treatments-xml/data/81/04/0A/81040A3BFF97FFEC747FF98CFC42FB29.xml

<document id="5AC4B27876CC65DBDF07A440C1FDFE77" ID-CLB-Dataset="4319" ID-DOI="10.11646/zootaxa.792.1.1" ID-GBIF-Dataset="ca95180d-948a-4bff-a657-630525a56e4f" ID-ISSN="1175­5334" ID-Zenodo-Dep="5495868" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624554437426" checkinUser="felipe" docAuthor="Jewell, Tony R. &amp; Leschen, Richard A. B." docDate="2004" docId="81040A3BFF97FFEC747FF98CFC42FB29" docLanguage="en" docName="zt00792.pdf" docOrigin="Zootaxa 792 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.792.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hoplodactylus cryptozoicus Jewell &amp; Leschen 2004, n. sp." docType="treatment" docUuid="27214474-7D98-4B68-B74B-7A4BC3C061FE" docUuidSource="ZooBank" docVersion="9" lastPageNumber="10" masterDocId="7D3D7243FF96FFE57577FFB2FFB4FFE7" masterDocTitle="A new species of Hoplodactylus (Reptilia: Pygopodidae) from the Takitimu Mountains, South Island, New Zealand" masterLastPageNumber="11" masterPageNumber="1" pageNumber="2" updateTime="1699020691533" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="82304112D56E0C89F8B1C27CF9305652">A new species of Hoplodactylus (Reptilia: Pygopodidae) from the Takitimu Mountains, South Island, New Zealand</mods:title>
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<mods:affiliation id="F02569746495D7EEDB6A0E84609B2DC6">216 Ripponvale Road, R. D., Cromwell, New Zealand New Zealand Arthropod Collection, Private Bag 92170, Auckland, New Zealand,</mods:affiliation>
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<mods:date id="3CD93CEB63658CE754E36173E4EB3212">2004</mods:date>
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<heading id="525A0C41FF97FFE4747FF98CFD29F9BF" bold="true" box="[264,669,1598,1624]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="3BD9673FFF97FFE4747FF98CFD29F9BF" bold="true" box="[264,669,1598,1624]" pageId="1" pageNumber="2">
<taxonomicName id="CEADC0AEFF97FFE4747FF98CFDE6F9BF" authority="Jewell &amp; Leschen, 2004" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[264,594,1598,1624]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="cryptozoicus" status="sp. nov.">
<emphasis id="3BD9673FFF97FFE4747FF98CFDE6F9BF" bold="true" box="[264,594,1598,1624]" italics="true" pageId="1" pageNumber="2">Hoplodactylus cryptozoicus</emphasis>
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<taxonomicNameLabel id="20EADA44FF97FFE4772CF98CFD29F9BF" box="[603,669,1598,1624]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="0912BB2DFF97FFE4747FF9D3FE09F99C" blockId="1.[264,669,1598,1659]" box="[264,445,1633,1659]" pageId="1" pageNumber="2">
<figureCitation id="9196A7A8FF97FFE4747FF9D3FE21F99C" box="[264,405,1633,1659]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[264,367,1819,1843]" captionStartId-1="4.[264,368,1749,1773]" captionTargetBox-0="[338,1289,284,1803]" captionTargetBox-1="[331,1256,1239,1739]" captionTargetId-0="figure-53@2.[498,1021,775,1181]" captionTargetId-1="figure-387@4.[330,1276,1237,1739]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. Scalation of the head and feet of H. cryptozoicus (specimen: holotype). A) head, lateral view; B) head, frontal view; C) underside of hind foot. Scale bar = 4 mm." captionText-1="FIGURE 2. An adult male Hoplodactylus cryptozoicus, collected live in January 2000 from the type locality in the Spence Burn, Takitimu Mountains. This specimen illustrates the typical primary markings (i.e., the herringbone­shaped patches connected longitudinally), and the two types of ground colour which occur in adults, grey (in this case restricted to the flanks) and two­tone orange. (Photograph by R. Morris)." figureDoi-0="http://doi.org/10.5281/zenodo.5495870" figureDoi-1="http://doi.org/10.5281/zenodo.5495872" httpUri-0="https://zenodo.org/record/5495870/files/figure.png" httpUri-1="https://zenodo.org/record/5495872/files/figure.png" pageId="1" pageNumber="2">Figures 1–2</figureCitation>
, 
<figureCitation id="9196A7A8FF97FFE474D5F9D3FE09F99C" box="[418,445,1633,1659]" captionStart="FIGURE 4" captionStartId="7.[264,368,1711,1735]" captionTargetBox="[367,1221,1165,1704]" captionTargetPageId="7" captionText="FIGURE 4. Sloughed skins of Hoplodactylus cryptozoicus (A) and sister­species H. nebulosus (B), illustrating differences in body colour pattern. The H. cryptozoicus skin is from the specimen featured in Fig. 2; the H. nebulosus skin is from a typical example of the species (collected from Big Island). Note the browner colouration of the H. cryptozoicus skin (resulting from the orange pigmentation) and an absence of the dark flecks which embellish H. nebulosus. (Photographs by T. Jewell)." figureDoi="http://doi.org/10.5281/zenodo.5495876" httpUri="https://zenodo.org/record/5495876/files/figure.png" pageId="1" pageNumber="2">4a</figureCitation>
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<emphasis id="3BD9673FFF97FFE4747FF916FE18F95A" box="[264,428,1700,1725]" italics="true" pageId="1" pageNumber="2">Type Material</emphasis>
</paragraph>
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<paragraph id="0912BB2DFF97FFE4744FF97EFEFEF8B9" blockId="1.[264,1323,1700,1886]" pageId="1" pageNumber="2">
<materialsCitation id="B9C5B170FF97FFE4744FF97EFB35F8D1" ID-GBIF-Occurrence="3344508302" collectingDate="1996-03-14" collectionCode="NMNZ" country="New Zealand" elevation="1140" latitude="-45.7" location="Spence Burn" longLatPrecision="1128" longitude="167.85" municipality="Takitimu Mts" pageId="1" pageNumber="2" specimenCode="NZMG 21099" specimenCount="1" typeStatus="holotype">
<typeStatus id="D616058FFF97FFE4744FF97EFE67F901" box="[312,467,1740,1766]" pageId="1" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
(juvenile female, 
<collectionCode id="6FBC23E8FF97FFE477DEF97EFCB6F901" box="[681,770,1740,1766]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="1" pageNumber="2" type="Museum">NMNZ</collectionCode>
, Re. 2514): 
<collectingCountry id="71BAFBBDFF97FFE476E5F97EFB86F901" box="[914,1074,1740,1766]" name="New Zealand" pageId="1" pageNumber="2">New Zealand</collectingCountry>
, SL, southern branch of 
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, 
<collectingMunicipality id="E9762157FF97FFE474A6F946FDC0F8E9" box="[465,628,1780,1806]" pageId="1" pageNumber="2">Takitimu Mts</collectingMunicipality>
, grid ref. 
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57879, 
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, 
<geoCoordinate id="6C99DDEAFF97FFE4712BF946FB71F8E9" box="[1116,1221,1780,1806]" degrees="167" direction="east" minutes="51" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="167.85">167°51E</geoCoordinate>
, 
<quantity id="CE5516C8FF97FFE471A4F946FA92F8E9" box="[1235,1318,1780,1806]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="1" pageNumber="2" unit="m" value="1140.0">
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, 
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<collectingDate id="6D576405FF97FFE4747FF8AEFE08F8D1" box="[264,444,1820,1846]" pageId="1" pageNumber="2" value="1996-03-14">14 March 1996</collectingDate>
</date>
, Tony Jewell, 
<taxonomicName id="CEADC0AEFF97FFE47713F8AEFB35F8D1" authority="Jewell &amp; Leschen" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[612,1153,1820,1846]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF97FFE47713F8AEFC1EF8D2" box="[612,938,1820,1845]" italics="true" pageId="1" pageNumber="2">Hoplodactylus cryptozoicus</emphasis>
Jewell &amp; Leschen
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<typeStatus id="D616058FFF97FFE471FFF8AEFA92F8D1" box="[1160,1318,1820,1846]" pageId="1" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
, 2004.
</paragraph>
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<caption id="5DD2EBA5FF94FFE7747FF8A9FBA3F8B4" ID-DOI="http://doi.org/10.5281/zenodo.5495870" ID-Zenodo-Dep="5495870" httpUri="https://zenodo.org/record/5495870/files/figure.png" pageId="2" pageNumber="3" startId="2.[264,367,1819,1843]" targetBox="[338,1289,284,1803]" targetPageId="2">
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<emphasis id="3BD9673FFF94FFE7747FF8A9FE3DF8D4" bold="true" box="[264,393,1819,1843]" pageId="2" pageNumber="3">FIGURE 1.</emphasis>
Scalation of the head and feet of 
<taxonomicName id="CEADC0AEFF94FFE7779BF8AEFC26F8D4" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[748,914,1820,1843]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF94FFE7779BF8AEFC26F8D4" box="[748,914,1820,1843]" italics="true" pageId="2" pageNumber="3">H. cryptozoicus</emphasis>
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(specimen: holotype). A) head, lateral view; B) head, frontal view; C) underside of hind foot. Scale bar = 4 mm.
</paragraph>
</caption>
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<emphasis id="3BD9673FFF95FFE6747FFEAEFDDAFED2" box="[264,622,284,309]" italics="true" pageId="3" pageNumber="4">Additional Material Examined</emphasis>
</paragraph>
<paragraph id="0912BB2DFF95FFE6744FFEF6FD6CFD91" blockId="3.[264,1325,284,630]" pageId="3" pageNumber="4">
<taxonomicName id="CEADC0AEFF95FFE6744FFEF6FD36FEBA" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[312,642,324,349]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF95FFE6744FFEF6FD36FEBA" box="[312,642,324,349]" italics="true" pageId="3" pageNumber="4">Hoplodactylus cryptozoicus</emphasis>
</taxonomicName>
(
<emphasis id="3BD9673FFF95FFE677E0FEF6FD12FEBA" box="[663,678,324,349]" italics="true" pageId="3" pageNumber="4">n</emphasis>
= 6): 
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<emphasis id="3BD9673FFF95FFE6778EFEF1FB36FEBA" bold="true" box="[761,1154,323,349]" pageId="3" pageNumber="4">
<collectingCountry id="71BAFBBDFF95FFE6778EFEF1FC10FEBA" box="[761,932,323,349]" name="New Zealand" pageId="3" pageNumber="4">New Zealand</collectingCountry>
, SL (
<collectingRegion id="CB6975CFFF95FFE6769AFEF1FBC1FEBA" box="[1005,1141,323,349]" country="New Zealand" name="Southland" pageId="3" pageNumber="4">Southland</collectingRegion>
):
</emphasis>
same data as Holotype, 1 live (captive), 5 live (wild). 
<taxonomicName id="CEADC0AEFF95FFE6779CFEDEFBA0FE62" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[747,1044,364,389]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF95FFE6779CFEDEFBA0FE62" box="[747,1044,364,389]" italics="true" pageId="3" pageNumber="4">Hoplodactylus nebulosus</emphasis>
</taxonomicName>
(
<emphasis id="3BD9673FFF95FFE67151FEDEFB80FE62" box="[1062,1076,364,389]" italics="true" pageId="3" pageNumber="4">n</emphasis>
=14)
</materialsCitation>
: 
<materialsCitation id="B9C5B170FF95FFE6710AFED9FCEAFE49" ID-GBIF-Occurrence="3344508301" collectorName="Big Island" country="New Zealand" latitude="-47.133335" location="Stewart Island" longLatPrecision="1119" longitude="167.51666" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Stewart Island">
<emphasis id="3BD9673FFF95FFE6710AFED9FE41FE4A" bold="true" pageId="3" pageNumber="4">
<collectingCountry id="71BAFBBDFF95FFE6710AFED9FA90FE62" box="[1149,1316,363,389]" name="New Zealand" pageId="3" pageNumber="4">New Zealand</collectingCountry>
, SI (
<collectingRegion id="CB6975CFFF95FFE67444FE21FE59FE4A" box="[307,493,403,429]" country="New Zealand" name="Stewart Island" pageId="3" pageNumber="4">Stewart Island</collectingRegion>
)
</emphasis>
: 
<collectorName id="A458DEFBFF95FFE67772FE26FD35FE49" box="[517,641,404,430]" pageId="3" pageNumber="4">Big Island</collectorName>
, 
<geoCoordinate id="6C99DDEAFF95FFE677F8FE26FD53FE49" box="[655,743,404,430]" degrees="47" direction="south" minutes="08" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-47.133335">47°08S</geoCoordinate>
, 
<geoCoordinate id="6C99DDEAFF95FFE67783FE26FCEAFE49" box="[756,862,404,430]" degrees="167" direction="east" minutes="31" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="167.51666">167°31E</geoCoordinate>
</materialsCitation>
, 
<materialsCitation id="B9C5B170FF95FFE6761CFE26FCBDFE19" ID-GBIF-Occurrence="3344508304" collectionCode="NMNZ" country="New Zealand" latitude="-47.116665" location="Re." longLatPrecision="1119" longitude="167.55" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Kundy Island">
D49 20945 53275, 5 (1 live captive; 4 live wild, examined from photographs); 
<collectionCode id="6FBC23E8FF95FFE67781FE0EFCF8FE31" box="[758,844,444,470]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="3" pageNumber="4" type="Museum">NMNZ</collectionCode>
<location id="0C72EDF6FF95FFE6762FFE0EFC34FE31" LSID="urn:lsid:plazi:treatment:81040A3BFF97FFEC747FF98CFC42FB29:0C72EDF6FF95FFE6762FFE0EFC34FE31" box="[856,896,444,470]" country="New Zealand" latitude="-47.116665" longLatPrecision="1119" longitude="167.55" name="Re." pageId="3" pageNumber="4" stateProvince="Kundy Island">Re.</location>
1569, 3; 
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G.549, 1; 
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G.550, 1; 
<collectingRegion id="CB6975CFFF95FFE674F6FE56FD92FE19" box="[385,550,484,510]" inferredBySuffix="true" pageId="3" pageNumber="4">Kundy Island</collectingRegion>
, 
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, 
<geoCoordinate id="6C99DDEAFF95FFE677D7FE56FCBDFE19" box="[672,777,484,510]" degrees="167" direction="east" minutes="33" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="167.55">167°33E</geoCoordinate>
</materialsCitation>
, 
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D49 20965 53295, 
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<collectorName id="A458DEFBFF95FFE6711EFE56FB25FE19" box="[1129,1169,484,510]" pageId="3" pageNumber="4">Re.</collectorName>
93, 1 (
<typeStatus id="D616058FFF95FFE6719AFE56FE8BFDC1" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
); 
<collectingRegion id="CB6975CFFF95FFE67422FDBEFE4BFDC1" box="[341,511,524,550]" inferredBySuffix="true" pageId="3" pageNumber="4">Stewart Island</collectingRegion>
, 
<collectionCode id="6FBC23E8FF95FFE6777AFDBEFD88FDC1" box="[525,572,524,550]" country="New Zealand" name="Otago Museum" pageId="3" pageNumber="4">OM</collectionCode>
<specimenCode id="590B1356FF95FFE67733FDBEFD2FFDC1" box="[580,667,524,550]" collectionCode="OM" country="New Zealand" name="Otago Museum" pageId="3" pageNumber="4">A98.91</specimenCode>
1, (
<typeStatus id="D616058FFF95FFE677BDFDBEFC80FDC1" box="[714,820,524,550]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
, examined from photographs); 
<location id="0C72EDF6FF95FFE671D1FDBEFA93FDC1" LSID="urn:lsid:plazi:treatment:81040A3BFF97FFEC747FF98CFC42FB29:0C72EDF6FF95FFE671D1FDBEFA93FDC1" box="[1190,1319,524,550]" country="New Zealand" latitude="-46.8" longLatPrecision="1121" longitude="168.23334" name="Zero Rock" pageId="3" pageNumber="4" stateProvince="Stewart Island">Zero Rock</location>
, 
<geoCoordinate id="6C99DDEAFF95FFE6747FFD86FED5FDA9" box="[264,353,564,590]" degrees="46" direction="south" minutes="48" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-46.8">46°48S</geoCoordinate>
, 
<geoCoordinate id="6C99DDEAFF95FFE67404FD86FE68FDA9" box="[371,476,564,590]" degrees="168" direction="east" minutes="14" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="168.23334">168°14E</geoCoordinate>
</materialsCitation>
, 
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E48 21466 53666, 1 (live captive); 
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<location id="0C72EDF6FF95FFE676D8FD86FBD8FDA9" LSID="urn:lsid:plazi:treatment:81040A3BFF97FFEC747FF98CFC42FB29:0C72EDF6FF95FFE676D8FD86FBD8FDA9" box="[943,1132,564,590]" country="New Zealand" latitude="-46.816666" longLatPrecision="1121" longitude="168.23334" name="Womens Island" pageId="3" pageNumber="4" stateProvince="Womens Island">Womens Island</location>
</collectingRegion>
, 
<collectionCode id="6FBC23E8FF95FFE6710AFD86FB67FDA9" box="[1149,1235,564,590]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="3" pageNumber="4" type="Museum">NMNZ</collectionCode>
G.203, 
<geoCoordinate id="6C99DDEAFF95FFE6747FFDEEFED5FD91" box="[264,353,604,630]" degrees="46" direction="south" minutes="49" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="-46.816666">46°49S</geoCoordinate>
, 
<geoCoordinate id="6C99DDEAFF95FFE6741AFDEEFE63FD91" box="[365,471,604,630]" degrees="168" direction="east" minutes="14" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="168.23334">168°14E</geoCoordinate>
</materialsCitation>
, E48 21470 53655, 1.
</paragraph>
<paragraph id="0912BB2DFF95FFE6747FFD2DFECBFD5F" blockId="3.[264,1324,671,937]" box="[264,383,671,696]" pageId="3" pageNumber="4">
<emphasis id="3BD9673FFF95FFE6747FFD2DFECBFD5F" box="[264,383,671,696]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
</paragraph>
<paragraph id="0912BB2DFF95FFE6744FFD75FEEDFC4E" blockId="3.[264,1324,671,937]" pageId="3" pageNumber="4">
A medium­sized 
<taxonomicName id="CEADC0AEFF95FFE67773FD75FD1BFD07" authorityName="Fitzinger" authorityYear="1843" box="[516,687,711,736]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="3BD9673FFF95FFE67773FD75FD1BFD07" box="[516,687,711,736]" italics="true" pageId="3" pageNumber="4">Hoplodactylus</emphasis>
</taxonomicName>
(
<quantity id="CE5516C8FF95FFE677B0FD75FCF4FD06" box="[711,832,711,737]" metricMagnitude="-2" metricUnit="m" metricValue="8.65" metricValueMax="8.7" metricValueMin="8.6" pageId="3" pageNumber="4" unit="mm" value="86.5" valueMax="87.0" valueMin="86.0">86–87mm</quantity>
SVL); mouth lining bright orange with a pink tongue; tail shorter than SVL; body scales minute (average length 0.002 x SVL); 1–2 internasal scales present; colour pattern grey with variable markings involving an overall herringboned pattern, often decorated with (or replaced by) irregular orange patches. Distinguished from all described congeners by the combination of mouth colour and tail length.
</paragraph>
<paragraph id="0912BB2DFF95FFE6747FFC63FE26FC0D" blockId="3.[264,1324,977,1889]" box="[264,402,977,1002]" pageId="3" pageNumber="4">
<emphasis id="3BD9673FFF95FFE6747FFC63FE26FC0D" box="[264,402,977,1002]" italics="true" pageId="3" pageNumber="4">Description</emphasis>
</paragraph>
<paragraph id="0912BB2DFF95FFE6744FFC4BFDF3FBF5" blockId="3.[264,1324,977,1889]" box="[312,583,1017,1042]" pageId="3" pageNumber="4">
<emphasis id="3BD9673FFF95FFE6744FFC4BFDF3FBF5" box="[312,583,1017,1042]" italics="true" pageId="3" pageNumber="4">Shape and Proportions</emphasis>
</paragraph>
<paragraph id="0912BB2DFF95FFE6744FFB93FC43FB54" blockId="3.[264,1324,977,1889]" pageId="3" pageNumber="4">
Measurements of 
<typeStatus id="D616058FFF95FFE6776AFB93FD3BFBDC" box="[541,655,1057,1083]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
: live, 
<quantity id="CE5516C8FF95FFE67790FB93FCFCFBDC" box="[743,840,1057,1083]" metricMagnitude="-2" metricUnit="m" metricValue="6.36" pageId="3" pageNumber="4" unit="mm" value="63.6">63.6mm</quantity>
SVL, 
<quantity id="CE5516C8FF95FFE676DEFB93FBBEFBDC" box="[937,1034,1057,1083]" metricMagnitude="-2" metricUnit="m" metricValue="5.62" pageId="3" pageNumber="4" unit="mm" value="56.2">56.2mm</quantity>
tail length; preserved, 60.0mm SVL, 55.0mm tail length. Body stout, limbs slender, pentadactyl; SVL of fully grown adults 
<quantity id="CE5516C8FF95FFE674DAFBC3FD93FB6C" box="[429,551,1137,1163]" metricMagnitude="-2" metricUnit="m" metricValue="8.65" metricValueMax="8.7" metricValueMin="8.6" pageId="3" pageNumber="4" unit="mm" value="86.5" valueMax="87.0" valueMin="86.0">86–87mm</quantity>
; intact tail cyclindrical, moderately stout but tapering, distinctly shorter (82–91%) than SVL; adult live weight 
<quantity id="CE5516C8FF95FFE6765CFB2BFC40FB54" box="[811,1012,1177,1203]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7299999999999998" metricValueMax="1.8399999999999999" metricValueMin="1.6199999999999999" pageId="3" pageNumber="4" unit="g" value="17.299999999999997" valueMax="18.4" valueMin="16.2">16.2–18.4 grams</quantity>
.
</paragraph>
<paragraph id="0912BB2DFF95FFE6744FFB73FAAFFA44" blockId="3.[264,1324,977,1889]" pageId="3" pageNumber="4">Head oviform, 0.87–1.08x longer than broad, forehead flat or slightly convex; snout flat or slightly concave; eye relatively small (diameter about 0.2x head width); not prominent in profile; pupil vertical with slightly serrated margins, eyebrows (= brillar fringe) prominent (width at widest point about 0.4x eye diameter); ear opening of moderate size, 1.5x longer than deep, length 0.5x eye diameter, axis horizontal pointing slightly up and back; distance from snout to eye equal to or slightly longer than distance from eye to ear.</paragraph>
<paragraph id="0912BB2DFF95FFE6744FFA03FBCDF996" blockId="3.[264,1324,977,1889]" pageId="3" pageNumber="4">
Digits tapering very gradually from sole to claw with faint webbing connecting bases of digits 2–4; proximal portion slightly dilated, the margin between proximal and distal portions indistinct on 1 
<superScript id="FED81665FF95FFE67757F9B3FD99F9F5" attach="left" box="[544,557,1537,1554]" fontSize="7" pageId="3" pageNumber="4">st</superScript>
digit; 4 
<superScript id="FED81665FF95FFE677F8F9B3FD2AF9F5" attach="left" box="[655,670,1537,1554]" fontSize="7" pageId="3" pageNumber="4">th</superScript>
digit the longest (0.08–0.10x SVL) on all limbs, first digit the shortest; 4 
<superScript id="FED81665FF95FFE6749DF99EFE4DF9DA" attach="left" box="[490,505,1580,1597]" fontSize="7" pageId="3" pageNumber="4">th</superScript>
toe of hind foot 3.7x longer than broad, length of dilated portion 0.6x total length of toe, width of dilated portion 1.7x width of distal attenuation.
</paragraph>
<paragraph id="0912BB2DFF95FFE6744FF9CDFE37F97F" blockId="3.[264,1324,977,1889]" box="[312,387,1663,1688]" pageId="3" pageNumber="4">
<emphasis id="3BD9673FFF95FFE6744FF9CDFE37F97F" box="[312,387,1663,1688]" italics="true" pageId="3" pageNumber="4">Scales</emphasis>
</paragraph>
<paragraph id="0912BB2DFF95FFE1744FF915FC7DFE3E" blockId="3.[264,1324,977,1889]" lastBlockId="4.[264,1324,284,1196]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Mouth bordered by large and prominent labial scales, each equal or less than as wide as long, 11–13 upper and 10–12 lower, separated at the tip of the snout by a large rostral and mental respectively; rostral 2x wider than deep, subrectangular, touching the nostrils in the upper corners, partially divided in the centre by a short cleft (0.4x rostral depth) extending from the upper margin; nostril also bordered by 4 (usual) or 5 nasals of which the anterior, inner­most is the largest, and the first labial; there are one large (usual) or two small primary internasals followed by 4, 5 (usual) or 6 secondary internasals of varying size; mental about 1.7x wider than deep, trapezium­shaped, usually partially divided by a median cleft (up to half as long as the mental is deep) extending from the posterior margin; one medium­sized postmental separating the 1 
<superScript id="FED81665FF92FFE1765EFE0EFC82FE2A" attach="left" box="[809,822,444,461]" fontSize="7" pageId="4" pageNumber="5">st</superScript>
infralabials.
</paragraph>
<paragraph id="0912BB2DFF92FFE1744FFE55FCC9FDB3" blockId="4.[264,1324,284,1196]" pageId="4" pageNumber="5">
All digits have straight lamellae on the undersurface of dilated proximal portion, with between 7–12 (usually 9–11) unbroken lamellae on the 4 
<superScript id="FED81665FF92FFE176B2FDBDFC66FDF9" attach="left" box="[965,978,527,542]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
toe of the hind foot; apical plates undivided, slightly enlargened on 1 
<superScript id="FED81665FF92FFE17785FD85FD4BFDAF" attach="left" box="[754,767,567,584]" fontSize="7" pageId="4" pageNumber="5">st</superScript>
digit only.
</paragraph>
<paragraph id="0912BB2DFF92FFE1744FFDD0FD6EFB4B" blockId="4.[264,1324,284,1196]" pageId="4" pageNumber="5">
Dorsal body scales minute, regular, juxtaposed, conical with domed tips, slightly broader than long, average length 0.002 x SVL, becoming slightly larger laterally; dorsal snout scales minute, similar in size to dorsal body scales, becoming slightly larger towards labials and nasals, regular; throat scales slightly smaller than dorsal scales; ventral body scales approximately twice the size of dorsal scales (average length 0.005x SVL), sub­flattened, mostly juxtaposed but a few subimbricate, distinctively broader than long; precloacal scales about twice the size of belly scales, each perforated with a single pore (consistently in males, inconsistently in females), forming a subtriangular patch 6–7 deep x 23–26 wide in males (up to 28 wide in females), extending onto femora for about a quarter to a third of their length; postcloacal scales slightly larger again, flattened, hexagonal or with rounded posterior margin, base of tail with 2–3 cloacal spurs (slightly enlargened in females, greatly so in males) with pointed to chisel­edged tips; tail scales 2–3x size of dorsal body scales, subimbricate, roughly pentagonal, arranged in whorls around the tail, becoming larger and more regular ventrally, with visible autonomy septa every 8–9 rows across basal half of tail (
<quantity id="CE5516C8FF92FFE17751FB20FD39FB4B" box="[550,653,1170,1196]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="4" pageNumber="5" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10–11 in</quantity>
total).
</paragraph>
<caption id="5DD2EBA5FF92FFE1747FF967FD9FF897" ID-DOI="http://doi.org/10.5281/zenodo.5495872" ID-Zenodo-Dep="5495872" httpUri="https://zenodo.org/record/5495872/files/figure.png" pageId="4" pageNumber="5" startId="4.[264,368,1749,1773]" targetBox="[331,1256,1239,1739]" targetPageId="4">
<paragraph id="0912BB2DFF92FFE1747FF967FD9FF897" blockId="4.[264,1323,1749,1904]" pageId="4" pageNumber="5">
<emphasis id="3BD9673FFF92FFE1747FF967FE39F90A" bold="true" box="[264,397,1749,1773]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
An adult male 
<taxonomicName id="CEADC0AEFF92FFE1774BF964FCDCF90A" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[572,872,1750,1773]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF92FFE1774BF964FCDCF90A" box="[572,872,1750,1773]" italics="true" pageId="4" pageNumber="5">Hoplodactylus cryptozoicus</emphasis>
</taxonomicName>
, collected live in January 2000 from the type locality in the Spence Burn, Takitimu Mountains. This specimen illustrates the typical primary markings (i.e., the herringbone­shaped patches connected longitudinally), and the two types of ground colour which occur in adults, grey (in this case restricted to the flanks) and two­tone orange. (Photograph by R. Morris).
</paragraph>
</caption>
<paragraph id="0912BB2DFF93FFE0744FFEAEFE3FFED2" blockId="5.[264,1325,284,1870]" box="[312,395,284,309]" pageId="5" pageNumber="6">
<emphasis id="3BD9673FFF93FFE0744FFEAEFE3FFED2" box="[312,395,284,309]" italics="true" pageId="5" pageNumber="6">Colour</emphasis>
</paragraph>
<paragraph id="0912BB2DFF93FFE0744FFEF6FE1DFD79" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">
Skin opaque in all colour phases. Ground colour light grey; 
<specimenCount id="1FAB70A4FF93FFE07171FEF6FB29FEB9" box="[1030,1181,324,350]" pageId="5" pageNumber="6" type="subadult">one subadult</specimenCount>
male had a pale olive tint, and the juvenile 
<typeStatus id="D616058FFF93FFE077F6FEDEFD53FE61" box="[641,743,364,390]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
ranged between light grey and dark brown. Frequently marked with two­tone orange colouration which may consist of irregularly scattered spots, broad expanses around the primary markings (
<figureCitation id="9196A7A8FF93FFE076CAFE0EFBB3FE31" box="[957,1031,444,470]" captionStart="FIGURE 2" captionStartId="4.[264,368,1749,1773]" captionTargetBox="[331,1256,1239,1739]" captionTargetId="figure-387@4.[330,1276,1237,1739]" captionTargetPageId="4" captionText="FIGURE 2. An adult male Hoplodactylus cryptozoicus, collected live in January 2000 from the type locality in the Spence Burn, Takitimu Mountains. This specimen illustrates the typical primary markings (i.e., the herringbone­shaped patches connected longitudinally), and the two types of ground colour which occur in adults, grey (in this case restricted to the flanks) and two­tone orange. (Photograph by R. Morris)." figureDoi="http://doi.org/10.5281/zenodo.5495872" httpUri="https://zenodo.org/record/5495872/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
), or may almost entirely replace the ground colour and primary markings; orange markings less extensive on ventral surfaces, and so far only recorded from males (present on 4 of 
<specimenCount id="1FAB70A4FF93FFE07158FDBEFB25FDC1" box="[1071,1169,524,550]" pageId="5" pageNumber="6" type="male">5 males</specimenCount>
). Individuals may change colour between lighter and darker tones, and in its darkest brown phase the 
<typeStatus id="D616058FFF93FFE0747FFDEEFEDAFD91" box="[264,366,604,630]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
developed a deep reddish­pink overtone along the lower lateral margins of the body and tail.
</paragraph>
<paragraph id="0912BB2DFF93FFE0744FFD1EFE1CFC39" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">Primary dorsal markings a paler shade of the ground colour and sometimes outlined with blackish shadings, consisting of a row of patches arranged in a herringbone configuration (angling forwards from on or near the midline) along either side of the back, sometimes meeting to create a sharp chevron across the midline, and sometimes with a large, paler spot on the lateral extremity; these patches range in shape from narrow bars to narrow triangles, broad triangles, or irregular blotches and are frequently linked longitudinally to create rough­edged dorso­lateral stripes. Usually a series of pale latero­ventral spots present.</paragraph>
<paragraph id="0912BB2DFF93FFE0744FFC5EFD58FAF9" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">On the top of the head the primary markings form a single complex marking on either side, leaving a broad dark median patch on the back of the head, a dark V­shaped mark linking the eyes, and a dark double­triangle (sometimes indistinct or merging into a single mark) linking the front of the eyes across the base of the snout; snout patterned with narrow or broad, pale canthal stripes commonly link in the middle with a backward­pointing chevron to create an irregular pale M­shaped patch across the snout. A dull to bright stripe connects the eye and ear. The jaws have pale colouration extending forwards from below the eye, sometimes almost to the nostrils.</paragraph>
<paragraph id="0912BB2DFF93FFE0744FFA9EFDF7FA59" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">Tail patterned with a continuation of the body markings but usually less regular (especially towards the tip) and commonly with two narrowly separated stripes along the anterior 1/4–3/4. Limbs and upper surfaces of feet and toes coloured as for body but markings small and mostly irregular.</paragraph>
<paragraph id="0912BB2DFF93FFE0744FFA7EFD2AF961" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">
Venter pale grey, sometimes with a pale yellow or pink wash, or grey­brown (
<typeStatus id="D616058FFF93FFE0719AFA7EFE8AF9E9" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
), with pale and indistinct grey­white mottling and flecks, sometimes also with indistinct mid­grey speckles; markings more prominent on tail; throat uniform or with barely discernable speckling; undersurface of feet and toes orange­brown to light orange; precloacal scales sometimes pale yellow.
</paragraph>
<paragraph id="0912BB2DFF93FFE0744FF926FC1CF8A9" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">Iris ranges from mid to dark brown with a dark filigree pattern, sometimes becoming grey­brown at the edges, and bordered (just beneath the surrounding skin, but visible when the skin is gently eased back) by a thin black ring. Mouth lining bright orange; throat lining blackish­grey. Tongue pink, usually with a large grey or indigo patch across the anterior half, anterior edges sometimes speckled with orange.</paragraph>
<paragraph id="0912BB2DFF90FFE3747FFEAEFE31FED2" blockId="6.[264,1324,284,550]" box="[264,389,284,309]" pageId="6" pageNumber="7">
<emphasis id="3BD9673FFF90FFE3747FFEAEFE31FED2" box="[264,389,284,309]" italics="true" pageId="6" pageNumber="7">Etymology</emphasis>
</paragraph>
<paragraph id="0912BB2DFF90FFE3744FFEF6FA92FDC1" blockId="6.[264,1324,284,550]" pageId="6" pageNumber="7">
The name 
<taxonomicName id="CEADC0AEFF90FFE374C2FEF6FDFDFEBA" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[437,585,324,349]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF90FFE374C2FEF6FDFDFEBA" box="[437,585,324,349]" italics="true" pageId="6" pageNumber="7">cryptozoicus</emphasis>
</taxonomicName>
, masculine and used as an adjective, is derived from a combination of the Greek words 
<emphasis id="3BD9673FFF90FFE37734FEDEFD2DFE62" box="[579,665,364,389]" italics="true" pageId="6" pageNumber="7">kryptos</emphasis>
(hidden) and 
<emphasis id="3BD9673FFF90FFE3764AFEDEFC32FE62" box="[829,902,364,389]" italics="true" pageId="6" pageNumber="7">zoikos</emphasis>
(living), referring to the subsurface scree habitat. The common name, Takitimu gecko, is derived from the 
<typeStatus id="D616058FFF90FFE37126FE26FB37FE49" box="[1105,1155,404,430]" pageId="6" pageNumber="7">type</typeStatus>
locality. Takitimu is derived from Maori folklore and is the name of the ancestral waka (canoe) that was swamped by waves and grounded at the mouth of the Waiau River. According to the mythology of the 
<collectingRegion id="CB6975CFFF90FFE374AEFDBEFDFCFDC1" box="[473,584,524,550]" country="New Zealand" name="Southland" pageId="6" pageNumber="7">Murihiku</collectingRegion>
people, the Takitimu Mountains is the waka turned into stone.
</paragraph>
<paragraph id="0912BB2DFF90FFE3747FFDEEFE37FD92" blockId="6.[264,1323,604,1670]" box="[264,387,604,629]" pageId="6" pageNumber="7">
<emphasis id="3BD9673FFF90FFE3747FFDEEFE37FD92" box="[264,387,604,629]" italics="true" pageId="6" pageNumber="7">Comments</emphasis>
</paragraph>
<paragraph id="0912BB2DFF90FFE3744FFD36FB2FFB29" blockId="6.[264,1323,604,1670]" pageId="6" pageNumber="7">
Among the described 
<taxonomicName id="CEADC0AEFF90FFE37735FD36FD59FD7A" authorityName="Fitzinger" authorityYear="1843" box="[578,749,644,669]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="3BD9673FFF90FFE37735FD36FD59FD7A" box="[578,749,644,669]" italics="true" pageId="6" pageNumber="7">Hoplodactylus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CEADC0AEFF90FFE37789FD36FC0DFD7A" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[766,953,644,669]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF90FFE37789FD36FC0DFD7A" box="[766,953,644,669]" italics="true" pageId="6" pageNumber="7">H. cryptozoicus</emphasis>
</taxonomicName>
differs clearly from most species in morphology and its terrestrial habits and presence in the alpine zone. It is most similar to the Stewart Island species 
<taxonomicName id="CEADC0AEFF90FFE377FDFD66FC16FD09" authority="(McCann)" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[650,930,724,750]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF90FFE377FDFD66FC97FD0A" box="[650,803,724,749]" italics="true" pageId="6" pageNumber="7">H. nebulosus</emphasis>
(McCann)
</taxonomicName>
and the more widespread species 
<taxonomicName id="CEADC0AEFF90FFE3747FFD4EFE18FCF2" baseAuthorityName="Gray" baseAuthorityYear="1845" box="[264,428,764,789]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="granulatus">
<emphasis id="3BD9673FFF90FFE3747FFD4EFE18FCF2" box="[264,428,764,789]" italics="true" pageId="6" pageNumber="7">H. granulatus</emphasis>
</taxonomicName>
by the sharing of similar colouration and the presence of orange or yellow mouth lining, a character which is also shared with the species 
<emphasis id="3BD9673FFF90FFE37684FC96FB21FCDA" box="[1011,1173,804,829]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="CEADC0AEFF90FFE37684FC96FB25FCDA" authorityName="Whitaker" authorityYear="1985" box="[1011,1169,804,829]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="kahutarae">H. kahutarae</taxonomicName>
,
</emphasis>
a group that appears to form a monophyletic group based on this character and 16Sr RNA data (
<bibRefCitation id="6D3CC6DCFF90FFE371ACFCFEFE17FC69" author="Hitchmough, R. A." pageId="6" pageNumber="7" refId="ref4726" refString="Hitchmough, R. A. (1997) A systematic review of the New Zealand Gekkonidae. Ph. D. Thesis, Victoria University of Wellington, 370 pp." type="book" year="1997">Hitchmough 1997</bibRefCitation>
; 
<bibRefCitation id="6D3CC6DCFF90FFE374C4FCC6FD0AFC69" author="Chambers, G. K. &amp; Boon, W. M. &amp; Buckley, T. R. &amp; Hitchmough, R. A." box="[435,702,884,910]" pageId="6" pageNumber="7" pagination="377 - 387" refId="ref4549" refString="Chambers, G. K., Boon, W. M., Buckley, T. R. &amp; Hitchmough, R. A. (2001) Using new molecular methods to understand the Gondwanan affinities of the New Zealand biota: Three case studies. Australian Journal of Botany, 49, 377 - 387." type="journal article" year="2001">Chambers et al. 2001</bibRefCitation>
). 
<taxonomicName id="CEADC0AEFF90FFE37796FCC6FB9FFC6A" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[737,1067,884,909]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF90FFE37796FCC6FB9FFC6A" box="[737,1067,884,909]" italics="true" pageId="6" pageNumber="7">Hoplodactylus cryptozoicus</emphasis>
</taxonomicName>
, though, is probably most closely related to 
<taxonomicName id="CEADC0AEFF90FFE37754FC2EFD0AFC52" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[547,702,924,949]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF90FFE37754FC2EFD0AFC52" box="[547,702,924,949]" italics="true" pageId="6" pageNumber="7">H. nebulosus</emphasis>
</taxonomicName>
based on the sharing of grey pigment on tongue, a unique character in the narrow­toed group of the genus, as well as a lightly mottled undersurface. 
<tableCitation id="442F8E96FF90FFE37406FC5EFE72FBE1" box="[369,454,1004,1030]" captionStart="TABLE 1" captionStartId="8.[264,343,283,304]" captionTargetPageId="8" captionText="TABLE 1: Distinguishing features of Hoplodactylus cryptozoicus n. sp. and H. nebulosus (McCann)." httpUri="http://table.plazi.org/id/5DD2EBA5FF9EFFED747FFEA9FB62FED6" pageId="6" pageNumber="7" tableUuid="5DD2EBA5FF9EFFED747FFEA9FB62FED6">Table 1</tableCitation>
provides a full list of differences between the species and 
<figureCitation id="9196A7A8FF90FFE3711BFC5EFB04FBE1" box="[1132,1200,1004,1030]" captionStart="FIGURE 4" captionStartId="7.[264,368,1711,1735]" captionTargetBox="[367,1221,1165,1704]" captionTargetPageId="7" captionText="FIGURE 4. Sloughed skins of Hoplodactylus cryptozoicus (A) and sister­species H. nebulosus (B), illustrating differences in body colour pattern. The H. cryptozoicus skin is from the specimen featured in Fig. 2; the H. nebulosus skin is from a typical example of the species (collected from Big Island). Note the browner colouration of the H. cryptozoicus skin (resulting from the orange pigmentation) and an absence of the dark flecks which embellish H. nebulosus. (Photographs by T. Jewell)." figureDoi="http://doi.org/10.5281/zenodo.5495876" httpUri="https://zenodo.org/record/5495876/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
illustrates some of the difference in colour pattern. 
<taxonomicName id="CEADC0AEFF90FFE3779EFBA6FB9AFBCA" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[745,1070,1044,1069]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF90FFE3779EFBA6FB9AFBCA" box="[745,1070,1044,1069]" italics="true" pageId="6" pageNumber="7">Hoplodactylus cryptozoicus</emphasis>
</taxonomicName>
is larger than 
<taxonomicName id="CEADC0AEFF90FFE371A4FBA6FEE7FBB2" baseAuthorityName="McCann" baseAuthorityYear="1955" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF90FFE371A4FBA6FEE7FBB2" italics="true" pageId="6" pageNumber="7">H. nebulosus</emphasis>
</taxonomicName>
, and has a distinctly shorter tail. The body scales are much smaller, so much so that specimens of 
<taxonomicName id="CEADC0AEFF90FFE374DEFBD6FDD5FB9A" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[425,609,1124,1149]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF90FFE374DEFBD6FDD5FB9A" box="[425,609,1124,1149]" italics="true" pageId="6" pageNumber="7">H. cryptozoicus</emphasis>
</taxonomicName>
have a noticeably smoother texture than similar­sized examples of 
<taxonomicName id="CEADC0AEFF90FFE37428FB3EFE4DFB42" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[351,505,1164,1189]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF90FFE37428FB3EFE4DFB42" box="[351,505,1164,1189]" italics="true" pageId="6" pageNumber="7">H. nebulosus</emphasis>
</taxonomicName>
. Adult 
<taxonomicName id="CEADC0AEFF90FFE37725FB3EFCBFFB42" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[594,779,1164,1189]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF90FFE37725FB3EFCBFFB42" box="[594,779,1164,1189]" italics="true" pageId="6" pageNumber="7">H. cryptozoicus</emphasis>
</taxonomicName>
are slate grey or orange in colour, whereas 
<taxonomicName id="CEADC0AEFF90FFE37079FB3EFEC9FB2A" baseAuthorityName="McCann" baseAuthorityYear="1955" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF90FFE37079FB3EFEC9FB2A" italics="true" pageId="6" pageNumber="7">H. nebulosus</emphasis>
</taxonomicName>
are typically olive­grey or brown, and never have orange markings.
</paragraph>
<paragraph id="0912BB2DFF90FFE3744FFB6EFD46F961" blockId="6.[264,1323,604,1670]" pageId="6" pageNumber="7">
<materialsCitation id="B9C5B170FF90FFE3744FFB6EFD59F961" ID-GBIF-Occurrence="3344508305" collectingDate="2000-01" country="New Zealand" county="Perhaps" location="Museum" municipality="Takitimu" pageId="6" pageNumber="7" specimenCount="2" specimenCount-adult="1" specimenCount-subadult="1" stateProvince="Southland">
The original specimen of Takitimu gecko, a large juvenile female (
<figureCitation id="9196A7A8FF90FFE37121FB6EFB2FFB11" box="[1110,1179,1244,1270]" captionStart="FIGURE 1" captionStartId="2.[264,367,1819,1843]" captionTargetBox="[338,1289,284,1803]" captionTargetId="figure-53@2.[498,1021,775,1181]" captionTargetPageId="2" captionText="FIGURE 1. Scalation of the head and feet of H. cryptozoicus (specimen: holotype). A) head, lateral view; B) head, frontal view; C) underside of hind foot. Scale bar = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5495870" httpUri="https://zenodo.org/record/5495870/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
), died after five months in captivity and was preserved in good condition. In 
<date id="7D139DEDFF90FFE37168FAB6FB0AFAF9" box="[1055,1214,1284,1310]" pageId="6" pageNumber="7" value="2000-01">
<collectingDate id="6D576405FF90FFE37168FAB6FB0AFAF9" box="[1055,1214,1284,1310]" pageId="6" pageNumber="7" value="2000-01">January 2000</collectingDate>
</date>
an adult male (
<figureCitation id="9196A7A8FF90FFE37427FA9EFE22FAA1" box="[336,406,1324,1350]" captionStart="FIGURE 2" captionStartId="4.[264,368,1749,1773]" captionTargetBox="[331,1256,1239,1739]" captionTargetId="figure-387@4.[330,1276,1237,1739]" captionTargetPageId="4" captionText="FIGURE 2. An adult male Hoplodactylus cryptozoicus, collected live in January 2000 from the type locality in the Spence Burn, Takitimu Mountains. This specimen illustrates the typical primary markings (i.e., the herringbone­shaped patches connected longitudinally), and the two types of ground colour which occur in adults, grey (in this case restricted to the flanks) and two­tone orange. (Photograph by R. Morris)." figureDoi="http://doi.org/10.5281/zenodo.5495872" httpUri="https://zenodo.org/record/5495872/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
) was collected live to facilitate behavioural studies and is currently being held at the 
<collectingRegion id="CB6975CFFF90FFE37427FAE6FE72FA89" box="[336,454,1364,1390]" country="New Zealand" name="Southland" pageId="6" pageNumber="7">Southland</collectingRegion>
<location id="0C72EDF6FF90FFE374B9FAE6FD80FA89" LSID="urn:lsid:plazi:treatment:81040A3BFF97FFEC747FF98CFC42FB29:0C72EDF6FF90FFE374B9FAE6FD80FA89" box="[462,564,1364,1390]" country="New Zealand" county="Perhaps" municipality="Takitimu" name="Museum" pageId="6" pageNumber="7" stateProvince="Southland">Museum</location>
and 
<location id="0C72EDF6FF90FFE37719FAE6FD41FA89" LSID="urn:lsid:plazi:treatment:81040A3BFF97FFEC747FF98CFC42FB29:0C72EDF6FF90FFE37719FAE6FD41FA89" box="[622,757,1364,1390]" country="New Zealand" county="Perhaps" municipality="Takitimu" name="Art Gallery" pageId="6" pageNumber="7" stateProvince="Southland">Art Gallery</location>
, Invercargill. Another five examples (
<specimenCount id="1FAB70A4FF90FFE371CDFAE6FA9CFA89" box="[1210,1320,1364,1390]" pageId="6" pageNumber="7" type="adult">one adult</specimenCount>
female and 
<specimenCount id="1FAB70A4FF90FFE374E6FACEFD9FFA71" box="[401,555,1404,1430]" pageId="6" pageNumber="7" type="subadult">four subadult</specimenCount>
males) were examined live by the author and released at the 
<typeStatus id="D616058FFF90FFE37181FACEFA9CFA71" box="[1270,1320,1404,1430]" pageId="6" pageNumber="7">type</typeStatus>
locality. 
<location id="0C72EDF6FF90FFE37419FA16FE01FA59" LSID="urn:lsid:plazi:treatment:81040A3BFF97FFEC747FF98CFC42FB29:0C72EDF6FF90FFE37419FA16FE01FA59" box="[366,437,1444,1470]" country="New Zealand" county="Perhaps" municipality="Takitimu" name="While" pageId="6" pageNumber="7" stateProvince="Southland">While</location>
a larger series of specimens would normally be preferable before attempting a taxonomic description, the 
<collectingMunicipality id="E9762157FF90FFE377E3FA7EFD49FA01" box="[660,765,1484,1510]" pageId="6" pageNumber="7">Takitimu</collectingMunicipality>
gecko is listed as “nationally critical” (
<bibRefCitation id="6D3CC6DCFF90FFE371ACFA7EFE29F9E9" author="Hitchmough, R. A." pageId="6" pageNumber="7" pagination="1 - 210" refId="ref4758" refString="Hitchmough, R. A. (2002) New Zealand threat classification system lists. Department of Conservation, Wellington, Threatened Species Occasional Publication, 23, 1 - 210." type="journal article" year="2002">Hitchmough 2002</bibRefCitation>
) and further collection to obtain museum specimens is illegal. 
<collectingCounty id="E073C3A1FF90FFE371F2FA46FB55F9E9" box="[1157,1249,1524,1550]" pageId="6" pageNumber="7">Perhaps</collectingCounty>
when a more thorough revision of the geckos of 
<collectingCountry id="71BAFBBDFF90FFE3778AF9AEFC2EF9D1" box="[765,922,1564,1590]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
is initated, with the intention of a more critical evaluation of morphological characters (including osteology), then permission to collect material could be obtained
</materialsCitation>
.
</paragraph>
<paragraph id="0912BB2DFF90FFE3747FF90EFDDAF932" blockId="6.[264,1321,1724,1870]" box="[264,622,1724,1749]" pageId="6" pageNumber="7">
<emphasis id="3BD9673FFF90FFE3747FF90EFDDAF932" box="[264,622,1724,1749]" italics="true" pageId="6" pageNumber="7">Natural History and Evolution</emphasis>
</paragraph>
<paragraph id="0912BB2DFF90FFE2744FF956FD98FE49" blockId="6.[264,1321,1724,1870]" lastBlockId="7.[264,1323,284,430]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
The type locality (
<figureCitation id="9196A7A8FF90FFE37766F956FDECF919" box="[529,600,1764,1790]" captionStart="FIGURE 3" captionStartId="7.[264,368,1010,1034]" captionTargetBox="[367,1216,473,992]" captionTargetId="figure-210@7.[367,1217,470,992]" captionTargetPageId="7" captionText="FIGURE 3. Habitat of H. cryptozoicus, showing the scree in which the species occurs, and the alpine vegetation, which surrounds it. The holotype was collected 1.5m uphill from the person shown here and the captive male shown in Fig. 2 was collected near the horizon of the scree in line with the tarns, which are visible in the background. (Photograph by T. Jewell)" figureDoi="http://doi.org/10.5281/zenodo.5495874" httpUri="https://zenodo.org/record/5495874/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
) is a steep, south­facing scree slope, which is surrounded by alpine herbfield/short tussock communities, and is isolated from other scree systems in the mountain range. Specimens were found in channels of medium sized rocks (about 
<quantity id="CE5516C8FF90FFE371B8F886FAAFF8A9" box="[1231,1307,1844,1870]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="cm" value="20.0">20 cm</quantity>
), often close to small islands of vegetation, between 
<quantity id="CE5516C8FF91FFE27617FEAEFC48FED1" box="[864,1020,284,310]" metricMagnitude="3" metricUnit="m" metricValue="1.14" metricValueMax="1.15" metricValueMin="1.13" pageId="7" pageNumber="8" unit="m" value="1140.0" valueMax="1150.0" valueMin="1130.0">1130–1150m</quantity>
. The climate is characterized by a long cold winter, a high rate of precipitation, exceeding 
<quantity id="CE5516C8FF91FFE27172FEF6FBC4FEB9" box="[1029,1136,324,350]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="2500.0">2500mm</quantity>
, much of which falls as snow, frequent cloud and fog cover, low sunshine hours, and frequent strong winds (
<bibRefCitation id="6D3CC6DCFF91FFE27467FE26FE2BFE49" author="Evans, G. R." box="[272,415,404,430]" pageId="7" pageNumber="8" refId="ref4656" refString="Evans, G. R. (1969) Alpine grasslands of the Takitimu Mountains. Protection Forestry Branch Report No. 54, New Zealand Forestry Service, Rangiora. 23 pp." type="book" year="1969">Evans 1969</bibRefCitation>
; pers. obs.).
</paragraph>
<caption id="5DD2EBA5FF91FFE2747FFC40FBF2FB8D" ID-DOI="http://doi.org/10.5281/zenodo.5495874" ID-Zenodo-Dep="5495874" httpUri="https://zenodo.org/record/5495874/files/figure.png" pageId="7" pageNumber="8" startId="7.[264,368,1010,1034]" targetBox="[367,1216,473,992]" targetPageId="7">
<paragraph id="0912BB2DFF91FFE2747FFC40FBF2FB8D" blockId="7.[264,1323,1010,1130]" pageId="7" pageNumber="8">
<emphasis id="3BD9673FFF91FFE2747FFC40FE39FBED" bold="true" box="[264,397,1010,1034]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
Habitat of 
<taxonomicName id="CEADC0AEFF91FFE27778FC41FD0EFBED" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[527,698,1011,1034]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF91FFE27778FC41FD0EFBED" box="[527,698,1011,1034]" italics="true" pageId="7" pageNumber="8">H. cryptozoicus</emphasis>
</taxonomicName>
, showing the scree in which the species occurs, and the alpine vegetation, which surrounds it. The holotype was collected 1.5m uphill from the person shown here and the captive male shown in Fig. 2 was collected near the horizon of the scree in line with the tarns, which are visible in the background. (Photograph by T. Jewell)
</paragraph>
</caption>
<caption id="5DD2EBA5FF91FFE2747FF91DFEE8F88D" ID-DOI="http://doi.org/10.5281/zenodo.5495876" ID-Zenodo-Dep="5495876" httpUri="https://zenodo.org/record/5495876/files/figure.png" pageId="7" pageNumber="8" startId="7.[264,368,1711,1735]" targetBox="[367,1221,1165,1704]" targetPageId="7">
<paragraph id="0912BB2DFF91FFE2747FF91DFEE8F88D" blockId="7.[264,1323,1711,1898]" pageId="7" pageNumber="8">
<emphasis id="3BD9673FFF91FFE2747FF91DFE3EF920" bold="true" box="[264,394,1711,1735]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
Sloughed skins of 
<taxonomicName id="CEADC0AEFF91FFE27722F902FCC9F920" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[597,893,1712,1735]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF91FFE27722F902FCC9F920" box="[597,893,1712,1735]" italics="true" pageId="7" pageNumber="8">Hoplodactylus cryptozoicus</emphasis>
</taxonomicName>
(A) and sister­species 
<taxonomicName id="CEADC0AEFF91FFE27118F902FB4EF920" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[1135,1274,1712,1735]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF91FFE27118F902FB4EF920" box="[1135,1274,1712,1735]" italics="true" pageId="7" pageNumber="8">H. nebulosus</emphasis>
</taxonomicName>
(B), illustrating differences in body colour pattern. The 
<taxonomicName id="CEADC0AEFF91FFE27645F962FC6DF900" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[818,985,1744,1767]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF91FFE27645F962FC6DF900" box="[818,985,1744,1767]" italics="true" pageId="7" pageNumber="8">H. cryptozoicus</emphasis>
</taxonomicName>
skin is from the specimen featured in Fig. 2; the 
<taxonomicName id="CEADC0AEFF91FFE274ADF942FDD1F8E0" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[474,613,1776,1799]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF91FFE274ADF942FDD1F8E0" box="[474,613,1776,1799]" italics="true" pageId="7" pageNumber="8">H. nebulosus</emphasis>
</taxonomicName>
skin is from a typical example of the species (collected from Big Island). Note the browner colouration of the 
<taxonomicName id="CEADC0AEFF91FFE27784F8A2FC28F8C0" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[755,924,1808,1831]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF91FFE27784F8A2FC28F8C0" box="[755,924,1808,1831]" italics="true" pageId="7" pageNumber="8">H. cryptozoicus</emphasis>
</taxonomicName>
skin (resulting from the orange pigmentation) and an absence of the dark flecks which embellish 
<taxonomicName id="CEADC0AEFF91FFE276CBF882FBFEF8A0" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[956,1098,1840,1863]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF91FFE276CBF882FBFEF8A0" box="[956,1098,1840,1863]" italics="true" pageId="7" pageNumber="8">H. nebulosus</emphasis>
</taxonomicName>
. (Photographs by T. Jewell).
</paragraph>
</caption>
<caption id="5DD2EBA5FF9EFFED747FFEA9FB62FED6" ID-Table-UUID="5DD2EBA5FF9EFFED747FFEA9FB62FED6" box="[264,1238,283,305]" httpUri="http://table.plazi.org/id/5DD2EBA5FF9EFFED747FFEA9FB62FED6" pageId="8" pageNumber="9" startId="8.[264,343,283,304]" targetBox="[264,1247,343,1638]" targetIsTable="true" targetPageId="8">
<paragraph id="0912BB2DFF9EFFED747FFEA9FB62FED6" blockId="8.[264,1238,283,305]" box="[264,1238,283,305]" pageId="8" pageNumber="9">
<emphasis id="3BD9673FFF9EFFED747FFEA9FEC5FED7" bold="true" box="[264,369,283,304]" pageId="8" pageNumber="9">TABLE 1:</emphasis>
Distinguishing features of 
<taxonomicName id="CEADC0AEFF9EFFED7701FEAEFC35FED6" authority="Jewell &amp; Leschen, 2004" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[630,897,284,305]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cryptozoicus" status="sp. nov.">
<emphasis id="3BD9673FFF9EFFED7701FEAEFC35FED6" box="[630,897,284,305]" italics="true" pageId="8" pageNumber="9">Hoplodactylus cryptozoicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20EADA44FF9EFFED76F0FEAEFC08FED6" box="[903,956,284,305]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
and 
<taxonomicName id="CEADC0AEFF9EFFED769EFEAEFB67FED6" authority="(McCann)" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[1001,1235,284,305]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF9EFFED769EFEAEFBD3FED6" box="[1001,1127,284,305]" italics="true" pageId="8" pageNumber="9">H. nebulosus</emphasis>
(McCann)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="0912BB2DFF9EFFED77F7FEE5FBDDF981" pageId="8" pageNumber="9">
<table id="7BAD498DFF9E001A747FFEE5FB6BF981" box="[264,1247,343,1638]" gridcols="3" gridrows="32" pageId="8" pageNumber="9">
<tr id="B79DB96FFF9E001A747FFEE5FB6BFE89" box="[264,1247,343,366]" gridrow="0" pageId="8" pageNumber="9" rowspan-0="1">
<th id="F44CD013FF9E001A77F7FEE5FC30FE89" box="[640,900,343,366]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">
<taxonomicName id="CEADC0AEFF9EFFED77F7FEE5FCB3FE89" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[640,775,343,366]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF9EFFED77F7FEE5FCB3FE89" box="[640,775,343,366]" italics="true" pageId="8" pageNumber="9">cryptozoicus</emphasis>
</taxonomicName>
</th>
<th id="F44CD013FF9E001A76C9FEE5FB6BFE89" box="[958,1247,343,366]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">
<taxonomicName id="CEADC0AEFF9EFFED76B7FEE5FB9FFE89" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[960,1067,343,366]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF9EFFED76B7FEE5FB9FFE89" box="[960,1067,343,366]" italics="true" pageId="8" pageNumber="9">nebulosus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="B79DB96FFF9E001A747FFE13FB6BFE51" box="[264,1247,417,438]" gridrow="1" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFE13FDD2FE51" box="[264,614,417,438]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">SVL of fully grown adult specimens</th>
<td id="F44CD013FF9E001A77F7FE13FC30FE51" box="[640,900,417,438]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">86–87mm</td>
<td id="F44CD013FF9E001A76C9FE13FB6BFE51" box="[958,1247,417,438]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">76–80mm</td>
</tr>
<tr id="B79DB96FFF9E001A747FFE7BFB6BFE39" box="[264,1247,457,478]" gridrow="2" pageId="8" pageNumber="9" rowspan-0="1" rowspan-1="1">
<td id="F44CD013FF9E001A76C9FE7BFB6BFE39" box="[958,1247,457,478]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">(to 85mm in the Boat Islands)</td>
</tr>
<tr id="B79DB96FFF9E001A747FFE43FB6BFDE1" box="[264,1247,497,518]" gridrow="3" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFE43FDD2FDE1" box="[264,614,497,518]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">Tail length</th>
<td id="F44CD013FF9E001A77F7FE43FC30FDE1" box="[640,900,497,518]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">0.82–0.91x SVL</td>
<td id="F44CD013FF9E001A76C9FE43FB6BFDE1" box="[958,1247,497,518]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">1.01–1.06x SVL</td>
</tr>
<tr id="B79DB96FFF9E001A747FFDABFB6BFDC9" box="[264,1247,537,558]" gridrow="4" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFDABFB6BFDC9" box="[264,1247,537,558]" colspan="3" colspanRight="2" gridcol="0" gridrow="4" pageId="8" pageNumber="9">Dorsal body scales:</th>
</tr>
<tr id="B79DB96FFF9E001A747FFDF3FB6BFDB1" box="[264,1247,577,598]" gridrow="5" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFDF3FDD2FDB1" box="[264,614,577,598]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">shape (average)</th>
<td id="F44CD013FF9E001A77F7FDF3FC30FDB1" box="[640,900,577,598]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">Broader than long</td>
<td id="F44CD013FF9E001A76C9FDF3FB6BFDB1" box="[958,1247,577,598]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">Longer than broad</td>
</tr>
<tr id="B79DB96FFF9E001A747FFDDBFB6BFD99" box="[264,1247,617,638]" gridrow="6" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFDDBFDD2FD99" box="[264,614,617,638]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">length (range)</th>
<td id="F44CD013FF9E001A77F7FDDBFC30FD99" box="[640,900,617,638]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">0.001–0.004x SVL</td>
<td id="F44CD013FF9E001A76C9FDDBFB6BFD99" box="[958,1247,617,638]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">0.004–0.006x SVL</td>
</tr>
<tr id="B79DB96FFF9E001A747FFD23FB6BFD41" box="[264,1247,657,678]" gridrow="7" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFD23FDD2FD41" box="[264,614,657,678]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">length (average)</th>
<td id="F44CD013FF9E001A77F7FD23FC30FD41" box="[640,900,657,678]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">0.002x SVL</td>
<td id="F44CD013FF9E001A76C9FD23FB6BFD41" box="[958,1247,657,678]" gridcol="2" gridrow="7" pageId="8" pageNumber="9">0.005x SVL</td>
</tr>
<tr id="B79DB96FFF9E001A747FFD0BFB6BFD29" box="[264,1247,697,718]" gridrow="8" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFD0BFB6BFD29" box="[264,1247,697,718]" colspan="3" colspanRight="2" gridcol="0" gridrow="8" pageId="8" pageNumber="9">Ventral body scales</th>
</tr>
<tr id="B79DB96FFF9E001A747FFD53FB6BFD11" box="[264,1247,737,758]" gridrow="9" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFD53FDD2FD11" box="[264,614,737,758]" gridcol="0" gridrow="9" pageId="8" pageNumber="9">shape (average)</th>
<td id="F44CD013FF9E001A77F7FD53FC30FD11" box="[640,900,737,758]" gridcol="1" gridrow="9" pageId="8" pageNumber="9">Broader than long</td>
<td id="F44CD013FF9E001A76C9FD53FB6BFD11" box="[958,1247,737,758]" gridcol="2" gridrow="9" pageId="8" pageNumber="9">Longer than broad</td>
</tr>
<tr id="B79DB96FFF9E001A747FFCBBFB6BFCF9" box="[264,1247,777,798]" gridrow="10" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFCBBFDD2FCF9" box="[264,614,777,798]" gridcol="0" gridrow="10" pageId="8" pageNumber="9">length (range)</th>
<td id="F44CD013FF9E001A77F7FCBBFC30FCF9" box="[640,900,777,798]" gridcol="1" gridrow="10" pageId="8" pageNumber="9">0.004–0.005x SVL</td>
<td id="F44CD013FF9E001A76C9FCBBFB6BFCF9" box="[958,1247,777,798]" gridcol="2" gridrow="10" pageId="8" pageNumber="9">0.007–0.011x SVL</td>
</tr>
<tr id="B79DB96FFF9E001A747FFC83FB6BFCA1" box="[264,1247,817,838]" gridrow="11" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFC83FDD2FCA1" box="[264,614,817,838]" gridcol="0" gridrow="11" pageId="8" pageNumber="9">length (average)</th>
<td id="F44CD013FF9E001A77F7FC83FC30FCA1" box="[640,900,817,838]" gridcol="1" gridrow="11" pageId="8" pageNumber="9">0.005x SVL</td>
<td id="F44CD013FF9E001A76C9FC83FB6BFCA1" box="[958,1247,817,838]" gridcol="2" gridrow="11" pageId="8" pageNumber="9">0.010x SVL</td>
</tr>
<tr id="B79DB96FFF9E001A747FFCEBFB6BFC89" box="[264,1247,857,878]" gridrow="12" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFCEBFDD2FC89" box="[264,614,857,878]" gridcol="0" gridrow="12" pageId="8" pageNumber="9">Eye profile</th>
<td id="F44CD013FF9E001A77F7FCEBFC30FC89" box="[640,900,857,878]" gridcol="1" gridrow="12" pageId="8" pageNumber="9">Small, not prominent</td>
<td id="F44CD013FF9E001A76C9FCEBFB6BFC89" box="[958,1247,857,878]" gridcol="2" gridrow="12" pageId="8" pageNumber="9">Usually large and prominent</td>
</tr>
<tr id="B79DB96FFF9E001A747FFC33FB6BFC71" box="[264,1247,897,918]" gridrow="13" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFC33FDD2FC71" box="[264,614,897,918]" gridcol="0" gridrow="13" pageId="8" pageNumber="9">Eye colour</th>
<td id="F44CD013FF9E001A77F7FC33FC30FC71" box="[640,900,897,918]" gridcol="1" gridrow="13" pageId="8" pageNumber="9">Brown</td>
<td id="F44CD013FF9E001A76C9FC33FB6BFC71" box="[958,1247,897,918]" gridcol="2" gridrow="13" pageId="8" pageNumber="9">Olive­grey</td>
</tr>
<tr id="B79DB96FFF9E001A747FFC1BFB6BFC59" box="[264,1247,937,958]" gridrow="14" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFC1BFDD2FC59" box="[264,614,937,958]" gridcol="0" gridrow="14" pageId="8" pageNumber="9">Black edge to iris</th>
<td id="F44CD013FF9E001A77F7FC1BFC30FC59" box="[640,900,937,958]" gridcol="1" gridrow="14" pageId="8" pageNumber="9">Prominent</td>
<td id="F44CD013FF9E001A76C9FC1BFB6BFC59" box="[958,1247,937,958]" gridcol="2" gridrow="14" pageId="8" pageNumber="9">Indistinct or absent</td>
</tr>
<tr id="B79DB96FFF9E001A747FFC63FB6BFC01" box="[264,1247,977,998]" gridrow="15" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFC63FDD2FC01" box="[264,614,977,998]" gridcol="0" gridrow="15" pageId="8" pageNumber="9">Orange pigmentation on tongue</th>
<td id="F44CD013FF9E001A77F7FC63FC30FC01" box="[640,900,977,998]" gridcol="1" gridrow="15" pageId="8" pageNumber="9">Restricted to tip or absent</td>
<td id="F44CD013FF9E001A76C9FC63FB6BFC01" box="[958,1247,977,998]" gridcol="2" gridrow="15" pageId="8" pageNumber="9">Widespread</td>
</tr>
<tr id="B79DB96FFF9E001A747FFC4BFB6BFBE9" box="[264,1247,1017,1038]" gridrow="16" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFC4BFDD2FBE9" box="[264,614,1017,1038]" gridcol="0" gridrow="16" pageId="8" pageNumber="9">Ground colour of adults</th>
<td id="F44CD013FF9E001A77F7FC4BFC30FBE9" box="[640,900,1017,1038]" gridcol="1" gridrow="16" pageId="8" pageNumber="9">Pale grey (sometimes</td>
<td id="F44CD013FF9E001A76C9FC4BFB6BFBE9" box="[958,1247,1017,1038]" gridcol="2" gridrow="16" pageId="8" pageNumber="9">Dark brown or olive­grey</td>
</tr>
<tr id="B79DB96FFF9E001A747FFB93FB6BFBD1" box="[264,1247,1057,1078]" gridrow="17" pageId="8" pageNumber="9" rowspan-0="1">
<td id="F44CD013FF9E001A77F7FB93FC30FBD1" box="[640,900,1057,1078]" gridcol="1" gridrow="17" pageId="8" pageNumber="9">replaced by orange)</td>
<td id="F44CD013FF9E001A76C9FB93FB6BFBD1" box="[958,1247,1057,1078]" gridcol="2" gridrow="17" pageId="8" pageNumber="9">(rarely grey or dark green)</td>
</tr>
<tr id="B79DB96FFF9E001A747FFBFBFB6BFBB9" box="[264,1247,1097,1118]" gridrow="18" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFBFBFDD2FBB9" box="[264,614,1097,1118]" gridcol="0" gridrow="18" pageId="8" pageNumber="9">Orange patches</th>
<td id="F44CD013FF9E001A77F7FBFBFC30FBB9" box="[640,900,1097,1118]" gridcol="1" gridrow="18" pageId="8" pageNumber="9">Frequently present, usually</td>
<td id="F44CD013FF9E001A76C9FBFBFB6BFBB9" box="[958,1247,1097,1118]" gridcol="2" gridrow="18" pageId="8" pageNumber="9">Absent</td>
</tr>
<tr id="B79DB96FFF9E001A747FFBC3FB6BFB61" box="[264,1247,1137,1158]" gridrow="19" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFBC3FB6BFB61" box="[264,1247,1137,1158]" colspan="3" colspanRight="2" gridcol="0" gridrow="19" pageId="8" pageNumber="9">highly prominent</th>
</tr>
<tr id="B79DB96FFF9E001A747FFB2BFB6BFB49" box="[264,1247,1177,1198]" gridrow="20" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFB2BFDD2FB49" box="[264,614,1177,1198]" gridcol="0" gridrow="20" pageId="8" pageNumber="9">Basic patterning</th>
<td id="F44CD013FF9E001A77F7FB2BFC30FB49" box="[640,900,1177,1198]" gridcol="1" gridrow="20" pageId="8" pageNumber="9">Very variable, bright or</td>
<td id="F44CD013FF9E001A76C9FB2BFB6BFB49" box="[958,1247,1177,1198]" gridcol="2" gridrow="20" pageId="8" pageNumber="9">Little variation, consistently</td>
</tr>
<tr id="B79DB96FFF9E001A747FFB73FB6BFB31" box="[264,1247,1217,1238]" gridrow="21" pageId="8" pageNumber="9" rowspan-0="1">
<td id="F44CD013FF9E001A77F7FB73FC30FB31" box="[640,900,1217,1238]" gridcol="1" gridrow="21" pageId="8" pageNumber="9">drab in appearance</td>
<td id="F44CD013FF9E001A76C9FB73FB6BFB31" box="[958,1247,1217,1238]" gridcol="2" gridrow="21" pageId="8" pageNumber="9">very drab</td>
</tr>
<tr id="B79DB96FFF9E001A747FFB5BFB6BFB19" box="[264,1247,1257,1278]" gridrow="22" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFB5BFDD2FB19" box="[264,614,1257,1278]" gridcol="0" gridrow="22" pageId="8" pageNumber="9">Dorsal neck stripes</th>
<td id="F44CD013FF9E001A77F7FB5BFC30FB19" box="[640,900,1257,1278]" gridcol="1" gridrow="22" pageId="8" pageNumber="9">Narrow or broken</td>
<td id="F44CD013FF9E001A76C9FB5BFB6BFB19" box="[958,1247,1257,1278]" gridcol="2" gridrow="22" pageId="8" pageNumber="9">Broad, complete</td>
</tr>
<tr id="B79DB96FFF9E001A747FFAA3FB6BFAC1" box="[264,1247,1297,1318]" gridrow="23" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFAA3FDD2FAC1" box="[264,614,1297,1318]" gridcol="0" gridrow="23" pageId="8" pageNumber="9">Dark speckles on dorsum</th>
<td id="F44CD013FF9E001A77F7FAA3FC30FAC1" box="[640,900,1297,1318]" gridcol="1" gridrow="23" pageId="8" pageNumber="9">Absent</td>
<td id="F44CD013FF9E001A76C9FAA3FB6BFAC1" box="[958,1247,1297,1318]" gridcol="2" gridrow="23" pageId="8" pageNumber="9">Prominent</td>
</tr>
<tr id="B79DB96FFF9E001A747FFA8BFB6BFAA9" box="[264,1247,1337,1358]" gridrow="24" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFA8BFDD2FAA9" box="[264,614,1337,1358]" gridcol="0" gridrow="24" pageId="8" pageNumber="9">Dark speckles on venter</th>
<td id="F44CD013FF9E001A77F7FA8BFC30FAA9" box="[640,900,1337,1358]" gridcol="1" gridrow="24" pageId="8" pageNumber="9">Indistinct mid­grey,</td>
<td id="F44CD013FF9E001A76C9FA8BFB6BFAA9" box="[958,1247,1337,1358]" gridcol="2" gridrow="24" pageId="8" pageNumber="9">Blackish­grey, prominent</td>
</tr>
<tr id="B79DB96FFF9E001A747FFAD3FB6BFA91" box="[264,1247,1377,1398]" gridrow="25" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFAD3FB6BFA91" box="[264,1247,1377,1398]" colspan="3" colspanRight="2" gridcol="0" gridrow="25" pageId="8" pageNumber="9">or absent</th>
</tr>
<tr id="B79DB96FFF9E001A747FFA3BFB6BFA79" box="[264,1247,1417,1438]" gridrow="26" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFA3BFDD2FA79" box="[264,614,1417,1438]" gridcol="0" gridrow="26" pageId="8" pageNumber="9">Throat patterning or speckled</th>
<td id="F44CD013FF9E001A77F7FA3BFC30FA79" box="[640,900,1417,1438]" gridcol="1" gridrow="26" pageId="8" pageNumber="9">Essentially uniform</td>
<td id="F44CD013FF9E001A76C9FA3BFB6BFA79" box="[958,1247,1417,1438]" gridcol="2" gridrow="26" pageId="8" pageNumber="9">Distinctly mottled</td>
</tr>
<tr id="B79DB96FFF9E001A747FFA03FB6BFA21" box="[264,1247,1457,1478]" gridrow="27" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFA03FDD2FA21" box="[264,614,1457,1478]" gridcol="0" gridrow="27" pageId="8" pageNumber="9">Vocalization</th>
<td id="F44CD013FF9E001A77F7FA03FC30FA21" box="[640,900,1457,1478]" gridcol="1" gridrow="27" pageId="8" pageNumber="9">Rare</td>
<td id="F44CD013FF9E001A76C9FA03FB6BFA21" box="[958,1247,1457,1478]" gridcol="2" gridrow="27" pageId="8" pageNumber="9">Common</td>
</tr>
<tr id="B79DB96FFF9E001A747FFA6BFB6BFA09" box="[264,1247,1497,1518]" gridrow="28" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FFA6BFDD2FA09" box="[264,614,1497,1518]" gridcol="0" gridrow="28" pageId="8" pageNumber="9">Island</th>
<td id="F44CD013FF9E001A77F7FA6BFC30FA09" box="[640,900,1497,1518]" gridcol="1" gridrow="28" pageId="8" pageNumber="9">South Island</td>
<td id="F44CD013FF9E001A76C9FA6BFB6BFA09" box="[958,1247,1497,1518]" gridcol="2" gridrow="28" pageId="8" pageNumber="9">Stewart Island archipelago</td>
</tr>
<tr id="B79DB96FFF9E001A747FF9B3FB6BF9F1" box="[264,1247,1537,1558]" gridrow="29" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FF9B3FDD2F9F1" box="[264,614,1537,1558]" gridcol="0" gridrow="29" pageId="8" pageNumber="9">Habitat</th>
<td id="F44CD013FF9E001A77F7F9B3FC30F9F1" box="[640,900,1537,1558]" gridcol="1" gridrow="29" pageId="8" pageNumber="9">Scree</td>
<td id="F44CD013FF9E001A76C9F9B3FB6BF9F1" box="[958,1247,1537,1558]" gridcol="2" gridrow="29" pageId="8" pageNumber="9">Forest and scrub</td>
</tr>
<tr id="B79DB96FFF9E001A747FF99BFB6BF9D9" box="[264,1247,1577,1598]" gridrow="30" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FF99BFDD2F9D9" box="[264,614,1577,1598]" gridcol="0" gridrow="30" pageId="8" pageNumber="9">Altitude range</th>
<td id="F44CD013FF9E001A77F7F99BFC30F9D9" box="[640,900,1577,1598]" gridcol="1" gridrow="30" pageId="8" pageNumber="9">1130–1150m</td>
<td id="F44CD013FF9E001A76C9F99BFB6BF9D9" box="[958,1247,1577,1598]" gridcol="2" gridrow="30" pageId="8" pageNumber="9">Sea level to about 500m</td>
</tr>
<tr id="B79DB96FFF9E001A747FF9E3FB6BF981" box="[264,1247,1617,1638]" gridrow="31" pageId="8" pageNumber="9">
<th id="F44CD013FF9E001A747FF9E3FDD2F981" box="[264,614,1617,1638]" gridcol="0" gridrow="31" pageId="8" pageNumber="9">Climate</th>
<td id="F44CD013FF9E001A77F7F9E3FC30F981" box="[640,900,1617,1638]" gridcol="1" gridrow="31" pageId="8" pageNumber="9">Alpine, subcontinental</td>
<td id="F44CD013FF9E001A76C9F9E3FB6BF981" box="[958,1247,1617,1638]" gridcol="2" gridrow="31" pageId="8" pageNumber="9">Montane, oceanic</td>
</tr>
</table>
</paragraph>
<paragraph id="0912BB2DFF9EFFED744FF906FB5FF889" blockId="8.[264,1323,1716,1902]" pageId="8" pageNumber="9">
Field observations of specimens found on the scree surface suggest 
<taxonomicName id="CEADC0AEFF9EFFED7120F906FABBF92A" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[1111,1295,1716,1741]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF9EFFED7120F906FABBF92A" box="[1111,1295,1716,1741]" italics="true" pageId="8" pageNumber="9">H. cryptozoicus</emphasis>
</taxonomicName>
is saxicolous (rock­inhabiting) and is at least partially nocturnal; 
<specimenCount id="1FAB70A4FF9EFFED7150F96EFB66F911" box="[1063,1234,1756,1782]" pageId="8" pageNumber="9" type="generic">one specimen</specimenCount>
, found beneath a rock by day, may have been sun basking. However, excepting these occasional examples found on the scree surface, it has proven virtually impossible to study the species; the extent and nature of its activity beneath the surface of the scree is unknown.
</paragraph>
<paragraph id="0912BB2DFF9FFFEC744FFEAEFEF8FE49" blockId="9.[264,1323,284,1230]" pageId="9" pageNumber="10">
The single adult female found in mid 
<date id="7D139DEDFF9FFFEC7785FEAEFC26FED1" box="[754,914,284,310]" pageId="9" pageNumber="10" value="2000-01">January 2000</date>
, was palpated and found to be carrying two, small, but clearly discernable, embryos. Vocalization has been recorded on rare occasions, and only among males when being handled. The call is in the form of a soft chirp.
</paragraph>
<paragraph id="0912BB2DFF9FFFEC744FFE0EFCB6FD21" blockId="9.[264,1323,284,1230]" pageId="9" pageNumber="10">
<taxonomicName id="CEADC0AEFF9FFFEC744FFE0EFD34FE32" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[312,640,444,469]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF9FFFEC744FFE0EFD34FE32" box="[312,640,444,469]" italics="true" pageId="9" pageNumber="10">Hoplodactylus cryptozoicus</emphasis>
</taxonomicName>
is the likely sister species of 
<taxonomicName id="CEADC0AEFF9FFFEC769BFE0EFB3CFE32" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[1004,1160,444,469]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF9FFFEC769BFE0EFB3CFE32" box="[1004,1160,444,469]" italics="true" pageId="9" pageNumber="10">H. nebulosus</emphasis>
</taxonomicName>
based on the morphological characters listed above, and this hypothesis is supported by unpublished phylogenetic data based on mitochondrial 16S rRNA and cytochrome­b genes (R.A. Hitchmough pers. comm. 2000). This pair of taxa form a group with 
<taxonomicName id="CEADC0AEFF9FFFEC714DFD86FB61FDAA" authorityName="Whitaker" authorityYear="1985" box="[1082,1237,564,589]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="kahutarae">
<emphasis id="3BD9673FFF9FFFEC714DFD86FB61FDAA" box="[1082,1237,564,589]" italics="true" pageId="9" pageNumber="10">H. kahutarae</emphasis>
</taxonomicName>
, a species restricted to the Seaward and Inland Kaikoura Ranges in 
<collectingRegion id="CB6975CFFF9FFFEC76A0FDEEFBC5FD91" box="[983,1137,604,630]" country="New Zealand" name="Marlborough" pageId="9" pageNumber="10">Marlborough</collectingRegion>
and Mt. Arthur in Northwest 
<collectingRegion id="CB6975CFFF9FFFEC74DFFD36FE4FFD79" box="[424,507,644,670]" country="New Zealand" name="Nelson" pageId="9" pageNumber="10">Nelson</collectingRegion>
(
<bibRefCitation id="6D3CC6DCFF9FFFEC777DFD36FCB5FD79" author="Whitaker, A. H. &amp; Shaw, T. &amp; Hitchmough, R." box="[522,769,644,670]" pageId="9" pageNumber="10" pagination="1 - 17" refId="ref5054" refString="Whitaker, A. H., Shaw, T. &amp; Hitchmough, R. (1999) Black-eyed geckos (Hoplodactylus kahutarae) on Mt. Arthur. Kahurangi National Park. Department of Conservation, Wellington, Conservation Advisory Notes, 230, 1 - 17." type="journal article" year="1999">
Whitaker 
<emphasis id="3BD9673FFF9FFFEC77F7FD36FD00FD7A" box="[640,692,644,669]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. 1999
</bibRefCitation>
), and the widespread species 
<taxonomicName id="CEADC0AEFF9FFFEC712BFD36FB4BFD7A" baseAuthorityName="Gray" baseAuthorityYear="1845" box="[1116,1279,644,669]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="granulatus">
<emphasis id="3BD9673FFF9FFFEC712BFD36FB4BFD7A" box="[1116,1279,644,669]" italics="true" pageId="9" pageNumber="10">H. granulatus</emphasis>
</taxonomicName>
, by the sharing of an orange­pigmented mouth.
</paragraph>
<paragraph id="0912BB2DFF9FFFEC744FFD66FC42FB29" blockId="9.[264,1323,284,1230]" pageId="9" pageNumber="10">
The two sister species occur on separate islands: 
<taxonomicName id="CEADC0AEFF9FFFEC7605FD66FB9EFD0A" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[882,1066,724,749]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF9FFFEC7605FD66FB9EFD0A" box="[882,1066,724,749]" italics="true" pageId="9" pageNumber="10">H. cryptozoicus</emphasis>
</taxonomicName>
occurs well inland on the South Island, while 
<taxonomicName id="CEADC0AEFF9FFFEC776CFD4EFD07FCF2" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[539,691,764,789]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF9FFFEC776CFD4EFD07FCF2" box="[539,691,764,789]" italics="true" pageId="9" pageNumber="10">H. nebulosus</emphasis>
</taxonomicName>
occurs 
<quantity id="CE5516C8FF9FFFEC767BFD4EFC23FCF1" box="[780,919,764,790]" metricMagnitude="5" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="9" pageNumber="10" unit="km" value="140.0" valueMax="180.0" valueMin="100.0">100–180km</quantity>
further south, on the opposite side of the Foveaux Strait, in the Stewart Island archipelago. These two species may be derived from a more widespread ancestor that was once located in the southern portion of the south island. Though relatively recent, the development of the Foveaux Strait 11,000 years ago (M. Marra, pers. com) may have acted as a barrier to restrict gene flow in the ancestral population that resulted in the formation of two species. Another consideration is that the divergence between 
<taxonomicName id="CEADC0AEFF9FFFEC748EFC5EFD05FBE2" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[505,689,1004,1029]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF9FFFEC748EFC5EFD05FBE2" box="[505,689,1004,1029]" italics="true" pageId="9" pageNumber="10">H. cryptozoicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="CEADC0AEFF9FFFEC779CFC5EFC31FBE2" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[747,901,1004,1029]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF9FFFEC779CFC5EFC31FBE2" box="[747,901,1004,1029]" italics="true" pageId="9" pageNumber="10">H. nebulosus</emphasis>
</taxonomicName>
could have been associated with an ecological shift from an ancestral forest­dwelling and arboreal species to the alpine and saxicolous habit of 
<taxonomicName id="CEADC0AEFF9FFFEC777BFB8EFD7AFBB2" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[524,718,1084,1109]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF9FFFEC777BFB8EFD7AFBB2" box="[524,718,1084,1109]" italics="true" pageId="9" pageNumber="10">H. cryptozoicus</emphasis>
</taxonomicName>
. The morphological and ecological differences between 
<taxonomicName id="CEADC0AEFF9FFFEC7404FBD6FD98FB9A" authorityName="Jewell &amp; Leschen" authorityYear="2004" box="[371,556,1124,1149]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cryptozoicus">
<emphasis id="3BD9673FFF9FFFEC7404FBD6FD98FB9A" box="[371,556,1124,1149]" italics="true" pageId="9" pageNumber="10">H. cryptozoicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="CEADC0AEFF9FFFEC771FFBD6FCB6FB9A" baseAuthorityName="McCann" baseAuthorityYear="1955" box="[616,770,1124,1149]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="3BD9673FFF9FFFEC771FFBD6FCB6FB9A" box="[616,770,1124,1149]" italics="true" pageId="9" pageNumber="10">H. nebulosus</emphasis>
</taxonomicName>
(
<tableCitation id="442F8E96FF9FFFEC7664FBD6FCDFFB99" box="[787,875,1124,1150]" captionStart="TABLE 1" captionStartId="8.[264,343,283,304]" captionTargetPageId="8" captionText="TABLE 1: Distinguishing features of Hoplodactylus cryptozoicus n. sp. and H. nebulosus (McCann)." httpUri="http://table.plazi.org/id/5DD2EBA5FF9EFFED747FFEA9FB62FED6" pageId="9" pageNumber="10" tableUuid="5DD2EBA5FF9EFFED747FFEA9FB62FED6">Table 1</tableCitation>
) suggests a more ancient divergence, but any questions about the rates of speciation and ecological shifts in the genus requires a robust phylogeny that includes all of the 
<taxonomicName id="CEADC0AEFF9FFFEC7790FB06FC42FB29" box="[743,1014,1204,1230]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="3BD9673FFF9FFFEC7790FB06FC26FB2A" box="[743,914,1204,1229]" italics="true" pageId="9" pageNumber="10">Hoplodactylus</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>