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<document id="6D2F81654A8E8D5F0147E62931DA0904" ID-CLB-Dataset="29285" ID-DOI="10.11646/zootaxa.4454.1.12" ID-GBIF-Dataset="8bc1819c-b047-44a1-86f1-eb7ae63b94ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446560" ID-ZooBank="72FED011-9838-46EC-A846-D4993F288339" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538746535996" checkinUser="plazi" docAuthor="Smith, David G. &amp; Ho, Hsuan-Ching" docDate="2018" docId="03CEC81768031A17E5EA8AFAFC9400EF" docLanguage="en" docName="zootaxa.4454.1.12.pdf" docOrigin="Zootaxa 4454 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bathycongrus macroporis" docType="treatment" docVersion="8" lastPageNumber="140" masterDocId="FFF7B06F68171A01E57D8834FFAF0302" masterDocTitle="The congrid eel genus Bathycongrus of Taiwan, with descriptions of three new species (Anguilliformes: Congridae)" masterLastPageNumber="146" masterPageNumber="118" pageNumber="138" updateTime="1698565288078" updateUser="ExternalLinkService">
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<mods:title id="DDA8E18B545D5E13F066D5C95EF5ED35">The congrid eel genus Bathycongrus of Taiwan, with descriptions of three new species (Anguilliformes: Congridae)</mods:title>
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<mods:namePart id="4D08719D1405553A7304D3B557ABD904">Smith, David G.</mods:namePart>
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<mods:namePart id="977541D39E891EE8948CBB33D8571EEF">Ho, Hsuan-Ching</mods:namePart>
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<treatment id="03CEC81768031A17E5EA8AFAFC9400EF" ID-DOI="http://doi.org/10.5281/zenodo.5960255" ID-GBIF-Taxon="148639524" ID-Zenodo-Dep="5960255" LSID="urn:lsid:plazi:treatment:03CEC81768031A17E5EA8AFAFC9400EF" httpUri="http://treatment.plazi.org/id/03CEC81768031A17E5EA8AFAFC9400EF" lastPageId="22" lastPageNumber="140" pageId="20" pageNumber="138">
<subSubSection id="C37D2A8A68031A15E5EA8AFAFDAE0008" pageId="20" pageNumber="138" type="nomenclature">
<paragraph id="8BD8790168031A15E5EA8AFAFCFB01EB" blockId="20.[151,852,718,779]" box="[151,852,718,745]" pageId="20" pageNumber="138">
<heading id="D090CE6D68031A15E5EA8AFAFCFB01EB" bold="true" box="[151,852,718,745]" fontSize="11" level="1" pageId="20" pageNumber="138" reason="1">
<emphasis id="B913A51368031A15E5EA8AFAFCFB01EB" bold="true" box="[151,852,718,745]" pageId="20" pageNumber="138">
<taxonomicName id="4C67028268031A15E5EA8AFAFD0701EB" authority="(Kotthaus, 1968)" baseAuthorityName="Kotthaus" baseAuthorityYear="1968" box="[151,680,718,745]" class="Actinopterygii" family="Congridae" genus="Bathycongrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="20" pageNumber="138" phylum="Chordata" rank="species" species="macroporis">
<emphasis id="B913A51368031A15E5EA8AFAFE7F01EA" bold="true" box="[151,464,718,744]" italics="true" pageId="20" pageNumber="138">Bathycongrus macroporis</emphasis>
(
<bibRefCitation id="EFF604F068031A15E4A28AFBFD0F01EB" box="[479,672,719,745]" pageId="20" pageNumber="138" refId="ref16194" refString="Kotthaus, A. (1968) Fische des Indischen Ozeans. A. Systematischer Teil. III. Ostariophysi und Apodes. Meteor Forschungsergebnisse, Reihe D, Biologie, 3, 14 - 56." type="journal article">Kotthaus, 1968</bibRefCitation>
)
</taxonomicName>
(New record)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD8790168031A15E5EA8AC6FDAE0008" blockId="20.[151,852,718,779]" box="[151,513,754,779]" pageId="20" pageNumber="138">
<figureCitation id="135C658468031A15E5EA8AC6FF5C0008" box="[151,243,754,779]" captionStart="FIGURE 3" captionStartId="5.[151,250,1429,1451]" captionTargetBox="[169,1400,300,1396]" captionTargetId="figure@5.[151,1436,265,1407]" captionTargetPageId="5" captionText="FIGURE 3. Tooth pattern on upper jaw. A. Bathycongrus bimaculatus sp. nov., NMMB-P7012, paratype. B. B. bleekeri, NMMB-P26214. C. B. castlei sp. nov., NMMB-P21740, paratype. D. B. guttulatus, NMMB-P7607. E. B. graciliceps sp. nov., holotype. F. B. macroporis, NMMB-P29128. Not to scale." httpUri="https://zenodo.org/record/1446566/files/figure.png" pageId="20" pageNumber="138">Figs. 3F</figureCitation>
,
<figureCitation id="135C658468031A15E5838AC6FEEA0008" box="[254,325,754,778]" captionStart="FIGURE 9" captionStartId="21.[151,250,1665,1687]" captionTargetBox="[151,1436,193,1638]" captionTargetId="figure@21.[151,1436,193,1644]" captionTargetPageId="21" captionText="FIGURE 9. Bathycongrus macroporis (Kotthaus, 1968), USNM 398471, 415+ mm TL. Bar indicates locality of posterior nostril. Arrow indicates the ST pore." httpUri="https://zenodo.org/record/1446578/files/figure.png" pageId="20" pageNumber="138">9AC</figureCitation>
; Tables 13, 78
</paragraph>
</subSubSection>
<subSubSection id="C37D2A8A68031A15E5EA8B0EFD5500DA" pageId="20" pageNumber="138" type="reference_group">
<paragraph id="8BD8790168031A15E5EA8B0EFC98006C" blockId="20.[151,1436,826,878]" pageId="20" pageNumber="138">
<materialsCitation id="3B0F735C68031A15E5EA8B0EFC98006C" ID-GBIF-Occurrence="1914264732" elevation="175" latitude="11.633333" location="Sokotra Island" longLatPrecision="1295" longitude="52.866665" municipality="Indian Ocean" pageId="20" pageNumber="138" specimenCount="1" typeStatus="holotype">
<treatmentCitationGroup id="AB775E2F68031A15E5EA8B0EFC98006C" infered="true" pageId="20" pageNumber="138">
<taxonomicName id="4C67028268031A15E5EA8B0EFD370052" authority="Kotthaus, 1968: 35" authorityName="Kotthaus" authorityPageNumber="35" authorityYear="1968" box="[151,664,826,848]" class="Actinopterygii" family="Congridae" genus="Pseudophichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="20" pageNumber="138" phylum="Chordata" rank="species" species="macroporis">
<emphasis id="B913A51368031A15E5EA8B0EFE6A0052" box="[151,453,826,848]" italics="true" pageId="20" pageNumber="138">Pseudophichthys macroporis</emphasis>
<treatmentCitation id="0AC65F1068031A15E4AE8B0EFD370052" author="" box="[467,664,826,848]" page="35" pageId="20" pageNumber="138">
<bibRefCitation id="EFF604F068031A15E4AE8B0EFDD50052" box="[467,634,826,848]" pageId="20" pageNumber="138" refId="ref16194" refString="Kotthaus, A. (1968) Fische des Indischen Ozeans. A. Systematischer Teil. III. Ostariophysi und Apodes. Meteor Forschungsergebnisse, Reihe D, Biologie, 3, 14 - 56." type="journal article">Kotthaus, 1968</bibRefCitation>
:35
</treatmentCitation>
</taxonomicName>
, figs. 138140, 152 (
<typeStatus id="54DCC7A368031A15E6F38B0EFC140052" box="[910,955,826,848]" pageId="20" pageNumber="138">type</typeStatus>
locality: off
<location id="8EB82FDA68031A15E12B8B0EFB5C0052" LSID="urn:lsid:plazi:treatment:03CEC81768031A17E5EA8AFAFC9400EF:8EB82FDA68031A15E12B8B0EFB5C0052" box="[1110,1267,826,848]" latitude="11.633333" longLatPrecision="1295" longitude="52.866665" municipality="Indian Ocean" name="Sokotra Island" pageId="20" pageNumber="138">Sokotra Island</location>
,
<collectingMunicipality id="6BBCE37B68031A15E07B8B0EFA370052" box="[1286,1432,826,848]" pageId="20" pageNumber="138">Indian Ocean</collectingMunicipality>
,
<geoCoordinate id="EE531FC668031A15E5BA8B6CFE9C006C" box="[199,307,856,878]" direction="north" orientation="latitude" pageId="20" pageNumber="138" precision="92" value="11.565">11°33.9N</geoCoordinate>
,
<geoCoordinate id="EE531FC668031A15E4408B6CFE3E006C" box="[317,401,856,878]" direction="east" orientation="longitude" pageId="20" pageNumber="138" precision="925" value="52.9">52°54E</geoCoordinate>
to
<geoCoordinate id="EE531FC668031A15E4CF8B6CFDA5006C" box="[434,522,856,878]" direction="north" orientation="latitude" pageId="20" pageNumber="138" precision="925" value="11.633333">11°38N</geoCoordinate>
,
<geoCoordinate id="EE531FC668031A15E7688B6CFDC3006C" box="[533,620,856,878]" direction="east" orientation="longitude" pageId="20" pageNumber="138" precision="925" value="52.866665">52°52E</geoCoordinate>
, depth 337
<elevation id="004A9E3268031A15E7948B6CFC84006C" box="[745,811,856,878]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="20" pageNumber="138" unit="m" value="175.0">
<quantity id="4C9FD4E468031A15E7948B6CFC84006C" box="[745,811,856,878]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="20" pageNumber="138" unit="m" value="175.0">175 m</quantity>
</elevation>
).
</treatmentCitationGroup>
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</paragraph>
<paragraph id="8BD8790168031A15E5EA8BAFFD5500DA" blockId="20.[151,1437,923,1920]" pageId="20" pageNumber="138">
<emphasis id="B913A51368031A15E5EA8BAFFE3500B6" bold="true" box="[151,410,923,948]" pageId="20" pageNumber="138">Specimens examined.</emphasis>
NMMB-P29128 (1, 395),
<date id="FFD95FC168031A15E7A88BAFFCDE00B6" box="[725,881,923,948]" pageId="20" pageNumber="138" value="2011-09-15">15 Sep. 2011</date>
;
<materialsCitation id="3B0F735C68031A15E6FD8BA8FD5500DA" ID-GBIF-Occurrence="1914264719" collectingDate="2008-09-23" collectionCode="USNM" country="Taiwan" location="Chang Bin" pageId="20" pageNumber="138" specimenCode="USNM 398471" specimenCount="1" stateProvince="Taitung">
<specimenCode id="DBC1D17A68031A15E6FD8BA8FB9F00B6" box="[896,1072,924,948]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="138">USNM 398471</specimenCode>
(1, 415+),
<collectingDate id="EF9DA62968031A15E1C48BA8FAF800B6" box="[1209,1367,924,948]" pageId="20" pageNumber="138" value="2008-09-23">
<date id="FFD95FC168031A15E1C48BA8FAF800B6" box="[1209,1367,924,948]" pageId="20" pageNumber="138" value="2008-09-23">23 Sep. 2008</date>
</collectingDate>
; from
<location id="8EB82FDA68031A15E5EA8B8BFEBD00DA" LSID="urn:lsid:plazi:treatment:03CEC81768031A17E5EA8AFAFC9400EF:8EB82FDA68031A15E5EA8B8BFEBD00DA" box="[151,274,959,984]" country="Taiwan" name="Chang Bin" pageId="20" pageNumber="138" stateProvince="Taitung">Chang Bin</location>
,
<collectingRegion id="49A3B7E368031A15E4608B8BFEDA00DA" box="[285,373,959,984]" country="Taiwan" name="Taitung" pageId="20" pageNumber="138">Taitung</collectingRegion>
, eastern
<collectingCountry id="F370399168031A15E4AA8B8BFD8300DA" box="[471,556,959,984]" name="Taiwan" pageId="20" pageNumber="138">Taiwan</collectingCountry>
, by hook and line.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37D2A8A68031A15E5BA8BD7FD9E076A" pageId="20" pageNumber="138" type="diagnosis">
<paragraph id="8BD8790168031A15E5BA8BD7FD9E076A" blockId="20.[151,1437,923,1920]" pageId="20" pageNumber="138">
<emphasis id="B913A51368031A15E5BA8BD7FEEE00FE" bold="true" box="[199,321,995,1020]" pageId="20" pageNumber="138">Diagnosis.</emphasis>
A large, robust species of
<taxonomicName id="4C67028268031A15E70D8BD7FCA400F9" authorityName="Ogilby" authorityYear="1898" box="[624,779,995,1019]" class="Actinopterygii" family="Congridae" genus="Bathycongrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="20" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="B913A51368031A15E70D8BD7FCA400F9" box="[624,779,995,1019]" italics="true" pageId="20" pageNumber="138">Bathycongrus</emphasis>
</taxonomicName>
with posterior nostril above level of upper margin of eye; band of granular teeth on jaws and vomer; vomerine teeth forming a long and broad patch; trunk 1.31.4 times head length; predorsal vertebrae 910, preanal vertebrae 3436, precaudal vertebrae 3940, total vertebrae 148+; and preanal lateral-line pores 3032.
</paragraph>
</subSubSection>
<subSubSection id="C37D2A8A68031A17E5BA8C47FAE901AA" lastPageId="22" lastPageNumber="140" pageId="20" pageNumber="138" type="description">
<paragraph id="8BD8790168031A15E5BA8C47FB72061E" blockId="20.[151,1437,923,1920]" pageId="20" pageNumber="138">
<emphasis id="B913A51368031A15E5BA8C47FEF6078E" bold="true" box="[199,345,1139,1164]" pageId="20" pageNumber="138">Description.</emphasis>
Proportional measurements and meristic data are shown in
<tableCitation id="C6E54CBA68031A15E17A8C47FB2A078E" box="[1031,1157,1139,1164]" captionStart-0="TABLE 7" captionStart-1="TABLE 8" captionStartId-0="16.[151,239,1036,1059]" captionStartId-1="17.[151,239,554,577]" captionText-0="TABLE 7. Morphometric data of five Bathycongrus species in present study. * means data not available." captionText-1="TABLE 8. Meristic data of five Bathycongrus species in present study. Values in brackets are those of comparative specimens." pageId="20" pageNumber="138">Tables 78</tableCitation>
. Head length 2.3 (
<quantity id="4C9FD4E468031A15E0248C40FA33078E" box="[1369,1436,1139,1164]" metricMagnitude="-2" metricUnit="m" metricValue="6.096" pageId="20" pageNumber="138" unit="in" value="2.4">2.4 in</quantity>
USNM 398471) in PAL, 6.4 (*) in TL; preanal length 2.8 (*) in TL; predorsal length 2.2 (2.3) in PAL, 6.0 (*) in TL; trunk length 1.8 (1.7) in PAL, 4.8 (*) in TL; tail length 1.6 (*) in TL; depth at head 6.2 (6.7) in PAL, width at head 7.8 (7.1) in PAL. Snout length 4.0 (4.4) in HL; eye diameter 5.6 (7.0); interorbital width 7.1 (7.3); upper jaw 2.9 (3.1); gill opening width 7.2 (6.7); interbranchial width 5.1 (5.0); pectoral-fin length 2.9 (3.1).
</paragraph>
<paragraph id="8BD8790168031A15E5BA8D13FC9D068A" blockId="20.[151,1437,923,1920]" pageId="20" pageNumber="138">Body moderately elongate, robust, more or less uniform in depth over head and trunk, round in cross section anteriorly, becoming compressed posteriorly; trunk stout, 1.31.4 times of head length; tip of tail tapering, moderately filiform; anus near anterior third of total length.</paragraph>
<paragraph id="8BD8790168031A15E5BA8DA7FD17051A" blockId="20.[151,1437,923,1920]" pageId="20" pageNumber="138">Dorsal fin begins over pectoral fin, slightly behind fin base; continuous around tip of tail with caudal and anal fins. Anal fin begins immediately behind anus. Pectoral fin well developed, long and broad. Gill opening moderately large, about same size as eye diameter, its upper end opposite middle of pectoral-fin base; interbranchial broader than gill opening and eye.</paragraph>
<paragraph id="8BD8790168031A15E5BA8E17FBDB05AA" blockId="20.[151,1437,923,1920]" pageId="20" pageNumber="138">Head relatively large, its length 15.5% TL, deepest about occiput, slightly tapering anteriorly from this point; dorsal profile nearly flat from occiput to internasal space; snout broadly rounded in dorsal view, its length 1.41.6 eye diameter, projecting beyond lower jaw; fleshy part of snout with a slight median keel on underside, projecting anteriorly beyond anterior end of intermaxillary tooth patch; rictus below middle of eye.</paragraph>
<paragraph id="8BD8790168031A15E5BA8E87FA8C0416" blockId="20.[151,1437,923,1920]" pageId="20" pageNumber="138">Anterior nostril tubular, near tip of snout, directed ventrolaterally. Posterior nostril elliptical, with a slightly raised rim, in front of eye above level of upper margin of eye and behind anterior margin of eye. Upper lip with flange greatly reduced; lower lip with well-developed downturned flange. Tongue free, long, and broad.</paragraph>
<paragraph id="8BD8790168031A15E5BA8F2BFEA40482" blockId="20.[151,1437,923,1920]" pageId="20" pageNumber="138">Lateral line complete, first pore on each side slightly enlarged, the canal extended to caudal-fin base; 5 (5) before pectoral-fin base, 6 (7) pores before dorsal-fin origin, 30 (32) before anal-fin origin, total pores not countable.</paragraph>
<caption id="DF18298968021A14E5EA8EB5FDA505B4" httpUri="https://zenodo.org/record/1446578/files/figure.png" pageId="21" pageNumber="139" startId="21.[151,250,1665,1687]" targetBox="[151,1436,193,1638]" targetPageId="21">
<paragraph id="8BD8790168021A14E5EA8EB5FDA505B4" blockId="21.[151,1436,1665,1718]" pageId="21" pageNumber="139">
<emphasis id="B913A51368021A14E5EA8EB5FEBA0594" bold="true" box="[151,277,1665,1687]" pageId="21" pageNumber="139">FIGURE 9.</emphasis>
<taxonomicName id="4C67028268021A14E4628EB5FD4C0595" authority="(Kotthaus, 1968)" authorityName="Kotthaus" authorityYear="1968" baseAuthorityName="Kotthaus" baseAuthorityYear="1968" box="[287,739,1665,1687]" class="Actinopterygii" family="Congridae" genus="Bathycongrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="21" pageNumber="139" phylum="Chordata" rank="species" species="macroporis">
<emphasis id="B913A51368021A14E4628EB5FD860595" box="[287,553,1665,1687]" italics="true" pageId="21" pageNumber="139">Bathycongrus macroporis</emphasis>
(Kotthaus, 1968)
</taxonomicName>
, USNM 398471, 415+ mm TL. Bar indicates locality of posterior nostril. Arrow indicates the ST pore.
</paragraph>
</caption>
<paragraph id="8BD8790168021A17E5BA8ED6FCF803F5" blockId="21.[151,1436,1762,2003]" lastBlockId="22.[151,1436,151,1005]" lastPageId="22" lastPageNumber="140" pageId="21" pageNumber="139">
Head pores vary in size, most enlarged to some extent (
<figureCitation id="135C658468021A14E62E8ED6FC0105F9" box="[851,942,1762,1787]" captionStart="FIGURE 9" captionStartId="21.[151,250,1665,1687]" captionTargetBox="[151,1436,193,1638]" captionTargetId="figure@21.[151,1436,193,1644]" captionTargetPageId="21" captionText="FIGURE 9. Bathycongrus macroporis (Kotthaus, 1968), USNM 398471, 415+ mm TL. Bar indicates locality of posterior nostril. Arrow indicates the ST pore." httpUri="https://zenodo.org/record/1446578/files/figure.png" pageId="21" pageNumber="139">Fig. 9C</figureCitation>
). Supraorbital canal with 3 pores; the first (ethmoidal pore) small, on ventral surface of snout tip, just above lip; the second enlarged, about twice the size of first, and immediately in front of anterior nostril, the third greatly enlarged and immediately above anterior nostril, about same size as anterior nostril. Infraorbital canal with 5 pores, the first 3 pores enlarged; the first at posterodorsal corner of anterior nostril; second behind and below the first, on upper lip; third behind second, on upper lip, about midway between anterior nostril and eye; fourth slightly smaller, below anterior part of eye; fifth behind and slightly below rictus; no pores behind eye. Preoperculomandibular canal with 10 pores; 7 pores in mandibular section, 6 along lower jaw and 1 behind rictus; first pore small, near tip of lower jaw, second and third pores progressively larger, the third the largest in the series; fourth through seventh smaller; 3 pores in preopercular section. Supratemporal commissure with a single median pore.
</paragraph>
<paragraph id="8BD8790168011A17E5BA8930FD100267" blockId="22.[151,1436,151,1005]" pageId="22" pageNumber="140">
Predorsal vertebrae 9 (10); preanal vertebrae 34 (36); precaudal vertebrae 40 (39). Total vertebrae not available due to the damaged tails in both specimens, the incomplete counts are 148+ (117+). The
<typeStatus id="54DCC7A368011A17E1B48913FA840242" box="[1225,1323,295,320]" pageId="22" pageNumber="140" type="holotype">holotype</typeStatus>
was also incomplete, with 126 vertebrae (
<bibRefCitation id="EFF604F068011A17E77F8978FD1C0267" box="[514,691,332,357]" pageId="22" pageNumber="140" refId="ref16194" refString="Kotthaus, A. (1968) Fische des Indischen Ozeans. A. Systematischer Teil. III. Ostariophysi und Apodes. Meteor Forschungsergebnisse, Reihe D, Biologie, 3, 14 - 56." type="journal article">Kotthaus, 1968</bibRefCitation>
).
</paragraph>
<paragraph id="8BD8790168011A17E5BA895BFF5C013F" blockId="22.[151,1436,151,1005]" pageId="22" pageNumber="140">
Teeth on jaws moderately stout to blunt (
<figureCitation id="135C658468011A17E7EF895BFD48028A" box="[658,743,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1429,1451]" captionTargetBox="[169,1400,300,1396]" captionTargetId="figure@5.[151,1436,265,1407]" captionTargetPageId="5" captionText="FIGURE 3. Tooth pattern on upper jaw. A. Bathycongrus bimaculatus sp. nov., NMMB-P7012, paratype. B. B. bleekeri, NMMB-P26214. C. B. castlei sp. nov., NMMB-P21740, paratype. D. B. guttulatus, NMMB-P7607. E. B. graciliceps sp. nov., holotype. F. B. macroporis, NMMB-P29128. Not to scale." httpUri="https://zenodo.org/record/1446566/files/figure.png" pageId="22" pageNumber="140">Fig. 3F</figureCitation>
). Intermaxillary with 5 or 6 transverse rows of teeth, those on outer 2 rows large, gradually smaller posteriorly, not exposed when mouth closed. No clear gap between intermaxillary and vomerine teeth. Vomerine teeth conical, forming a long patch, extending about half length of maxillary tooth band. Maxillary teeth in about 5 irregular rows, gradually narrowing to 1 or 2 rows posteriorly, outer teeth longest, inner teeth gradually smaller. Mandibular teeth similar in size and arrangement to those on maxilla.
</paragraph>
<paragraph id="8BD8790168011A17E5B38A73FAE901AA" blockId="22.[151,1436,151,1005]" pageId="22" pageNumber="140">
<emphasis id="B913A51368011A17E5B38A73FEF90162" bold="true" box="[206,342,583,608]" pageId="22" pageNumber="140">Coloration.</emphasis>
Light brown dorsally, paler ventrally. Dorsal and anal fin pale with broad black margin; posterior portion entirely black. Pectoral fin pale. Stomach, intestine, and peritoneum pale. Mouth cavity pale; gill chamber unpigmented on outer side, inner side with small, closely spaced melanophores forming a slight darkening.
</paragraph>
</subSubSection>
<subSubSection id="C37D2A8A68011A17E5BA8A80FB0801CF" box="[199,1191,692,717]" pageId="22" pageNumber="140" type="distribution">
<paragraph id="8BD8790168011A17E5BA8A80FB0801CF" blockId="22.[151,1436,151,1005]" box="[199,1191,692,717]" pageId="22" pageNumber="140">
<emphasis id="B913A51368011A17E5BA8A80FECF01CF" bold="true" box="[199,352,692,717]" pageId="22" pageNumber="140">Distribution.</emphasis>
Known from off Sokotra Is. in the western Indian Ocean and now
<collectingCountry id="F370399168011A17E1308A80FB0C01CF" box="[1101,1187,692,717]" name="Taiwan" pageId="22" pageNumber="140">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D2A8A68011A17E5BA8AE3FC9400EF" pageId="22" pageNumber="140" type="discussion">
<paragraph id="8BD8790168011A17E5BA8AE3FC9400EF" blockId="22.[151,1436,151,1005]" pageId="22" pageNumber="140">
<emphasis id="B913A51368011A17E5BA8AE3FE9401F2" bold="true" box="[199,315,727,752]" pageId="22" pageNumber="140">Remarks.</emphasis>
This species is placed in
<taxonomicName id="4C67028268011A17E7198AECFCAF01F2" authorityName="Ogilby" authorityYear="1898" box="[612,768,728,752]" class="Actinopterygii" family="Congridae" genus="Bathycongrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="22" pageNumber="140" phylum="Chordata" rank="genus">
<emphasis id="B913A51368011A17E7198AECFCAF01F2" box="[612,768,728,752]" italics="true" pageId="22" pageNumber="140">Bathycongrus</emphasis>
</taxonomicName>
provisionally. The lack of any notably enlarged teeth, and the long and broad vomerine tooth patch differ from the condition found in most species of
<taxonomicName id="4C67028268011A17E1D18AC8FAE70016" authorityName="Ogilby" authorityYear="1898" box="[1196,1352,764,788]" class="Actinopterygii" family="Congridae" genus="Bathycongrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="22" pageNumber="140" phylum="Chordata" rank="genus">
<emphasis id="B913A51368011A17E1D18AC8FAE70016" box="[1196,1352,764,788]" italics="true" pageId="22" pageNumber="140">Bathycongrus</emphasis>
</taxonomicName>
, which have a short patch with at least a few teeth enlarged. The elevated position of the posterior nostril, above the dorsal margin and behind the anterior margin of the eye, differs markedly from the other species, which have the nostril in front of the eye near mid-eye level.
<bibRefCitation id="EFF604F068011A17E75B8B53FD730082" box="[550,732,871,896]" pageId="22" pageNumber="140" refId="ref16194" refString="Kotthaus, A. (1968) Fische des Indischen Ozeans. A. Systematischer Teil. III. Ostariophysi und Apodes. Meteor Forschungsergebnisse, Reihe D, Biologie, 3, 14 - 56." type="journal article">Kotthaus (1968)</bibRefCitation>
placed it in the genus
<taxonomicName id="4C67028268011A17E6AA8B5CFB3A0082" authorityName="Roule" authorityYear="1915" box="[983,1173,872,896]" class="Actinopterygii" family="Congridae" genus="Pseudophichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="22" pageNumber="140" phylum="Chordata" rank="genus">
<emphasis id="B913A51368011A17E6AA8B5CFB3A0082" box="[983,1173,872,896]" italics="true" pageId="22" pageNumber="140">Pseudophichthys</emphasis>
</taxonomicName>
, but the
<typeStatus id="54DCC7A368011A17E18B8B5CFA890082" box="[1270,1318,872,896]" pageId="22" pageNumber="140">type</typeStatus>
species of that genus,
<taxonomicName id="4C67028268011A17E4658BB9FD8000A6" authority="(Lea, 1913)" baseAuthorityName="Lea" baseAuthorityYear="1913" box="[280,559,908,932]" class="Actinopterygii" family="Congridae" genus="Pseudophichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="22" pageNumber="140" phylum="Chordata" rank="species" species="splendens">
<emphasis id="B913A51368011A17E4658BB9FE0C00A6" box="[280,419,908,932]" italics="true" pageId="22" pageNumber="140">P. splendens</emphasis>
(Lea, 1913)
</taxonomicName>
, is more slender and has the posterior nostril in front of mid-eye. This species was known previously only from the
<typeStatus id="54DCC7A368011A17E7468B9BFD3200CA" box="[571,669,943,968]" pageId="22" pageNumber="140" type="holotype">holotype</typeStatus>
collected in the northwestern Indian Ocean. Its discovery in
<collectingCountry id="F370399168011A17E0378B9BFA3300CA" box="[1354,1436,943,968]" name="Taiwan" pageId="22" pageNumber="140">Taiwan</collectingCountry>
suggests that it may be present in intermediate areas as well.
</paragraph>
</subSubSection>
</treatment>
</document>