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<document id="FDCBFF3C16F21245B76AC50506124E0D" ID-CLB-Dataset="3390" ID-DOI="10.11646/zootaxa.5068.4.6" ID-GBIF-Dataset="79b5ca84-737c-4722-9994-9f5cdefbebb3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5709642" ID-ZooBank="E4A3743F-14A9-4292-AC55-56B7E7F0905C" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637219973168" checkinUser="plazi" docAuthor="Golovatch, Sergei I." docDate="2021" docId="5038879DCE2A550390A12981FC67FE1E" docLanguage="en" docName="zootaxa.5068.4.6.pdf" docOrigin="Zootaxa 5068 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pyrgodesmus Pocock 1892" docType="treatment" docVersion="6" lastPageNumber="577" masterDocId="AC01FFE5CE2B550690362C03FFB5FFEB" masterDocTitle="Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida)" masterLastPageNumber="578" masterPageNumber="572" pageNumber="573" updateTime="1699468885165" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0734D245A323B3D749861E7FB62078A0">Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="4537D810D5F808DC5C0D42F4AA869610">Golovatch, Sergei I.</mods:namePart>
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<mods:date id="36656C51F444DEF4154374FC22F09B3C">2021</mods:date>
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<treatment id="5038879DCE2A550390A12981FC67FE1E" ID-DOI="http://doi.org/10.5281/zenodo.5714972" ID-GBIF-Taxon="190310263" ID-Zenodo-Dep="5714972" LSID="urn:lsid:plazi:treatment:5038879DCE2A550390A12981FC67FE1E" httpUri="http://treatment.plazi.org/id/5038879DCE2A550390A12981FC67FE1E" lastPageId="5" lastPageNumber="577" pageId="1" pageNumber="573">
<subSubSection id="908B6500CE2A550790A12981FD8FFA77" box="[151,570,1410,1437]" pageId="1" pageNumber="573" type="nomenclature">
<paragraph id="D82E368BCE2A550790A12981FD8FFA77" blockId="1.[151,570,1410,1437]" box="[151,570,1410,1437]" pageId="1" pageNumber="573">
<heading id="836681E7CE2A550790A12981FD8FFA77" bold="true" box="[151,570,1410,1437]" fontSize="11" level="1" pageId="1" pageNumber="573" reason="1">
<emphasis id="EAE5EA99CE2A550790A12981FD8FFA77" bold="true" box="[151,570,1410,1437]" pageId="1" pageNumber="573">
Genus
<taxonomicName id="1F914D08CE2A550790D82981FD8FFA77" ID-CoL="7PX6N" authority="Pocock, 1892" authorityName="Pocock" authorityYear="1892" box="[238,570,1410,1437]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="EAE5EA99CE2A550790D82981FE3AFA76" bold="true" box="[238,399,1410,1437]" italics="true" pageId="1" pageNumber="573">Pyrgodesmus</emphasis>
<bibRefCitation id="BC004B7ACE2A550791A12980FD8FFA77" author="Pocock, R. I." box="[407,570,1410,1437]" pageId="1" pageNumber="573" pagination="131 - 174" refId="ref3370" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural History Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock, 1892</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="908B6500CE2A550790A129C4FC80FA0B" box="[151,821,1479,1504]" pageId="1" pageNumber="573" type="type_taxon">
<paragraph id="D82E368BCE2A550790A129C4FC80FA0B" blockId="1.[151,939,1479,1534]" box="[151,821,1479,1504]" pageId="1" pageNumber="573">
Type-species:
<taxonomicName id="1F914D08CE2A5507911E29C4FD2BFA34" ID-CoL="4QT8C" authority="Pocock 1892" authorityName="Pocock" authorityYear="1892" box="[296,670,1479,1503]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="EAE5EA99CE2A5507911E29C4FDA4FA34" box="[296,529,1479,1503]" italics="true" pageId="1" pageNumber="573">Pyrgodesmus obscurus</emphasis>
<bibRefCitation id="BC004B7ACE2A5507922E29C4FD2BFA34" author="Pocock, R. I." box="[536,670,1479,1503]" pageId="1" pageNumber="573" pagination="131 - 174" refId="ref3370" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural History Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock 1892</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="908B6500CE2A550790A129E5FC1EFA15" box="[151,939,1510,1534]" pageId="1" pageNumber="573" type="synonymic_list">
<paragraph id="D82E368BCE2A550790A129E5FC1EFA15" blockId="1.[151,939,1479,1534]" box="[151,939,1510,1534]" pageId="1" pageNumber="573">
Other species included:
<taxonomicName id="1F914D08CE2A550791BD29E5FC98FA15" authority="(Attems, 1936)" authorityName="Golovatch" authorityYear="2021" baseAuthorityName="Attems" baseAuthorityYear="1936" box="[395,813,1510,1534]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="permutatus" status="comb. nov.">
<emphasis id="EAE5EA99CE2A550791BD29E5FD3EFA15" box="[395,651,1510,1534]" italics="true" pageId="1" pageNumber="573">Pyrgodesmus permutatus</emphasis>
(
<bibRefCitation id="BC004B7ACE2A550792AF29E5FC93FA15" author="Attems, C." box="[665,806,1510,1534]" pageId="1" pageNumber="573" pagination="133 - 323" refId="ref2812" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11 (4), 133 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="EAE5EA99CE2A5507930E29E5FC1EFA15" bold="true" box="[824,939,1510,1534]" pageId="1" pageNumber="573">
<taxonomicNameLabel id="F1D657E2CE2A5507930E29E5FC1EFA15" box="[824,939,1510,1534]" pageId="1" pageNumber="573" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<caption id="8CEE6603CE29550490A12B42FE3BF801" ID-DOI="http://doi.org/10.5281/zenodo.5709646" ID-Zenodo-Dep="5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="2" pageNumber="574" startId="2.[151,264,1857,1881]" targetBox="[186,1401,181,1831]" targetPageId="2">
<paragraph id="D82E368BCE29550490A12B42FE3BF801" blockId="2.[151,1437,1857,2026]" pageId="2" pageNumber="574">
<emphasis id="EAE5EA99CE29550490A12B42FE89F8B2" bold="true" box="[151,316,1857,1881]" pageId="2" pageNumber="574">FIGURES 18.</emphasis>
<taxonomicName id="1F914D08CE29550491752B41FD76F8B1" authority="Pocock, 1892" authorityName="Pocock" authorityYear="1892" box="[323,707,1858,1882]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="EAE5EA99CE29550491752B41FD9BF8B1" box="[323,558,1858,1882]" italics="true" pageId="2" pageNumber="574">Pyrgodesmus obscurus</emphasis>
<bibRefCitation id="BC004B7ACE29550492002B41FD76F8B1" author="Pocock, R. I." box="[566,707,1858,1882]" pageId="2" pageNumber="574" pagination="131 - 174" refId="ref3370" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural History Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock, 1892</bibRefCitation>
</taxonomicName>
, male syntypes.
<emphasis id="EAE5EA99CE29550493592B42FCC9F8B2" bold="true" box="[879,892,1857,1881]" pageId="2" pageNumber="574">1</emphasis>
. Original labels.
<emphasis id="EAE5EA99CE29550494192B42FB89F8B2" bold="true" box="[1071,1084,1857,1881]" pageId="2" pageNumber="574">2</emphasis>
. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin.
<emphasis id="EAE5EA99CE29550492F32B66FD67F896" bold="true" box="[709,722,1893,1917]" pageId="2" pageNumber="574">3</emphasis>
. Habitus of a syntype, lateral view (after
<bibRefCitation id="BC004B7ACE29550494402B65FAA3F895" author="Pocock, R. I." box="[1142,1302,1894,1918]" pageId="2" pageNumber="574" pagination="131 - 174" refId="ref3370" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural History Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock (1892))</bibRefCitation>
.
<emphasis id="EAE5EA99CE29550495162B66FA98F896" bold="true" box="[1312,1325,1893,1917]" pageId="2" pageNumber="574">4</emphasis>
. Picture of the same, lateral view.
<emphasis id="EAE5EA99CE29550491B62B89FE38F849" bold="true" box="[384,397,1930,1954]" pageId="2" pageNumber="574">5</emphasis>
. Body rings 57 and gonopods
<emphasis id="EAE5EA99CE29550492E62B89FCBAF84A" box="[720,783,1930,1953]" italics="true" pageId="2" pageNumber="574">in situ</emphasis>
, ventral view.
<emphasis id="EAE5EA99CE29550493932B8AFC07F84A" bold="true" box="[933,946,1929,1953]" pageId="2" pageNumber="574">6</emphasis>
. Body rings 69 and gonopods
<emphasis id="EAE5EA99CE29550494C32B89FA81F84A" box="[1269,1332,1930,1953]" italics="true" pageId="2" pageNumber="574">in situ</emphasis>
, ventrolateral view.
<emphasis id="EAE5EA99CE29550491332BADFE8AF82E" bold="true" box="[261,319,1965,1990]" pageId="2" pageNumber="574">7 &amp; 8</emphasis>
. Enlarged gonopods
<emphasis id="EAE5EA99CE29550492252BADFDE4F82E" box="[531,593,1966,1989]" italics="true" pageId="2" pageNumber="574">in situ</emphasis>
, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov.
</paragraph>
</caption>
<caption id="8CEE6603CE28550590A12AB6FC63F91A" ID-DOI="http://doi.org/10.5281/zenodo.5709648" ID-Zenodo-Dep="5709648" httpUri="https://zenodo.org/record/5709648/files/figure.png" pageId="3" pageNumber="575" startId="3.[151,264,1717,1741]" targetBox="[156,1434,817,1690]" targetPageId="3">
<paragraph id="D82E368BCE28550590A12AB6FC63F91A" blockId="3.[151,1437,1717,1777]" pageId="3" pageNumber="575">
<emphasis id="EAE5EA99CE28550590A12AB6FEF2F926" bold="true" box="[151,327,1717,1741]" pageId="3" pageNumber="575">FIGURES 918.</emphasis>
<taxonomicName id="1F914D08CE285505917B2AB6FD5AF926" authority="(Attems, 1936)" authorityName="Golovatch" authorityYear="2021" baseAuthorityName="Attems" baseAuthorityYear="1936" box="[333,751,1717,1741]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="575" phylum="Arthropoda" rank="species" species="permutatus" status="comb. nov.">
<emphasis id="EAE5EA99CE285505917B2AB6FDF8F926" box="[333,589,1717,1741]" italics="true" pageId="3" pageNumber="575">Pyrgodesmus permutatus</emphasis>
(
<bibRefCitation id="BC004B7ACE285505926C2AB6FD5DF926" author="Attems, C." box="[602,744,1717,1741]" pageId="3" pageNumber="575" pagination="133 - 323" refId="ref2812" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11 (4), 133 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="EAE5EA99CE28550592CF2AB6FCD8F926" bold="true" box="[761,877,1717,1741]" pageId="3" pageNumber="575">
<taxonomicNameLabel id="F1D657E2CE28550592CF2AB6FCD8F926" box="[761,877,1717,1741]" pageId="3" pageNumber="575" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, habituses of two male syntypes, one coated with earth (1012) and the other clean (1318). All after
<bibRefCitation id="BC004B7ACE285505925D2ADAFCB9F91A" author="Silvestri, F." box="[619,780,1753,1777]" pageId="3" pageNumber="575" pagination="117 - 135" refId="ref3455" refString="Silvestri, F. (1920) Descriptions of some Oriental Diplopoda Polydesmoidea of the subfamily Pyrgodesminae. Records of the Indian Museum, 19 (4), 117 - 135." type="journal article" year="1920">Silvestri (1920)</bibRefCitation>
, drawn not to scale.
</paragraph>
</caption>
<caption id="8CEE6603CE2F550290A12B17FBB6F856" ID-DOI="http://doi.org/10.5281/zenodo.5709650" ID-Zenodo-Dep="5709650" httpUri="https://zenodo.org/record/5709650/files/figure.png" pageId="4" pageNumber="576" startId="4.[151,264,1812,1836]" targetBox="[176,1412,804,1775]" targetPageId="4">
<paragraph id="D82E368BCE2F550290A12B17FBB6F856" blockId="4.[151,1436,1812,1981]" pageId="4" pageNumber="576">
<emphasis id="EAE5EA99CE2F550290A12B17FEE3F8C6" bold="true" box="[151,342,1812,1837]" pageId="4" pageNumber="576">FIGURES 1930.</emphasis>
<taxonomicName id="1F914D08CE2F550291682B16FCB3F8C6" authority="(Attems, 1936)" authorityName="Golovatch" authorityYear="2021" baseAuthorityName="Attems" baseAuthorityYear="1936" box="[350,774,1813,1837]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="permutatus" status="comb. nov.">
<emphasis id="EAE5EA99CE2F550291682B16FDD5F8C6" box="[350,608,1813,1837]" italics="true" pageId="4" pageNumber="576">Pyrgodesmus permutatus</emphasis>
(
<bibRefCitation id="BC004B7ACE2F550292462B16FD4AF8C6" author="Attems, C." box="[624,767,1813,1837]" pageId="4" pageNumber="576" pagination="133 - 323" refId="ref2812" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11 (4), 133 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="EAE5EA99CE2F550293252B16FC3DF8C6" bold="true" box="[787,904,1813,1837]" pageId="4" pageNumber="576">
<taxonomicNameLabel id="F1D657E2CE2F550293252B16FC3DF8C6" box="[787,904,1813,1837]" pageId="4" pageNumber="576" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, male syntype.
<emphasis id="EAE5EA99CE2F550294192B16FBFDF8C6" bold="true" box="[1071,1096,1813,1837]" pageId="4" pageNumber="576">19</emphasis>
. Head, dorsal view.
<emphasis id="EAE5EA99CE2F550295162B16FA8CF8C6" bold="true" box="[1312,1337,1813,1837]" pageId="4" pageNumber="576">20</emphasis>
. Anterior part of body, ventral view.
<emphasis id="EAE5EA99CE2F550291982B3AFE72F8BA" bold="true" box="[430,455,1849,1873]" pageId="4" pageNumber="576">21</emphasis>
. Antenna, lateral view.
<emphasis id="EAE5EA99CE2F5502928B2B3AFD63F8BA" bold="true" box="[701,726,1849,1873]" pageId="4" pageNumber="576">22</emphasis>
. Gnathochilarium, ventral view.
<emphasis id="EAE5EA99CE2F5502941D2B3AFBF1F8BA" bold="true" box="[1067,1092,1849,1873]" pageId="4" pageNumber="576">23</emphasis>
. Anteromedial part of gnathochilarium, ventral view.
<emphasis id="EAE5EA99CE2F550291592B5EFE3DF89E" bold="true" box="[367,392,1885,1909]" pageId="4" pageNumber="576">24</emphasis>
. Right paraterga 11 and 12, dorsal view.
<emphasis id="EAE5EA99CE2F5502931C2B5EFCF6F89E" bold="true" box="[810,835,1885,1909]" pageId="4" pageNumber="576">25</emphasis>
. Cross-section of body ring 8, posterior view.
<emphasis id="EAE5EA99CE2F5502952D2B5EFA81F89E" bold="true" box="[1307,1332,1885,1909]" pageId="4" pageNumber="576">26</emphasis>
. Posterior part of body, ventral view.
<emphasis id="EAE5EA99CE2F5502919E2B82FE74F872" bold="true" box="[424,449,1921,1945]" pageId="4" pageNumber="576">27</emphasis>
. Fine structure of limbus, dorsal view.
<emphasis id="EAE5EA99CE2F550293662B82FCDCF872" bold="true" box="[848,873,1921,1945]" pageId="4" pageNumber="576">28</emphasis>
. Leg of body ring 8.
<emphasis id="EAE5EA99CE2F550294742B82FBEEF872" bold="true" box="[1090,1115,1921,1945]" pageId="4" pageNumber="576">29</emphasis>
. Left gonopod, mesal view.
<emphasis id="EAE5EA99CE2F5502954B2B82FA23F872" bold="true" box="[1405,1430,1921,1945]" pageId="4" pageNumber="576">30</emphasis>
. Telopodite of right gonopod, mesal view. All after
<bibRefCitation id="BC004B7ACE2F550292AE2BA6FC8CF856" author="Silvestri, F." box="[664,825,1957,1981]" pageId="4" pageNumber="576" pagination="117 - 135" refId="ref3455" refString="Silvestri, F. (1920) Descriptions of some Oriental Diplopoda Polydesmoidea of the subfamily Pyrgodesminae. Records of the Indian Museum, 19 (4), 117 - 135." type="journal article" year="1920">Silvestri (1920)</bibRefCitation>
, drawn not to scale.
</paragraph>
</caption>
<subSubSection id="908B6500CE2E550390A12C94FE6DFF59" box="[151,472,151,178]" pageId="5" pageNumber="577" type="multiple">
<paragraph id="D82E368BCE2E550390A12C94FE6DFF59" blockId="5.[151,472,151,178]" box="[151,472,151,178]" pageId="5" pageNumber="577">
<heading id="836681E7CE2E550390A12C94FE6DFF59" bold="true" box="[151,472,151,178]" fontSize="11" level="1" pageId="5" pageNumber="577" reason="1">
<emphasis id="EAE5EA99CE2E550390A12C94FE6DFF59" bold="true" box="[151,472,151,178]" pageId="5" pageNumber="577">
Diagnosis of
<taxonomicName id="1F914D08CE2E550391012C94FE6DFF59" authorityName="Pocock" authorityYear="1892" box="[311,472,151,178]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="EAE5EA99CE2E550391012C94FE6DFF59" bold="true" box="[311,472,151,178]" italics="true" pageId="5" pageNumber="577">Pyrgodesmus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="908B6500CE2E550390A12CDCFC67FE1E" pageId="5" pageNumber="577" type="diagnosis">
<paragraph id="D82E368BCE2E550390A12CDCFC67FE1E" blockId="5.[151,1436,223,502]" pageId="5" pageNumber="577">
<taxonomicName id="1F914D08CE2E550390A12CDCFE98FF12" authorityName="Pocock" authorityYear="1892" box="[151,301,223,249]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="EAE5EA99CE2E550390A12CDCFE98FF12" box="[151,301,223,249]" italics="true" pageId="5" pageNumber="577">Pyrgodesmus</emphasis>
</taxonomicName>
seems to be particularly similar to
<taxonomicName id="1F914D08CE2E5503928A2CDCFCD8FF12" authorityName="Carl" authorityYear="1932" box="[700,877,223,249]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="EAE5EA99CE2E5503928A2CDCFCD8FF12" box="[700,877,223,249]" italics="true" pageId="5" pageNumber="577">Klimakodesmus</emphasis>
</taxonomicName>
in most somatic characters, as correctly noted by
<bibRefCitation id="BC004B7ACE2E550390A12D00FEAFFEF6" author="Carl, J." box="[151,282,259,285]" pageId="5" pageNumber="577" pagination="411 - 529" refId="ref2894" refString="Carl, J. (1932) Diplopoden aus Sud-Indien und Ceylon. 1. Teil. Polydesmoidea. Revue suisse de Zoologie, 39 (17), 411 - 529. https: // doi. org / 10.5962 / bhl. part. 118948" type="journal article" year="1932">Carl (1932)</bibRefCitation>
and recently reinstated by
<bibRefCitation id="BC004B7ACE2E550392792D00FCFFFEF6" author="Aswathy, M. D. &amp; Golovatch, S. I. &amp; Sudhikumar, A. V." box="[591,842,259,285]" pageId="5" pageNumber="577" pagination="373 - 382" refId="ref2712" refString="Aswathy, M. D., Golovatch, S. I. &amp; Sudhikumar, A. V. (2021) The millipede genus Klimakodesmus Carl, 1932, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Pyrgodesmidae). Zootaxa, 4980 (2), 373 - 382. https: // doi. org / 10.11646 / zootaxa. 4980.2.8" type="journal article" year="2021">
Aswathy
<emphasis id="EAE5EA99CE2E5503928D2D07FD40FEF6" box="[699,757,259,285]" italics="true" pageId="5" pageNumber="577">et al.</emphasis>
(2021)
</bibRefCitation>
. Yet both genera are distinct primarily in gonopodal structure: hypertrophied coxites, a deep gonocoel and ribbon-shaped, unipartite, distally attenuating, but not acuminate telopodites (
<figureCitation id="40AA2A0ECE2E5503914D2D48FE51FE8D" box="[379,484,331,358]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="5" pageNumber="577">Figs 58</figureCitation>
,
<figureCitation id="40AA2A0ECE2E550391C52D48FDECFE8E" box="[499,601,331,357]" captionStart="FIGURES 1930" captionStartId="4.[151,264,1812,1836]" captionTargetBox="[176,1412,804,1775]" captionTargetId="figure-231@4.[159,1429,803,1775]" captionTargetPageId="4" captionText="FIGURES 1930. Pyrgodesmus permutatus (Attems, 1936), comb. nov., male syntype. 19. Head, dorsal view. 20. Anterior part of body, ventral view. 21. Antenna, lateral view. 22. Gnathochilarium, ventral view. 23. Anteromedial part of gnathochilarium, ventral view. 24. Right paraterga 11 and 12, dorsal view. 25. Cross-section of body ring 8, posterior view. 26. Posterior part of body, ventral view. 27. Fine structure of limbus, dorsal view. 28. Leg of body ring 8. 29. Left gonopod, mesal view. 30. Telopodite of right gonopod, mesal view. All after Silvestri (1920), drawn not to scale." figureDoi="http://doi.org/10.5281/zenodo.5709650" httpUri="https://zenodo.org/record/5709650/files/figure.png" pageId="5" pageNumber="577">29 &amp; 30</figureCitation>
),
<emphasis id="EAE5EA99CE2E550392472D4FFD25FE8E" box="[625,656,332,357]" italics="true" pageId="5" pageNumber="577">vs.</emphasis>
moderately incrassate, apparently smaller, subspherical or squarish coxites, each of which supports a strong and unipartite telopodite represented solely by a slender, strongly exposed, increasingly attenuating and acuminate solenomere. In addition,
<emphasis id="EAE5EA99CE2E550393412D90FC16FE46" bold="true" box="[887,931,403,429]" pageId="5" pageNumber="577">PM</emphasis>
crests, including the one on the penultimate ring that overhangs and conceals the epiproct from above, are much higher and stronger, whereas
<emphasis id="EAE5EA99CE2E550394C62DB4FAA2FE3A" bold="true" box="[1264,1303,439,465]" pageId="5" pageNumber="577">DL</emphasis>
reduced, in
<taxonomicName id="1F914D08CE2E550390A12DD8FE98FE1E" authorityName="Pocock" authorityYear="1892" box="[151,301,475,501]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="EAE5EA99CE2E550390A12DD8FE98FE1E" box="[151,301,475,501]" italics="true" pageId="5" pageNumber="577">Pyrgodesmus</emphasis>
</taxonomicName>
compared to
<taxonomicName id="1F914D08CE2E550391F02DD8FD65FE1E" box="[454,720,475,501]" class="Diplopoda" family="Pyrgodesmidae" genus="Klimakodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EAE5EA99CE2E550391F02DD8FDC2FE1E" box="[454,631,475,501]" italics="true" pageId="5" pageNumber="577">Klimakodesmus</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="BC004B7ACE2E550392E82DD8FC70FE1E" author="Aswathy, M. D. &amp; Golovatch, S. I. &amp; Sudhikumar, A. V." box="[734,965,475,501]" pageId="5" pageNumber="577" pagination="373 - 382" refId="ref2712" refString="Aswathy, M. D., Golovatch, S. I. &amp; Sudhikumar, A. V. (2021) The millipede genus Klimakodesmus Carl, 1932, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Pyrgodesmidae). Zootaxa, 4980 (2), 373 - 382. https: // doi. org / 10.11646 / zootaxa. 4980.2.8" type="journal article" year="2021">
Aswathy
<emphasis id="EAE5EA99CE2E5503937C2DDFFC36FE1E" box="[842,899,475,501]" italics="true" pageId="5" pageNumber="577">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>