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<document id="CBC590232B4465E4C186556FC3FFD59B" ID-DOI="10.5852/ejt.2020.626" ID-GBIF-Dataset="c906d9cd-cfcb-43db-a24c-b57b39d9184b" ID-Zenodo-Dep="3754989" ID-ZooBank="D91881D6-55D8-48FC-A383-069BC643A91E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587066773978" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2020" docId="5C11879EFFBCFFD1FDE1EFC8FD091685" docLanguage="en" docName="ejt-626_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 626" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor denticulatus Frederiksen 2013" docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="A028FFE6FFB6FFDAFFC1ED40FFC7143C" masterDocTitle="A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="11" updateTime="1698738618004" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DCBB1C78667AF75F118342617870CF70">A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="417E9D5F8DC947FE41B6811086803020">Enghoff, Henrik</mods:namePart>
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<mods:title id="0FBFB30A0536FC089DE255BDEA4F3EFC">European Journal of Taxonomy</mods:title>
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<mods:date id="664F76AABBAE89585BAB573680B8EB21">2020</mods:date>
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<mods:number id="E5075A5D8E03400028DF15D68F9099AA">2020-04-14</mods:number>
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<treatment id="5C11879EFFBCFFD1FDE1EFC8FD091685" ID-GBIF-Taxon="163642160" LSID="urn:lsid:plazi:treatment:5C11879EFFBCFFD1FDE1EFC8FD091685" httpUri="http://treatment.plazi.org/id/5C11879EFFBCFFD1FDE1EFC8FD091685" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="9CA26503FFBCFFD0FDE1EFC8FBD4169E" box="[544,1043,647,674]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="D4073688FFBCFFD0FDE1EFC8FBD4169E" blockId="10.[544,1043,647,709]" box="[544,1043,647,674]" pageId="10" pageNumber="11">
<heading id="8F4F81E4FFBCFFD0FDE1EFC8FBD4169E" box="[544,1043,647,674]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="13B84D0BFFBCFFD0FDE1EFC8FBD4169E" ID-CoL="FZKY" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[544,1043,647,674]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="E6CCEA9AFFBCFFD0FDE1EFC8FCFF169E" bold="true" box="[544,824,647,674]" italics="true" pageId="10" pageNumber="11">Aquattuor denticulatus</emphasis>
<bibRefCitation id="B0294B79FFBCFFD0FCFEEFC8FBD4169E" author="Frederiksen S. B." box="[831,1043,648,674]" pageId="10" pageNumber="11" pagination="59 - 66" refId="ref12038" refString="Frederiksen S. B. 2013. East African odontopygid millipedes 3: Two new genera, Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. https: // doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen, 2013</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFBCFFD0FD0CEFEBFCA016F9" box="[717,871,683,709]" pageId="10" pageNumber="11" type="description">
<paragraph id="D4073688FFBCFFD0FD0CEFEBFCA016F9" blockId="10.[544,1043,647,709]" box="[717,871,683,709]" pageId="10" pageNumber="11">
<figureCitation id="4C832A0DFFBCFFD0FD0CEFEBFCD116F9" box="[717,790,683,709]" captionStart="Fig" captionStartId="2.[189,232,1948,1974]" captionTargetBox="[192,1385,1023,1872]" captionTargetId="figure@2.[186,1391,1009,1876]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites for Aquattuor spp. in the Udzungwa Mts. Based on Marshall et al. (2010: fig. 1); inset by permission of the Eastern Arc Mountains Conservation Endowment Fund." figureDoi="http://doi.org/10.5281/zenodo.3754991" httpUri="https://zenodo.org/record/3754991/files/figure.png" pageId="10" pageNumber="11">Figs 1</figureCitation>
,
<figureCitation id="4C832A0DFFBCFFD0FCE5EFEBFCF516F9" box="[804,818,683,709]" captionStart="Fig" captionStartId="6.[189,232,1890,1916]" captionTargetBox="[226,1373,279,1795]" captionTargetId="figure@6.[189,1399,265,1821]" captionTargetPageId="6" captionText="Fig. 4. A–C. Aquattuor nguruensis sp. nov., holotype, ♂ (VMNH110617), first pair of legs. Notice mite larva (”hypopus”) on left femur. D–F. A. denticulatus Frederiksen, 2013, paratype, ♂ (NHMD 621640), left leg of first pair. A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviation: pfd = prefemoral depression. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754997" httpUri="https://zenodo.org/record/3754997/files/figure.png" pageId="10" pageNumber="11">4</figureCitation>
D–F
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFBCFFD0FF7CEFADFD1D173A" box="[189,730,748,775]" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="D4073688FFBCFFD0FF7CEFADFD1D173A" blockId="10.[189,730,748,775]" box="[189,730,748,775]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="F4A811A6FFBCFFD0FF7CEFADFD1D173A" box="[189,730,748,775]" pageId="10" pageNumber="11">
<treatmentCitation id="55191099FFBCFFD0FF7CEFADFD11173A" ID-DOI="http://doi.org/10.5281/zenodo.6161325" ID-Zenodo-Dep="6161325" author="Frederiksen S. B." box="[189,726,748,775]" httpUri="http://treatment.plazi.org/id/456F277EC46F1A05FF0BFB77FDF730F7" page="64" pageId="10" pageNumber="11" year="2013">
<taxonomicName id="13B84D0BFFBCFFD0FF7CEFADFD11173A" ID-CoL="FZKY" authority="Frederiksen, 2013: 64" authorityName="Frederiksen" authorityPageNumber="64" authorityYear="2013" box="[189,726,748,775]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="E6CCEA9AFFBCFFD0FF7CEFADFE0D173B" box="[189,458,749,775]" italics="true" pageId="10" pageNumber="11">Aquattuor denticulatus</emphasis>
<bibRefCitation id="B0294B79FFBCFFD0FE13EFACFD11173A" author="Frederiksen S. B." box="[466,726,748,775]" pageId="10" pageNumber="11" pagination="59 - 66" refId="ref12038" refString="Frederiksen S. B. 2013. East African odontopygid millipedes 3: Two new genera, Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. https: // doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen, 2013: 64</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFBCFFD1FF7CEE6EFB941616" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="D4073688FFBCFFD0FF7CEE6EFEF11774" blockId="10.[189,310,814,840]" box="[189,310,814,840]" pageId="10" pageNumber="11">
<heading id="8F4F81E4FFBCFFD0FF7CEE6EFEF11774" bold="true" box="[189,310,814,840]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="E6CCEA9AFFBCFFD0FF7CEE6EFEF11774" bold="true" box="[189,310,814,840]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFBCFFD0FF7CEE1CFC8817DC" blockId="10.[189,1399,860,993]" pageId="10" pageNumber="11">
Diameter
<quantity id="13409B6DFFBCFFD0FEF7EE1CFE2F174B" box="[310,488,860,887]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="1.77" metricValueMin="1.58" pageId="10" pageNumber="11" unit="mm" value="1.675" valueMax="1.77" valueMin="1.58">1.58–1.77 mm</quantity>
, 50–52 podous rings. Shares a short (as broad as long, or slightly longer) gonopodal palette with
<taxonomicName id="13B84D0BFFBCFFD0FE0AEEC0FD5A17A6" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[459,669,896,922]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBCFFD0FE0AEEC0FD5A17A6" box="[459,669,896,922]" italics="true" pageId="10" pageNumber="11">A. claudiahempae</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFBCFFD0FD68EEC0FCF717A6" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[681,816,896,922]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFBCFFD0FD68EEC0FCF717A6" box="[681,816,896,922]" italics="true" pageId="10" pageNumber="11">A. fasciatus</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFBCFFD0FCFDEEC0FC2717A6" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[828,992,896,922]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="E6CCEA9AFFBCFFD0FCFDEEC0FC2717A6" box="[828,992,896,922]" italics="true" pageId="10" pageNumber="11">A. stereosathe</emphasis>
</taxonomicName>
,
<taxonomicName id="13B84D0BFFBCFFD0FC2AEEC0FBBE17A6" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1003,1145,896,922]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="E6CCEA9AFFBCFFD0FC2AEEC0FBBE17A6" box="[1003,1145,896,922]" italics="true" pageId="10" pageNumber="11">A. submajor</emphasis>
</taxonomicName>
and
<taxonomicName id="13B84D0BFFBCFFD0FB6FEEC0FAA817A6" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1198,1391,896,922]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="udzungwensis">
<emphasis id="E6CCEA9AFFBCFFD0FB6FEEC0FAA817A6" box="[1198,1391,896,922]" italics="true" pageId="10" pageNumber="11">A. udzungwensis</emphasis>
</taxonomicName>
. Differs from these species by having the subdistal part of the gonopodal telomere almost straight, parallel to the coxa, the telomere tip being bent abruptly laterad.
</paragraph>
<caption id="80C76600FFBCFFD0FF7CEA16FAFD13E7" ID-DOI="http://doi.org/10.5281/zenodo.3755005" ID-Zenodo-Dep="3755005" httpUri="https://zenodo.org/record/3755005/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1878,1904]" targetBox="[205,1373,1111,1837]" targetPageId="10">
<paragraph id="D4073688FFBCFFD0FF7CEA16FAFD13E7" blockId="10.[189,1399,1877,2011]" pageId="10" pageNumber="11">
<emphasis id="E6CCEA9AFFBCFFD0FF7CEA16FED7134C" bold="true" box="[189,272,1878,1904]" pageId="10" pageNumber="11">Fig. 8.</emphasis>
<taxonomicName id="13B84D0BFFBCFFD0FEDCEA17FE53134D" authorityName="Frederiksen" authorityYear="2013" box="[285,404,1879,1905]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="E6CCEA9AFFBCFFD0FEDCEA17FE53134D" box="[285,404,1879,1905]" italics="true" pageId="10" pageNumber="11">Aquattuor</emphasis>
</taxonomicName>
spp., males, left leg of first pair.
<emphasis id="E6CCEA9AFFBCFFD0FC82EA16FC9F134C" bold="true" box="[835,856,1878,1904]" pageId="10" pageNumber="11">A</emphasis>
.
<taxonomicName id="13B84D0BFFBCFFD0FCACEA17FB11134D" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[877,1238,1878,1905]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="E6CCEA9AFFBCFFD0FCACEA17FBDF134C" box="[877,1048,1878,1905]" italics="true" pageId="10" pageNumber="11">A. stereosathe</emphasis>
<bibRefCitation id="B0294B79FFBCFFD0FBE5EA16FB11134D" author="Enghoff H. &amp; Frederiksen S. B." box="[1060,1238,1878,1905]" pageId="10" pageNumber="11" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
, paratype, ♂ (NHMD 621650): setae same length as in normal walking legs.
<emphasis id="E6CCEA9AFFBCFFD0FC31EA3AFBC313A8" bold="true" box="[1008,1028,1914,1940]" pageId="10" pageNumber="11">B</emphasis>
.
<taxonomicName id="13B84D0BFFBCFFD0FBD8EA3AFAB513A8" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1049,1394,1913,1940]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="longipala">
<emphasis id="E6CCEA9AFFBCFFD0FBD8EA3AFB7613A8" box="[1049,1201,1914,1940]" italics="true" pageId="10" pageNumber="11">A. longipala</emphasis>
<bibRefCitation id="B0294B79FFBCFFD0FB7EEA3AFAB513A8" author="Enghoff H. &amp; Frederiksen S. B." box="[1215,1394,1913,1940]" pageId="10" pageNumber="11" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
, specimen from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642): setae of femur shortened.
<emphasis id="E6CCEA9AFFBCFFD0FF7CEA80FF1513E6" bold="true" box="[189,210,1984,2010]" pageId="10" pageNumber="11">C</emphasis>
.
<taxonomicName id="13B84D0BFFBCFFD0FF20EA80FEBA13E6" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[225,381,1984,2010]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
<emphasis id="E6CCEA9AFFBCFFD0FF20EA80FEBA13E6" box="[225,381,1984,2010]" italics="true" pageId="10" pageNumber="11">A. mollilobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDFF57E1FFBCFFD0FE44EA81FE2513E7" box="[389,482,1985,2011]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (NHMD 621646): all setae shortened. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="D4073688FFBDFFD1FF7CEC4BFAB01568" blockId="11.[189,535,269,293]" lastBlockId="11.[189,1399,314,411]" pageId="11" pageNumber="12">
<emphasis id="E6CCEA9AFFBDFFD1FF7CEC4BFE4B1519" bold="true" box="[189,396,267,293]" pageId="11" pageNumber="12">Material studied</emphasis>
(total: 1 ♂)
<materialsCitation id="64D03CD5FFBDFFD1FF7CEC7AFAB01568" ID-GBIF-Occurrence="2598688697" box="[189,1399,312,340]" collectingDate="1977-02-07" collectionCode="NHMD, ZMUC" collectorName="H. Enghoff &amp; O. Lomhodlt &amp; O. Martin" country="Tanzania" county="Muheza District" elevation="500" location="Amani" municipality="East Usambara Mts" pageId="11" pageNumber="12" specimenCode="621640" specimenCount="1" specimenCount-male="1" stateProvince="Tanga Region" typeStatus="paratype">
<collectingCountry id="ACAF7618FFBDFFD1FF7CEC7AFE951568" box="[189,338,314,340]" name="Tanzania" pageId="11" pageNumber="12">TANZANIA</collectingCountry>
•
<specimenCount id="C2BEFD01FFBDFFD1FEB2EC7AFE601568" box="[371,423,312,340]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
,
<typeStatus id="0B03882AFFBDFFD1FE77EC7AFDE71568" box="[438,544,314,340]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
;
<collectingRegion id="167CF86AFFBDFFD1FDF1EC7AFD1B1568" box="[560,732,314,340]" country="Tanzania" name="Tanga" pageId="11" pageNumber="12">Tanga Region</collectingRegion>
,
<collectingCounty id="3D664E04FFBDFFD1FD2AEC7AFC751568" box="[747,946,314,340]" pageId="11" pageNumber="12">Muheza District</collectingCounty>
,
<collectingMunicipality id="3463ACF2FFBDFFD1FC03EC7AFB721568" box="[962,1205,314,340]" pageId="11" pageNumber="12">East Usambara Mts</collectingMunicipality>
,
<location id="D1676053FFBDFFD1FB02EC7AFAD01568" LSID="urn:lsid:plazi:treatment:5C11879EFFBCFFD1FDE1EFC8FD091685:D1676053FFBDFFD1FB02EC7AFAD01568" box="[1219,1303,314,340]" country="Tanzania" county="Muheza District" municipality="East Usambara Mts" name="Amani" pageId="11" pageNumber="12" stateProvince="Tanga Region">Amani</location>
, at
<location id="D1676053FFBDFFD1FA86EC7AFAB01568" LSID="urn:lsid:plazi:treatment:5C11879EFFBCFFD1FDE1EFC8FD091685:D1676053FFBDFFD1FA86EC7AFAB01568" box="[1351,1399,314,340]" country="Tanzania" county="Muheza District" municipality="East Usambara Mts" name="Sigi" pageId="11" pageNumber="12" stateProvince="Tanga Region">Sigi</location>
</materialsCitation>
</paragraph>
<paragraph id="D4073688FFBDFFD1FF7CEC1DFE7A15A7" blockId="11.[189,1399,314,411]" pageId="11" pageNumber="12">
River;
<quantity id="13409B6DFFBDFFD1FED5EC1DFEA21544" box="[276,357,349,376]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="500.0">
<elevation id="5F95D1BBFFBDFFD1FED5EC1DFEA21544" box="[276,357,349,376]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="500.0">500 m</elevation>
</quantity>
a.s.l.;
<date id="A0061048FFBDFFD1FE7FEC1EFD9D154B" box="[446,602,349,376]" pageId="11" pageNumber="12" value="1977-02-07">
<collectingDate id="B042E9A0FFBDFFD1FE7FEC1EFD9D154B" box="[446,602,349,376]" pageId="11" pageNumber="12" value="1977-02-07">7 Feb. 1977</collectingDate>
</date>
;
<collectorName id="794D535EFFBDFFD1FDADEC1EFD3E154B" box="[620,761,349,376]" pageId="11" pageNumber="12">H. Enghoff</collectorName>
,
<collectorName id="794D535EFFBDFFD1FCCBEC1DFC6E154B" box="[778,937,349,375]" pageId="11" pageNumber="12">O. Lomhodlt</collectorName>
and
<collectorName id="794D535EFFBDFFD1FC2FEC1DFBAE154B" box="[1006,1129,349,375]" pageId="11" pageNumber="12">O. Martin</collectorName>
leg.; NHMD
<specimenCode id="841E9EF3FFBDFFD1FADFEC1DFAB0154B" box="[1310,1399,349,375]" pageId="11" pageNumber="12">621640</specimenCode>
[=ZMUC 00020528].
</paragraph>
<paragraph id="D4073688FFBDFFD1FF7CECFEFE5415E4" blockId="11.[189,403,446,472]" box="[189,403,446,472]" pageId="11" pageNumber="12">
<heading id="8F4F81E4FFBDFFD1FF7CECFEFE5415E4" bold="true" box="[189,403,446,472]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="E6CCEA9AFFBDFFD1FF7CECFEFE5415E4" bold="true" box="[189,403,446,472]" pageId="11" pageNumber="12">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFBDFFD1FF7CECACFB941616" blockId="11.[189,1398,492,554]" pageId="11" pageNumber="12">
First pair of male legs (
<figureCitation id="4C832A0DFFBDFFD1FE29ECACFDF6163B" box="[488,561,492,519]" captionStart="Fig" captionStartId="6.[189,232,1890,1916]" captionTargetBox="[226,1373,279,1795]" captionTargetId="figure@6.[189,1399,265,1821]" captionTargetPageId="6" captionText="Fig. 4. A–C. Aquattuor nguruensis sp. nov., holotype, ♂ (VMNH110617), first pair of legs. Notice mite larva (”hypopus”) on left femur. D–F. A. denticulatus Frederiksen, 2013, paratype, ♂ (NHMD 621640), left leg of first pair. A, D. Anterior view. B, E. Sublateral view. C, F. Sub-ventral view. Abbreviation: pfd = prefemoral depression. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754997" httpUri="https://zenodo.org/record/3754997/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
D–F) each with a single, finger-shaped prefemoral process; setae of telopodite distal to prefemur similar in length to those of normal walking legs.
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFBDFFD1FF7CEF0DFD091685" pageId="11" pageNumber="12" type="distribution">
<paragraph id="D4073688FFBDFFD1FF7CEF0DFE91165B" blockId="11.[189,342,589,615]" box="[189,342,589,615]" pageId="11" pageNumber="12">
<heading id="8F4F81E4FFBDFFD1FF7CEF0DFE91165B" bold="true" box="[189,342,589,615]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="E6CCEA9AFFBDFFD1FF7CEF0DFE91165B" bold="true" box="[189,342,589,615]" pageId="11" pageNumber="12">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFBDFFD1FF7CEF3CFD091685" blockId="11.[189,1398,636,698]" pageId="11" pageNumber="12">
Known only from the
<typeStatus id="0B03882AFFBDFFD1FE7AEF3CFE2916AA" box="[443,494,636,662]" pageId="11" pageNumber="12">type</typeStatus>
locality,
<collectingRegion id="167CF86AFFBDFFD1FD97EF3CFD3C16AA" box="[598,763,636,662]" country="Tanzania" name="Tanga" pageId="11" pageNumber="12">Tanga Region</collectingRegion>
, Muheza District, E Usambara Mts, Amani.Altitudinal range:
<quantity id="13409B6DFFBDFFD1FECFEFDFFE661686" box="[270,417,671,698]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="11" pageNumber="12" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">
500–
<date id="A0061048FFBDFFD1FE88EFDFFE451686" box="[329,386,671,698]" pageId="11" pageNumber="12" value="1000">1000</date>
m
</quantity>
a.s.l. (
<bibRefCitation id="B0294B79FFBDFFD1FE2FEFDFFD051685" author="Frederiksen S. B." box="[494,706,671,697]" pageId="11" pageNumber="12" pagination="59 - 66" refId="ref12038" refString="Frederiksen S. B. 2013. East African odontopygid millipedes 3: Two new genera, Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. https: // doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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