311 lines
35 KiB
XML
311 lines
35 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3664837" ID-GBIF-Dataset="dd032848-7430-45d0-85d9-e528f25c83bb" ID-GBIF-Taxon="162054112" ID-Zenodo-Dep="3664837" LSID="urn:lsid:zoobank.org:act:D91488C6-3BEB-4A21-9F0E-3101BB9CCA85" checkinTime="1581022300054" checkinUser="plazi" docAuthor="Perrichot, Vincent, Wang, Bo & Barden, Phillip" docDate="2020" docId="03A3850A9F5BFFA7FCC9F8B90F4C7FAB" docLanguage="en" docName="createceousResearch.109.104.381.pdf" docOrigin="Cretaceous Research 109" docStyle="DocumentStyle{}" docTitle="Linguamyrmex brevicornis Perrichot & Wang & Barden 2020, sp. nov." docType="treatment" docVersion="5" lastPageId="15" lastPageNumber="16" masterDocId="FF9AFD729F57FFA8FFF6FFD90E4F796E" masterDocTitle="New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar" masterLastPageNumber="18" masterPageNumber="1" pageId="12" pageNumber="13" updateTime="1643457689448" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perrichot, Vincent</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wang, Bo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Barden, Phillip</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Cretaceous Research</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-01-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>109</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>18</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1016/j.cretres.2020.104381</mods:identifier>
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<mods:identifier type="GBIF-Dataset">dd032848-7430-45d0-85d9-e528f25c83bb</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3653072</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3664837" ID-GBIF-Taxon="162054112" ID-Zenodo-Dep="3664837" LSID="urn:lsid:zoobank.org:act:D91488C6-3BEB-4A21-9F0E-3101BB9CCA85" httpUri="http://treatment.plazi.org/id/03A3850A9F5BFFA7FCC9F8B90F4C7FAB" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
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<subSubSection pageId="12" pageNumber="13" type="nomenclature">
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||
<paragraph blockId="12.[831,1375,1888,1965]" box="[831,1192,1888,1909]" pageId="12" pageNumber="13">
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<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[831,1108,1888,1908]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brevicornis" status="sp. nov.">
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<emphasis bold="true" box="[831,1108,1888,1908]" italics="true" pageId="12" pageNumber="13">
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Linguamyrmex
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<taxonomicName box="[992,1108,1888,1908]" pageId="12" pageNumber="13" rank="species" species="brevicornis" status="sp. nov.">brevicornis</taxonomicName>
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</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel box="[1115,1192,1890,1909]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph blockId="12.[831,1375,1888,1965]" pageId="12" pageNumber="13">
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||
<uri pageId="12" pageNumber="13">
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||
urn:lsid:zoobank.org:act:
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||
<uuid pageId="12" pageNumber="13">D91488C6-3BEB-4A21-9F0E- 3101BB9CCA85</uuid>
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||
</uri>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[88,220,178,198]" pageId="13" pageNumber="14" type="description">
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<paragraph blockId="13.[88,220,178,198]" box="[88,220,178,198]" pageId="13" pageNumber="14">
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<figureCitation box="[88,195,178,198]" captionStart="Fig" captionStartId="12.[113,138,1380,1394]" captionTargetBox="[202,1412,178,1351]" captionTargetId="figure@12.[202,1412,178,1351]" captionTargetPageId="12" captionText="Fig. 6. Representative females of Protoceratomyrmex revelatus gen. et sp. nov. (AeB) and Linguamyrmex brevicornis gen. et sp. nov. (CeE). Habitus (A) and head (B) of worker holotype NIGP172002.Habitus of worker holotype NIGP172001 (C), head of gyne specimen LA01 in dorsal view (D), head of worker specimen RM4 (E) in lateral view. Abbreviations: cp, clypeal process; ft, frontal triangle; ts, trigger setae. Scale bars: 1 mm (A, C), 0.25 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.3653084" httpUri="https://zenodo.org/record/3653084/files/figure.png" pageId="13" pageNumber="14">Figs. 6CeE</figureCitation>
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,
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||
<figureCitation box="[205,216,179,198]" captionStart="Fig" captionStartId="13.[88,113,1945,1959]" captionTargetBox="[176,1386,890,1916]" captionTargetId="figure@13.[176,1386,890,1916]" captionTargetPageId="13" captionText="Fig. 7. Overview and details of females of Linguamyrmex brevicornis gen. et sp. nov. Gyne specimen LA01 (AeB), worker specimen RM4 (C), worker holotype NIGP172001 (D). Habitus (A, C), head and mesosoma in dorsal view (B), head in ventrolateral view (D). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3653086" httpUri="https://zenodo.org/record/3653086/files/figure.png" pageId="13" pageNumber="14">7</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="etymology">
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<paragraph blockId="13.[88,758,219,826]" pageId="13" pageNumber="14">
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<emphasis box="[88,189,219,239]" italics="true" pageId="13" pageNumber="14">Etymology</emphasis>
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. The specific epithet is a combination of
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<emphasis box="[635,684,219,239]" italics="true" pageId="13" pageNumber="14">brevi</emphasis>
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(Latin, meaning ‘short’) and
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<emphasis box="[308,373,247,267]" italics="true" pageId="13" pageNumber="14">cornus</emphasis>
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(Latin, meaning ‘horn’), and refers to the smaller clypeal horn compared to the
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<typeStatus box="[514,559,277,296]" pageId="13" pageNumber="14">type</typeStatus>
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species
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<taxonomicName box="[644,712,275,295]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vladi">
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<emphasis box="[644,712,275,295]" italics="true" pageId="13" pageNumber="14">
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L.
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<taxonomicName box="[665,712,275,295]" pageId="13" pageNumber="14" rank="species" species="vladi">vladi</taxonomicName>
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</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="materials_examined">
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<paragraph blockId="13.[88,758,219,826]" box="[88,537,303,324]" pageId="13" pageNumber="14">
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<materialsCitation ID-GBIF-Occurrence="2564234958" box="[88,537,303,324]" collectionCode="NIGP" pageId="13" pageNumber="14" specimenCode="NIGP172001" specimenCount="1" typeStatus="holotype">
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<emphasis box="[88,175,303,323]" italics="true" pageId="13" pageNumber="14">
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<typeStatus box="[88,175,303,323]" pageId="13" pageNumber="14">Holotype</typeStatus>
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</emphasis>
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.
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<specimenCode box="[186,310,304,323]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="13" pageNumber="14">NIGP172001</specimenCode>
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, worker (
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<figureCitation box="[407,485,304,323]" captionStart="Fig" captionStartId="12.[113,138,1380,1394]" captionTargetBox="[202,1412,178,1351]" captionTargetId="figure@12.[202,1412,178,1351]" captionTargetPageId="12" captionText="Fig. 6. Representative females of Protoceratomyrmex revelatus gen. et sp. nov. (AeB) and Linguamyrmex brevicornis gen. et sp. nov. (CeE). Habitus (A) and head (B) of worker holotype NIGP172002.Habitus of worker holotype NIGP172001 (C), head of gyne specimen LA01 in dorsal view (D), head of worker specimen RM4 (E) in lateral view. Abbreviations: cp, clypeal process; ft, frontal triangle; ts, trigger setae. Scale bars: 1 mm (A, C), 0.25 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.3653084" httpUri="https://zenodo.org/record/3653084/files/figure.png" pageId="13" pageNumber="14">Figs. 6C</figureCitation>
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,
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<figureCitation box="[495,527,304,324]" captionStart="Fig" captionStartId="13.[88,113,1945,1959]" captionTargetBox="[176,1386,890,1916]" captionTargetId="figure@13.[176,1386,890,1916]" captionTargetPageId="13" captionText="Fig. 7. Overview and details of females of Linguamyrmex brevicornis gen. et sp. nov. Gyne specimen LA01 (AeB), worker specimen RM4 (C), worker holotype NIGP172001 (D). Habitus (A, C), head and mesosoma in dorsal view (B), head in ventrolateral view (D). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3653086" httpUri="https://zenodo.org/record/3653086/files/figure.png" pageId="13" pageNumber="14">7D</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="13.[88,758,219,826]" pageId="13" pageNumber="14">
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<emphasis box="[88,292,331,351]" italics="true" pageId="13" pageNumber="14">Additional specimens</emphasis>
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. LA01, alate female (
|
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<figureCitation box="[497,577,332,351]" captionStart="Fig" captionStartId="12.[113,138,1380,1394]" captionTargetBox="[202,1412,178,1351]" captionTargetId="figure@12.[202,1412,178,1351]" captionTargetPageId="12" captionText="Fig. 6. Representative females of Protoceratomyrmex revelatus gen. et sp. nov. (AeB) and Linguamyrmex brevicornis gen. et sp. nov. (CeE). Habitus (A) and head (B) of worker holotype NIGP172002.Habitus of worker holotype NIGP172001 (C), head of gyne specimen LA01 in dorsal view (D), head of worker specimen RM4 (E) in lateral view. Abbreviations: cp, clypeal process; ft, frontal triangle; ts, trigger setae. Scale bars: 1 mm (A, C), 0.25 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.3653084" httpUri="https://zenodo.org/record/3653084/files/figure.png" pageId="13" pageNumber="14">Figs. 6D</figureCitation>
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,
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<figureCitation box="[586,646,332,351]" captionStart="Fig" captionStartId="13.[88,113,1945,1959]" captionTargetBox="[176,1386,890,1916]" captionTargetId="figure@13.[176,1386,890,1916]" captionTargetPageId="13" captionText="Fig. 7. Overview and details of females of Linguamyrmex brevicornis gen. et sp. nov. Gyne specimen LA01 (AeB), worker specimen RM4 (C), worker holotype NIGP172001 (D). Habitus (A, C), head and mesosoma in dorsal view (B), head in ventrolateral view (D). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3653086" httpUri="https://zenodo.org/record/3653086/files/figure.png" pageId="13" pageNumber="14">7AeB</figureCitation>
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||
), and
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<collectionCode box="[705,742,332,351]" country="Canada" name="McGill University, Redpath Museum" pageId="13" pageNumber="14" type="Museum">RM</collectionCode>
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4, worker (
|
||
<figureCitation box="[175,252,360,379]" captionStart="Fig" captionStartId="12.[113,138,1380,1394]" captionTargetBox="[202,1412,178,1351]" captionTargetId="figure@12.[202,1412,178,1351]" captionTargetPageId="12" captionText="Fig. 6. Representative females of Protoceratomyrmex revelatus gen. et sp. nov. (AeB) and Linguamyrmex brevicornis gen. et sp. nov. (CeE). Habitus (A) and head (B) of worker holotype NIGP172002.Habitus of worker holotype NIGP172001 (C), head of gyne specimen LA01 in dorsal view (D), head of worker specimen RM4 (E) in lateral view. Abbreviations: cp, clypeal process; ft, frontal triangle; ts, trigger setae. Scale bars: 1 mm (A, C), 0.25 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.3653084" httpUri="https://zenodo.org/record/3653084/files/figure.png" pageId="13" pageNumber="14">Figs. 6E</figureCitation>
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,
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<figureCitation box="[262,291,360,379]" captionStart="Fig" captionStartId="13.[88,113,1945,1959]" captionTargetBox="[176,1386,890,1916]" captionTargetId="figure@13.[176,1386,890,1916]" captionTargetPageId="13" captionText="Fig. 7. Overview and details of females of Linguamyrmex brevicornis gen. et sp. nov. Gyne specimen LA01 (AeB), worker specimen RM4 (C), worker holotype NIGP172001 (D). Habitus (A, C), head and mesosoma in dorsal view (B), head in ventrolateral view (D). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3653086" httpUri="https://zenodo.org/record/3653086/files/figure.png" pageId="13" pageNumber="14">7C</figureCitation>
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).
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</paragraph>
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<paragraph blockId="13.[88,758,219,826]" pageId="13" pageNumber="14">
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Horizon
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<emphasis box="[176,291,387,407]" italics="true" pageId="13" pageNumber="14">and locality</emphasis>
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. Upper Cretaceous, upper Albianelower Cenomanian (ca. 99 Ma); in amber from the Hukawng Valley,
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<collectingRegion country="Myanmar" name="Kachin" pageId="13" pageNumber="14">Kachin State</collectingRegion>
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,
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<collectingCountry box="[149,247,444,463]" name="Myanmar" pageId="13" pageNumber="14">Myanmar</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="diagnosis">
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<paragraph blockId="13.[88,758,219,826]" lastBlockId="13.[805,1475,179,337]" pageId="13" pageNumber="14">
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<emphasis box="[88,181,470,490]" italics="true" pageId="13" pageNumber="14">Diagnosis</emphasis>
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(females). Closely similar to
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<taxonomicName box="[473,542,470,490]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vladi">
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<emphasis box="[473,542,470,490]" italics="true" pageId="13" pageNumber="14">
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L.
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<taxonomicName box="[495,542,470,490]" pageId="13" pageNumber="14" rank="species" species="vladi">vladi</taxonomicName>
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</emphasis>
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</taxonomicName>
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, but differs from this species by having a shorter clypeal horn (horn index HoI
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<emphasis bold="true" box="[646,663,499,518]" pageId="13" pageNumber="14">¼</emphasis>
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31e
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||
<quantity box="[708,757,499,519]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="13" pageNumber="14" unit="in" value="34.0">34 in</quantity>
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||
<taxonomicName authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[88,221,526,546]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevicornis">
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<emphasis box="[88,221,526,546]" italics="true" pageId="13" pageNumber="14">
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L.
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<taxonomicName box="[114,221,526,546]" pageId="13" pageNumber="14" rank="species" species="brevicornis">brevicornis</taxonomicName>
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</emphasis>
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||
</taxonomicName>
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,
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||
<quantity box="[237,293,527,546]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="13" pageNumber="14" unit="in" value="56.0">56 in</quantity>
|
||
<taxonomicName box="[304,377,526,546]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vladi">
|
||
<emphasis box="[304,377,526,546]" italics="true" pageId="13" pageNumber="14">
|
||
L.
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<taxonomicName box="[330,377,526,546]" pageId="13" pageNumber="14" rank="species" species="vladi">vladi</taxonomicName>
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</emphasis>
|
||
</taxonomicName>
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), with stalk much reduced, slightly broader than long, with the setose pad more hexagonal and aligned with stalk (circular and with a distinct bend between stalk and clypeal pad in
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<taxonomicName box="[231,298,610,630]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vladi">
|
||
<emphasis box="[231,298,610,630]" italics="true" pageId="13" pageNumber="14">
|
||
L.
|
||
<taxonomicName box="[251,298,610,630]" pageId="13" pageNumber="14" rank="species" species="vladi">vladi</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
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||
), and with clypeal pad coated in stout setae. It is also differentiated by the mandibles with two teeth on ventral corner of medioventral blade (teeth absent in
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<taxonomicName box="[589,663,665,685]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vladi">
|
||
<emphasis box="[589,663,665,685]" italics="true" pageId="13" pageNumber="14">
|
||
L.
|
||
<taxonomicName box="[616,663,665,685]" pageId="13" pageNumber="14" rank="species" species="vladi">vladi</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
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||
) and, in workers, with the apical mandibular portion shorter than basal portion (apical portion longer in
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||
<taxonomicName box="[422,491,721,741]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vladi">
|
||
<emphasis box="[422,491,721,741]" italics="true" pageId="13" pageNumber="14">
|
||
L.
|
||
<taxonomicName box="[444,491,721,741]" pageId="13" pageNumber="14" rank="species" species="vladi">vladi</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
). In addition to the difference in the proportion of the mandible, the female castes of
|
||
<taxonomicName authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[88,219,777,797]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevicornis">
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||
<emphasis box="[88,219,777,797]" italics="true" pageId="13" pageNumber="14">
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L.
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<taxonomicName box="[112,219,777,797]" pageId="13" pageNumber="14" rank="species" species="brevicornis">brevicornis</taxonomicName>
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</emphasis>
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||
</taxonomicName>
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||
differ by their size (body length ca.
|
||
<quantity box="[604,665,779,798]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="mm" value="7.0">7 mm</quantity>
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||
in gyne,
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||
<quantity box="[88,148,806,826]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="14" unit="mm" value="4.0">4 mm</quantity>
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in worker), by the antennae with flagellomeres compact in worker, more elongate in gyne, and by subpetiolar process present in worker.
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</paragraph>
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</subSubSection>
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||
<subSubSection lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
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||
<paragraph blockId="13.[805,1475,179,337]" pageId="13" pageNumber="14">
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||
<emphasis box="[805,915,233,253]" italics="true" pageId="13" pageNumber="14">Description</emphasis>
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(gyne). Total length of body around
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<quantity box="[1323,1390,235,254]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="14" unit="mm" value="7.0">7 mm</quantity>
|
||
. Cuticle minutely shagreened, densely covered by adpressed, minute setae, the apical gastral segments additionally with sparse pilosity of long, fine, erect setae becoming gradually denser toward apex.
|
||
</paragraph>
|
||
<paragraph blockId="13.[805,1475,373,812]" lastBlockId="14.[113,784,1311,1972]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
Head. As in
|
||
<taxonomicName box="[978,1183,373,393]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vladi">
|
||
<emphasis box="[978,1183,373,393]" italics="true" pageId="13" pageNumber="14">
|
||
Linguamyrmex
|
||
<taxonomicName box="[1136,1183,373,393]" pageId="13" pageNumber="14" rank="species" species="vladi">vladi</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, but with compound eyes slightly reniform, emarginate in middle of posterior margin. Ocelli distinct, positioned on a prominent cuticular triangle, in dorsal view, with posterior margin of lateral ocelli aligned with posterior margin of eyes. Antenna filiform, with flagellomeres elongate, cylindrical except apical one, which has acute tip; third antennomere twice as long as following one. Antennal sockets immediately flanking a medial frontal triangle that is sharply expanded anteriorly above clypeal horn, and connecting ventrally with dorsal surface of horn by a cleared, laterally flattened cuticle; frontal triangle projection terminates anteriorly at approximately same length as clypeal horn with slight indentation just dorsad clypeal horn, giving the appearance of a second, small, broadly rounded horn. Clypeal horn originating at base of frontal triangle, short and straight, in profile view almost perpendicular to longest axis of head. Horn paddle-shaped, with short, compact stalk and hexagonal, setose pad; stalk apparently glabrous,
|
||
<quantity box="[448,539,1311,1330]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="mm" value="0.1">0.10 mm</quantity>
|
||
in length,
|
||
<quantity box="[661,752,1311,1330]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="15" unit="mm" value="0.12">0.12 mm</quantity>
|
||
in width; setose pad
|
||
<quantity box="[295,382,1339,1358]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="15" unit="mm" value="0.26">0.26 mm</quantity>
|
||
in length,
|
||
<quantity box="[486,574,1339,1358]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="mm" value="0.2">0.20 mm</quantity>
|
||
at greatest diameter, with anterior margin straight, transverse; dorsal surface of setose pad glabrous, ventral surface densely and uniformly coated with very short, velcro-like vestiture; trigger hairs not visible. Lateral clypeal margins extending from above mandibular insertions toward horn as gently curved lines. Anterior clypeal margin broadly concave. Labrum visible. Mandible scythe-shaped; basal portion linear, short, meeting apical curved portion nearly at right angle; medioventral triangular blade bearing one large apical tooth and one small preapical tooth, with concave dorsal surface coated with pointed setae; apical portion of mandible posterior to medioventral blade 1.6 as long as blade, with tip broadly rounded. Mandibles preserved in closed position (with tips of apical portion reaching nearby clypeal setose pad), almost entirely parallel and closely approximated, except ventral corners of blades overlapping and tips slightly diverging. Maxillary palps 5-segmented, labial palps 3- segmented.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3653086" ID-Zenodo-Dep="3653086" httpUri="https://zenodo.org/record/3653086/files/figure.png" pageId="13" pageNumber="14" startId="13.[88,113,1945,1959]" targetBox="[176,1386,890,1916]" targetPageId="13">
|
||
<paragraph blockId="13.[88,1476,1944,1982]" pageId="13" pageNumber="14">
|
||
<emphasis bold="true" box="[88,136,1945,1959]" pageId="13" pageNumber="14">Fig. 7.</emphasis>
|
||
Overview and details of females of
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[427,626,1945,1960]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevicornis" status="gen. et sp. nov.">
|
||
<emphasis box="[427,626,1945,1960]" italics="true" pageId="13" pageNumber="14">Linguamyrmex brevicornis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[633,755,1946,1960]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Gyne specimen LA01 (AeB), worker specimen RM4 (C), worker holotype NIGP172001 (D). Habitus (A, C), head and mesosoma in dorsal view (B), head in ventrolateral view (D). Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3653088" ID-Zenodo-Dep="3653088" httpUri="https://zenodo.org/record/3653088/files/figure.png" pageId="14" pageNumber="15" startId="14.[113,138,1215,1229]" targetBox="[282,1334,179,1185]" targetPageId="14">
|
||
<paragraph blockId="14.[113,1502,1215,1253]" pageId="14" pageNumber="15">
|
||
<emphasis bold="true" box="[113,160,1215,1229]" pageId="14" pageNumber="15">Fig. 8.</emphasis>
|
||
Artistic reconstructions of the heads of hell ants in lateral view, with mandibles in closed position. A,
|
||
<taxonomicName authorityName="Dlussky" authorityYear="1996" box="[947,1052,1215,1230]" class="Hexapoda" family="Formicidae" genus="Haidomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[947,1052,1215,1230]" italics="true" pageId="14" pageNumber="15">Haidomyrmex</emphasis>
|
||
</taxonomicName>
|
||
. B,
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[1078,1224,1215,1230]" class="Insecta" family="Formicidae" genus="Protoceratomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[1078,1224,1215,1230]" italics="true" pageId="14" pageNumber="15">Protoceratomyrmex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1229,1299,1216,1230]" pageId="14" pageNumber="15" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
C,
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[1319,1427,1215,1230]" class="Insecta" family="Formicidae" genus="Aquilomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[1319,1427,1215,1230]" italics="true" pageId="14" pageNumber="15">Aquilomyrmex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1432,1502,1216,1230]" pageId="14" pageNumber="15" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
D,
|
||
<taxonomicName authorityName="Barden & Grimaldi" authorityYear="2017" box="[135,246,1237,1252]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[135,246,1237,1252]" italics="true" pageId="14" pageNumber="15">Linguamyrmex</emphasis>
|
||
</taxonomicName>
|
||
. E,
|
||
<taxonomicName authorityName="Perrichot, Wang & Engel" authorityYear="2016" box="[275,384,1237,1252]" class="Hexapoda" family="Formicidae" genus="Ceratomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[275,384,1237,1252]" italics="true" pageId="14" pageNumber="15">Ceratomyrmex</emphasis>
|
||
</taxonomicName>
|
||
. F,
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[410,481,1237,1252]" class="Insecta" family="Formicidae" genus="Chonidris" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[410,481,1237,1252]" italics="true" pageId="14" pageNumber="15">Chonidris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[487,558,1239,1253]" pageId="14" pageNumber="15" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
G,
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[584,673,1237,1252]" class="Insecta" family="Formicidae" genus="Dhagnathos" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[584,673,1237,1252]" italics="true" pageId="14" pageNumber="15">Dhagnathos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[678,750,1239,1253]" pageId="14" pageNumber="15" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="14.[113,784,1311,1972]" lastBlockId="15.[88,758,179,1733]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">Mesosoma. Pronotal colar pronounced but not concealing entire propleuron in dorsal view; pronotal dorsum weakly convex. Promesonotal suture present, complete. In dorsal view, mesoscutum as long as pronotum (excluding neck), about as broad as long, with lateral and posterior margins carinate; posterior mesoscutal margin broadly concave; dorsal mesoscutal surface with long parapsidal furrows almost reaching anterior mesonotal margin, converging posteriorly but not touching. In profile view, pronotal dorsal outline feebly convex, mesoscutal dorsal outline nearly flat. Mesoscutellum prominent posteriorly; posterior mesoscutellar surfaces concave, their junction forming a sharp angle; dorsal mesoscutellar surface with a deep, broad, transverse groove immediately posterior to scuto-scutellar suture. Metanotum medially as high as long, with posterior surface forming distinct angle with pronotal dorsum. Propodeum 1.3 as high as long; in profile view, propodeal dorsum distinctly lower than pronotal dorsum; propodeal dorsum nearly flat, declivitous surface feebly convex, their junction broadly rounded. Propodeal spiracle slit-like, oriented posteriad. Metapleural gland opening circular, gaping. Legs long and robust, mesocoxa distinctly shorter than pro- and metacoxae; small trochantellus present on mid- and hind legs; all femora distinctly swollen in their basal half, tibiae swollen in their apical half; ventral margin of protibia apically with large calcar and two simple, stout setae less than half as long as calcar; mesotibia apically with two long, straight, pectinate spurs, and two short, straight, stout setae; metatibia apically with one long, pectinate spur and one long, simple spur. Tarsomeres IeIV of all legs with pairs of stout setae along entire ventral surface (8e10 pairs on tarsomere I, 4e5 pairs on tarsomere II, 3 pairs on tarsomere III, 2 pairs on tarsomere IV), and apically with 2 pairs of stout setae each flanking a spatulate spine; additionally the ventral surface of tarsomeres IeIV covered by dense brush of fine, erect setae; pretarsal claws thick, with a distinct subapical tooth.</paragraph>
|
||
<paragraph blockId="15.[88,758,179,1733]" pageId="15" pageNumber="16">
|
||
Fore wing with veins M∙f4 and Cu1 nebulous, not reaching wing margin; remaining veins tubular; cross-vein 1r-rs absent; Rs∙f1 faintly arched; M∙f1 distinctly arched, twice as long as Rs∙f1; Rs∙f2 nearly at right angle with Rs
|
||
<emphasis bold="true" box="[380,397,347,366]" pageId="15" pageNumber="16">þ</emphasis>
|
||
M, half as long as M∙f2; Rs
|
||
<emphasis bold="true" box="[674,691,347,366]" pageId="15" pageNumber="16">þ</emphasis>
|
||
M and M∙f2 not aligned, so that discal cell pentagonal, with vein 1m-cu distinctly longer than M∙f1; second submarginal cell long, crossvein 2rs-m situated far beyond apex of pterostigma; cross-vein cu-a arising from Cu, shortly distal to M∙f1. Hind wing with jugal lobe present.
|
||
</paragraph>
|
||
<paragraph blockId="15.[88,758,179,1733]" pageId="15" pageNumber="16">
|
||
Metasoma. Petiole short-pedunculate; petiolar tergite forming a broadly convex node, with anterior surface approximately twice as long as posterior surface; subpetiolar process present, in profile a high, transverse, tooth pointing ventrally, with anterior surface concave, posterior surface vertical; transverse sulcus visible across petiole near basal margin. First gastral segment with helcium pronounced, forming a post-petiolar peduncle, with anterior surface posterior to helcium high, oblique, and dorsal surface strongly convex, short; anteriormost part of first gastral sternite with a distinct mesal process pointing anteroventrally below helcium. Second gastral segment distinctly longer than first, with presclerite largely exposed to form a deep, broad constriction between first and second gastral segments (abdominal segments
|
||
<collectionCode box="[641,661,849,868]" pageId="15" pageNumber="16">III</collectionCode>
|
||
and IV). Posterior margin of gastral segments II onward with sparse, elongate setae. Pygidium nearly glabrous while hypopygium with sparse setae; pygidium apparently broadly acute towards sting shaft.
|
||
</paragraph>
|
||
<paragraph blockId="15.[88,758,179,1733]" pageId="15" pageNumber="16">
|
||
<emphasis box="[119,383,987,1007]" italics="true" pageId="15" pageNumber="16">Wingless female (worker).</emphasis>
|
||
Smaller than gyne (total length of body around
|
||
<quantity box="[219,279,1016,1035]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="mm" value="4.0">4 mm</quantity>
|
||
), and with the following differences: compound eyes oval, ocelli faintly visible, not positioned on a prominent cuticular triangle. Antenna more compact, with flagellomeres IIeIX gradually broadened apically, apical flagellomere with rounded apex. Clypeal setose pad more quadrangular, the ventral surface with a pair of trigger hairs originating at pad base. Mandible with medioventral triangular blade larger and apical portion (posterior to blade) reduced, 0.6 as long as blade.
|
||
</paragraph>
|
||
<paragraph blockId="15.[88,758,179,1733]" pageId="15" pageNumber="16">
|
||
Mesosoma and metasoma as in
|
||
<taxonomicName box="[474,548,1238,1258]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="vladi">
|
||
<emphasis box="[474,548,1238,1258]" italics="true" pageId="15" pageNumber="16">L. vladi</emphasis>
|
||
</taxonomicName>
|
||
except propodeum higher than long, petiole with subpetiolar process present, and girdling constriction deeper between abdominal segments III and IV (
|
||
<collectionCode box="[121,158,1323,1342]" pageId="15" pageNumber="16">AIII</collectionCode>
|
||
with an oblique posterior surface). Protibia with one large apical spur (calcar) and two short stout setae; mesotibia with two simple spurs and three conspicuous setae positioned along anterior margin of tibial apex; metatibia with one pectinate and one simple spur, additionally with a brush of 8e10 setae flanking the pectinate spur. Sparse, erect setae positioned on dorsum of pronotum, propodeum, petiolar tergite, and first and second gastral tergites and sternites. Pygidium not preserved.
|
||
</paragraph>
|
||
<paragraph blockId="15.[88,758,179,1733]" pageId="15" pageNumber="16">
|
||
<emphasis box="[119,260,1545,1565]" italics="true" pageId="15" pageNumber="16">Measurements</emphasis>
|
||
(in mm) (worker
|
||
<typeStatus box="[443,532,1546,1565]" pageId="15" pageNumber="16">holotype</typeStatus>
|
||
<collectionCode box="[538,595,1546,1565]" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="15" pageNumber="16">NIGP</collectionCode>
|
||
172001) [worker specimen
|
||
<collectionCode box="[191,229,1574,1593]" country="Canada" name="McGill University, Redpath Museum" pageId="15" pageNumber="16" type="Museum">RM</collectionCode>
|
||
4] {gyne specimen LA01}. HL [0.82] {0.82}; EL [0.21] {0.42}; HoL [0.17] {0.36}; MDL [0.59] {0.80}; length of antennomeres: I (scape) (0.57) [0.67] {1.04}, II (pedicel) (0.09) [0.12] {0.14}, III-XII (0.35, 0.17, 0.16, 0.15, 0.15, 0.13, 0.13, 0.12, 0.13, 0.17) {0.80, 0.40, 0.34, 0.32, 0.31, 0.30, 0.30, 0.32, 0.32, 0.35}; WL [1.29] {2.29};
|
||
<collectionCode box="[163,186,1714,1733]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="15" pageNumber="16" type="Museum">PL</collectionCode>
|
||
{0.70}.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |