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<document id="7C77BB41DB7B2E914A758338835F3F11" ID="10.11646/zootaxa.4258.1.5SLASH10900" ID-CLB-Dataset="33163" ID-DOI="10.11646/zootaxa.4258.1.5" ID-GBIF-Dataset="bdf2db84-05b5-4c94-bfa7-7cf9274fda9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="569169" ID-ZooBank="D1095E5E-2C9F-40C6-BA10-EADBEA56681C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493196110549" checkinUser="plazi" docAuthor="Fomichev, Alexander A. &amp; Marusik, Yuri M." docDate="2017" docId="039C9E749664FFFBFF40F89EDCCCF86E" docLanguage="en" docName="zootaxa.4258.1.5SLASH10900.pdf" docOrigin="Zootaxa 4258 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gnaphosa zonsteini Fomichev &amp; Marusik, 2017, sp. n." docType="treatment" docVersion="13" lastPageNumber="77" masterDocId="FFA5E60C9662FFF3FFD7FFE8D87AFFD4" masterDocTitle="A survey of East Palaearctic Gnaphosidae (Araneae). 8. New data on Berlandina and Gnaphosa from Mongolia" masterLastPageNumber="80" masterPageNumber="69" pageNumber="75" updateTime="1698439081613" updateUser="ExternalLinkService">
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<mods:title id="F36E78FC17CCB5066C73857A247124F4">A survey of East Palaearctic Gnaphosidae (Araneae). 8. New data on Berlandina and Gnaphosa from Mongolia</mods:title>
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<mods:namePart id="688B8461E37598C005657C51C276C2EC">Fomichev, Alexander A.</mods:namePart>
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<mods:namePart id="F46BDFAF8309A9F363CF58F4748A9C35">Marusik, Yuri M.</mods:namePart>
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<mods:title id="274C4BE94077CC8BE6E008407CB81244">Zootaxa</mods:title>
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<mods:date id="5DCB97970707012A4025E0A55841CE4A">2017</mods:date>
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<treatment id="039C9E749664FFFBFF40F89EDCCCF86E" ID-DOI="http://doi.org/10.5281/zenodo.6028589" ID-GBIF-Taxon="128699698" ID-Zenodo-Dep="6028589" LSID="urn:lsid:plazi:treatment:039C9E749664FFFBFF40F89EDCCCF86E" httpUri="http://treatment.plazi.org/id/039C9E749664FFFBFF40F89EDCCCF86E" lastPageId="8" lastPageNumber="77" pageId="6" pageNumber="75">
<subSubSection id="C32F7CE99664FFF5FF40F89ED9E8F866" pageId="6" pageNumber="75" type="nomenclature">
<paragraph id="8B8A2F629664FFF5FF40F89ED9B5F844" blockId="6.[151,463,1910,1970]" box="[151,463,1910,1936]" pageId="6" pageNumber="75">
<heading id="D0C2980E9664FFF5FF40F89ED9B5F844" bold="true" box="[151,463,1910,1936]" fontSize="11" level="1" pageId="6" pageNumber="75" reason="1">
<emphasis id="B941F3709664FFF5FF40F89ED9B5F844" bold="true" box="[151,463,1910,1936]" pageId="6" pageNumber="75">
<taxonomicName id="4C3554E19664FFF5FF40F89ED9FCF844" ID-CoL="6KNQQ" box="[151,390,1910,1936]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
<emphasis id="B941F3709664FFF5FF40F89ED9FCF844" bold="true" box="[151,390,1910,1936]" italics="true" pageId="6" pageNumber="75">Gnaphosa zonsteini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2724E0B9664FFF5FE5BF89ED9B5F844" box="[396,463,1910,1936]" pageId="6" pageNumber="75" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A2F629664FFF5FF40F871D9E8F866" blockId="6.[151,463,1910,1970]" box="[151,402,1945,1970]" pageId="6" pageNumber="75">
<figureCitation id="130E33E79664FFF5FF40F871D96CF865" box="[151,278,1945,1970]" captionStart="FIGURES 20 - 27" captionStartId="8.[151,264,1609,1632]" captionTargetBox="[156,1434,206,1584]" captionTargetId="figure@8.[151,1436,195,1588]" captionTargetPageId="8" captionText="FIGURES 20 - 27. Male of Gnaphosa zonsteini sp. n. 20 habitus, dorsal; 21 ‒ 23 palp, ventral, retro- and prolateral; 24 cymbium and tibia, retrolateral; 25 ‒ 27 bulb, prolateral, ventral and anterior. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569172/files/figure.png" pageId="6" pageNumber="75">Figs 20‒27</figureCitation>
,
<figureCitation id="130E33E79664FFF5FEF5F871D911F865" box="[290,363,1945,1969]" captionStart="FIGURES 28 ‒ 33" captionStartId="9.[151,264,1714,1737]" captionTargetBox="[151,1436,368,1692]" captionTargetId="figure@9.[151,1436,363,1692]" captionTargetPageId="9" captionText="FIGURES 28 ‒ 33. Female of Gnaphosa rasnitsyni (28 ‒ 31) and G. zonsteini sp. n. (32 ‒ 33). 28 habitus, dorsal; 29 intact epigyne, ventral; 30, 32 macerated epigyne, ventral; 31, 33 macerated epigyne, dorsal. Scale = 0.2 mm if not otherwise indicated. Abbreviations. Ab foveal bulges, Lp lateral pockets, Ps pocket of the scape; Rh head of receptacle." httpUri="https://zenodo.org/record/569173/files/figure.png" pageId="6" pageNumber="75">32‒33</figureCitation>
,
<figureCitation id="130E33E79664FFF5FEA1F871D9E8F866" box="[374,402,1945,1970]" captionStart="FIGURES 34 ‒ 35" captionStartId="10.[151,264,1665,1688]" captionTargetBox="[188,1398,1095,1643]" captionTargetId="figure@10.[188,1398,1095,1643]" captionTargetPageId="10" captionText="FIGURES 34 ‒ 35. Collecting localities of Berlandina and Gnaphosa species examined in this study. Solid symbol on Fig. 34 refers to new records, open symbols to the literature data, dotted square refers to the region shown on Fig. 35. Blue squares in Fig. 35 refer to the records of G. ustyuzhanini (= G. rasnitsyni). One symbol may designate more than one locality." httpUri="https://zenodo.org/record/569174/files/figure.png" pageId="6" pageNumber="75">35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99664FFF4FF40F833DB01FF00" lastPageId="7" lastPageNumber="76" pageId="6" pageNumber="75" type="materials_examined">
<paragraph id="8B8A2F629664FFF4FF40F833DB01FF00" blockId="6.[151,1436,2011,2036]" lastBlockId="7.[151,1437,151,645]" lastPageId="7" lastPageNumber="76" pageId="6" pageNumber="75">
<materialsCitation id="3B5D253F9664FFF4FF40F833D905FF00" ID-GBIF-Occurrence="1493295301" collectingDate="2015-05-18" collectionCode="ISEA" country="Mongolia" elevation="1280" lastPageId="7" lastPageNumber="76" latitude="45.766666" location="Bodonchiyn-Gol River Valley" longLatPrecision="1128" longitude="92.2" municipality="Altai Village" pageId="6" pageNumber="75" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Khovd" typeStatus="holotype">
<emphasis id="B941F3709664FFF5FF40F833D97CF820" bold="true" box="[151,262,2011,2036]" pageId="6" pageNumber="75">Material.</emphasis>
<collectingCountry id="F3226FF29664FFF5FED8F834D9D6F820" box="[271,428,2012,2036]" name="Mongolia" pageId="6" pageNumber="75">MONGOLIA</collectingCountry>
:
<collectingRegion id="49F1E1809664FFF5FE6FF833DA79F820" box="[440,515,2011,2036]" country="Mongolia" name="Hovd" pageId="6" pageNumber="75">
<emphasis id="B941F3709664FFF5FE6FF833DA79F820" bold="true" box="[440,515,2011,2036]" italics="true" pageId="6" pageNumber="75">Khovd</emphasis>
</collectingRegion>
Aimag:
<typeStatus id="548E91C09664FFF5FDBFF833DAB0F820" box="[616,714,2011,2036]" pageId="6" pageNumber="75" type="holotype">holotype</typeStatus>
<specimenCount id="9D33E4EB9664FFF5FD04F834DA92F827" box="[723,744,2012,2035]" pageId="6" pageNumber="75" type="male">♂</specimenCount>
(
<collectionCode id="ED24B7A79664FFF5FD2FF834DB43F820" box="[760,825,2012,2036]" pageId="6" pageNumber="75">ISEA</collectionCode>
) and
<typeStatus id="548E91C09664FFF5FCACF834DB9DF820" box="[891,999,2012,2036]" pageId="6" pageNumber="75" type="paratype">paratypes</typeStatus>
<specimenCount id="9D33E4EB9664FFF5FC38F834DC68F820" box="[1007,1042,2012,2036]" pageId="6" pageNumber="75" type="male">4♂</specimenCount>
<specimenCount id="9D33E4EB9664FFF5FBCCF834DC44F820" box="[1051,1086,2012,2036]" pageId="6" pageNumber="75" type="female">1♀</specimenCount>
(
<collectionCode id="ED24B7A79664FFF5FB98F834DCEAF820" box="[1103,1168,2012,2036]" pageId="6" pageNumber="75">ISEA</collectionCode>
)
<quantity id="4CCD82879664FFF5FB76F833DC90F820" box="[1185,1258,2011,2036]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="75" unit="km" value="36.0">36 km</quantity>
SW from
<collectingMunicipality id="6BEEB5189664FFF4FAB3F833D891FF64" lastPageId="7" lastPageNumber="76" pageId="6" pageNumber="75">Altai Village</collectingMunicipality>
,
<location id="8EEA79B99665FFF4FF21FF7FDA32FF64" LSID="urn:lsid:plazi:treatment:039C9E749664FFFBFF40F89EDCCCF86E:8EEA79B99665FFF4FF21FF7FDA32FF64" box="[246,584,151,176]" country="Mongolia" latitude="45.766666" longLatPrecision="1128" longitude="92.2" municipality="Altai Village" name="Bodonchiyn-Gol River Valley" pageId="7" pageNumber="76" stateProvince="Khovd">Bodonchiyn-Gol River Valley</location>
,
<geoCoordinate id="EE0149A59665FFF4FD83FF7FDACDFF64" box="[596,695,151,176]" direction="north" orientation="latitude" pageId="7" pageNumber="76" precision="925" value="45.766666">45°46’N</geoCoordinate>
,
<geoCoordinate id="EE0149A59665FFF4FD14FF70DB59FF64" box="[707,803,152,176]" direction="east" orientation="longitude" pageId="7" pageNumber="76" precision="925" value="92.2">92°12’E</geoCoordinate>
, tugay (gallery forest), night, attracted by light,
<elevation id="0018C8519665FFF4FA97FF70DDEDFF7B" box="[1344,1431,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="7" pageNumber="76" unit="m" value="1280.0">
<quantity id="4CCD82879665FFF4FA97FF70DDEDFF7B" box="[1344,1431,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="7" pageNumber="76" unit="m" value="1280.0">1280 m</quantity>
</elevation>
, 17‒
<collectingDate id="EFCFF04A9665FFF4FF69FF54D944FF01" box="[190,318,188,213]" pageId="7" pageNumber="76" value="2015-05-18">
<date id="FF8B09A29665FFF4FF69FF54D944FF01" box="[190,318,188,213]" pageId="7" pageNumber="76" value="2015-05-18">18.05.2015</date>
</collectingDate>
(AF)
</materialsCitation>
;
<materialsCitation id="3B5D253F9665FFF4FE5CFF54DB01FF00" ID-GBIF-Occurrence="1493295251" box="[395,891,188,213]" collectingDate="2015-05-18" collectionCode="ZMMU" country="Mongolia" elevation="1280" latitude="45.766666" location="Bodonchiyn-Gol River Valley" longLatPrecision="1128" longitude="92.2" municipality="Altai Village" pageId="7" pageNumber="76" specimenCount="1" specimenCount-male="1" stateProvince="Khovd" typeStatus="holotype">
<specimenCount id="9D33E4EB9665FFF4FE5CFF54D9D4FF00" box="[395,430,188,212]" pageId="7" pageNumber="76" type="male">1♂</specimenCount>
(
<collectionCode id="ED24B7A79665FFF4FE6AFF54DA6DFF00" box="[445,535,188,212]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="7" pageNumber="76">ZMMU</collectionCode>
) same locality, 1.06.2015 (AF).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99665FFF4FF10FF37DA83FEC9" pageId="7" pageNumber="76" type="etymology">
<paragraph id="8B8A2F629665FFF4FF10FF37DA83FEC9" blockId="7.[151,1437,151,645]" pageId="7" pageNumber="76">
<emphasis id="B941F3709665FFF4FF10FF37D934FF2C" bold="true" box="[199,334,223,248]" pageId="7" pageNumber="76">Etymology.</emphasis>
The species name is a patronym in honour of our friend and colleague Sergei L. Zonstein, on the occasion of his 60th birthday, the 8th of August, 2016.
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99665FFF4FF10FECFD9ADFDCC" pageId="7" pageNumber="76" type="diagnosis">
<paragraph id="8B8A2F629665FFF4FF10FECFD9ADFDCC" blockId="7.[151,1437,151,645]" pageId="7" pageNumber="76">
<emphasis id="B941F3709665FFF4FF10FECFD93BFE94" bold="true" box="[199,321,295,320]" pageId="7" pageNumber="76">Diagnosis.</emphasis>
Males of the new species resemble
<taxonomicName id="4C3554E19665FFF4FD3DFEC0DD65FE94" authority="Marusik, Fomichev &amp; Omelko, 2014" authorityName="Marusik, Fomichev &amp; Omelko" authorityYear="2014" box="[746,1311,295,320]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="esyunini">
<emphasis id="B941F3709665FFF4FD3DFEC0DB13FEEB" box="[746,873,296,319]" italics="true" pageId="7" pageNumber="76">G. esyunini</emphasis>
Marusik, Fomichev &amp; Omelko, 2014
</taxonomicName>
by having large embolic spine directed prolaterally, but can be distinguished from the former species by longer embolus without serration on the prolateral edge, lowered base of embolus and not bulged median apophysis (cf.
<figureCitation id="130E33E79665FFF4FAC8FE87DDE6FE5C" box="[1311,1436,367,392]" captionStart="FIGURES 20 - 27" captionStartId="8.[151,264,1609,1632]" captionTargetBox="[156,1434,206,1584]" captionTargetId="figure@8.[151,1436,195,1588]" captionTargetPageId="8" captionText="FIGURES 20 - 27. Male of Gnaphosa zonsteini sp. n. 20 habitus, dorsal; 21 ‒ 23 palp, ventral, retro- and prolateral; 24 cymbium and tibia, retrolateral; 25 ‒ 27 bulb, prolateral, ventral and anterior. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569172/files/figure.png" pageId="7" pageNumber="76">Figs 21–27</figureCitation>
and figs
<quantity id="4CCD82879665FFF4FF2EFE7CD934FE79" box="[249,334,404,429]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.794" metricValueMin="1.778" pageId="7" pageNumber="76" unit="in" value="9.0" valueMax="11.0" valueMin="7.0">7–11 in</quantity>
<bibRefCitation id="EFA452939665FFF4FE82FE7CDA3DFE79" author="Marusik" box="[341,583,404,429]" pageId="7" pageNumber="76" refString="Marusik, Y. M., Fomichev, A. A. &amp; Omelko, M. M. (2014 b) A survey of East Palaearctic Gnaphosidae (Araneae). 2. Two new Gnaphosa Latreille, 1804 species from Western Mongolia. ZooKeys, 426, 1 - 9." type="journal article" year="2014" yearSuffix="b">
Marusik
<emphasis id="B941F3709665FFF4FE6CFE7DD98EFE78" box="[443,500,404,428]" italics="true" pageId="7" pageNumber="76">et al.</emphasis>
2014b
</bibRefCitation>
). Females similar to
<taxonomicName id="4C3554E19665FFF4FCE4FE7CDCFCFE78" authority="Kulczynski, 1901" authorityName="Kulczynski" authorityYear="1901" box="[819,1158,404,429]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="gracilior">
<emphasis id="B941F3709665FFF4FCE4FE7CDBCCFE78" box="[819,950,404,428]" italics="true" pageId="7" pageNumber="76">G. gracilior</emphasis>
Kulczyński, 1901
</taxonomicName>
,
<taxonomicName id="4C3554E19665FFF4FB44FE7CD8A0FE04" authority="L. Koch, 1866" authorityName="L. Koch" authorityYear="1866" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="muscorum">
<emphasis id="B941F3709665FFF4FB44FE7CDD5DFE78" box="[1171,1319,404,428]" italics="true" pageId="7" pageNumber="76">G. muscorum</emphasis>
(L. Koch, 1866)
</taxonomicName>
,
<taxonomicName id="4C3554E19665FFF4FF3EFE50DA5DFE04" authority="Marusik, 1993" authorityName="Marusik" authorityYear="1993" box="[233,551,439,464]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="rasnitsyni">
<emphasis id="B941F3709665FFF4FF3EFE50D902FE1B" box="[233,376,440,463]" italics="true" pageId="7" pageNumber="76">G. rasnitsyni</emphasis>
<bibRefCitation id="EFA452939665FFF4FE55FE5FDA5DFE04" author="Marusik" box="[386,551,439,464]" pageId="7" pageNumber="76" refString="Marusik, Y. M. (1993) Three new wandering spider species (Aranei Lycosidae Gnaphosidae) from Mongolia. Arthropoda Selecta, 2, 77 - 81." type="journal article" year="1993">Marusik, 1993</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3554E19665FFF4FDB4FE50DBEAFE04" authority="Schenkel, 1963" authorityName="Schenkel" authorityYear="1963" box="[611,912,439,464]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="wiehlei">
<emphasis id="B941F3709665FFF4FDB4FE50DAAFFE04" box="[611,725,440,464]" italics="true" pageId="7" pageNumber="76">G. wiehlei</emphasis>
Schenkel, 1963
</taxonomicName>
, but differ from the later three species by the shape of atrium and septum and from
<taxonomicName id="4C3554E19665FFF4FDB9FE34DA8DFE20" box="[622,759,476,500]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="gracilior">
<emphasis id="B941F3709665FFF4FDB9FE34DA8DFE20" box="[622,759,476,500]" italics="true" pageId="7" pageNumber="76">G. gracilior</emphasis>
</taxonomicName>
by less concave posterior margin of scape and head of receptacles directed upward.
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE99665FFFBFF10FDCCD887F8D2" lastPageId="8" lastPageNumber="77" pageId="7" pageNumber="76" type="description">
<paragraph id="8B8A2F629665FFF4FF10FDCCDB93FD51" blockId="7.[151,1437,151,645]" pageId="7" pageNumber="76">
<emphasis id="B941F3709665FFF4FF10FDCCD923FDE9" bold="true" box="[199,345,548,573]" pageId="7" pageNumber="76">Description.</emphasis>
Male. Total length 6.6. Carapace: 3.0 long, 2.3 wide. Coloration. Prosoma and chelicerae darkbrown. Carapace with black edges and two dark stripes around cephalic area. Legs and palps brown. Abdomen dark-gray, almost black dorsally and yellow-gray ventrally. Spinnerets gray.
</paragraph>
<paragraph id="8B8A2F629665FFF4FF40FD5DD908FD18" blockId="7.[151,370,693,717]" box="[151,370,693,717]" pageId="7" pageNumber="76">Leg measurements:</paragraph>
<paragraph id="8B8A2F629665FFF4FEF9FD01D942FBD7" blockId="7.[159,1287,744,931]" lastBlockId="7.[151,312,1002,1027]" pageId="7" pageNumber="76">Fe Pt Ti Mt Ta Total I 2.1 1.2 1.65 1.5 1.25 7.7 II 1.9 1.1 1.45 1.35 1.2 7.0 III 1.75 0.95 1.2 1.5 1.05 6.45 IV 2.35 1.2 1.85 2.5 1.35 9.25 Leg spination:</paragraph>
<paragraph id="8B8A2F629665FFF4FF31FBF7DC53FBE0" blockId="7.[159,1320,1054,1242]" box="[230,1065,1054,1076]" pageId="7" pageNumber="76">Fe Pt Ti Mt</paragraph>
<paragraph id="8B8A2F629665FFF4FF48FBA1DC34FB8A" blockId="7.[159,1320,1054,1242]" box="[159,1102,1096,1118]" pageId="7" pageNumber="76">
I d1-1-0 p0-0-
<date id="FF8B09A29665FFF4FEBDFBA0DAFEFB8A" box="[362,644,1096,1118]" pageId="7" pageNumber="76">1 v</date>
1-1-
<date id="FF8B09A29665FFF4FD7FFBA0DC63FB8A" box="[680,1049,1096,1118]" pageId="7" pageNumber="76">2 v</date>
2-0-0
</paragraph>
<paragraph id="8B8A2F629665FFF4FF48FB9ADC34FB53" blockId="7.[159,1320,1054,1242]" box="[159,1102,1137,1159]" pageId="7" pageNumber="76">
II d1-1-0 p0-1-1 p0-0-
<date id="FF8B09A29665FFF4FD79FB99DAB7FB53" box="[686,717,1137,1159]" pageId="7" pageNumber="76">1 v</date>
1-2-
<date id="FF8B09A29665FFF4FD24FB99DC63FB53" box="[755,1049,1137,1159]" pageId="7" pageNumber="76">2 v</date>
2-0-0
</paragraph>
<paragraph id="8B8A2F629665FFF4FF48FB73DA4BFA1E" blockId="7.[159,1320,1054,1242]" lastBlockId="7.[151,1436,1313,1518]" pageId="7" pageNumber="76">
III d1-1-0 p0-1-1 r0-1-1 r1-0 d1-0-0 p2-1-1 r2-1-
<date id="FF8B09A29665FFF4FCEAFB72DB21FB64" box="[829,859,1178,1200]" pageId="7" pageNumber="76">1 v</date>
2-2-2 d0-1-0 p1-1-0 r1-1-0
<date id="FF8B09A29665FFF4FB33FB72DC85FB64" box="[1252,1279,1178,1200]" pageId="7" pageNumber="76">v2</date>
-0-
<date id="FF8B09A29665FFF4FACCFB72D8C0FB0E" pageId="7" pageNumber="76">2 IV</date>
d1-1-0 p0-1-1 r0-1-1 p2-1-1 r2-1-
<date id="FF8B09A29665FFF4FD24FB2CDB68FB0E" box="[755,786,1220,1242]" pageId="7" pageNumber="76">1 v</date>
2-2-2 d0-2-0 p1-1-0 r1-1-0
<date id="FF8B09A29665FFF4FB33FB2CDD69FB0E" box="[1252,1299,1220,1242]" pageId="7" pageNumber="76">v2-2</date>
-0 Palp as in
<figureCitation id="130E33E79665FFF4FE96FAC9D9BEFAEE" box="[321,452,1313,1338]" captionStart="FIGURES 20 - 27" captionStartId="8.[151,264,1609,1632]" captionTargetBox="[156,1434,206,1584]" captionTargetId="figure@8.[151,1436,195,1588]" captionTargetPageId="8" captionText="FIGURES 20 - 27. Male of Gnaphosa zonsteini sp. n. 20 habitus, dorsal; 21 ‒ 23 palp, ventral, retro- and prolateral; 24 cymbium and tibia, retrolateral; 25 ‒ 27 bulb, prolateral, ventral and anterior. Scale = 0.2 mm if not otherwise indicated." httpUri="https://zenodo.org/record/569172/files/figure.png" pageId="7" pageNumber="76">Figs 21‒27</figureCitation>
; tibial apophysis rather long, almost equal in length to tibia, with pointed tip; median apophysis large, hook-shaped; embolus long, filamentous, embolic base with strong spine directed prolaterally. Size variations. Total length 6.2–8.3; carapace 3.0–3.65 long and 2.3–2.9 wide (n=5). Female. Total length 12.3. Carapace: 4.35 long, 3.25 wide. Coloration as in male, but with lighter carapace and monochrome dirty-yellow abdomen.
</paragraph>
<paragraph id="8B8A2F629665FFF4FF40FA3ED908FA39" blockId="7.[151,1436,1313,1518]" box="[151,370,1494,1518]" pageId="7" pageNumber="76">Leg measurements:</paragraph>
<paragraph id="8B8A2F629665FFF4FEE0F9E2DC9CF9A6" blockId="7.[159,1279,1545,1732]" pageId="7" pageNumber="76">Fe Pt Ti Mt Ta Total I 2.65 1.6 1.95 1.55 1.3 9.05 II 2.4 1.45 1.7 1.45 1.2 8.2</paragraph>
<paragraph id="8B8A2F629665FFF4FF48F96EDC9CF94F" blockId="7.[159,1279,1545,1732]" box="[159,1254,1669,1691]" pageId="7" pageNumber="76">III 2.2 1.25 1.4 1.8 1.15 7.8</paragraph>
<paragraph id="8B8A2F629665FFF4FF48F947DC85F910" blockId="7.[159,1279,1545,1732]" box="[159,1279,1710,1732]" pageId="7" pageNumber="76">IV 2.95 1.6 2.15 2.85 1.4 10.95</paragraph>
<paragraph id="8B8A2F629665FFF4FF40F900D942F8D4" blockId="7.[151,312,1767,1792]" box="[151,312,1767,1792]" pageId="7" pageNumber="76">Leg spination:</paragraph>
<paragraph id="8B8A2F629665FFF4FF3FF8F5DB83F8E6" blockId="7.[159,1271,1820,2007]" box="[232,1017,1820,1842]" pageId="7" pageNumber="76">Fe Pt Ti Mt</paragraph>
<paragraph id="8B8A2F629665FFF4FF48F8AEDC64F88F" blockId="7.[159,1271,1820,2007]" box="[159,1054,1861,1883]" pageId="7" pageNumber="76">
I d1-0-0 p0-0-
<date id="FF8B09A29665FFF4FEBAF8ADDAF5F88F" box="[365,655,1861,1883]" pageId="7" pageNumber="76">1 v</date>
0-0-
<date id="FF8B09A29665FFF4FD6FF8ADDB92F88F" box="[696,1000,1861,1883]" pageId="7" pageNumber="76">2 v</date>
0-2-0
</paragraph>
<paragraph id="8B8A2F629665FFF4FF48F887DC64F850" blockId="7.[159,1271,1820,2007]" box="[159,1054,1902,1924]" pageId="7" pageNumber="76">
II d1-1-0 p0-0-
<date id="FF8B09A29665FFF4FEBAF886DAEBF850" box="[365,657,1902,1924]" pageId="7" pageNumber="76">1 v</date>
0-1-
<date id="FF8B09A29665FFF4FD60F886DB92F850" box="[695,1000,1902,1924]" pageId="7" pageNumber="76">2 v</date>
0-2-0
</paragraph>
<paragraph id="8B8A2F629665FFFBFF48F871D887F8D2" blockId="7.[159,1271,1820,2007]" lastBlockId="8.[151,1436,1701,1978]" lastPageId="8" lastPageNumber="77" pageId="7" pageNumber="76">
III d0-1-0 p0-1-1 r0-1-1 r1-0 p2-1-1 r1-1-
<date id="FF8B09A29665FFF4FCD7F870DB65F87A" box="[768,799,1944,1966]" pageId="7" pageNumber="76">1 v</date>
2-2-2 d0-1-0 p1-1-0 r1-1-0
<date id="FF8B09A29665FFF4FB63F870DC98F87A" box="[1204,1250,1944,1966]" pageId="7" pageNumber="76">v2-1</date>
-0 IV d1-1-0 p0-0-1 r0-0-1 p1-1-1 r2-1-
<date id="FF8B09A29665FFF4FCD6F829DB65F803" box="[769,799,1985,2007]" pageId="7" pageNumber="76">1 v</date>
2-2-2 d0-1-0 p1-1-0 r1-0-0
<date id="FF8B09A29665FFF4FB63F829DC99F803" box="[1204,1251,1985,2007]" pageId="7" pageNumber="76">v2-2</date>
-0 Epigyne as in
<figureCitation id="130E33E7966AFFFBFEB2F94DD99FF96A" box="[357,485,1701,1726]" captionStart="FIGURES 28 ‒ 33" captionStartId="9.[151,264,1714,1737]" captionTargetBox="[151,1436,368,1692]" captionTargetId="figure@9.[151,1436,363,1692]" captionTargetPageId="9" captionText="FIGURES 28 ‒ 33. Female of Gnaphosa rasnitsyni (28 ‒ 31) and G. zonsteini sp. n. (32 ‒ 33). 28 habitus, dorsal; 29 intact epigyne, ventral; 30, 32 macerated epigyne, ventral; 31, 33 macerated epigyne, dorsal. Scale = 0.2 mm if not otherwise indicated. Abbreviations. Ab foveal bulges, Lp lateral pockets, Ps pocket of the scape; Rh head of receptacle." httpUri="https://zenodo.org/record/569173/files/figure.png" pageId="8" pageNumber="77">Figs 32‒33</figureCitation>
; fovea diamond-shaped, with pair of clearly-delineated bulges (
<emphasis id="B941F370966AFFFBFB7DF94EDCB3F96A" box="[1194,1225,1702,1726]" italics="true" pageId="8" pageNumber="77">Ab</emphasis>
); scape wider than long with slightly concave posterior margin and stretched pocket of scape (
<emphasis id="B941F370966AFFFBFC27F922DC76F935" box="[1008,1036,1738,1761]" italics="true" pageId="8" pageNumber="77">Ps</emphasis>
); head of receptacles (
<emphasis id="B941F370966AFFFBFAC6F922DD4AF936" box="[1297,1328,1738,1762]" italics="true" pageId="8" pageNumber="77">Rh</emphasis>
) directed upwards.
</paragraph>
</subSubSection>
<caption id="DF4A7FEA966AFFFBFF40F9A1DC9FF9AA" httpUri="https://zenodo.org/record/569172/files/figure.png" pageId="8" pageNumber="77" targetBox="[156,1434,206,1584]" targetPageId="8">
<paragraph id="8B8A2F62966AFFFBFF40F9A1DC9FF9AA" blockId="8.[151,1436,1609,1662]" pageId="8" pageNumber="77">
<emphasis id="B941F370966AFFFBFF40F9A1D935F98B" bold="true" box="[151,335,1609,1632]" pageId="8" pageNumber="77">FIGURES 20-27.</emphasis>
Male of
<taxonomicName id="4C3554E1966AFFFBFE7DF9A2DA08F9B4" box="[426,626,1610,1632]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
<emphasis id="B941F370966AFFFBFE7DF9A2DA08F9B4" box="[426,626,1610,1632]" italics="true" pageId="8" pageNumber="77">Gnaphosa zonsteini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2724E0B966AFFFBFDAFF9A2DACAF98B" box="[632,688,1610,1631]" pageId="8" pageNumber="77" rank="species">
<emphasis id="B941F370966AFFFBFDAFF9A2DACAF98B" bold="true" box="[632,688,1610,1631]" pageId="8" pageNumber="77">sp. n.</emphasis>
</taxonomicNameLabel>
20 habitus, dorsal; 21‒23 palp, ventral, retro- and prolateral; 24 cymbium and tibia, retrolateral; 25‒27 bulb, prolateral, ventral and anterior. Scale = 0.2 mm if not otherwise indicated.
</paragraph>
</caption>
<subSubSection id="C32F7CE9966AFFFBFF10F8F9DA8BF8FE" box="[199,753,1809,1834]" pageId="8" pageNumber="77" type="distribution">
<paragraph id="8B8A2F62966AFFFBFF10F8F9DA8BF8FE" blockId="8.[151,1436,1701,1978]" box="[199,753,1809,1834]" pageId="8" pageNumber="77">
<emphasis id="B941F370966AFFFBFF10F8F9D91AF8FE" bold="true" box="[199,352,1809,1834]" pageId="8" pageNumber="77">Distribution.</emphasis>
Known from the
<typeStatus id="548E91C0966AFFFBFDF0F8FADA2DF8FE" box="[551,599,1810,1834]" pageId="8" pageNumber="77">type</typeStatus>
locality only.
</paragraph>
</subSubSection>
<subSubSection id="C32F7CE9966AFFFBFF10F8DDDCCCF86E" pageId="8" pageNumber="77" type="discussion">
<paragraph id="8B8A2F62966AFFFBFF10F8DDDCCCF86E" blockId="8.[151,1436,1701,1978]" pageId="8" pageNumber="77">
<emphasis id="B941F370966AFFFBFF10F8DDD935F89A" bold="true" box="[199,335,1845,1870]" pageId="8" pageNumber="77">Comments.</emphasis>
Judging from the combination of such features as base of embolus with spine in anterior part, filiform and rounded embolus and hook-like median apophysis it can be assumed that
<taxonomicName id="4C3554E1966AFFFBFB8DF8B2DCA6F8A5" box="[1114,1244,1882,1905]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="zonsteini" status="sp. nov.">
<emphasis id="B941F370966AFFFBFB8DF8B2DCA6F8A5" box="[1114,1244,1882,1905]" italics="true" pageId="8" pageNumber="77">G. zonsteini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2724E0B966AFFFBFB34F8B1DD58F8A6" box="[1251,1314,1881,1906]" pageId="8" pageNumber="77" rank="species">
<emphasis id="B941F370966AFFFBFB34F8B1DD58F8A6" bold="true" box="[1251,1314,1881,1906]" pageId="8" pageNumber="77">sp. n.</emphasis>
</taxonomicNameLabel>
belongs to a monophyletic species group formed by following species:
<taxonomicName id="4C3554E1966AFFFBFC9FF896DD02F842" authority="Marusik, Fomichev &amp; Omelko, 2014" authorityName="Marusik, Fomichev &amp; Omelko" authorityYear="2014" box="[840,1400,1917,1942]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="esyunini">
<emphasis id="B941F370966AFFFBFC9FF896DBBFF842" box="[840,965,1918,1942]" italics="true" pageId="8" pageNumber="77">G. esyunini</emphasis>
Marusik, Fomichev &amp; Omelko, 2014
</taxonomicName>
,
<taxonomicName id="4C3554E1966AFFFBFA53F896DAC2F86E" authority="Marusik, Fomichev &amp; Omelko, 2014" authorityName="Marusik, Fomichev &amp; Omelko" authorityYear="2014" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="khovdensis">
<emphasis id="B941F370966AFFFBFA53F896D968F86E" italics="true" pageId="8" pageNumber="77">G. khovdensis</emphasis>
Marusik, Fomichev &amp; Omelko, 2014
</taxonomicName>
and
<taxonomicName id="4C3554E1966AFFFBFD39F84ADC5FF86E" authority="Marusik, 1993" authorityName="Marusik" authorityYear="1993" box="[750,1061,1953,1978]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="rasnitsyni">
<emphasis id="B941F370966AFFFBFD39F84ADB01F86D" box="[750,891,1954,1977]" italics="true" pageId="8" pageNumber="77">G. rasnitsyni</emphasis>
<bibRefCitation id="EFA45293966AFFFBFC55F849DC5FF86E" author="Marusik" box="[898,1061,1953,1978]" pageId="8" pageNumber="77" refString="Marusik, Y. M. (1993) Three new wandering spider species (Aranei Lycosidae Gnaphosidae) from Mongolia. Arthropoda Selecta, 2, 77 - 81." type="journal article" year="1993">Marusik, 1993</bibRefCitation>
</taxonomicName>
(see below).
</paragraph>
</subSubSection>
</treatment>
</document>
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