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<mods:title id="29F43ACCF263C7FB56CB2E570F8AC597">A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<taxonomicName id="13B84D0BFFB2FFDEFE0DE875FBA01173" ID-CoL="FZKX" authority="Enghoff" authorityName="Enghoff" authorityYear="2015" box="[460,1127,1332,1359]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFB2FFDEFE0DE875FCC01173" bold="true" box="[460,775,1332,1359]" italics="true" pageId="4" pageNumber="5">Aquattuor claudiahempae</emphasis>
<bibRefCitation id="B0294B79FFB2FFDEFCCFE874FBA01173" author="Enghoff H. &amp; Frederiksen S. B." box="[782,1127,1332,1359]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen, 2015</bibRefCitation>
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<paragraph id="D4073688FFB2FFDEFD6AE818FC40114E" blockId="4.[460,1127,1332,1394]" box="[683,903,1368,1394]" pageId="4" pageNumber="5">
<figureCitation id="4C832A0DFFB2FFDEFD6AE818FD33114E" box="[683,756,1368,1394]" captionStart="Fig" captionStartId="2.[189,232,1948,1974]" captionTargetBox="[192,1385,1023,1872]" captionTargetId="figure@2.[186,1391,1009,1876]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites for Aquattuor spp. in the Udzungwa Mts. Based on Marshall et al. (2010: fig. 1); inset by permission of the Eastern Arc Mountains Conservation Endowment Fund." figureDoi="http://doi.org/10.5281/zenodo.3754991" httpUri="https://zenodo.org/record/3754991/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
, 
<figureCitation id="4C832A0DFFB2FFDEFCC3E818FCC8114E" box="[770,783,1368,1394]" captionStart="Fig" captionStartId="5.[189,232,1849,1875]" captionTargetBox="[205,1381,684,1788]" captionTargetId="figure@5.[189,1399,675,1801]" captionTargetPageId="5" captionText="Fig. 3. Aquattuor spp., first pair of male legs. A–C. A. fasciatus (Attems, 1896), non-type male from Zanzibar (NHMD 621641). D–F. A. claudiahempae Enghoff &amp; Frederiksen, 2015, paratype from Kilimanjaro (NHMD 621638). G–I. A. claudiahempae, specimen from Udzungwa Mts (NHMD 621639). A, D, G. Anterior view. B, E, H. Sublateral view. C, F, I. Ventral view. Abbreviation: pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754995" httpUri="https://zenodo.org/record/3754995/files/figure.png" pageId="4" pageNumber="5">3</figureCitation>
D–I, 9–10
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<taxonomicName id="13B84D0BFFB2FFDEFF7CE8DAFCB9118F" ID-CoL="FZKX" authority="Enghoff" authorityName="Enghoff" authorityPageNumber="17" authorityYear="2015" box="[189,894,1433,1460]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFB2FFDEFF7CE8DAFE29118F" box="[189,494,1433,1460]" italics="true" pageId="4" pageNumber="5">Aquattuor claudiahempae</emphasis>
<bibRefCitation id="B0294B79FFB2FFDEFE34E8D9FCB9118F" author="Enghoff H. &amp; Frederiksen S. B." box="[501,894,1433,1460]" pageId="4" pageNumber="5" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen, 2015: 17</bibRefCitation>
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.
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<emphasis id="E6CCEA9AFFB2FFDEFF7CE89AFEF111C8" bold="true" box="[189,310,1498,1524]" pageId="4" pageNumber="5">Diagnosis</emphasis>
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Diameter 
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, 50–53 podous rings (
<figureCitation id="4C832A0DFFB2FFDEFD17EB49FCDB121F" box="[726,796,1545,1571]" captionStart="Fig" captionStartId="11.[187,230,1893,1919]" captionTargetBox="[193,1391,1095,1856]" captionTargetId="figure@11.[192,1402,1089,1860]" captionTargetPageId="11" captionText="Fig. 9. Aquattuor spp. Body size (number of podous rings and midbody vertical diameter) in males of Aquattuor species from the Udzungwa Mts, and of A. claudiahempae Enghoff &amp; Frederiksen, 2015 from Mt Kilimanjaro. In case of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3755007" httpUri="https://zenodo.org/record/3755007/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
). Shares a short (as broad as long, or slightly longer) gonopodal palette with 
<taxonomicName id="13B84D0BFFB2FFDEFE10EB6DFD46127A" authorityName="Frederiksen" authorityYear="2013" box="[465,641,1580,1607]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="E6CCEA9AFFB2FFDEFE10EB6DFD46127A" box="[465,641,1580,1607]" italics="true" pageId="4" pageNumber="5">A. denticulatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="13B84D0BFFB2FFDEFD51EB6DFCDE127A" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[656,793,1580,1607]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFB2FFDEFD51EB6DFCDE127A" box="[656,793,1580,1607]" italics="true" pageId="4" pageNumber="5">A. fasciatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="13B84D0BFFB2FFDEFCE6EB6DFC0A127A" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[807,973,1580,1607]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="E6CCEA9AFFB2FFDEFCE6EB6DFC0A127A" box="[807,973,1580,1607]" italics="true" pageId="4" pageNumber="5">A. stereosathe</emphasis>
</taxonomicName>
, 
<taxonomicName id="13B84D0BFFB2FFDEFC1AEB6DFBAD127A" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[987,1130,1580,1607]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="E6CCEA9AFFB2FFDEFC1AEB6DFBAD127A" box="[987,1130,1580,1607]" italics="true" pageId="4" pageNumber="5">A. submajor</emphasis>
</taxonomicName>
, and 
<taxonomicName id="13B84D0BFFB2FFDEFB6AEB6DFAA8127A" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[1195,1391,1580,1607]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="udzungwensis">
<emphasis id="E6CCEA9AFFB2FFDEFB6AEB6DFAA8127A" box="[1195,1391,1580,1607]" italics="true" pageId="4" pageNumber="5">A. udzungwensis</emphasis>
</taxonomicName>
. Differs from these species, except 
<taxonomicName id="13B84D0BFFB2FFDEFD8FEB10FD111256" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[590,726,1616,1642]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFB2FFDEFD8FEB10FD111256" box="[590,726,1616,1642]" italics="true" pageId="4" pageNumber="5">A. fasciatus</emphasis>
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, by the combination of fully developed telopodites on the first pair of male legs, the gonopodal telomere describing a ca 270° curve, curving almost in one plane only, by lacking a spinose lid-like flap on the besal telomeral lamella, and by having a distinct mesobasal lobe of the gonopod palette. Very similar to, and perhaps not distinct from 
<taxonomicName id="13B84D0BFFB2FFDEFBACEBFAFB3112EF" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[1133,1270,1721,1748]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFB2FFDEFBACEBFAFB3112EF" box="[1133,1270,1721,1748]" italics="true" pageId="4" pageNumber="5">A. fasciatus</emphasis>
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, differs by being mostly smaller (male diameter &lt;
<quantity id="13409B6DFFB2FFDEFD49EB9CFD2312CB" box="[648,740,1756,1783]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="5" unit="mm" value="1.7">1.7 mm</quantity>
) and by having the prefemoral processes of the first pair of male legs more slender, finger-shaped.
</paragraph>
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<emphasis id="E6CCEA9AFFB2FFDEFF7CEA01FE4B1367" bold="true" box="[189,396,1857,1883]" pageId="4" pageNumber="5">Material studied</emphasis>
(total: 25 ♂♂, 
<specimenCount id="C2BEFD01FFB2FFDEFD81EA01FD561367" box="[576,657,1853,1883]" pageId="4" pageNumber="5" type="female">29 ♀♀</specimenCount>
) 
<collectingCountry id="ACAF7618FFB2FFDEFF7CEA30FE9613B6" box="[189,337,1904,1930]" name="Tanzania" pageId="4" pageNumber="5">TANZANIA</collectingCountry>
• 
<materialsCitation id="64D03CD5FFB2FFDEFEB7EA30FAB113B6" ID-GBIF-Occurrence="2598688708" box="[374,1398,1902,1930]" collectingDate="2013-11-20" collectionCode="NHMD" collectorName="S. B. Frederiksen" country="Tanzania" elevation="1345" latitude="-3.2331583" location="Hai District" longLatPrecision="1" longitude="37.269245" municipality="Mt Kilimanjaro" pageId="4" pageNumber="5" specimenCode="621638" specimenCount="1" specimenCount-male="1" stateProvince="Kilimanjaro Region" typeStatus="paratype">
<specimenCount id="C2BEFD01FFB2FFDEFEB7EA30FE6E13B6" box="[374,425,1904,1930]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
, 
<typeStatus id="0B03882AFFB2FFDEFE7CEA30FDE113B6" box="[445,550,1904,1930]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
; 
<collectingRegion id="167CF86AFFB2FFDEFDF9EA2FFCEC13B5" box="[568,811,1903,1929]" country="Tanzania" name="Kilimanjaro" pageId="4" pageNumber="5">Kilimanjaro Region</collectingRegion>
, 
<collectingMunicipality id="3463ACF2FFB2FFDEFCFCEA30FC3913B5" box="[829,1022,1903,1930]" pageId="4" pageNumber="5">Mt Kilimanjaro</collectingMunicipality>
, 
<location id="D1676053FFB2FFDEFBD1EA2FFB6313B5" LSID="urn:lsid:plazi:treatment:5C11879EFFB2FFD0FE0DE875FAB01676:D1676053FFB2FFDEFBD1EA2FFB6313B5" box="[1040,1188,1903,1929]" country="Tanzania" latitude="-3.2331583" longLatPrecision="1" longitude="37.269245" municipality="Mt Kilimanjaro" name="Hai District" pageId="4" pageNumber="5" stateProvince="Kilimanjaro Region">Hai District</location>
; 
<geoCoordinate id="B18C504FFFB2FFDEFB77EA30FAB713B6" box="[1206,1392,1902,1930]" degrees="03" direction="south" minutes="13" orientation="latitude" pageId="4" pageNumber="5" precision="1" seconds="59.37" value="-3.2331583">03°13′59.37″ S</geoCoordinate>
,
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</paragraph>
<paragraph id="D4073688FFB2FFDEFF7CEAD3FAB113EC" blockId="4.[189,1398,1903,2000]" pageId="4" pageNumber="5">
<geoCoordinate id="B18C504FFFB2FFDEFF7CEAD3FEBA1391" box="[189,381,1937,1965]" degrees="37" direction="east" minutes="16" orientation="longitude" pageId="4" pageNumber="5" precision="1" seconds="09.28" value="37.269245">37°16′09.28″ E</geoCoordinate>
; 
<quantity id="13409B6DFFB2FFDEFE53EAD2FE341391" box="[402,499,1938,1965]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="4" pageNumber="5" unit="m" value="1345.0">
<elevation id="5F95D1BBFFB2FFDEFE53EAD2FE341391" box="[402,499,1938,1965]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="4" pageNumber="5" unit="m" value="1345.0">1345 m</elevation>
</quantity>
a.s.l.; 
<date id="A0061048FFB2FFDEFD93EAD3FCC21391" box="[594,773,1939,1965]" pageId="4" pageNumber="5" value="2013-11-20">
<collectingDate id="B042E9A0FFB2FFDEFD93EAD3FCC21391" box="[594,773,1939,1965]" pageId="4" pageNumber="5" value="2013-11-20">20 Nov. 2013</collectingDate>
</date>
; 
<collectorName id="794D535EFFB2FFDEFCDBEAD3FC201391" box="[794,999,1939,1965]" pageId="4" pageNumber="5">S.B. Frederiksen</collectorName>
leg.; coffee plantation, Plot 2; NHMD 
<specimenCode id="841E9EF3FFB2FFDEFEE0EAF6FEBE13EC" box="[289,377,1974,2000]" pageId="4" pageNumber="5">621638</specimenCode>
• 
<materialsCitation id="64D03CD5FFB2FFDEFE59EAF6FAB113EC" ID-GBIF-Occurrence="2598688694" box="[408,1398,1974,2000]" collectingDate="2014-10-24" collectingMethod="pitfall trapping (summed catch from eight traps)" collectionCode="NHMD" county="Udzungwa Mts National Park" elevation="650" latitude="-7.6874723" location="Kidatu" longLatPrecision="1" longitude="36.940197" municipality="Kidatu" pageId="4" pageNumber="5" specimenCode="621639" specimenCount="53" specimenCount-female="29" specimenCount-male="24" stateProvince="Morogoro Region">
<specimenCount id="C2BEFD01FFB2FFDEFE59EAF6FE2913EC" box="[408,494,1974,2000]" pageId="4" pageNumber="5" type="male">24 ♂♂</specimenCount>
, 
<specimenCount id="C2BEFD01FFB2FFDEFE3CEAF6FD9513EC" box="[509,594,1974,2000]" pageId="4" pageNumber="5" type="female">29 ♀♀</specimenCount>
; 
<collectingRegion id="167CF86AFFB2FFDEFDA5EAF6FCFA13EC" box="[612,829,1974,2000]" country="Tanzania" name="Morogoro" pageId="4" pageNumber="5">Morogoro Region</collectingRegion>
, 
<collectingCounty id="3D664E04FFB2FFDEFC8AEAF6FB7213EC" box="[843,1205,1974,2000]" pageId="4" pageNumber="5">Udzungwa Mts National Park</collectingCounty>
, 
<collectingMunicipality id="3463ACF2FFB2FFDEFB05EAF6FAD013EC" box="[1220,1303,1974,2000]" pageId="4" pageNumber="5">Kidatu</collectingMunicipality>
, Plot 2;
</materialsCitation>
</paragraph>
<paragraph id="D4073688FFB3FFDFFF7CEC4BFCCB1574" blockId="5.[189,1399,265,329]" pageId="5" pageNumber="6">
<geoCoordinate id="B18C504FFFB3FFDFFF7CEC4BFEA11519" box="[189,358,265,293]" degrees="07" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="14.9" value="-7.6874723">07°41′14.9″ S</geoCoordinate>
, 
<geoCoordinate id="B18C504FFFB3FFDFFEB7EC4BFDE5151A" box="[374,546,265,294]" degrees="36" direction="east" minutes="56" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="24.7" value="36.940197">36°56′24.7″ E</geoCoordinate>
; 
<quantity id="13409B6DFFB3FFDFFDF3EC4BFD47151A" box="[562,640,267,294]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="m" value="650.0">
<elevation id="5F95D1BBFFB3FFDFFDF3EC4BFD47151A" box="[562,640,267,294]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="m" value="650.0">650 m</elevation>
</quantity>
a.s.l.; 
<date id="A0061048FFB3FFDFFD15EC4BFCB11519" box="[724,886,267,293]" pageId="5" pageNumber="6" value="2014-10-24">
<collectingDate id="B042E9A0FFB3FFDFFD15EC4BFCB11519" box="[724,886,267,293]" pageId="5" pageNumber="6" value="2014-10-24">24 Oct. 2014</collectingDate>
</date>
; J. Malumbres-Olarte leg.; 
<collectingMethod id="0DF94E9FFFB3FFDFFB05EC4BFDF81575" pageId="5" pageNumber="6">pitfall trapping (summed catch from eight traps)</collectingMethod>
; NHMD 
<specimenCode id="841E9EF3FFB3FFDFFD6DEC6EFCCF1574" box="[684,776,302,328]" pageId="5" pageNumber="6">621639</specimenCode>
.
</paragraph>
<paragraph id="D4073688FFB3FFDFFF7CEC30FE5415B6" blockId="5.[189,403,368,394]" box="[189,403,368,394]" pageId="5" pageNumber="6">
<heading id="8F4F81E4FFB3FFDFFF7CEC30FE5415B6" bold="true" box="[189,403,368,394]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="E6CCEA9AFFB3FFDFFF7CEC30FE5415B6" bold="true" box="[189,403,368,394]" pageId="5" pageNumber="6">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFB3FFDFFF7CECDEFA831655" blockId="5.[189,1399,414,617]" pageId="5" pageNumber="6">
The specimens from Udzungwa Mts National Park very much resemble those from Mt 
<collectingRegion id="167CF86AFFB3FFDFFB6BECDEFAF21584" box="[1194,1333,414,440]" country="Tanzania" name="Kilimanjaro" pageId="5" pageNumber="6">Kilimanjaro</collectingRegion>
(
<typeStatus id="0B03882AFFB3FFDFFA82ECDFFAB11585" box="[1347,1398,415,441]" pageId="5" pageNumber="6">type</typeStatus>
locality of 
<taxonomicName id="13B84D0BFFB3FFDFFEFEEC82FDD715E0" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[319,528,450,476]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFB3FFDFFEFEEC82FDD715E0" box="[319,528,450,476]" italics="true" pageId="5" pageNumber="6">A. claudiahempae</emphasis>
</taxonomicName>
), in size (
<figureCitation id="4C832A0DFFB3FFDFFD4AEC82FD1515E0" box="[651,722,450,476]" captionStart="Fig" captionStartId="11.[187,230,1893,1919]" captionTargetBox="[193,1391,1095,1856]" captionTargetId="figure@11.[192,1402,1089,1860]" captionTargetPageId="11" captionText="Fig. 9. Aquattuor spp. Body size (number of podous rings and midbody vertical diameter) in males of Aquattuor species from the Udzungwa Mts, and of A. claudiahempae Enghoff &amp; Frederiksen, 2015 from Mt Kilimanjaro. In case of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3755007" httpUri="https://zenodo.org/record/3755007/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
) as well as in gonopod structure (
<figureCitation id="4C832A0DFFB3FFDFFBABEC82FB0515E0" box="[1130,1218,450,476]" captionStart="Fig" captionStartId="12.[189,232,1839,1865]" captionTargetBox="[241,1350,283,1781]" captionTargetId="figure@12.[217,1371,265,1806]" captionTargetPageId="12" captionText="Fig. 10. Aquattuor claudiahempae Enghoff &amp; Frederiksen, 2015, male, specimen from the Udzungwa Mts (NHMD 621639), left gonopod. A–C. Coxa. A. Anterior view. B. Mesal view. C. Posterior view. D–G. Telopodite. D. Tip of telomere. E. Anterior view. F. Posterior view. G. Mesal view. Abbreviations: btl = basal telomeral lamella; li = lateral incision; mbl = meso-basal lobe of palette; mi = mesal incision; mpl = mesal-posterior telomeral lamella; pa = palette. Scale bars: A–C, E–G = 0.1 mm; D = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3755009" httpUri="https://zenodo.org/record/3755009/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
, cf. 
<bibRefCitation id="B0294B79FFB3FFDFFB39EC82FE4C163C" author="Enghoff H. &amp; Frederiksen S. B." pageId="5" pageNumber="6" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">
Enghoff &amp; Frederiksen 
<date id="A0061048FFB3FFDFFE8CECA5FE4C163C" box="[333,395,485,512]" pageId="5" pageNumber="6" value="2015">2015</date>
</bibRefCitation>
: fig. 10). In particular, the two samples agree in having the distal part of the telomere compressed, i.e., instead of forming a gutter, the lamella constituting the telomere is folded flat such that the inside of the gutter becomes invisible; the outside is strongly microspinose on one side and the free margin of the other side is characteristically wavy-spinose (
<figureCitation id="4C832A0DFFB3FFDFFCCFEF0EFCBF1655" box="[782,888,590,617]" captionStart="Fig" captionStartId="12.[189,232,1839,1865]" captionTargetBox="[241,1350,283,1781]" captionTargetId="figure@12.[217,1371,265,1806]" captionTargetPageId="12" captionText="Fig. 10. Aquattuor claudiahempae Enghoff &amp; Frederiksen, 2015, male, specimen from the Udzungwa Mts (NHMD 621639), left gonopod. A–C. Coxa. A. Anterior view. B. Mesal view. C. Posterior view. D–G. Telopodite. D. Tip of telomere. E. Anterior view. F. Posterior view. G. Mesal view. Abbreviations: btl = basal telomeral lamella; li = lateral incision; mbl = meso-basal lobe of palette; mi = mesal incision; mpl = mesal-posterior telomeral lamella; pa = palette. Scale bars: A–C, E–G = 0.1 mm; D = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3755009" httpUri="https://zenodo.org/record/3755009/files/figure.png" pageId="5" pageNumber="6">Fig. 10D</figureCitation>
, 
<bibRefCitation id="B0294B79FFB3FFDFFC42EF0EFB151655" author="Enghoff H. &amp; Frederiksen S. B." box="[899,1234,590,617]" pageId="5" pageNumber="6" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">
Enghoff &amp; Frederiksen 
<date id="A0061048FFB3FFDFFB55EF0EFB151655" box="[1172,1234,590,617]" pageId="5" pageNumber="6" value="2015">2015</date>
</bibRefCitation>
: fig. 10d).
</paragraph>
<caption id="80C76600FFB3FFDFFF7CEA79FDDB13DC" ID-DOI="http://doi.org/10.5281/zenodo.3754995" ID-Zenodo-Dep="3754995" httpUri="https://zenodo.org/record/3754995/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1849,1875]" targetBox="[205,1381,684,1788]" targetPageId="5">
<paragraph id="D4073688FFB3FFDFFF7CEA79FDDB13DC" blockId="5.[189,1399,1849,2016]" pageId="5" pageNumber="6">
<emphasis id="E6CCEA9AFFB3FFDFFF7CEA79FECB136F" bold="true" box="[189,268,1849,1875]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="13B84D0BFFB3FFDFFED7EA7AFE4A1368" authorityName="Frederiksen" authorityYear="2013" box="[278,397,1850,1876]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="E6CCEA9AFFB3FFDFFED7EA7AFE4A1368" box="[278,397,1850,1876]" italics="true" pageId="5" pageNumber="6">Aquattuor</emphasis>
</taxonomicName>
spp., first pair of male legs. 
<emphasis id="E6CCEA9AFFB3FFDFFD29EA79FCE6136F" bold="true" box="[744,801,1849,1876]" pageId="5" pageNumber="6">A–C</emphasis>
. 
<taxonomicName id="13B84D0BFFB3FFDFFCF3EA7AFBBB1368" authority="(Attems, 1896)" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[818,1148,1849,1876]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFB3FFDFFCF3EA7AFC7B136F" box="[818,956,1849,1876]" italics="true" pageId="5" pageNumber="6">A. fasciatus</emphasis>
(
<bibRefCitation id="B0294B79FFB3FFDFFC0FEA79FBB41368" author="Attems C. G." box="[974,1139,1849,1876]" pageId="5" pageNumber="6" pagination="23 - 42" refId="ref11215" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13, Suppl.: 23 - 42. Available from: https: // biodiversitylibrary. org / page / 28745426 [accessed 1 Oct. 2019]." type="book chapter" year="1896">Attems, 1896</bibRefCitation>
)
</taxonomicName>
, non-type male from Zanzibar (NHMD 621641). 
<emphasis id="E6CCEA9AFFB3FFDFFDDBEA1CFD97134A" bold="true" box="[538,592,1884,1911]" pageId="5" pageNumber="6">D–F</emphasis>
. 
<taxonomicName id="13B84D0BFFB3FFDFFDA5EA1DFB7D134B" authority="Enghoff &amp; Frederiksen, 2015" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[612,1210,1884,1911]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFB3FFDFFDA5EA1DFCF9134A" box="[612,830,1884,1911]" italics="true" pageId="5" pageNumber="6">A. claudiahempae</emphasis>
<bibRefCitation id="B0294B79FFB3FFDFFC8AEA1CFB7D134B" author="Enghoff H. &amp; Frederiksen S. B." box="[843,1210,1884,1911]" pageId="5" pageNumber="6" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff &amp; Frederiksen, 2015</bibRefCitation>
</taxonomicName>
, paratype from Kilimanjaro (NHMD 621638). 
<emphasis id="E6CCEA9AFFB3FFDFFDE4EA3FFD9213A6" bold="true" box="[549,597,1919,1946]" pageId="5" pageNumber="6">G–I</emphasis>
. 
<taxonomicName id="13B84D0BFFB3FFDFFDA3EAC0FCF513A5" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[610,818,1919,1946]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFB3FFDFFDA3EAC0FCF513A5" box="[610,818,1919,1946]" italics="true" pageId="5" pageNumber="6">A. claudiahempae</emphasis>
</taxonomicName>
, specimen from Udzungwa Mts (NHMD 621639). 
<emphasis id="E6CCEA9AFFB3FFDFFF7CEAE2FEDB1380" bold="true" box="[189,284,1954,1981]" pageId="5" pageNumber="6">A, D, G</emphasis>
. Anterior view. 
<emphasis id="E6CCEA9AFFB3FFDFFE1BEAE3FDF11381" bold="true" box="[474,566,1955,1981]" pageId="5" pageNumber="6">B, E, H</emphasis>
. Sublateral view. 
<emphasis id="E6CCEA9AFFB3FFDFFCC8EAE2FC901381" bold="true" box="[777,855,1954,1981]" pageId="5" pageNumber="6">C, F, I</emphasis>
. Ventral view. Abbreviation: 
<emphasis id="E6CCEA9AFFB3FFDFFB70EAE3FB111381" box="[1201,1238,1955,1981]" italics="true" pageId="5" pageNumber="6">pfp</emphasis>
= prefemoral process. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="80C76600FFB0FFDCFF7CEA22FCC913DB" ID-DOI="http://doi.org/10.5281/zenodo.3754997" ID-Zenodo-Dep="3754997" httpUri="https://zenodo.org/record/3754997/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1890,1916]" targetBox="[226,1373,279,1795]" targetPageId="6">
<paragraph id="D4073688FFB0FFDCFF7CEA22FCC913DB" blockId="6.[189,1399,1890,2023]" pageId="6" pageNumber="7">
<emphasis id="E6CCEA9AFFB0FFDCFF7CEA22FE8E1340" bold="true" box="[189,329,1890,1917]" pageId="6" pageNumber="7">Fig. 4. A–C</emphasis>
. 
<taxonomicName id="13B84D0BFFB0FFDCFE96EA23FD911341" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[343,598,1891,1917]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nguruensis" status="sp. nov.">
<emphasis id="E6CCEA9AFFB0FFDCFE96EA23FD911341" box="[343,598,1891,1917]" italics="true" pageId="6" pageNumber="7">Aquattuor nguruensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDFF57E1FFB0FFDCFD9DEA23FD7E1341" box="[604,697,1891,1917]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (VMNH110617), first pair of legs. Notice mite larva (”hypopus”) on left femur. 
<emphasis id="E6CCEA9AFFB0FFDCFDFBEAC6FDA8139C" bold="true" box="[570,623,1926,1952]" pageId="6" pageNumber="7">D–F</emphasis>
. 
<taxonomicName id="13B84D0BFFB0FFDCFDBCEAC6FBCF139C" authority="Frederiksen, 2013" authorityName="Frederiksen" authorityYear="2013" box="[637,1032,1926,1952]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="E6CCEA9AFFB0FFDCFDBCEAC6FCEB139C" box="[637,812,1926,1952]" italics="true" pageId="6" pageNumber="7">A. denticulatus</emphasis>
<bibRefCitation id="B0294B79FFB0FFDCFCF2EAC6FBCF139C" author="Frederiksen S. B." box="[819,1032,1926,1952]" pageId="6" pageNumber="7" pagination="59 - 66" refId="ref12038" refString="Frederiksen S. B. 2013. East African odontopygid millipedes 3: Two new genera, Lamelloramus and Aquattuor proposed to contain three new species (Diplopoda, Spirostreptida, Odontopygidae). Zootaxa 3694: 59 - 66. https: // doi. org / 10.11646 / zootaxa. 3694.1.4" type="journal article" year="2013">Frederiksen, 2013</bibRefCitation>
</taxonomicName>
, paratype, ♂ (NHMD 621640), left leg of first pair. 
<emphasis id="E6CCEA9AFFB0FFDCFE6DEAE9FE2113FF" bold="true" box="[428,486,1961,1987]" pageId="6" pageNumber="7">A, D</emphasis>
. Anterior view. 
<emphasis id="E6CCEA9AFFB0FFDCFD69EAE9FD2713FF" bold="true" box="[680,736,1961,1987]" pageId="6" pageNumber="7">B, E</emphasis>
. Sublateral view. 
<emphasis id="E6CCEA9AFFB0FFDCFC77EAE9FC2A13FF" bold="true" box="[950,1005,1961,1987]" pageId="6" pageNumber="7">C, F</emphasis>
. Sub-ventral view. Abbreviation: 
<emphasis id="E6CCEA9AFFB0FFDCFF7CEA8CFF2513DA" box="[189,226,1996,2022]" italics="true" pageId="6" pageNumber="7">pfd</emphasis>
= prefemoral depression. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="80C76600FFB1FFDDFF7CEA1FFB3313DF" ID-DOI="http://doi.org/10.5281/zenodo.3754999" ID-Zenodo-Dep="3754999" httpUri="https://zenodo.org/record/3754999/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1887,1913]" targetBox="[234,1343,270,1842]" targetPageId="7">
<paragraph id="D4073688FFB1FFDDFF7CEA1FFB3313DF" blockId="7.[189,1399,1887,2019]" pageId="7" pageNumber="8">
<emphasis id="E6CCEA9AFFB1FFDDFF7CEA1FFEC01345" bold="true" box="[189,263,1887,1913]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
<taxonomicName id="13B84D0BFFB1FFDDFECDEA1FFE441345" authorityName="Frederiksen" authorityYear="2013" box="[268,387,1887,1913]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="E6CCEA9AFFB1FFDDFECDEA1FFE441345" box="[268,387,1887,1913]" italics="true" pageId="7" pageNumber="8">Aquattuor</emphasis>
</taxonomicName>
spp., first pair of male legs. 
<emphasis id="E6CCEA9AFFB1FFDDFD7DEA1FFD321345" bold="true" box="[700,757,1887,1913]" pageId="7" pageNumber="8">A–C</emphasis>
. 
<taxonomicName id="13B84D0BFFB1FFDDFCC0EA1FFBD51346" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[769,1042,1887,1914]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="major">
<emphasis id="E6CCEA9AFFB1FFDDFCC0EA1FFCA31345" box="[769,868,1887,1913]" italics="true" pageId="7" pageNumber="8">A. major</emphasis>
<bibRefCitation id="B0294B79FFB1FFDDFCA8EA1FFBD51346" author="Enghoff H. &amp; Frederiksen S. B." box="[873,1042,1887,1914]" pageId="7" pageNumber="8" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
, paratype, ♂ (NHMD 621644), first pair of legs 
<emphasis id="E6CCEA9AFFB1FFDDFE46EAC3FE1513A0" box="[391,466,1923,1948]" italics="true" pageId="7" pageNumber="8">in situ</emphasis>
. 
<emphasis id="E6CCEA9AFFB1FFDDFE22EAC2FDDE13A0" bold="true" box="[483,537,1922,1949]" pageId="7" pageNumber="8">D–F</emphasis>
. 
<taxonomicName id="13B84D0BFFB1FFDDFDEBEAC3FCB013A1" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[554,887,1922,1949]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="longipala">
<emphasis id="E6CCEA9AFFB1FFDDFDEBEAC3FD7A13A0" box="[554,701,1922,1949]" italics="true" pageId="7" pageNumber="8">A. longipala</emphasis>
<bibRefCitation id="B0294B79FFB1FFDDFD09EAC2FCB013A1" author="Enghoff H. &amp; Frederiksen S. B." box="[712,887,1922,1949]" pageId="7" pageNumber="8" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
, male from Udzungwa Mts National Park, Mito Mitatu (NHMD 621642). 
<emphasis id="E6CCEA9AFFB1FFDDFDFCEAE5FDBE1383" bold="true" box="[573,633,1957,1983]" pageId="7" pageNumber="8">A, D</emphasis>
. Anterior view. 
<emphasis id="E6CCEA9AFFB1FFDDFC83EAE5FCBC1383" bold="true" box="[834,891,1957,1983]" pageId="7" pageNumber="8">B, E</emphasis>
. Sublateral view. 
<emphasis id="E6CCEA9AFFB1FFDDFB98EAE5FB551383" bold="true" box="[1113,1170,1957,1983]" pageId="7" pageNumber="8">C, F</emphasis>
. Sub-ventral view. Abbreviations: 
<emphasis id="E6CCEA9AFFB1FFDDFEB2EA88FE5F13DE" box="[371,408,1992,2018]" italics="true" pageId="7" pageNumber="8">dpl</emphasis>
= distal prefemoral lobe; 
<emphasis id="E6CCEA9AFFB1FFDDFD09EA89FD2A13DF" box="[712,749,1993,2019]" italics="true" pageId="7" pageNumber="8">pfp</emphasis>
= prefemoral process. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="80C76600FFBEFFD2FF7CEA1FFDDB13DF" ID-DOI="http://doi.org/10.5281/zenodo.3755001" ID-Zenodo-Dep="3755001" httpUri="https://zenodo.org/record/3755001/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1887,1913]" targetBox="[200,1384,275,1839]" targetPageId="8">
<paragraph id="D4073688FFBEFFD2FF7CEA1FFDDB13DF" blockId="8.[189,1399,1887,2019]" pageId="8" pageNumber="9">
<emphasis id="E6CCEA9AFFBEFFD2FF7CEA1FFECE1345" bold="true" box="[189,265,1887,1913]" pageId="8" pageNumber="9">Fig. 6.</emphasis>
<taxonomicName id="13B84D0BFFBEFFD2FECEEA1FFE411345" authorityName="Frederiksen" authorityYear="2013" box="[271,390,1887,1913]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="E6CCEA9AFFBEFFD2FECEEA1FFE411345" box="[271,390,1887,1913]" italics="true" pageId="8" pageNumber="9">Aquattuor</emphasis>
</taxonomicName>
spp., first pair of male legs. 
<emphasis id="E6CCEA9AFFBEFFD2FD08EA1FFCC51345" bold="true" box="[713,770,1887,1913]" pageId="8" pageNumber="9">A–C</emphasis>
. 
<taxonomicName id="13B84D0BFFBEFFD2FCCEEA1FFC6C1345" authority="Enghoff, 2020" authorityName="Enghoff" authorityYear="2020" box="[783,939,1887,1913]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="mollilobus" status="sp. nov.">
<emphasis id="E6CCEA9AFFBEFFD2FCCEEA1FFC6C1345" box="[783,939,1887,1913]" italics="true" pageId="8" pageNumber="9">A. mollilobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDFF57E1FFBEFFD2FC71EA1FFBCB1345" box="[944,1036,1887,1913]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (NHMD 621647). 
<emphasis id="E6CCEA9AFFBEFFD2FF7CEAC2FF3413A0" bold="true" box="[189,243,1922,1949]" pageId="8" pageNumber="9">D–F</emphasis>
. 
<taxonomicName id="13B84D0BFFBEFFD2FEC2EAC3FDA513A1" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[259,610,1922,1949]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="stereosathe">
<emphasis id="E6CCEA9AFFBEFFD2FEC2EAC3FE6C13A0" box="[259,427,1922,1949]" italics="true" pageId="8" pageNumber="9">A. stereosathe</emphasis>
<bibRefCitation id="B0294B79FFBEFFD2FE75EAC2FDA513A1" author="Enghoff H. &amp; Frederiksen S. B." box="[436,610,1922,1949]" pageId="8" pageNumber="9" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
, paratype, ♂ (NHMD 621650). 
<emphasis id="E6CCEA9AFFBEFFD2FC25EAC2FBD813A0" bold="true" box="[996,1055,1922,1948]" pageId="8" pageNumber="9">A, D</emphasis>
. Anterior view. 
<emphasis id="E6CCEA9AFFBEFFD2FB22EAC2FADC13A0" bold="true" box="[1251,1307,1922,1948]" pageId="8" pageNumber="9">B, E</emphasis>
. (Sub)- lateral view. 
<emphasis id="E6CCEA9AFFBEFFD2FE98EAE5FE561383" bold="true" box="[345,401,1957,1983]" pageId="8" pageNumber="9">C, F</emphasis>
. (Sub)-ventral view. Abbreviations: 
<emphasis id="E6CCEA9AFFBEFFD2FC89EAE5FCAA1383" box="[840,877,1957,1983]" italics="true" pageId="8" pageNumber="9">dpl</emphasis>
= distal prefemoral lobe; 
<emphasis id="E6CCEA9AFFBEFFD2FB6DEAE5FB161383" box="[1196,1233,1957,1983]" italics="true" pageId="8" pageNumber="9">pfp</emphasis>
= prefemoral process. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="D4073688FFBFFFD3FF7CEC4BFB941550" blockId="9.[189,1399,267,364]" pageId="9" pageNumber="10">
First pair of male legs (
<figureCitation id="4C832A0DFFBFFFD3FE1FEC4BFDE21519" box="[478,549,267,293]" captionStart="Fig" captionStartId="5.[189,232,1849,1875]" captionTargetBox="[205,1381,684,1788]" captionTargetId="figure@5.[189,1399,675,1801]" captionTargetPageId="5" captionText="Fig. 3. Aquattuor spp., first pair of male legs. A–C. A. fasciatus (Attems, 1896), non-type male from Zanzibar (NHMD 621641). D–F. A. claudiahempae Enghoff &amp; Frederiksen, 2015, paratype from Kilimanjaro (NHMD 621638). G–I. A. claudiahempae, specimen from Udzungwa Mts (NHMD 621639). A, D, G. Anterior view. B, E, H. Sublateral view. C, F, I. Ventral view. Abbreviation: pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754995" httpUri="https://zenodo.org/record/3754995/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
D–I) each with a single, finger-shaped prefemoral process; prefemoral processes more slender than those of the otherwise very similar 
<taxonomicName id="13B84D0BFFBFFFD3FC16EC6FFBA31575" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[983,1124,303,329]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFBFFFD3FC16EC6FFBA31575" box="[983,1124,303,329]" italics="true" pageId="9" pageNumber="10">A. fasciatus</emphasis>
</taxonomicName>
(
<figureCitation id="4C832A0DFFBFFFD3FBB8EC6EFB061575" box="[1145,1217,302,329]" captionStart="Fig" captionStartId="5.[189,232,1849,1875]" captionTargetBox="[205,1381,684,1788]" captionTargetId="figure@5.[189,1399,675,1801]" captionTargetPageId="5" captionText="Fig. 3. Aquattuor spp., first pair of male legs. A–C. A. fasciatus (Attems, 1896), non-type male from Zanzibar (NHMD 621641). D–F. A. claudiahempae Enghoff &amp; Frederiksen, 2015, paratype from Kilimanjaro (NHMD 621638). G–I. A. claudiahempae, specimen from Udzungwa Mts (NHMD 621639). A, D, G. Anterior view. B, E, H. Sublateral view. C, F, I. Ventral view. Abbreviation: pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754995" httpUri="https://zenodo.org/record/3754995/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
A–C); setae of telopodite distal to prefemur similar in length to those of normal walking legs.
</paragraph>
</subSubSection>
<subSubSection id="9CA26503FFBFFFD3FF7CECD3FAC915C3" pageId="9" pageNumber="10" type="distribution">
<paragraph id="D4073688FFBFFFD3FF7CECD3FE911591" blockId="9.[189,342,403,429]" box="[189,342,403,429]" pageId="9" pageNumber="10">
<heading id="8F4F81E4FFBFFFD3FF7CECD3FE911591" bold="true" box="[189,342,403,429]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="E6CCEA9AFFBFFFD3FF7CECD3FE911591" bold="true" box="[189,342,403,429]" pageId="9" pageNumber="10">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFBFFFD3FF7CEC82FAC915C3" blockId="9.[189,1399,449,512]" pageId="9" pageNumber="10">
Described from Mt 
<collectingRegion id="167CF86AFFBFFFD3FE5FEC82FDED15E0" box="[414,554,450,476]" country="Tanzania" name="Kilimanjaro" pageId="9" pageNumber="10">Kilimanjaro</collectingRegion>
(
<quantity id="13409B6DFFBFFFD3FDF9EC81FD1F15E0" box="[568,728,449,476]" metricMagnitude="3" metricUnit="m" metricValue="1.257" metricValueMax="1.345" metricValueMin="1.169" pageId="9" pageNumber="10" unit="m" value="1257.0" valueMax="1345.0" valueMin="1169.0">1169–1345 m</quantity>
a.s.l., in habitats disturbed by human activities; 
<bibRefCitation id="B0294B79FFBFFFD3FB3AEC82FE4B163C" author="Enghoff H. &amp; Frederiksen S. B." pageId="9" pageNumber="10" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">
Enghoff &amp; Frederiksen 
<date id="A0061048FFBFFFD3FE8CECA5FE4B163C" box="[333,396,485,512]" pageId="9" pageNumber="10" value="2015">2015</date>
</bibRefCitation>
); now also known from one site at 
<quantity id="13409B6DFFBFFFD3FCE0ECA5FCAC15C3" box="[801,875,485,512]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="10" unit="m" value="650.0">650 m</quantity>
a.s.l. in the Udzungwa Mts (
<figureCitation id="4C832A0DFFBFFFD3FB78ECA5FB3A15C3" box="[1209,1277,485,511]" captionStart="Fig" captionStartId="2.[189,232,1948,1974]" captionTargetBox="[192,1385,1023,1872]" captionTargetId="figure@2.[186,1391,1009,1876]" captionTargetPageId="2" captionText="Fig. 1. Collecting sites for Aquattuor spp. in the Udzungwa Mts. Based on Marshall et al. (2010: fig. 1); inset by permission of the Eastern Arc Mountains Conservation Endowment Fund." figureDoi="http://doi.org/10.5281/zenodo.3754991" httpUri="https://zenodo.org/record/3754991/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="80C76600FFBFFFD3FF7CEA15FE3C13E6" ID-DOI="http://doi.org/10.5281/zenodo.3755003" ID-Zenodo-Dep="3755003" httpUri="https://zenodo.org/record/3755003/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1877,1903]" targetBox="[206,1353,584,1811]" targetPageId="9">
<paragraph id="D4073688FFBFFFD3FF7CEA15FE3C13E6" blockId="9.[189,1399,1877,2010]" pageId="9" pageNumber="10">
<emphasis id="E6CCEA9AFFBFFFD3FF7CEA15FED5134C" bold="true" box="[189,274,1877,1904]" pageId="9" pageNumber="10">Fig. 7.</emphasis>
<taxonomicName id="13B84D0BFFBFFFD3FEE0EA16FE5F134C" authorityName="Frederiksen" authorityYear="2013" box="[289,408,1878,1904]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="E6CCEA9AFFBFFFD3FEE0EA16FE5F134C" box="[289,408,1878,1904]" italics="true" pageId="9" pageNumber="10">Aquattuor</emphasis>
</taxonomicName>
spp., first pair of male legs. 
<emphasis id="E6CCEA9AFFBFFFD3FCDAEA15FC941353" bold="true" box="[795,851,1877,1904]" pageId="9" pageNumber="10">A–C</emphasis>
. 
<taxonomicName id="13B84D0BFFBFFFD3FCABEA16FB15134C" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" box="[874,1234,1877,1904]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="submajor">
<emphasis id="E6CCEA9AFFBFFFD3FCABEA16FBC61353" box="[874,1025,1877,1904]" italics="true" pageId="9" pageNumber="10">A. submajor</emphasis>
<bibRefCitation id="B0294B79FFBFFFD3FBD1EA15FB15134C" author="Enghoff H. &amp; Frederiksen S. B." box="[1040,1234,1877,1904]" pageId="9" pageNumber="10" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
, paratype, ♂ (NHMD 621651). 
<emphasis id="E6CCEA9AFFBFFFD3FE54EA38FE6D13AE" bold="true" box="[405,426,1912,1938]" pageId="9" pageNumber="10">A</emphasis>
. Anterior view. 
<emphasis id="E6CCEA9AFFBFFFD3FDA3EA39FDB113AF" bold="true" box="[610,630,1913,1939]" pageId="9" pageNumber="10">B</emphasis>
. Sublateral view. 
<emphasis id="E6CCEA9AFFBFFFD3FC85EA38FC9E13AE" bold="true" box="[836,857,1912,1938]" pageId="9" pageNumber="10">C</emphasis>
. Ventral view. 
<emphasis id="E6CCEA9AFFBFFFD3FBC5EA39FBFC13AF" bold="true" box="[1028,1083,1913,1939]" pageId="9" pageNumber="10">D–E</emphasis>
. 
<taxonomicName id="13B84D0BFFBFFFD3FB89EA39FF3F138B" authority="Enghoff, 2015" authorityName="Enghoff, in Enghoff &amp; Frederiksen" authorityYear="2015" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="udzungwensis">
<emphasis id="E6CCEA9AFFBFFFD3FB89EA39FACD13AF" box="[1096,1290,1913,1939]" italics="true" pageId="9" pageNumber="10">A. udzungwensis</emphasis>
<bibRefCitation id="B0294B79FFBFFFD3FAD1EA39FF3F138B" author="Enghoff H. &amp; Frederiksen S. B." pageId="9" pageNumber="10" pagination="1 - 25" refId="ref11852" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. https: // doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
, male from Udzungwa Mts National Park, Kidatu (NHMD 621662). 
<emphasis id="E6CCEA9AFFBFFFD3FBEAEADCFB87138A" bold="true" box="[1067,1088,1948,1974]" pageId="9" pageNumber="10">D</emphasis>
. Anterior view. 
<emphasis id="E6CCEA9AFFBFFFD3FB3EEADCFAD4138A" bold="true" box="[1279,1299,1948,1974]" pageId="9" pageNumber="10">E</emphasis>
. Ventral view. Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="9CA26503FFBCFFD0FF7CEC4BFAB01676" pageId="10" pageNumber="11" type="discussion">
<paragraph id="D4073688FFBCFFD0FF7CEC4BFEE81519" blockId="10.[189,303,267,293]" box="[189,303,267,293]" pageId="10" pageNumber="11">
<heading id="8F4F81E4FFBCFFD0FF7CEC4BFEE81519" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="E6CCEA9AFFBCFFD0FF7CEC4BFEE81519" bold="true" box="[189,303,267,293]" pageId="10" pageNumber="11">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="D4073688FFBCFFD0FF7CEC79FAB01676" blockId="10.[189,1399,313,586]" pageId="10" pageNumber="11">
<bibRefCitation id="B0294B79FFBCFFD0FF7CEC79FE431568" author="Enghoff H." box="[189,388,313,340]" pageId="10" pageNumber="11" pagination="1 - 23" refId="ref11582" refString="Enghoff H. 2016 b. A mountain of millipedes IV: Species of Prionopetalum Attems, 1909, from the Udzungwa Mountains, Tanzania. With notes on &quot; P. &quot; fasciatum (Attems, 1896) and a revised species key (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 215: 1 - 23. https: // doi. org / 10.5852 / ejt. 2016.215" type="journal article" year="2016">
Enghoff (
<date id="A0061048FFBCFFD0FEEEEC79FEAA1568" box="[303,365,313,340]" pageId="10" pageNumber="11" value="2016">2016</date>
b)
</bibRefCitation>
discussed 
<taxonomicName id="13B84D0BFFBCFFD0FDC8EC7AFC931568" authority="(Attems, 1896)" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[521,852,313,340]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFBCFFD0FDC8EC7AFD521568" box="[521,661,314,340]" italics="true" pageId="10" pageNumber="11">A. fasciatus</emphasis>
(
<bibRefCitation id="B0294B79FFBCFFD0FD69EC7AFC8B1568" author="Attems C. G." box="[680,844,313,340]" pageId="10" pageNumber="11" pagination="23 - 42" refId="ref11215" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13, Suppl.: 23 - 42. Available from: https: // biodiversitylibrary. org / page / 28745426 [accessed 1 Oct. 2019]." type="book chapter" year="1896">Attems, 1896</bibRefCitation>
)
</taxonomicName>
(see below), emphasizing its great similarity with 
<taxonomicName id="13B84D0BFFBCFFD0FF39EC1DFE0C154B" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[248,459,349,375]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBCFFD0FF39EC1DFE0C154B" box="[248,459,349,375]" italics="true" pageId="10" pageNumber="11">A. claudiahempae</emphasis>
</taxonomicName>
and suggesting that the latter might be a synonym of 
<taxonomicName id="13B84D0BFFBCFFD0FBFDEC1DFB03154B" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[1084,1220,349,375]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFBCFFD0FBFDEC1DFB03154B" box="[1084,1220,349,375]" italics="true" pageId="10" pageNumber="11">A. fasciatus</emphasis>
</taxonomicName>
. The similarity between both nominal species also extends to the compressed distal part of the telomere (
<bibRefCitation id="B0294B79FFBCFFD0FAD4ECC0FECC1581" author="Enghoff H." pageId="10" pageNumber="11" pagination="1 - 23" refId="ref11582" refString="Enghoff H. 2016 b. A mountain of millipedes IV: Species of Prionopetalum Attems, 1909, from the Udzungwa Mountains, Tanzania. With notes on &quot; P. &quot; fasciatum (Attems, 1896) and a revised species key (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 215: 1 - 23. https: // doi. org / 10.5852 / ejt. 2016.215" type="journal article" year="2016">
Enghoff 
<date id="A0061048FFBCFFD0FF7CECE3FF3B1581" box="[189,252,419,445]" pageId="10" pageNumber="11" value="2016">2016</date>
b
</bibRefCitation>
: fig. 7c, f). The only differences between the two seem to be size (cf. the identification key below, couplet 9) and the shape of the prefemoral process of the first pair of legs (
<figureCitation id="4C832A0DFFBCFFD0FB6BEC86FB3315DC" box="[1194,1268,454,480]" captionStart="Fig" captionStartId="5.[189,232,1849,1875]" captionTargetBox="[205,1381,684,1788]" captionTargetId="figure@5.[189,1399,675,1801]" captionTargetPageId="5" captionText="Fig. 3. Aquattuor spp., first pair of male legs. A–C. A. fasciatus (Attems, 1896), non-type male from Zanzibar (NHMD 621641). D–F. A. claudiahempae Enghoff &amp; Frederiksen, 2015, paratype from Kilimanjaro (NHMD 621638). G–I. A. claudiahempae, specimen from Udzungwa Mts (NHMD 621639). A, D, G. Anterior view. B, E, H. Sublateral view. C, F, I. Ventral view. Abbreviation: pfp = prefemoral process. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3754995" httpUri="https://zenodo.org/record/3754995/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
). The new specimens of 
<taxonomicName id="13B84D0BFFBCFFD0FE9FECAAFDF5163F" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[350,562,489,516]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBCFFD0FE9FECAAFDF5163F" box="[350,562,489,516]" italics="true" pageId="10" pageNumber="11">A. claudiahempae</emphasis>
</taxonomicName>
from Kidatu, Udzungwa Mts are of the same size as 
<taxonomicName id="13B84D0BFFBCFFD0FB62ECAAFAB1163F" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[1187,1398,489,516]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBCFFD0FB62ECAAFAB1163F" box="[1187,1398,489,516]" italics="true" pageId="10" pageNumber="11">A. claudiahempae</emphasis>
</taxonomicName>
from Mt 
<collectingRegion id="167CF86AFFBCFFD0FEEAEF4CFE7E161A" box="[299,441,524,550]" country="Tanzania" name="Kilimanjaro" pageId="10" pageNumber="11">Kilimanjaro</collectingRegion>
(
<figureCitation id="4C832A0DFFBCFFD0FE0AEF4CFDD2161B" box="[459,533,524,551]" captionStart="Fig" captionStartId="11.[187,230,1893,1919]" captionTargetBox="[193,1391,1095,1856]" captionTargetId="figure@11.[192,1402,1089,1860]" captionTargetPageId="11" captionText="Fig. 9. Aquattuor spp. Body size (number of podous rings and midbody vertical diameter) in males of Aquattuor species from the Udzungwa Mts, and of A. claudiahempae Enghoff &amp; Frederiksen, 2015 from Mt Kilimanjaro. In case of (almost) coinciding values, symbols have been slightly displaced horizontally." figureDoi="http://doi.org/10.5281/zenodo.3755007" httpUri="https://zenodo.org/record/3755007/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
), and their prefemoral processes strongly resemble those of 
<collectingRegion id="167CF86AFFBCFFD0FB28EF4CFAB0161A" box="[1257,1399,524,550]" country="Tanzania" name="Kilimanjaro" pageId="10" pageNumber="11">Kilimanjaro</collectingRegion>
specimens. 
<taxonomicName id="13B84D0BFFBCFFD0FE85EF70FDB31676" authorityName="Enghoff &amp; Frederiksen" authorityYear="2015" box="[324,628,560,586]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="claudiahempae">
<emphasis id="E6CCEA9AFFBCFFD0FE85EF70FDB31676" box="[324,628,560,586]" italics="true" pageId="10" pageNumber="11">Aquattuor claudiahempae</emphasis>
</taxonomicName>
and 
<taxonomicName id="13B84D0BFFBCFFD0FD6BEF70FCF61676" baseAuthorityName="Attems" baseAuthorityYear="1896" box="[682,817,560,586]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="E6CCEA9AFFBCFFD0FD6BEF70FCF61676" box="[682,817,560,586]" italics="true" pageId="10" pageNumber="11">A. fasciatus</emphasis>
</taxonomicName>
are, for the time being, upheld as separate species.
</paragraph>
</subSubSection>
</treatment>
</document>