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<document id="FE84F309D8B46932A1C75F8308E556E5" ID-DOI="10.11646/zootaxa.2104.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5315428" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629623837903" checkinUser="felipe" docAuthor="Barnich, Ruth &amp; Fiege, Dieter" docDate="2009" docId="03A61979C216FFEEFF18FE8AFE831238" docLanguage="en" docName="zt02104p076.pdf" docOrigin="Zootaxa 2104 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2104.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Harmothoe mariannae Barnich &amp; Fiege 2009, n. sp." docType="treatment" docVersion="5" lastPageNumber="56" masterDocId="FF9F6101C223FFD9FF8FFD4EFFA41036" masterDocTitle="Revision of the genus Harmothoe Kinberg, 1856 (Polychaeta: Polynoidae) in the Northeast Atlantic" masterLastPageNumber="76" masterPageNumber="1" pageNumber="54" updateTime="1699085409460" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="05EA7DFACE1C1783D61C6166CA1338D1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="394097B8C06A3A1595754C2519E878BD">Revision of the genus Harmothoe Kinberg, 1856 (Polychaeta: Polynoidae) in the Northeast Atlantic</mods:title>
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<mods:namePart id="F96DD462A91A45FE996177C22A276602">Barnich, Ruth</mods:namePart>
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<mods:namePart id="E2C68A6EC53A24EC03E1FDBBDF09D5B3">Fiege, Dieter</mods:namePart>
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<mods:title id="2ECC3959F34ADBB2D28E9A716C28D3C4">Zootaxa</mods:title>
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<mods:date id="F992E973A41EDE15B0910942088A2F10">2009</mods:date>
<mods:detail id="52403984BED14C759249AA540360BFAE" type="pubDate">
<mods:number id="E734A3F8FE9CF2AE0E02F1A6FFCE9E65">2009-05-11</mods:number>
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<mods:detail id="98A8FD7F91A9C469FBEFE42F160F0EF8" type="volume">
<mods:number id="E3B59FEF6B5742795E07BD5ADEB15381">2104</mods:number>
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<mods:url id="0D43295B5CAE85FC357908F7CDC127DB">https://biotaxa.org/Zootaxa/article/view/zootaxa.2104.1.1</mods:url>
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<mods:classification id="00E46B09AB2C87DBD2B52F58E62B3126">journal article</mods:classification>
<mods:identifier id="16CCB4C34429FAB95223B4ECC827E34B" type="DOI">10.11646/zootaxa.2104.1.1</mods:identifier>
<mods:identifier id="DBAA19D8382E7A37AFE3F88777B8EF7C" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="853A69CE54E051744D5877B93ED834C5" type="Zenodo-Dep">5315428</mods:identifier>
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<treatment id="03A61979C216FFEEFF18FE8AFE831238" ID-DOI="http://doi.org/10.5281/zenodo.5323862" ID-Zenodo-Dep="5323862" LSID="urn:lsid:plazi:treatment:03A61979C216FFEEFF18FE8AFE831238" httpUri="http://treatment.plazi.org/id/03A61979C216FFEEFF18FE8AFE831238" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54">
<subSubSection id="C315FBE4C216FFECFF18FE8AFE5213EB" box="[151,502,963,990]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="8BB0A86FC216FFECFF18FE8AFE5213EB" blockId="53.[151,502,963,1024]" box="[151,502,963,990]" pageId="53" pageNumber="54">
<heading id="D0F81F03C216FFECFF18FE8AFE5213EB" bold="true" box="[151,502,963,990]" fontSize="11" level="1" pageId="53" pageNumber="54" reason="1">
<emphasis id="B97B747DC216FFECFF18FE8AFE5213EB" bold="true" box="[151,502,963,990]" pageId="53" pageNumber="54">
<taxonomicName id="4C0FD3ECC216FFECFF18FE8AFE0913E8" ID-CoL="3JRGD" authority="Barnich &amp; Fiege, 2009" authorityName="Barnich &amp; Fiege" authorityYear="2009" box="[151,429,964,990]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="53" pageNumber="54" phylum="Annelida" rank="species" species="mariannae" status="sp. nov.">
<emphasis id="B97B747DC216FFECFF18FE8AFE0913E8" bold="true" box="[151,429,964,990]" italics="true" pageId="53" pageNumber="54">Harmothoe mariannae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248C906C216FFECFE3BFE8DFE5213EB" box="[436,502,963,989]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C315FBE4C216FFEEFF18FEA8FE831238" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="description">
<paragraph id="8BB0A86FC216FFECFF18FEA8FEF21436" blockId="53.[151,502,963,1024]" box="[151,342,998,1024]" pageId="53" pageNumber="54">
(
<figureCitation id="1334B4EAC216FFECFF10FEA8FF571436" box="[159,243,998,1024]" captionStart="FIGURE 3" captionStartId="11.[151,257,1938,1962]" captionTargetBox="[264,1405,270,1885]" captionTargetId="figure-83@11.[151,1436,209,2008]" captionTargetPageId="11" captionText="FIGURE 3. Identification characters of Harmothoe impar, H. clavigera, H. rarispina, H. extenuata, H. fragilis, H. vesiculosa, H. abyssicola, and H. mariannae n. sp." figureDoi="http://doi.org/10.5281/zenodo.5315438" httpUri="https://zenodo.org/record/5315438/files/figure.png" pageId="53" pageNumber="54">Figs. 3</figureCitation>
;
<figureCitation id="1334B4EAC216FFECFE8FFEA8FEE91436" box="[256,333,998,1024]" captionStart="FIGURE 26" captionStartId="54.[151,257,1842,1866]" captionTargetBox="[184,1389,332,1800]" captionTargetId="figure-162@54.[151,1436,313,1813]" captionTargetPageId="54" captionText="FIGURE 26. Harmothoe mariannae n. sp. (holotype, SMF 17290). (A) anterior end; (B) second right elytron from segment 4; (C) detail of posterior margin of same; (D) left cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus not figured; (E) distal part of long notochaeta; (F) tip of same; (G) distal part of middle neurochaeta; (H) tip of same; (I) tip of upper neurochaeta; (J) tip of lower neurochaeta. (Scales: A, B = 1 mm; C = 250 µm; D = 500 µm; E, G = 100 µm; F, H–J = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361493" httpUri="https://zenodo.org/record/5361493/files/figure.png" pageId="53" pageNumber="54">26A–J</figureCitation>
)
</paragraph>
<paragraph id="8BB0A86FC216FFECFF18F97AFDD814AD" blockId="53.[151,1436,1076,2030]" pageId="53" pageNumber="54">
<emphasis id="B97B747DC216FFECFF18F97AFE9A147B" box="[151,318,1076,1101]" italics="true" pageId="53" pageNumber="54">Type material</emphasis>
.
<taxonomicName id="4C0FD3ECC216FFECFEC0F97AFE53147B" authority="Barnich &amp; Fiege, 2009" authorityName="Barnich &amp; Fiege" authorityYear="2009" box="[335,503,1076,1101]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="53" pageNumber="54" phylum="Annelida" rank="species" species="mariannae" status="sp. nov.">
<emphasis id="B97B747DC216FFECFEC0F97AFE53147B" box="[335,503,1076,1101]" italics="true" pageId="53" pageNumber="54">H. mariannae</emphasis>
</taxonomicName>
<materialsCitation id="3B67A232C216FFECFD8EF97AFDDC14AD" accessionNumber="SMF 17290" collectingDate="2006-05-10" collectorName="C. d'Udekem d'Acoz" country="Norway" elevation="251" latitude="69.872665" location="Varangerfjord" longLatPrecision="9" longitude="30.134834" municipality="Bokfjord Kjelmsoy" pageId="53" pageNumber="54" specimenCount="1" typeStatus="holotype">
<taxonomicNameLabel id="A248C906C216FFECFD8EF97AFDE31478" box="[513,583,1076,1102]" pageId="53" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
:
<typeStatus id="54B416CDC216FFECFDDBF97AFD1A1478" box="[596,702,1076,1102]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
(cs in three fragments), SMF 17290, &quot;Johan Ruud&quot; St. 482,
<geoCoordinate id="EE3BCEA8C216FFECFF18F914FE851442" box="[151,289,1114,1140]" degrees="69" direction="north" minutes="52.36" orientation="latitude" pageId="53" pageNumber="54" precision="9" value="69.872665">69°52.36’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C216FFECFEA7F914FE151442" box="[296,433,1114,1140]" degrees="30" direction="east" minutes="08.09" orientation="longitude" pageId="53" pageNumber="54" precision="9" value="30.134834">30°08.09’E</geoCoordinate>
, N
<collectingCountry id="F318E8FFC216FFECFE55F914FD9E1442" box="[474,570,1114,1140]" name="Norway" pageId="53" pageNumber="54">Norway</collectingCountry>
,
<location id="8ED0FEB4C216FFECFDCAF914FD4F1442" LSID="urn:lsid:plazi:treatment:03A61979C216FFEEFF18FE8AFE831238:8ED0FEB4C216FFECFDCAF914FD4F1442" box="[581,747,1114,1140]" country="Norway" latitude="69.872665" longLatPrecision="9" longitude="30.134834" municipality="Bokfjord Kjelmsoy" name="Varangerfjord" pageId="53" pageNumber="54">Varangerfjord</location>
,
<collectingMunicipality id="6BD43215C216FFECFD79F914FC7E1442" box="[758,986,1114,1140]" pageId="53" pageNumber="54">Bøkfjord Kjelmsøy</collectingMunicipality>
lykt,
<date id="FFB18EAFC216FFECFB92F914FB181442" box="[1053,1212,1114,1140]" pageId="53" pageNumber="54" value="2006-05-10">
<collectingDate id="EFF57747C216FFECFB92F914FB181442" box="[1053,1212,1114,1140]" pageId="53" pageNumber="54" value="2006-05-10">10 May 2006</collectingDate>
</date>
, epibenthic sledge,
<quantity id="4CF7058AC216FFECFF18F9CFFF4014AD" box="[151,228,1153,1179]" metricMagnitude="2" metricUnit="m" metricValue="2.51" pageId="53" pageNumber="54" unit="m" value="251.0">
<elevation id="00224F5CC216FFECFF18F9CFFF4014AD" box="[151,228,1153,1179]" metricMagnitude="2" metricUnit="m" metricValue="2.51" pageId="53" pageNumber="54" unit="m" value="251.0">251 m</elevation>
</quantity>
, leg. &amp; ded.
<collectorName id="26FACDB9C216FFECFEF6F9CFFDDC14AD" box="[377,632,1153,1179]" pageId="53" pageNumber="54">C. d’Udekem d’Acoz</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB0A86FC216FFECFF4AF9E6FDD114DE" blockId="53.[151,1436,1076,2030]" pageId="53" pageNumber="54">
<materialsCitation id="3B67A232C216FFECFF4AF9E6FDD514DE" accessionNumber="SMF 17293" collectingDate="1999-05-30" collectorName="A. Sikorski" elevation="280" latitude="61.381344" longLatPrecision="1" longitude="2.1103303" pageId="53" pageNumber="54" specimenCount="1" typeStatus="paratype">
<typeStatus id="54B416CDC216FFECFF4AF9E6FE9514F4" box="[197,305,1192,1218]" pageId="53" pageNumber="54" type="paratype">Paratype</typeStatus>
(cs in two fragments), SMF 17293, Vigdis 1999, St. 17-1,
<date id="FFB18EAFC216FFECFB94F9E6FB6014F4" box="[1051,1220,1192,1218]" pageId="53" pageNumber="54" value="1999-05-30">
<collectingDate id="EFF57747C216FFECFB94F9E6FB6014F4" box="[1051,1220,1192,1218]" pageId="53" pageNumber="54" value="1999-05-30">30 May 1999</collectingDate>
</date>
,
<geoCoordinate id="EE3BCEA8C216FFECFB5CF9E6FA3314F4" box="[1235,1431,1192,1218]" degrees="61" direction="north" minutes="22" orientation="latitude" pageId="53" pageNumber="54" precision="1" seconds="52.84" value="61.381344">61°22'52.84'' N</geoCoordinate>
,
<geoCoordinate id="EE3BCEA8C216FFECFF18F980FEEE14DE" box="[151,330,1230,1256]" degrees="02" direction="east" minutes="06" orientation="longitude" pageId="53" pageNumber="54" precision="1" seconds="37.19" value="2.1103303">02°06'37.19'' E</geoCoordinate>
,
<quantity id="4CF7058AC216FFECFED8F980FE0714DE" box="[343,419,1230,1256]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="53" pageNumber="54" unit="m" value="280.0">
<elevation id="00224F5CC216FFECFED8F980FE0714DE" box="[343,419,1230,1256]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="53" pageNumber="54" unit="m" value="280.0">280 m</elevation>
</quantity>
, ded.
<collectorName id="26FACDB9C216FFECFE67F980FDD514DE" box="[488,625,1230,1256]" pageId="53" pageNumber="54">A. Sikorski</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB0A86FC216FFECFF4AF9BBFA38156A" blockId="53.[151,1436,1076,2030]" pageId="53" pageNumber="54">
<emphasis id="B97B747DC216FFECFF4AF9BBFE991538" box="[197,317,1269,1294]" italics="true" pageId="53" pageNumber="54">Diagnosis</emphasis>
. Anterior pair of eyes dorsolateral at widest part of prostomium. Elytral margin and adjacent surface papillate, papillae long at outer lateral margin, becoming shorter towards posterior margin; surface with conical microtubercles; macrotubercles conical to low, globose, scattered in a row near posterior margin.
</paragraph>
<paragraph id="8BB0A86FC216FFECFF4AF827FDE315B5" blockId="53.[151,1436,1076,2030]" box="[197,583,1385,1411]" pageId="53" pageNumber="54">
<emphasis id="B97B747DC216FFECFF4AF827FEEB15B4" box="[197,335,1385,1410]" italics="true" pageId="53" pageNumber="54">Description</emphasis>
(based on
<typeStatus id="54B416CDC216FFECFE40F827FD9F15B5" box="[463,571,1385,1411]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BB0A86FC216FFECFF4AF8DEFC0C1628" blockId="53.[151,1436,1076,2030]" pageId="53" pageNumber="54">
Body with 36 segments. At anterior end (
<figureCitation id="1334B4EAC216FFECFD35F8DEFC8E159C" box="[698,810,1424,1450]" captionStart="FIGURE 26" captionStartId="54.[151,257,1842,1866]" captionTargetBox="[184,1389,332,1800]" captionTargetId="figure-162@54.[151,1436,313,1813]" captionTargetPageId="54" captionText="FIGURE 26. Harmothoe mariannae n. sp. (holotype, SMF 17290). (A) anterior end; (B) second right elytron from segment 4; (C) detail of posterior margin of same; (D) left cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus not figured; (E) distal part of long notochaeta; (F) tip of same; (G) distal part of middle neurochaeta; (H) tip of same; (I) tip of upper neurochaeta; (J) tip of lower neurochaeta. (Scales: A, B = 1 mm; C = 250 µm; D = 500 µm; E, G = 100 µm; F, H–J = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361493" httpUri="https://zenodo.org/record/5361493/files/figure.png" pageId="53" pageNumber="54">Fig. 26A</figureCitation>
), prostomium bilobed, with distinct cephalic peaks; ceratophore of median antenna in anterior notch, lateral antennae inserted ventrally, styles of antennae papillate, tapering; anterior pair of eyes situated dorsolaterally at widest part of prostomium, posterior pair dorsally near hind margin of prostomium; palps papillate, tapering.
</paragraph>
<paragraph id="8BB0A86FC216FFECFF4AFB64FAF916A4" blockId="53.[151,1436,1076,2030]" pageId="53" pageNumber="54">Tentaculophores inserted laterally to prostomium, each with two notochaetae and a dorsal and ventral tentacular cirrus, styles of cirri papillate, tapering. Second segment with first pair of elytra, biramous parapodia, and long buccal cirri. Following segments with tapering, short, slightly papillate ventral cirri.</paragraph>
<paragraph id="8BB0A86FC216FFECFF4AFBD0FAF21765" blockId="53.[151,1436,1076,2030]" pageId="53" pageNumber="54">
Fifteen pairs of elytra, covering dorsum, on segments 2, 4, 5, 7, then on every second segment to 23, 26, 29, 32, last four segments cirrigerous; elytral margin and adjacent surface papillate, papillae long at outer lateral margin, becoming shorter towards posterior margin; surface with conical microtubercles; macrotubercles conical to low, globose, scattered in a row near posterior margin. (
<figureCitation id="1334B4EAC216FFECFBFBFA5CFB5B171A" box="[1140,1279,1810,1836]" captionStart="FIGURE 26" captionStartId="54.[151,257,1842,1866]" captionTargetBox="[184,1389,332,1800]" captionTargetId="figure-162@54.[151,1436,313,1813]" captionTargetPageId="54" captionText="FIGURE 26. Harmothoe mariannae n. sp. (holotype, SMF 17290). (A) anterior end; (B) second right elytron from segment 4; (C) detail of posterior margin of same; (D) left cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus not figured; (E) distal part of long notochaeta; (F) tip of same; (G) distal part of middle neurochaeta; (H) tip of same; (I) tip of upper neurochaeta; (J) tip of lower neurochaeta. (Scales: A, B = 1 mm; C = 250 µm; D = 500 µm; E, G = 100 µm; F, H–J = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361493" httpUri="https://zenodo.org/record/5361493/files/figure.png" pageId="53" pageNumber="54">Fig. 26B,C</figureCitation>
). Cirrigerous segments with distinct dorsal tubercles; dorsal cirri with cylindrical cirrophore, style papillate, tapering.
</paragraph>
<paragraph id="8BB0A86FC216FFEFFF4AFA2EFA9D1084" blockId="53.[151,1436,1076,2030]" lastBlockId="54.[151,1435,152,255]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
Parapodia biramous; notopodia with elongate acicular lobe; neuropodia with elongate prechaetal acicular lobe with long, digitiform supra-acicular process; neuropodial postchaetal lobe shorter than prechaetal lobe, rounded; tips of noto- and neuroacicula penetrating epidermis (
<figureCitation id="1334B4EAC216FFECFC44FAE3FBE017F1" box="[971,1092,1965,1991]" captionStart="FIGURE 26" captionStartId="54.[151,257,1842,1866]" captionTargetBox="[184,1389,332,1800]" captionTargetId="figure-162@54.[151,1436,313,1813]" captionTargetPageId="54" captionText="FIGURE 26. Harmothoe mariannae n. sp. (holotype, SMF 17290). (A) anterior end; (B) second right elytron from segment 4; (C) detail of posterior margin of same; (D) left cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus not figured; (E) distal part of long notochaeta; (F) tip of same; (G) distal part of middle neurochaeta; (H) tip of same; (I) tip of upper neurochaeta; (J) tip of lower neurochaeta. (Scales: A, B = 1 mm; C = 250 µm; D = 500 µm; E, G = 100 µm; F, H–J = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361493" httpUri="https://zenodo.org/record/5361493/files/figure.png" pageId="53" pageNumber="54">Fig. 26D</figureCitation>
). Notochaetae stouter than neurochaetae, with distinct rows of spines and blunt tip (
<figureCitation id="1334B4EAC216FFECFCDBFA9AFC7817D8" box="[852,988,2004,2030]" captionStart="FIGURE 26" captionStartId="54.[151,257,1842,1866]" captionTargetBox="[184,1389,332,1800]" captionTargetId="figure-162@54.[151,1436,313,1813]" captionTargetPageId="54" captionText="FIGURE 26. Harmothoe mariannae n. sp. (holotype, SMF 17290). (A) anterior end; (B) second right elytron from segment 4; (C) detail of posterior margin of same; (D) left cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus not figured; (E) distal part of long notochaeta; (F) tip of same; (G) distal part of middle neurochaeta; (H) tip of same; (I) tip of upper neurochaeta; (J) tip of lower neurochaeta. (Scales: A, B = 1 mm; C = 250 µm; D = 500 µm; E, G = 100 µm; F, H–J = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361493" httpUri="https://zenodo.org/record/5361493/files/figure.png" pageId="53" pageNumber="54">Fig. 26E,F</figureCitation>
); neurochaetae with distinct rows of spines, upper group bidentate with short, stout secondary tooth, lower group unidentate (
<figureCitation id="1334B4EAC215FFEFFB2AFDD6FA891084" box="[1189,1325,152,178]" captionStart="FIGURE 26" captionStartId="54.[151,257,1842,1866]" captionTargetBox="[184,1389,332,1800]" captionTargetId="figure-162@54.[151,1436,313,1813]" captionTargetPageId="54" captionText="FIGURE 26. Harmothoe mariannae n. sp. (holotype, SMF 17290). (A) anterior end; (B) second right elytron from segment 4; (C) detail of posterior margin of same; (D) left cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus not figured; (E) distal part of long notochaeta; (F) tip of same; (G) distal part of middle neurochaeta; (H) tip of same; (I) tip of upper neurochaeta; (J) tip of lower neurochaeta. (Scales: A, B = 1 mm; C = 250 µm; D = 500 µm; E, G = 100 µm; F, H–J = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361493" httpUri="https://zenodo.org/record/5361493/files/figure.png" pageId="54" pageNumber="55">Fig. 26G–J</figureCitation>
).
</paragraph>
<paragraph id="8BB0A86FC215FFEFFF4AFDF1FCB410C9" blockId="54.[151,1435,152,255]" pageId="54" pageNumber="55">
<emphasis id="B97B747DC215FFEFFF4AFDF1FED410EE" box="[197,368,191,216]" italics="true" pageId="54" pageNumber="55">Measurements</emphasis>
.
<typeStatus id="54B416CDC215FFEFFE0FFDF0FE4A10EE" box="[384,494,190,216]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
(cs), SMF 17290 (
<figureCitation id="1334B4EAC215FFEFFD5FFDF0FCFF10EE" box="[720,859,190,216]" captionStart="FIGURE 26" captionStartId="54.[151,257,1842,1866]" captionTargetBox="[184,1389,332,1800]" captionTargetId="figure-162@54.[151,1436,313,1813]" captionTargetPageId="54" captionText="FIGURE 26. Harmothoe mariannae n. sp. (holotype, SMF 17290). (A) anterior end; (B) second right elytron from segment 4; (C) detail of posterior margin of same; (D) left cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus not figured; (E) distal part of long notochaeta; (F) tip of same; (G) distal part of middle neurochaeta; (H) tip of same; (I) tip of upper neurochaeta; (J) tip of lower neurochaeta. (Scales: A, B = 1 mm; C = 250 µm; D = 500 µm; E, G = 100 µm; F, H–J = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361493" httpUri="https://zenodo.org/record/5361493/files/figure.png" pageId="54" pageNumber="55">Fig. 26A–J</figureCitation>
): L
<quantity id="4CF7058AC215FFEFFC05FDF0FC4510EE" box="[906,993,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="mm" value="16.0">16 mm</quantity>
, W
<quantity id="4CF7058AC215FFEFFB9EFDF0FBF210EE" box="[1041,1110,190,216]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="54" pageNumber="55" unit="mm" value="4.0">4 mm</quantity>
for 36 segments.
<typeStatus id="54B416CDC215FFEFFAA2FDF0FA3310EE" box="[1325,1431,190,216]" pageId="54" pageNumber="55" type="paratype">Paratype</typeStatus>
, SMF 17293 (cs): L
<quantity id="4CF7058AC215FFEFFEF1FDABFE7610C9" box="[382,466,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="54" pageNumber="55" unit="mm" value="13.0">13 mm</quantity>
, W
<quantity id="4CF7058AC215FFEFFD8DFDABFDE210C9" box="[514,582,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="54" pageNumber="55" unit="mm" value="4.0">4 mm</quantity>
for 33 segments.
</paragraph>
<caption id="DF70F8E7C215FFEFFF18FA7CFE5817E2" ID-DOI="http://doi.org/10.5281/zenodo.5361493" ID-Zenodo-Dep="5361493" httpUri="https://zenodo.org/record/5361493/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,257,1842,1866]" targetBox="[184,1389,332,1800]" targetPageId="54">
<paragraph id="8BB0A86FC215FFEFFF18FA7CFE5817E2" blockId="54.[151,1436,1842,2004]" pageId="54" pageNumber="55">
<emphasis id="B97B747DC215FFEFFF18FA7CFE8F177C" bold="true" box="[151,299,1842,1866]" pageId="54" pageNumber="55">FIGURE 26.</emphasis>
<taxonomicName id="4C0FD3ECC215FFEFFEBCFA7CFD8B177F" authority="Barnich &amp; Fiege, 2009" authorityName="Barnich &amp; Fiege" authorityYear="2009" box="[307,559,1842,1865]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="54" pageNumber="55" phylum="Annelida" rank="species" species="mariannae" status="sp. nov.">
<emphasis id="B97B747DC215FFEFFEBCFA7CFD8B177F" box="[307,559,1842,1865]" italics="true" pageId="54" pageNumber="55">Harmothoe mariannae</emphasis>
</taxonomicName>
<emphasis id="B97B747DC215FFEFFDB9FA7CFDD1177C" bold="true" box="[566,629,1842,1866]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="A248C906C215FFEFFDB9FA7CFDD1177C" box="[566,629,1842,1866]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype, SMF 17290). (A) anterior end; (B) second right elytron from segment 4; (C) detail of posterior margin of same; (D) left cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus not figured; (E) distal part of long notochaeta; (F) tip of same; (G) distal part of middle neurochaeta; (H) tip of same; (I) tip of upper neurochaeta; (J) tip of lower neurochaeta. (Scales: A, B = 1 mm; C = 250 µm; D = 500 µm; E, G = 100 µm; F, H–J = 50 µm).
</paragraph>
</caption>
<paragraph id="8BB0A86FC214FFEEFF4AFDD6FEFD1145" blockId="55.[151,1436,151,526]" pageId="55" pageNumber="56">
<emphasis id="B97B747DC214FFEEFF4AFDD6FDF61087" box="[197,594,152,177]" italics="true" pageId="55" pageNumber="56">
Remarks.
<taxonomicName id="4C0FD3ECC214FFEEFEB1FDD6FDF61087" authority="Barnich &amp; Fiege, 2009" authorityName="Barnich &amp; Fiege" authorityYear="2009" box="[318,594,152,177]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="55" pageNumber="56" phylum="Annelida" rank="species" species="mariannae" status="sp. nov.">Harmothoe mariannae</taxonomicName>
</emphasis>
<emphasis id="B97B747DC214FFEEFDD3FDD9FD051087" bold="true" box="[604,673,151,177]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A248C906C214FFEEFDD3FDD9FD051087" box="[604,673,151,177]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C0FD3ECC214FFEEFCCCFDD6FB171084" authority="Ditlevsen, 1917" authorityName="Ditlevsen" authorityYear="1917" box="[835,1203,152,178]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="55" pageNumber="56" phylum="Annelida" rank="species" species="vesiculosa">
<emphasis id="B97B747DC214FFEEFCCCFDD6FC431087" box="[835,999,152,177]" italics="true" pageId="55" pageNumber="56">H. vesiculosa</emphasis>
<bibRefCitation id="EF9ED59EC214FFEEFC7FFDD6FB171084" author="Ditlevsen, H." box="[1008,1203,152,178]" pageId="55" pageNumber="56" pagination="1 - 71" refId="ref37221" refString="Ditlevsen, H. (1917) Annelids. The Danish Ingolf Expedition, 4 (4), 1 - 71." type="journal article" year="1917">Ditlevsen, 1917</bibRefCitation>
</taxonomicName>
, but in this species elytra show shorter marginal papillae and macrotubercles are smaller and arranged in a dense row near the posterior margin. Confusion might also be possible with
<taxonomicName id="4C0FD3ECC214FFEEFCDBFDABFB0F10C9" authority="Moore, 1902" authorityName="Moore" authorityYear="1902" box="[852,1195,229,255]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="55" pageNumber="56" phylum="Annelida" rank="species" species="multisetosa">
<emphasis id="B97B747DC214FFEEFCDBFDABFBA110C8" box="[852,1029,229,254]" italics="true" pageId="55" pageNumber="56">H. multisetosa</emphasis>
<bibRefCitation id="EF9ED59EC214FFEEFB81FDABFB0F10C9" author="Moore, J. P." box="[1038,1195,229,255]" pageId="55" pageNumber="56" pagination="258 - 278" refId="ref39353" refString="Moore, J. P. (1902) Description of some new Polynoidae, with a list of other Polychaeta from North Greenland waters. Proceedings of the Academy of Natural Sciences Philadelphia, 258 - 278." type="book chapter" year="1902">Moore, 1902</bibRefCitation>
</taxonomicName>
from the Northeast Pacific. Here elytra have shorter marginal papillae, macrotubercles are not only present near the posterior margin, but also scattered on the surface, and microtubercles are conical to pointed (cf.
<bibRefCitation id="EF9ED59EC214FFEEFB5DFC7CFAC3117A" author="Moore, J. P." box="[1234,1383,306,332]" pageId="55" pageNumber="56" pagination="258 - 278" refId="ref39353" refString="Moore, J. P. (1902) Description of some new Polynoidae, with a list of other Polychaeta from North Greenland waters. Proceedings of the Academy of Natural Sciences Philadelphia, 258 - 278." type="book chapter" year="1902">Moore 1902</bibRefCitation>
and
<bibRefCitation id="EF9ED59EC214FFEEFF18FC17FEE81145" author="Pettibone, M. H." box="[151,332,345,371]" pageId="55" pageNumber="56" refId="ref39612" refString="Pettibone, M. H. (1953) Some scale-bearing polychaetes of Puget Sound and adjacent waters. University of Washington Press, Seattle, 89 pp." type="book" year="1953">Pettibone 1953</bibRefCitation>
).
</paragraph>
<paragraph id="8BB0A86FC214FFEEFF4AFCCEFB2F11AC" blockId="55.[151,1436,151,526]" box="[197,1163,384,410]" pageId="55" pageNumber="56">
<emphasis id="B97B747DC214FFEEFF4AFCCEFEF811AF" box="[197,348,384,409]" italics="true" pageId="55" pageNumber="56">Distribution.</emphasis>
Northeast Atlantic, from northern
<collectingCountry id="F318E8FFC214FFEEFD7CFCCEFCF611AC" box="[755,850,384,410]" name="Norway" pageId="55" pageNumber="56">Norway</collectingCountry>
to the northern North Sea.
</paragraph>
<paragraph id="8BB0A86FC214FFEEFF4AFCE9FDD411F6" blockId="55.[151,1436,151,526]" box="[197,624,422,448]" pageId="55" pageNumber="56">
<emphasis id="B97B747DC214FFEEFF4AFCE9FEBA11F6" box="[197,286,423,448]" italics="true" pageId="55" pageNumber="56">Habitat</emphasis>
. Unknown, in
<quantity id="4CF7058AC214FFEEFE42FCE8FDC811F6" box="[461,620,422,448]" metricMagnitude="2" metricUnit="m" metricValue="2.655" metricValueMax="2.8" metricValueMin="2.51" pageId="55" pageNumber="56" unit="m" value="265.5" valueMax="280.0" valueMin="251.0">251 to 280 m</quantity>
.
</paragraph>
<paragraph id="8BB0A86FC214FFEEFF4AFC83FE831238" blockId="55.[151,1436,151,526]" pageId="55" pageNumber="56">
<emphasis id="B97B747DC214FFEEFF4AFC83FEE411D0" box="[197,320,461,486]" italics="true" pageId="55" pageNumber="56">Etymology</emphasis>
. The species is named after Marianne Barnich-Brimaire for her continuous support and interest in our work.
</paragraph>
</subSubSection>
</treatment>
</document>
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