FD7B1676" blockId="13.[189,1399,313,586]" pageId="13" pageNumber="14">
Diameter
<quantity id="13409B6DFFBBFFD7FEF0EC79FE191568" box="[305,478,313,340]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" metricValueMax="1.91" metricValueMin="1.55" pageId="13" pageNumber="14" unit="mm" value="1.73" valueMax="1.91" valueMin="1.55">1.55–1.91 mm</quantity>
, 51–53 (55?) podous rings. Shares a short (as broad as long, or slightly longer) gonopodal palette with
<taxonomicName id="13B84D0BFFBBFFD7FE0AEC1DFD5A154B" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[459,669,349,375]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBBFFD7FE0AEC1DFD5A154B" box="[459,669,349,375]" italics="true" pageId="13" pageNumber="14">A. claudiahempae</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFBBFFD7FD68EC1DFCF7154B" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[681,816,349,375]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFBBFFD7FD68EC1DFCF7154B" box="[681,816,349,375]" italics="true" pageId="13" pageNumber="14">A. fasciatus</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFBBFFD7FCFDEC1DFC27154B" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[828,992,349,375]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="E6CCEA9AFFBBFFD7FCFDEC1DFC27154B" box="[828,992,349,375]" italics="true" pageId="13" pageNumber="14">A. stereosathe</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFBBFFD7FC2AEC1DFBBE154B" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1003,1145,349,375]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="E6CCEA9AFFBBFFD7FC2AEC1DFBBE154B" box="[1003,1145,349,375]" italics="true" pageId="13" pageNumber="14">A. submajor</emphasis>
</taxonomicName>
and
<taxonomicName id="13B84D0BFFBBFFD7FB6FEC1DFAA8154B" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1198,1391,349,375]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="udzungwensis">
<emphasis id="E6CCEA9AFFBBFFD7FB6FEC1DFAA8154B" box="[1198,1391,349,375]" italics="true" pageId="13" pageNumber="14">A. udzungwensis</emphasis>
</taxonomicName>
. Differs from these species, except
<taxonomicName id="13B84D0BFFBBFFD7FD8EECC0FCE515A6" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[591,802,384,410]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBBFFD7FD8EECC0FCE515A6" box="[591,802,384,410]" italics="true" pageId="13" pageNumber="14">A. claudiahempae</emphasis>
</taxonomicName>
, by the combination of fully developed telopodites on the first pair of male legs, the gonopodal telomere describing a ca 270° curve, curving almost in one plane only, by lacking a spinose lid-like flap on the basal telomeral lamella, and by having a distinct meso-basal lobe of the gonopod palette. Very similar to, and perhaps not distinct from
<taxonomicName id="13B84D0BFFBBFFD7FB5CECAAFAA8163F" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[1181,1391,489,516]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBBFFD7FB5CECAAFAA8163F" box="[1181,1391,489,516]" italics="true" pageId="13" pageNumber="14">A. claudiahempae</emphasis>
</taxonomicName>
, differs by being mostly larger (male diameter&gt;
<quantity id="13409B6DFFBBFFD7FD31EF4CFC8B161B" box="[752,844,524,551]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="14" unit="mm" value="1.5">1.5 mm</quantity>
) and by having the prefemoral processes of the first pair of male legs stouter, subtriangular.
</paragraph>
<paragraph id="D4073688FFBBFFD7FF7CEF2DFDD016B4" blockId="13.[189,535,621,648]" box="[189,535,621,648]" pageId="13" pageNumber="14">
<emphasis id="E6CCEA9AFFBBFFD7FF7CEF2DFE4B16BB" bold="true" box="[189,396,621,647]" pageId="13" pageNumber="14">Material studied</emphasis>
(total: 1 ♂)
</paragraph>
<paragraph id="D4073688FFBBFFD7FF7CEFDCFAE3168A" blockId="13.[189,1316,668,695]" box="[189,1316,668,695]" pageId="13" pageNumber="14">
<materialsCitation id="64D03CD5FFBBFFD7FF7CEFDCFAE6168A" ID-GBIF-Occurrence="2598688689" box="[189,1313,668,695]" collectingDate="1979-06-16" collectionCode="NHMD" collectorName="M. Stoltze" country="Tanzania" location="TANZANIA" pageId="13" pageNumber="14" specimenCode="NHMD 621641" specimenCount="1" specimenCount-male="1" stateProvince="Kaskazini Unguja">
<collectingCountry id="ACAF7618FFBBFFD7FF7CEFDCFE96168A" box="[189,337,668,694]" name="Tanzania" pageId="13" pageNumber="14">TANZANIA</collectingCountry>
•
<specimenCount id="C2BEFD01FFBBFFD7FEA8EFDCFE50168B" box="[361,407,668,695]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
;
<collectingCountry id="ACAF7618FFBBFFD7FE64EFDCFDC8168A" box="[421,527,668,694]" name="Tanzania" pageId="13" pageNumber="14">Zanzibar</collectingCountry>
, the sultan’s palace;
<date id="A0061048FFBBFFD7FCC0EFDCFC5C168A" box="[769,923,668,695]" pageId="13" pageNumber="14" value="1979-06-16">
<collectingDate id="B042E9A0FFBBFFD7FCC0EFDCFC5C168A" box="[769,923,668,695]" pageId="13" pageNumber="14" value="1979-06-16">16 Jun. 1979</collectingDate>
</date>
;
<collectorName id="794D535EFFBBFFD7FC66EFDDFBE6168A" box="[935,1057,668,695]" pageId="13" pageNumber="14">M. Stoltze</collectorName>
leg.;
<specimenCode id="841E9EF3FFBBFFD7FBA5EFDDFAE6168A" box="[1124,1313,668,695]" collectionCode="NHMD" pageId="13" pageNumber="14">NHMD 621641</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="D4073688FFBBFFD7FF7CEF99FE5416C8" blockId="13.[189,403,729,756]" box="[189,403,729,756]" pageId="13" pageNumber="14">
<heading id="8F4F81E4FFBBFFD7FF7CEF99FE5416C8" bold="true" box="[189,403,729,756]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="E6CCEA9AFFBBFFD7FF7CEF99FE5416C8" bold="true" box="[189,403,729,756]" pageId="13" pageNumber="14">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFBBFFD7FF7CEE48FB941754" blockId="13.[189,1398,776,873]" pageId="13" pageNumber="14">
First pair of male legs (
<figureCitation id="4C832A0DFFBBFFD7FE16EE48FDDC171E" box="[471,539,776,802]" captionStart="Fig" captionStartId="5.[189,232,1849,1875]" captionTargetBox="[205,1381,684,1788]" captionTargetId="figure@5.[189,1399,675,1801]" captionTargetPageId="5" captionText="Fig. 3. Aquattuor spp., first pair of male legs. A–C. A. fasciatus (Attems, 1896), non-type male from Zanzibar (NHMD 621641). D–F. A. claudiahempae Enghoff &amp; Frederiksen, 2015, paratype from Kilimanjaro (NHMD 621638). G–I. A. claudiahempae, specimen from Udzungwa Mts (NHMD 621639). A, D, G. Anterior view. B, E, H. Sublateral view. C, F, I. Ventral view. Abbreviation: pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754995" httpUri="https://zenodo.org/record/3754995/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
A–C) each with a single, relatively stout, triangular prefemoral process; prefemoral processes stouter than those of the otherwise very similar
<taxonomicName id="13B84D0BFFBBFFD7FBE6EE6CFAC71779" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[1063,1280,811,838]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBBFFD7FBE6EE6CFAC71779" box="[1063,1280,811,838]" italics="true" pageId="13" pageNumber="14">A. claudiahempae</emphasis>
</taxonomicName>
; setae of telopodite distal to prefemur similar in length to those of normal walking legs.
</paragraph>
<paragraph id="D4073688FFBBFFD7FF7CEECCFDBD179A" blockId="13.[189,634,908,935]" box="[189,634,908,935]" pageId="13" pageNumber="14">
See remarks under
<emphasis id="E6CCEA9AFFBBFFD7FE5EEECDFDBD179A" box="[415,634,908,935]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="13B84D0BFFBBFFD7FE5EEECDFDB1179A" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[415,630,908,935]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="claudiahempae">A. claudiahempae</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFBBFFD7FF7CEE8AFB281065" pageId="13" pageNumber="14" type="distribution">
<paragraph id="D4073688FFBBFFD7FF7CEE8AFE9117D8" blockId="13.[189,342,970,996]" box="[189,342,970,996]" pageId="13" pageNumber="14">
<heading id="8F4F81E4FFBBFFD7FF7CEE8AFE9117D8" bold="true" box="[189,342,970,996]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="E6CCEA9AFFBBFFD7FF7CEE8AFE9117D8" bold="true" box="[189,342,970,996]" pageId="13" pageNumber="14">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFBBFFD7FF7CEEB8FB281065" blockId="13.[189,1399,1016,1114]" pageId="13" pageNumber="14">
Recorded from
<collectingCountry id="ACAF7618FFBBFFD7FEBEEEB8FE20102E" box="[383,487,1016,1042]" name="Tanzania" pageId="13" pageNumber="14">Zanzibar</collectingCountry>
(
<typeStatus id="0B03882AFFBBFFD7FE3DEEB9FDF7102F" box="[508,560,1017,1043]" pageId="13" pageNumber="14">type</typeStatus>
locality),
<collectingRegion id="167CF86AFFBBFFD7FD73EEB8FC98102E" box="[690,863,1016,1042]" country="Tanzania" name="Pwani" pageId="13" pageNumber="14">Pwani Region</collectingRegion>
, Bagamoyo Distr., Vula Mountain, Pongwe (
<bibRefCitation id="B0294B79FFBBFFD7FF04E95CFD94100A" author="VandenSpiegel D. &amp; Pierrard G." box="[197,595,1052,1078]" pageId="13" pageNumber="14" pagination="149 - 163" refId="ref12440" refString="VandenSpiegel D. &amp; Pierrard G. 2009. Revision du genre Prionopetalum (Odontopygidae, Diplopoda) et descriptions de nouvelles especes d'Afrique de l'Est. Journal of Afrotropical Zoology 5: 149 - 163." type="journal article" year="2009">VandenSpiegel &amp; Pierrard, 2009</bibRefCitation>
, as
<taxonomicName id="13B84D0BFFBBFFD7FD44E95CFC75100A" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[645,946,1052,1078]" class="Diplopoda" family="Odontopygidae" genus="Prionopetalum" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fasciatum">
<emphasis id="E6CCEA9AFFBBFFD7FD44E95CFC75100A" box="[645,946,1052,1078]" italics="true" pageId="13" pageNumber="14">Prionopetalum fasciatum</emphasis>
</taxonomicName>
) and from
<collectingRegion id="167CF86AFFBBFFD7FB80E95CFADC100A" box="[1089,1307,1052,1078]" country="Tanzania" name="Morogoro" pageId="13" pageNumber="14">Morogoro Region</collectingRegion>
, Kilosa District, Rubeho Mts,
<quantity id="13409B6DFFBBFFD7FE02E97FFDCA1065" box="[451,525,1087,1113]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="14" unit="m" value="900.0">900 m</quantity>
a.s.l. (
<bibRefCitation id="B0294B79FFBBFFD7FD9BE97FFC761066" author="Enghoff H. &amp; Frederiksen S. B." box="[602,945,1087,1114]" pageId="13" pageNumber="14" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen 2015</bibRefCitation>
, as
<taxonomicName id="13B84D0BFFBBFFD7FC1DE97FFB191065" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[988,1246,1087,1113]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBBFFD7FC1DE97FFC321065" box="[988,1013,1087,1113]" italics="true" pageId="13" pageNumber="14">A.</emphasis>
aff.
<emphasis id="E6CCEA9AFFBBFFD7FBEAE97FFB191065" box="[1067,1246,1087,1113]" italics="true" pageId="13" pageNumber="14">claudiahempae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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