857 lines
240 KiB
XML
857 lines
240 KiB
XML
<document id="60ED04114D621E0B8B90CD9F4675812D" ID-DOI="10.1016/j.jcz.2022.09.001" ID-ISSN="1873-2674" ID-Zenodo-Dep="10375202" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702544282064" checkinUser="felipe" docAuthor="Cepeda, Diego, González-Casarrubios, Alberto, Sánchez, Nuria, Spedicato, Adriana, Michaud, Emma & Zeppilli, Daniela" docDate="2022" docId="616487EBFFFB7755FFF2CEF3CA7BFD93" docLanguage="en" docName="ZoolAnz.301.23-41.pdf" docOrigin="Zoologischer Anzeiger 301" docSource="http://dx.doi.org/10.1016/j.jcz.2022.09.001" docStyle="DocumentStyle:32483D6D135DE2FF927BA3F1400E0FFD.1:ZoolAnz.2022-.journal_article" docStyleId="32483D6D135DE2FF927BA3F1400E0FFD" docStyleName="ZoolAnz.2022-.journal_article" docStyleVersion="1" docTitle="Echinoderes parthenope Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli 2022, sp. nov" docType="treatment" docVersion="2" lastPageNumber="33" masterDocId="9D5DFF93FFFD775FFF96C814CF14FF9E" masterDocTitle="Two new species of mud dragons (Scalidophora: Kinorhyncha) inhabiting a human-impacted mangrove from Mayotte (Southwestern Indian Ocean)" masterLastPageNumber="41" masterPageNumber="23" pageNumber="29" updateTime="1702546782698" updateUser="ExternalLinkService">
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<mods:titleInfo id="539958CCF938A2DB55B7609D6DAC6150">
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<mods:title id="43E4EEA54B4DB3CEC98F7BDFC9D84C49">Two new species of mud dragons (Scalidophora: Kinorhyncha) inhabiting a human-impacted mangrove from Mayotte (Southwestern Indian Ocean)</mods:title>
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</mods:titleInfo>
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<mods:name id="AD9D62BECDDB5C72477A99EC8A335FF5" type="personal">
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<mods:roleTerm id="7A82AF03284CED227E681C5AAA455E98">Author</mods:roleTerm>
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<mods:namePart id="25E316A988FDCE61AB786F6A38945E6C">Cepeda, Diego</mods:namePart>
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</mods:name>
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<mods:name id="E03D798608BF65DF8C19EFD0F7B5F1DD" type="personal">
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<mods:roleTerm id="3E84A3CB445A90070B5234B51319D665">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D1146027ED7D4DC0C03A1209571291CB">González-Casarrubios, Alberto</mods:namePart>
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</mods:name>
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<mods:name id="0EA83BF7D5106470F9DFB6C5CB549445" type="personal">
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<mods:roleTerm id="ACF20900189BACD19042245A12871E05">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CFC872CAC88618406BEB2866416C8683">Sánchez, Nuria</mods:namePart>
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</mods:name>
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<mods:name id="7485D11C0FA90C7A9018EF2D516A783B" type="personal">
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<mods:roleTerm id="D6C44F5FEE1FED64424B1B8D7A01B6B5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3CF7AB0EEB85C302BCB5A6DD13B3D331">Spedicato, Adriana</mods:namePart>
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</mods:name>
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<mods:name id="573A0D8A8D830E979E0282FF7AB9DC38" type="personal">
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</mods:role>
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<mods:namePart id="E4F9F2050699F7CFA4A6FBE3D50EAE6F">Michaud, Emma</mods:namePart>
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</mods:name>
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<mods:name id="CAE0BA6456F09ECCA4DDB36C49C8AAD3" type="personal">
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<mods:roleTerm id="6E62CD40E8008D2972C525CF888D0A27">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A7354221AE9ACFF18485D7729BDBDD4A">Zeppilli, Daniela</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="D832296CABE29098AAFAA3909947FE84">text</mods:typeOfResource>
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<mods:titleInfo id="1D6256867D93A5A31CDEBEFECAD95A02">
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<mods:title id="98CF59018C15A40A87A1EF746AD466C3">Zoologischer Anzeiger</mods:title>
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</mods:titleInfo>
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<mods:part id="837F960A512E88167FD28015BB730A55">
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<mods:date id="D892211276D38035751D4652CAD3DCDE">2022</mods:date>
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<mods:detail id="3810C695B15856ACF3875B4D0A699695" type="pubDate">
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<mods:number id="6862163A3355D0F350A2412B47308C42">2022-11-30</mods:number>
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</mods:detail>
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<mods:detail id="34A1D355A6CAC0147113C3E27274E7A3" type="volume">
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<mods:number id="C4B18ACAFEC38126606EFB2D4091F275">301</mods:number>
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<mods:start id="82AB0720255FF19377604F9B07C58714">23</mods:start>
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<mods:end id="FFE8C71D0710E52CF0B719C4C1C3C8D5">41</mods:end>
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<mods:location id="02F0737D51CC6C1935502A9CE15EBB8F">
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<mods:url id="25613D36FFA069A87C44B0389D04F711">http://dx.doi.org/10.1016/j.jcz.2022.09.001</mods:url>
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</mods:location>
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<mods:classification id="78409F8E79F70DB857B2DEF55D288AE4">journal article</mods:classification>
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<mods:identifier id="501031F7EF0285DFAC125B1418891116" type="DOI">10.1016/j.jcz.2022.09.001</mods:identifier>
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<mods:identifier id="4B43FCA81AA48472B377A5BEC28CC682" type="ISSN">1873-2674</mods:identifier>
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<mods:identifier id="1E3980358E774FE3D9053874CD3AE67F" type="Zenodo-Dep">10375202</mods:identifier>
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</mods:mods>
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<treatment id="616487EBFFFB7755FFF2CEF3CA7BFD93" LSID="urn:lsid:plazi:treatment:616487EBFFFB7755FFF2CEF3CA7BFD93" httpUri="http://treatment.plazi.org/id/616487EBFFFB7755FFF2CEF3CA7BFD93" lastPageId="10" lastPageNumber="33" pageId="6" pageNumber="29">
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<subSubSection id="A1D76576FFFB7759FFF2CEF3CC16F8AC" pageId="6" pageNumber="29" type="nomenclature">
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<paragraph id="E97236FDFFFB7759FFF2CEF3CEBAF965" blockId="6.[100,430,1767,1787]" box="[100,430,1767,1787]" pageId="6" pageNumber="29">
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<heading id="B23A8191FFFB7759FFF2CEF3CEBAF965" box="[100,430,1767,1787]" fontSize="36" level="2" pageId="6" pageNumber="29" reason="3">
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<emphasis id="DBB9EAEFFFFB7759FFF2CEF3CEBAF965" bold="true" box="[100,430,1767,1787]" italics="true" pageId="6" pageNumber="29">
|
||
3.2.
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||
<taxonomicName id="2ECD4D7EFFFB7759FF03CEF3CE7CF964" ID-CoL="9WJ8Y" authority="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli, 2022" authorityName="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli" authorityYear="2022" box="[149,360,1767,1786]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="6" pageNumber="29" phylum="Kinorhyncha" rank="species" species="parthenope" status="sp. nov.">Echinoderes parthenope</taxonomicName>
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<taxonomicNameLabel id="C08A5794FFFB7759FEF9CEFCCEBAF965" box="[367,430,1768,1787]" pageId="6" pageNumber="29" rank="species">sp. nov</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="E97236FDFFFB7759FF12CF0BCC16F8AC" blockId="6.[100,770,1823,1870]" box="[132,770,1823,1842]" pageId="6" pageNumber="29">Zoobank code: urn:lsid:zoobank.org:act:4E6BFE9E-D25F-4A91-</paragraph>
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</subSubSection>
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||
<subSubSection id="A1D76576FFFB7759FFF2CF2FCE25F8D0" box="[100,305,1851,1870]" pageId="6" pageNumber="29" type="description">
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<paragraph id="E97236FDFFFB7759FFF2CF2FCE25F8D0" blockId="6.[100,770,1823,1870]" box="[100,305,1851,1870]" pageId="6" pageNumber="29">99F0E5D47E37BA91.</paragraph>
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</subSubSection>
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||
<subSubSection id="A1D76576FFFB7759FFF2CF67CAC4F963" pageId="6" pageNumber="29" type="materials_examined">
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<paragraph id="E97236FDFFFB7759FFF2CF67CE5AF81B" blockId="6.[100,771,1906,1982]" box="[100,334,1906,1926]" pageId="6" pageNumber="29">
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<emphasis id="DBB9EAEFFFFB7759FFF2CF67CE5AF81B" bold="true" box="[100,334,1906,1926]" italics="true" pageId="6" pageNumber="29">3.2.1. Material examined</emphasis>
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</paragraph>
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<paragraph id="E97236FDFFFB7759FF12CF9BCC17F820" blockId="6.[100,771,1906,1982]" pageId="6" pageNumber="29">
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<materialsCitation id="59A53CA0FFFB7759FF12CF9BCC17F820" collectingDate="2018-10-06" collectorName="Archipelago, SW" country="Mayotte" location="Chirongui Bay" municipality="Grande-Terre Island" pageId="6" pageNumber="29" specimenCount="1" typeStatus="holotype">
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<typeStatus id="3676885FFFFB7759FF12CF9BCFC9F83C" box="[132,221,1935,1954]" pageId="6" pageNumber="29" type="holotype">Holotype</typeStatus>
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, adult male, collected on
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<date id="9D73103DFFFB7759FE47CF9BCD95F83C" box="[465,641,1935,1954]" pageId="6" pageNumber="29" value="2018-10-06">
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<collectingDate id="8D37E9D5FFFB7759FE47CF9BCD95F83C" box="[465,641,1935,1954]" pageId="6" pageNumber="29" value="2018-10-06">October 6th, 2018</collectingDate>
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</date>
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, at
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<location id="EC126026FFFB7759FD32CF9BCF9DF820" LSID="urn:lsid:plazi:treatment:616487EBFFFB7755FFF2CEF3CA7BFD93:EC126026FFFB7759FD32CF9BCF9DF820" country="Mayotte" municipality="Grande-Terre Island" name="Chirongui Bay" pageId="6" pageNumber="29">Chirongui Bay</location>
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,
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<collectingMunicipality id="0916AC87FFFB7759FF0CCFBFCE4BF820" box="[154,351,1963,1982]" pageId="6" pageNumber="29">Grande-Terre Island</collectingMunicipality>
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,
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<collectingCountry id="91DA766DFFFB7759FEF9CFBFCEA8F820" box="[367,444,1963,1982]" name="Mayotte" pageId="6" pageNumber="29">Mayotte</collectingCountry>
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<collectorName id="4438532BFFFB7759FE5FCFBFCD78F820" box="[457,620,1963,1982]" pageId="6" pageNumber="29">Archipelago, SW</collectorName>
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Indian Ocean:
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</materialsCitation>
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||
</paragraph>
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<paragraph id="E97236FDFFFB7759FCA4CDFACAC4F963" blockId="6.[818,1488,1515,1789]" pageId="6" pageNumber="29">
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<materialsCitation id="59A53CA0FFFB7759FCA4CDFACAD9F963" collectingDate="2018-10-06" collectionCode="MNHN, UCM" collectorName="Fluoromount G & Ma." country="Mayotte" location="Chirongui Bay" municipality="Grande-Terre Island" pageId="6" pageNumber="29" specimenCount="17" specimenCount-adult="7" specimenCount-juvenile="10" typeStatus="paratype">
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12
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<superScript id="1EB89BB5FFFB7759FCDFCDF8CC45FA66" attach="none" box="[841,849,1516,1528]" fontSize="6" pageId="6" pageNumber="29">Ǫ</superScript>
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55
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<superScript id="1EB89BB5FFFB7759FCFCCDFFCC7BFA66" attach="none" box="[874,879,1515,1528]" fontSize="6" pageId="6" pageNumber="29">′</superScript>
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17.5
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<superScript id="1EB89BB5FFFB7759FC0FCDFFCCB6FA66" attach="right" box="[921,930,1515,1528]" fontSize="6" pageId="6" pageNumber="29">′′</superScript>
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S, 45
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<superScript id="1EB89BB5FFFB7759FC48CDF8CCF2FA66" attach="none" box="[990,998,1516,1528]" fontSize="6" pageId="6" pageNumber="29">Ǫ</superScript>
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09
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<superScript id="1EB89BB5FFFB7759FC69CDFFCB10FA66" attach="none" box="[1023,1028,1515,1528]" fontSize="6" pageId="6" pageNumber="29">′</superScript>
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10.4
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<superScript id="1EB89BB5FFFB7759FBB8CDFFCB23FA66" attach="right" box="[1070,1079,1515,1528]" fontSize="6" pageId="6" pageNumber="29">′′</superScript>
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E 12
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<superScript id="1EB89BB5FFFB7759FBFACDF8CB60FA66" attach="none" box="[1132,1140,1516,1528]" fontSize="6" pageId="6" pageNumber="29">Ǫ</superScript>
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55
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<superScript id="1EB89BB5FFFB7759FB1BCDFFCB86FA66" attach="none" box="[1165,1170,1515,1528]" fontSize="6" pageId="6" pageNumber="29">′</superScript>
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22.8
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<superScript id="1EB89BB5FFFB7759FB2ACDFFCBD1FA66" attach="right" box="[1212,1221,1515,1528]" fontSize="6" pageId="6" pageNumber="29">′′</superScript>
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S, 45
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<superScript id="1EB89BB5FFFB7759FA96CDF8CA1CFA66" attach="none" box="[1280,1288,1516,1528]" fontSize="6" pageId="6" pageNumber="29">Ǫ</superScript>
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09
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<superScript id="1EB89BB5FFFB7759FAB7CDFFCA32FA66" attach="none" box="[1313,1318,1515,1528]" fontSize="6" pageId="6" pageNumber="29">′</superScript>
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09.1
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<superScript id="1EB89BB5FFFB7759FAC6CDFFCA4DFA66" attach="right" box="[1360,1369,1515,1528]" fontSize="6" pageId="6" pageNumber="29">′′</superScript>
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E in black organic mud at the intertidal zone; mounted in
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<collectorName id="4438532BFFFB7759FAB2CE1ECAADF983" box="[1316,1465,1546,1565]" pageId="6" pageNumber="29">Fluoromount G</collectorName>
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®, deposited at
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<collectionCode id="8FDCAE38FFFB7759FC3BCE32CCFAF9A7" box="[941,1006,1574,1593]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="29">MNHN</collectionCode>
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under catalogue number: 613
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<collectorName id="4438532BFFFB7759FA8ECE32CA29F9A7" box="[1304,1341,1574,1593]" pageId="6" pageNumber="29">Ma.</collectorName>
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<typeStatus id="3676885FFFFB7759FAD5CE32CAB5F9A7" box="[1347,1441,1574,1593]" pageId="6" pageNumber="29" type="paratype">Paratypes</typeStatus>
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,
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<specimenCount id="FFCBFD74FFFB7759FA3ACE32CC76F9CB" count="5" pageId="6" pageNumber="29" type="adult">five adult</specimenCount>
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males and
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<specimenCount id="FFCBFD74FFFB7759FC41CE56CB26F9CB" box="[983,1074,1602,1621]" count="1" pageId="6" pageNumber="29" type="adult">one adult</specimenCount>
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female, same collecting data as holotype; mounted in
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<collectorName id="4438532BFFFB7759FC38CE4ACB57F9EF" box="[942,1091,1630,1649]" pageId="6" pageNumber="29">Fluoromount G</collectorName>
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||
®, deposited at
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<collectionCode id="8FDCAE38FFFB7759FB71CE4ACA3CF9EF" box="[1255,1320,1630,1649]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="29">MNHN</collectionCode>
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under catalogue numbers: 637 Ma, 641 Ma-643 Ma, 649 Ma and 651 Ma.
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<typeStatus id="3676885FFFFB7759FAFACE6ECC97F937" isOtherTypeStatus="true" pageId="6" pageNumber="29">Additional material</typeStatus>
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,
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<specimenCount id="FFCBFD74FFFB7759FC1CCE82CCF5F937" box="[906,993,1686,1705]" count="1" pageId="6" pageNumber="29" type="adult">one adult</specimenCount>
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male, same collecting data as type material; mounted for SEM, stored at the Meiofauna collection of the
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<collectionCode id="8FDCAE38FFFB7759FA8CCEA6CA58F95B" box="[1306,1356,1714,1733]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" lsid="urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="6" pageNumber="29" type="University or college">UCM</collectionCode>
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||
;
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<specimenCount id="FFCBFD74FFFB7759FAC0CEA6CAD8F95B" box="[1366,1484,1714,1733]" count="10" pageId="6" pageNumber="29" type="juvenile">10 juveniles</specimenCount>
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, same collecting data as type material, deposited at
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<collectionCode id="8FDCAE38FFFB7759FA8DCEDACA48F97F" box="[1307,1372,1742,1761]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="29">MNHN</collectionCode>
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under catalogue numbers: 638 Ma-640 Ma, 644 Ma-648 Ma, 650 Ma and 652 Ma
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="A1D76576FFFB7758FCA4CF35CE6EF894" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29" type="diagnosis">
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||
<paragraph id="E97236FDFFFB7759FCA4CF35CCDCF8AA" blockId="6.[818,1487,1825,1956]" box="[818,968,1825,1844]" pageId="6" pageNumber="29">
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||
<emphasis id="DBB9EAEFFFFB7759FCA4CF35CCDCF8AA" bold="true" box="[818,968,1825,1844]" italics="true" pageId="6" pageNumber="29">3.2.2. Diagnosis</emphasis>
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</paragraph>
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<paragraph id="E97236FDFFFB7758FCC7CF29CE6EF894" blockId="6.[818,1487,1825,1956]" lastBlockId="7.[100,770,1448,1802]" lastPageId="7" lastPageNumber="30" pageId="6" pageNumber="29">
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<taxonomicName id="2ECD4D7EFFFB7759FCC7CF29CCAEF8CE" authorityName="Claparede" authorityYear="1863" box="[849,954,1853,1872]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="6" pageNumber="29" phylum="Kinorhyncha" rank="genus">
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<emphasis id="DBB9EAEFFFFB7759FCC7CF29CCAEF8CE" bold="true" box="[849,954,1853,1872]" italics="true" pageId="6" pageNumber="29">Echinoderes</emphasis>
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</taxonomicName>
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with spines that are short, poorly sclerotized, weakly articulated in middorsal position on segment 4, lateroventral position on segment 6, sublateral position on segment 7 and lateral accessory position on segment 8, plus large tubes in lateroventral position on segment 5, sublateral position on segment 8 and laterodorsal position on segment 10 (only in males). Minute, rounded
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<typeStatus id="3676885FFFFA7758FDA5CDD0CD48FA49" box="[563,604,1476,1495]" pageId="7" pageNumber="30">type</typeStatus>
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2 glandular cell outlets in subdorsal position on segments 5–6, in laterodorsal position on segment 9, in midlateral position on segment 4, in sublateral position on segment 7, and in lateral accessory position on segment 6. Enlarged, triangular sieve plates in sublateral position on segment 9, consisting of a slightly convex region bearing numerous pores followed by a small, slightly concave area bearing a single, enlarged pore. Males with tergal plate of segment 10 forming a pair of cuticular, rectangular extensions in subdorsal position, with straight, abruptly tapering, slightly curved tips; females with tufts of hairs instead. Males with tergal plate of segment 11 forming a middorsal, triangular, bulged process that extends slightly beyond the tergal extensions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="BDB26675FFFA7758FFF2CCFACB8DFAE1" ID-DOI="http://doi.org/10.5281/zenodo.10375210" ID-Zenodo-Dep="10375210" httpUri="https://zenodo.org/record/10375210/files/figure.png" pageId="7" pageNumber="30" startId="7.[100,130,1262,1279]" targetBox="[208,1379,150,1233]" targetPageId="7" targetType="figure">
|
||
<paragraph id="E97236FDFFFA7758FFF2CCFACB8DFAE1" blockId="7.[100,1488,1262,1407]" pageId="7" pageNumber="30">
|
||
<emphasis id="DBB9EAEFFFFA7758FFF2CCFACF88FA9E" bold="true" box="[100,156,1262,1280]" pageId="7" pageNumber="30">Fig. 4.</emphasis>
|
||
Light micrographs of male holotype MNHN-623 Ma (B–C, E), male paratype MNHN-599 Ma (A), and female paratypes MNHN-602 Ma (D) and MNHN-616 Ma (F) of
|
||
<taxonomicName id="2ECD4D7EFFFA7758FF0FCD1CCE75FA87" authority="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli, 2022" authorityName="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli" authorityYear="2022" box="[153,353,1288,1305]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="30" phylum="Kinorhyncha" rank="species" species="cyaneafictus" status="sp. nov.">
|
||
<emphasis id="DBB9EAEFFFFA7758FF0FCD1CCE75FA87" bold="true" box="[153,353,1288,1305]" italics="true" pageId="7" pageNumber="30">Echinoderes cyaneafictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C08A5794FFFA7758FEF0CD1CCEBFFA86" box="[358,427,1288,1304]" pageId="7" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, showing trunk overview and details on the nephridiopore and segments 5–11. A: Ventral trunk overview; B: Dorsal view on segments 5–8; C; Dorsal view on segments 8–10; D: Lateral terminal and lateral accessory terminal spines; E: Ventral view on segments 10–11; F: Overview of nephridiopore in midlateral view. Abbreviations: epo, enlarged pore; f, female condition of sexually dimorphic character; ltas, lateral terminal accessory spines; lts, lateral terminal spines; m, male condition of sexually dimorphic character; mlne, midlateral nephridiopore; ppf, primary pectinate fringe; ps, penile spine; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A1D76576FFFA7755FFF2CF3BCA7BFD93" lastPageId="10" lastPageNumber="33" pageId="7" pageNumber="30" type="description">
|
||
<paragraph id="E97236FDFFFA7758FFF2CF3BCE17F8DF" blockId="7.[100,770,1838,1970]" box="[100,259,1838,1858]" pageId="7" pageNumber="30">
|
||
<emphasis id="DBB9EAEFFFFA7758FFF2CF3BCE17F8DF" bold="true" box="[100,259,1838,1858]" italics="true" pageId="7" pageNumber="30">3.2.3. Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFFA7758FF12CF5FCCACF991" blockId="7.[100,770,1838,1970]" lastBlockId="7.[818,1488,1448,1551]" pageId="7" pageNumber="30">The species is dedicated to the Greek, mythological mermaid Parthenope (from the Ancient Greek Παρθενόπη). Parthenope gave name to the first Greek settlement now part of the Italian city of Naples. According to legend, Parthenope despaired after failing to lure Odysseus to her island, cast herself into the sea and drowned, her body washed ashore near Naples. The mermaid is usually represented in art with two tails, which resembles in shape the characteristic segment 11 tergal plate of the species.</paragraph>
|
||
<paragraph id="E97236FDFFFA7758FCA4CE20CCC3F9D9" blockId="7.[818,1488,1588,1970]" box="[818,983,1588,1607]" pageId="7" pageNumber="30">
|
||
<emphasis id="DBB9EAEFFFFA7758FCA4CE20CCC3F9D9" bold="true" box="[818,983,1588,1607]" italics="true" pageId="7" pageNumber="30">3.2.4. Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFFA7758FCC7CE44CCF9F928" blockId="7.[818,1488,1588,1970]" pageId="7" pageNumber="30">
|
||
See
|
||
<tableCitation id="A44F0346FFFA7758FCEFCE5BCCD4F9FD" box="[889,960,1615,1635]" captionStart="Table 4" captionStartId="8.[100,150,150,166]" captionText="Table 4 Morphological measurements and dimensions (in μm) of adult specimens of Echinoderes parthenope sp. nov. from the type locality, including number of measured specimens (n), mean value and standard deviation (Sd) of each feature, depicted for the holotype and all the type series. Abbreviations: ac, acicular spine; LA, lateral accessory; LD, laterodorsal; LTAS, lateral terminal accessory spine length; LTS, lateral terminal spine length; LV, lateroventral; MD, middorsal; MSW,maximum sternal width; S, segment length; SL, sublateral; SW, standard sternal width; t, tube; TL, total trunk length; numbers after abbreviations indicate the corresponding segment." pageId="7" pageNumber="30">Table 4</tableCitation>
|
||
for measurements of selected morphological traits and dimensions, and
|
||
<tableCitation id="A44F0346FFFA7758FC41CE7FCB35F9E0" box="[983,1057,1643,1662]" captionStart="Table 5" captionStartId="8.[100,150,1526,1542]" captionText="Table 5 Summary of nature and arrangement of spines, tubes, sensory spots, glandular cell outlets, nephridiopores and other cuticular structures in adults of Echinoderes parthenope sp. nov. Abbreviations: ac, acicular spine; ce, cuticular extension; cp, cuticular process; dss, droplet-shaped sensory spot; gcoI/II, glandular cell outlet type 1/2; LA, lateral accessory; LD, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; LV, lateroventral; MD, middorsal; ML, midlateral; ne, nephridiopore; PD, paradorsal; ps, penile spines; rss, rounded sensory spot; SD, subdorsal; SL, sublateral; th, tuft of hairs; t, tube; VL, ventrolateral; VM, ventromedial; Ƌ/♀ indicate sexually dimorphic structures." pageId="7" pageNumber="30">Table 5</tableCitation>
|
||
for summary of acicular spine, tube, nephridiopore, glandular cell outlet, sensory spot, hair tuft and cuticular extension locations.
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFFA7757FCC7CEABCE8DFBF3" blockId="7.[818,1488,1588,1970]" lastBlockId="8.[100,770,1114,1468]" lastPageId="8" lastPageNumber="31" pageId="7" pageNumber="30">
|
||
Head: a single specimen for LM was found with a completely everted head, hence only a few details in some structures can be provided. Ring 00 of mouth cone with nine outer oral styles alternating in size between slightly larger and smaller ones. Outer oral styles composed of two jointed subunits: a rectangular, basal piece with two proximal fringes located at their lateral edges with a couple of tips each one; and a triangular, hooked, distally pointed and curved end-piece (
|
||
<figureCitation id="71F62A78FFFA7758FAEDCF72CAD0F8E4" box="[1403,1476,1894,1914]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="7" pageNumber="30">Fig. 6B</figureCitation>
|
||
). Triangular, poorly sclerotized cuticular thickenings flanking the outer oral styles. Outer oral styles located anterior to each introvert sector, except in the middorsal sector 6.
|
||
</paragraph>
|
||
<caption id="BDB26675FFF57757FFF2C882CEDDFE12" pageId="8" pageNumber="31" startId="8.[100,150,150,166]" targetType="table">
|
||
<paragraph id="E97236FDFFF57757FFF2C882CFB3FF39" blockId="8.[100,771,150,397]" box="[100,167,150,167]" pageId="8" pageNumber="31">
|
||
<emphasis id="DBB9EAEFFFF57757FFF2C882CFB3FF39" bold="true" box="[100,167,150,167]" pageId="8" pageNumber="31">Table 4</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FFF2C8A4CEDDFE12" blockId="8.[100,771,150,397]" pageId="8" pageNumber="31">
|
||
Morphological measurements and dimensions (in μm) of adult specimens of
|
||
<taxonomicName id="2ECD4D7EFFF57757FFF2C8DDCE30FF44" authority="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli, 2022" authorityName="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli" authorityYear="2022" box="[100,292,201,218]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="31" phylum="Kinorhyncha" rank="species" species="parthenope" status="sp. nov.">
|
||
<emphasis id="DBB9EAEFFFF57757FFF2C8DDCE30FF44" bold="true" box="[100,292,201,218]" italics="true" pageId="8" pageNumber="31">Echinoderes parthenope</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C08A5794FFF57757FEB8C8DECE61FF44" box="[302,373,202,218]" pageId="8" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the type locality, including number of measured specimens (
|
||
<emphasis id="DBB9EAEFFFF57757FEBEC8F7CE26FF6D" bold="true" box="[296,306,227,243]" italics="true" pageId="8" pageNumber="31">n</emphasis>
|
||
), mean value and standard deviation (Sd) of each feature, depicted for the holotype and all the type series. Abbreviations: ac, acicular spine; LA, lateral accessory; LD, laterodorsal; LTAS, lateral terminal accessory spine length; LTS, lateral terminal spine length; LV, lateroventral; MD, middorsal; MSW,maximum sternal width; S, segment length; SL, sublateral; SW, standard sternal width; t, tube; TL, total trunk length; numbers after abbreviations indicate the corresponding segment.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E97236FDFFF57757FFE2C9B5CDFDFBBA" pageId="8" pageNumber="31">
|
||
<table id="9BCDC45DFFF588A0FFE2C9B5CDE6FBBA" box="[116,754,417,1060]" gridcols="5" gridrows="28" pageId="8" pageNumber="31">
|
||
<tr id="57FD34BFFFF588A0FFE2C9B5CDE6FE31" box="[116,754,417,431]" gridrow="0" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2C9B5CFF3FE31" box="[116,231,417,431]" gridcol="0" gridrow="0" pageId="8" pageNumber="31">Character</th>
|
||
<th id="142C5DC3FFF588A0FEBFC9B5CE7AFE31" box="[297,366,417,431]" gridcol="1" gridrow="0" pageId="8" pageNumber="31">Holotype</th>
|
||
<th id="142C5DC3FFF588A0FE39C9B5CD1AFE31" box="[431,526,417,431]" gridcol="2" gridrow="0" pageId="8" pageNumber="31">Range</th>
|
||
<th id="142C5DC3FFF588A0FDC6C9B5CD6FFE31" box="[592,635,417,431]" gridcol="3" gridrow="0" pageId="8" pageNumber="31">Mean</th>
|
||
<th id="142C5DC3FFF588A0FD2BC9B5CDE6FE31" box="[701,754,417,431]" gridcol="4" gridrow="0" pageId="8" pageNumber="31">
|
||
Sd,
|
||
<emphasis id="DBB9EAEFFFF57757FD4FC9B5CDF6FE31" bold="true" box="[729,738,417,431]" italics="true" pageId="8" pageNumber="31">n</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2C9D0CDE6FE4C" box="[116,754,452,466]" gridrow="1" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2C9D0CFF3FE4C" box="[116,231,452,466]" gridcol="0" gridrow="1" pageId="8" pageNumber="31">TL</th>
|
||
<td id="142C5DC3FFF588A0FEBFC9D0CE7AFE4C" box="[297,366,452,466]" gridcol="1" gridrow="1" pageId="8" pageNumber="31">310.7</td>
|
||
<td id="142C5DC3FFF588A0FE39C9D0CD1AFE4C" box="[431,526,452,466]" gridcol="2" gridrow="1" pageId="8" pageNumber="31">217.0–312.9</td>
|
||
<td id="142C5DC3FFF588A0FDC6C9D0CD6FFE4C" box="[592,635,452,466]" gridcol="3" gridrow="1" pageId="8" pageNumber="31">285.5</td>
|
||
<td id="142C5DC3FFF588A0FD2BC9D0CDE6FE4C" box="[701,754,452,466]" gridcol="4" gridrow="1" pageId="8" pageNumber="31">30.4, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2C9CECDE6FE77" box="[116,754,474,489]" gridrow="2" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2C9CECFF3FE77" box="[116,231,474,489]" gridcol="0" gridrow="2" pageId="8" pageNumber="31">MSW-8</th>
|
||
<td id="142C5DC3FFF588A0FEBFC9CECE7AFE77" box="[297,366,474,489]" gridcol="1" gridrow="2" pageId="8" pageNumber="31">57.0</td>
|
||
<td id="142C5DC3FFF588A0FE39C9CECD1AFE77" box="[431,526,474,489]" gridcol="2" gridrow="2" pageId="8" pageNumber="31">57.0–66.7</td>
|
||
<td id="142C5DC3FFF588A0FDC6C9CECD6FFE77" box="[592,635,474,489]" gridcol="3" gridrow="2" pageId="8" pageNumber="31">61.5</td>
|
||
<td id="142C5DC3FFF588A0FD2BC9CECDE6FE77" box="[701,754,474,489]" gridcol="4" gridrow="2" pageId="8" pageNumber="31">2.4, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2C9E6CDE6FD9E" box="[116,754,498,512]" gridrow="3" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2C9E6CFF3FD9E" box="[116,231,498,512]" gridcol="0" gridrow="3" pageId="8" pageNumber="31">MSW-8/TL (%)</th>
|
||
<td id="142C5DC3FFF588A0FEBFC9E6CE7AFD9E" box="[297,366,498,512]" gridcol="1" gridrow="3" pageId="8" pageNumber="31">18.3</td>
|
||
<td id="142C5DC3FFF588A0FE39C9E6CD1AFD9E" box="[431,526,498,512]" gridcol="2" gridrow="3" pageId="8" pageNumber="31">18.3–28.3</td>
|
||
<td id="142C5DC3FFF588A0FDC6C9E6CD6FFD9E" box="[592,635,498,512]" gridcol="3" gridrow="3" pageId="8" pageNumber="31">21.9</td>
|
||
<td id="142C5DC3FFF588A0FD2BC9E6CDE6FD9E" box="[701,754,498,512]" gridcol="4" gridrow="3" pageId="8" pageNumber="31">2.2, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CA1CCDE6FD89" box="[116,754,520,535]" gridrow="4" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CA1CCFF3FD89" box="[116,231,520,535]" gridcol="0" gridrow="4" pageId="8" pageNumber="31">SW-10</th>
|
||
<td id="142C5DC3FFF588A0FEBFCA1CCE7AFD89" box="[297,366,520,535]" gridcol="1" gridrow="4" pageId="8" pageNumber="31">35.5</td>
|
||
<td id="142C5DC3FFF588A0FE39CA1CCD1AFD89" box="[431,526,520,535]" gridcol="2" gridrow="4" pageId="8" pageNumber="31">35.5–55.3</td>
|
||
<td id="142C5DC3FFF588A0FDC6CA1CCD6FFD89" box="[592,635,520,535]" gridcol="3" gridrow="4" pageId="8" pageNumber="31">48.5</td>
|
||
<td id="142C5DC3FFF588A0FD2BCA1CCDE6FD89" box="[701,754,520,535]" gridcol="4" gridrow="4" pageId="8" pageNumber="31">5.3, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CA0BCDE6FDB3" box="[116,754,543,557]" gridrow="5" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CA0BCFF3FDB3" box="[116,231,543,557]" gridcol="0" gridrow="5" pageId="8" pageNumber="31">SW-10/TL (%)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCA0BCE7AFDB3" box="[297,366,543,557]" gridcol="1" gridrow="5" pageId="8" pageNumber="31">11.4</td>
|
||
<td id="142C5DC3FFF588A0FE39CA0BCD1AFDB3" box="[431,526,543,557]" gridcol="2" gridrow="5" pageId="8" pageNumber="31">11.4–21.4</td>
|
||
<td id="142C5DC3FFF588A0FDC6CA0BCD6FFDB3" box="[592,635,543,557]" gridcol="3" gridrow="5" pageId="8" pageNumber="31">17.3</td>
|
||
<td id="142C5DC3FFF588A0FD2BCA0BCDE6FDB3" box="[701,754,543,557]" gridcol="4" gridrow="5" pageId="8" pageNumber="31">2.6, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CA22CDE6FDDA" box="[116,754,566,580]" gridrow="6" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CA22CFF3FDDA" box="[116,231,566,580]" gridcol="0" gridrow="6" pageId="8" pageNumber="31">S1</th>
|
||
<td id="142C5DC3FFF588A0FEBFCA22CE7AFDDA" box="[297,366,566,580]" gridcol="1" gridrow="6" pageId="8" pageNumber="31">31.7</td>
|
||
<td id="142C5DC3FFF588A0FE39CA22CD1AFDDA" box="[431,526,566,580]" gridcol="2" gridrow="6" pageId="8" pageNumber="31">26.4–37.8</td>
|
||
<td id="142C5DC3FFF588A0FDC6CA22CD6FFDDA" box="[592,635,566,580]" gridcol="3" gridrow="6" pageId="8" pageNumber="31">33.1</td>
|
||
<td id="142C5DC3FFF588A0FD2BCA22CDE6FDDA" box="[701,754,566,580]" gridcol="4" gridrow="6" pageId="8" pageNumber="31">2.8, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CA59CDE6FDC5" box="[116,754,589,603]" gridrow="7" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CA59CFF3FDC5" box="[116,231,589,603]" gridcol="0" gridrow="7" pageId="8" pageNumber="31">S2</th>
|
||
<td id="142C5DC3FFF588A0FEBFCA59CE7AFDC5" box="[297,366,589,603]" gridcol="1" gridrow="7" pageId="8" pageNumber="31">25.2</td>
|
||
<td id="142C5DC3FFF588A0FE39CA59CD1AFDC5" box="[431,526,589,603]" gridcol="2" gridrow="7" pageId="8" pageNumber="31">19.9–28.4</td>
|
||
<td id="142C5DC3FFF588A0FDC6CA59CD6FFDC5" box="[592,635,589,603]" gridcol="3" gridrow="7" pageId="8" pageNumber="31">23.9</td>
|
||
<td id="142C5DC3FFF588A0FD2BCA59CDE6FDC5" box="[701,754,589,603]" gridcol="4" gridrow="7" pageId="8" pageNumber="31">2.7, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CA70CDE6FDEC" box="[116,754,612,626]" gridrow="8" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CA70CFF3FDEC" box="[116,231,612,626]" gridcol="0" gridrow="8" pageId="8" pageNumber="31">S3</th>
|
||
<td id="142C5DC3FFF588A0FEBFCA70CE7AFDEC" box="[297,366,612,626]" gridcol="1" gridrow="8" pageId="8" pageNumber="31">25.3</td>
|
||
<td id="142C5DC3FFF588A0FE39CA70CD1AFDEC" box="[431,526,612,626]" gridcol="2" gridrow="8" pageId="8" pageNumber="31">19.7–29.9</td>
|
||
<td id="142C5DC3FFF588A0FDC6CA70CD6FFDEC" box="[592,635,612,626]" gridcol="3" gridrow="8" pageId="8" pageNumber="31">25.1</td>
|
||
<td id="142C5DC3FFF588A0FD2BCA70CDE6FDEC" box="[701,754,612,626]" gridcol="4" gridrow="8" pageId="8" pageNumber="31">2.8, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CA6FCDE6FD17" box="[116,754,635,649]" gridrow="9" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CA6FCFF3FD17" box="[116,231,635,649]" gridcol="0" gridrow="9" pageId="8" pageNumber="31">S4</th>
|
||
<td id="142C5DC3FFF588A0FEBFCA6FCE7AFD17" box="[297,366,635,649]" gridcol="1" gridrow="9" pageId="8" pageNumber="31">27.9</td>
|
||
<td id="142C5DC3FFF588A0FE39CA6FCD1AFD17" box="[431,526,635,649]" gridcol="2" gridrow="9" pageId="8" pageNumber="31">22.2–38.4</td>
|
||
<td id="142C5DC3FFF588A0FDC6CA6FCD6FFD17" box="[592,635,635,649]" gridcol="3" gridrow="9" pageId="8" pageNumber="31">28.7</td>
|
||
<td id="142C5DC3FFF588A0FD2BCA6FCDE6FD17" box="[701,754,635,649]" gridcol="4" gridrow="9" pageId="8" pageNumber="31">3.9, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CA85CDE6FD3E" box="[116,754,657,672]" gridrow="10" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CA85CFF3FD3E" box="[116,231,657,672]" gridcol="0" gridrow="10" pageId="8" pageNumber="31">S5</th>
|
||
<td id="142C5DC3FFF588A0FEBFCA85CE7AFD3E" box="[297,366,657,672]" gridcol="1" gridrow="10" pageId="8" pageNumber="31">29.6</td>
|
||
<td id="142C5DC3FFF588A0FE39CA85CD1AFD3E" box="[431,526,657,672]" gridcol="2" gridrow="10" pageId="8" pageNumber="31">22.5–32.5</td>
|
||
<td id="142C5DC3FFF588A0FDC6CA85CD6FFD3E" box="[592,635,657,672]" gridcol="3" gridrow="10" pageId="8" pageNumber="31">28.7</td>
|
||
<td id="142C5DC3FFF588A0FD2BCA85CDE6FD3E" box="[701,754,657,672]" gridcol="4" gridrow="10" pageId="8" pageNumber="31">3.0, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CABCCDE6FD29" box="[116,754,680,695]" gridrow="11" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CABCCFF3FD29" box="[116,231,680,695]" gridcol="0" gridrow="11" pageId="8" pageNumber="31">S6</th>
|
||
<td id="142C5DC3FFF588A0FEBFCABCCE7AFD29" box="[297,366,680,695]" gridcol="1" gridrow="11" pageId="8" pageNumber="31">31.7</td>
|
||
<td id="142C5DC3FFF588A0FE39CABCCD1AFD29" box="[431,526,680,695]" gridcol="2" gridrow="11" pageId="8" pageNumber="31">21.8–35.1</td>
|
||
<td id="142C5DC3FFF588A0FDC6CABCCD6FFD29" box="[592,635,680,695]" gridcol="3" gridrow="11" pageId="8" pageNumber="31">30.8</td>
|
||
<td id="142C5DC3FFF588A0FD2BCABCCDE6FD29" box="[701,754,680,695]" gridcol="4" gridrow="11" pageId="8" pageNumber="31">4.8, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CAABCDE6FD53" box="[116,754,703,717]" gridrow="12" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CAABCFF3FD53" box="[116,231,703,717]" gridcol="0" gridrow="12" pageId="8" pageNumber="31">S7</th>
|
||
<td id="142C5DC3FFF588A0FEBFCAABCE7AFD53" box="[297,366,703,717]" gridcol="1" gridrow="12" pageId="8" pageNumber="31">36.3</td>
|
||
<td id="142C5DC3FFF588A0FE39CAABCD1AFD53" box="[431,526,703,717]" gridcol="2" gridrow="12" pageId="8" pageNumber="31">29.8–40.8</td>
|
||
<td id="142C5DC3FFF588A0FDC6CAABCD6FFD53" box="[592,635,703,717]" gridcol="3" gridrow="12" pageId="8" pageNumber="31">36.7</td>
|
||
<td id="142C5DC3FFF588A0FD2BCAABCDE6FD53" box="[701,754,703,717]" gridcol="4" gridrow="12" pageId="8" pageNumber="31">3.5, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CAC2CDE6FD7A" box="[116,754,726,740]" gridrow="13" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CAC2CFF3FD7A" box="[116,231,726,740]" gridcol="0" gridrow="13" pageId="8" pageNumber="31">S8</th>
|
||
<td id="142C5DC3FFF588A0FEBFCAC2CE7AFD7A" box="[297,366,726,740]" gridcol="1" gridrow="13" pageId="8" pageNumber="31">44.1</td>
|
||
<td id="142C5DC3FFF588A0FE39CAC2CD1AFD7A" box="[431,526,726,740]" gridcol="2" gridrow="13" pageId="8" pageNumber="31">44.1–49.5</td>
|
||
<td id="142C5DC3FFF588A0FDC6CAC2CD6FFD7A" box="[592,635,726,740]" gridcol="3" gridrow="13" pageId="8" pageNumber="31">46.7</td>
|
||
<td id="142C5DC3FFF588A0FD2BCAC2CDE6FD7A" box="[701,754,726,740]" gridcol="4" gridrow="13" pageId="8" pageNumber="31">1.9, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CAF9CDE6FD65" box="[116,754,749,763]" gridrow="14" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CAF9CFF3FD65" box="[116,231,749,763]" gridcol="0" gridrow="14" pageId="8" pageNumber="31">S9</th>
|
||
<td id="142C5DC3FFF588A0FEBFCAF9CE7AFD65" box="[297,366,749,763]" gridcol="1" gridrow="14" pageId="8" pageNumber="31">51.2</td>
|
||
<td id="142C5DC3FFF588A0FE39CAF9CD1AFD65" box="[431,526,749,763]" gridcol="2" gridrow="14" pageId="8" pageNumber="31">51.2–59.8</td>
|
||
<td id="142C5DC3FFF588A0FDC6CAF9CD6FFD65" box="[592,635,749,763]" gridcol="3" gridrow="14" pageId="8" pageNumber="31">54.9</td>
|
||
<td id="142C5DC3FFF588A0FD2BCAF9CDE6FD65" box="[701,754,749,763]" gridcol="4" gridrow="14" pageId="8" pageNumber="31">2.8, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CB10CDE6FC8C" box="[116,754,772,786]" gridrow="15" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CB10CFF3FC8C" box="[116,231,772,786]" gridcol="0" gridrow="15" pageId="8" pageNumber="31">S10</th>
|
||
<td id="142C5DC3FFF588A0FEBFCB10CE7AFC8C" box="[297,366,772,786]" gridcol="1" gridrow="15" pageId="8" pageNumber="31">26.2</td>
|
||
<td id="142C5DC3FFF588A0FE39CB10CD1AFC8C" box="[431,526,772,786]" gridcol="2" gridrow="15" pageId="8" pageNumber="31">24.0–36.9</td>
|
||
<td id="142C5DC3FFF588A0FDC6CB10CD6FFC8C" box="[592,635,772,786]" gridcol="3" gridrow="15" pageId="8" pageNumber="31">29.1</td>
|
||
<td id="142C5DC3FFF588A0FD2BCB10CDE6FC8C" box="[701,754,772,786]" gridcol="4" gridrow="15" pageId="8" pageNumber="31">4.8, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CB0ECDE6FCB7" box="[116,754,794,809]" gridrow="16" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CB0ECFF3FCB7" box="[116,231,794,809]" gridcol="0" gridrow="16" pageId="8" pageNumber="31">S11</th>
|
||
<td id="142C5DC3FFF588A0FEBFCB0ECE7AFCB7" box="[297,366,794,809]" gridcol="1" gridrow="16" pageId="8" pageNumber="31">24.5</td>
|
||
<td id="142C5DC3FFF588A0FE39CB0ECD1AFCB7" box="[431,526,794,809]" gridcol="2" gridrow="16" pageId="8" pageNumber="31">20.2–26.5</td>
|
||
<td id="142C5DC3FFF588A0FDC6CB0ECD6FFCB7" box="[592,635,794,809]" gridcol="3" gridrow="16" pageId="8" pageNumber="31">23.4</td>
|
||
<td id="142C5DC3FFF588A0FD2BCB0ECDE6FCB7" box="[701,754,794,809]" gridcol="4" gridrow="16" pageId="8" pageNumber="31">1.4, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CB25CDE6FCDE" box="[116,754,817,832]" gridrow="17" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CB25CFF3FCDE" box="[116,231,817,832]" gridcol="0" gridrow="17" pageId="8" pageNumber="31">MD4 (ac)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCB25CE7AFCDE" box="[297,366,817,832]" gridcol="1" gridrow="17" pageId="8" pageNumber="31">6.1</td>
|
||
<td id="142C5DC3FFF588A0FE39CB25CD1AFCDE" box="[431,526,817,832]" gridcol="2" gridrow="17" pageId="8" pageNumber="31">6.1–10.6</td>
|
||
<td id="142C5DC3FFF588A0FDC6CB25CD6FFCDE" box="[592,635,817,832]" gridcol="3" gridrow="17" pageId="8" pageNumber="31">8.6</td>
|
||
<td id="142C5DC3FFF588A0FD2BCB25CDE6FCDE" box="[701,754,817,832]" gridcol="4" gridrow="17" pageId="8" pageNumber="31">1.6, 6</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CB5CCDE6FCC9" box="[116,754,840,855]" gridrow="18" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CB5CCFF3FCC9" box="[116,231,840,855]" gridcol="0" gridrow="18" pageId="8" pageNumber="31">LV5 (t)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCB5CCE7AFCC9" box="[297,366,840,855]" gridcol="1" gridrow="18" pageId="8" pageNumber="31">21.4</td>
|
||
<td id="142C5DC3FFF588A0FE39CB5CCD1AFCC9" box="[431,526,840,855]" gridcol="2" gridrow="18" pageId="8" pageNumber="31">14.7–28.5</td>
|
||
<td id="142C5DC3FFF588A0FDC6CB5CCD6FFCC9" box="[592,635,840,855]" gridcol="3" gridrow="18" pageId="8" pageNumber="31">20.3</td>
|
||
<td id="142C5DC3FFF588A0FD2BCB5CCDE6FCC9" box="[701,754,840,855]" gridcol="4" gridrow="18" pageId="8" pageNumber="31">3.6, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CB4BCDE6FCF3" box="[116,754,863,877]" gridrow="19" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CB4BCFF3FCF3" box="[116,231,863,877]" gridcol="0" gridrow="19" pageId="8" pageNumber="31">LV6 (ac)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCB4BCE7AFCF3" box="[297,366,863,877]" gridcol="1" gridrow="19" pageId="8" pageNumber="31">4.3</td>
|
||
<td id="142C5DC3FFF588A0FE39CB4BCD1AFCF3" box="[431,526,863,877]" gridcol="2" gridrow="19" pageId="8" pageNumber="31">4.3–9.0</td>
|
||
<td id="142C5DC3FFF588A0FDC6CB4BCD6FFCF3" box="[592,635,863,877]" gridcol="3" gridrow="19" pageId="8" pageNumber="31">6.2</td>
|
||
<td id="142C5DC3FFF588A0FD2BCB4BCDE6FCF3" box="[701,754,863,877]" gridcol="4" gridrow="19" pageId="8" pageNumber="31">1.1, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CB62CDE6FC1A" box="[116,754,886,900]" gridrow="20" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CB62CFF3FC1A" box="[116,231,886,900]" gridcol="0" gridrow="20" pageId="8" pageNumber="31">SL7 (ac)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCB62CE7AFC1A" box="[297,366,886,900]" gridcol="1" gridrow="20" pageId="8" pageNumber="31">3.4</td>
|
||
<td id="142C5DC3FFF588A0FE39CB62CD1AFC1A" box="[431,526,886,900]" gridcol="2" gridrow="20" pageId="8" pageNumber="31">3.4–8.2</td>
|
||
<td id="142C5DC3FFF588A0FDC6CB62CD6FFC1A" box="[592,635,886,900]" gridcol="3" gridrow="20" pageId="8" pageNumber="31">6.1</td>
|
||
<td id="142C5DC3FFF588A0FD2BCB62CDE6FC1A" box="[701,754,886,900]" gridcol="4" gridrow="20" pageId="8" pageNumber="31">1.2, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CB99CDE6FC05" box="[116,754,909,923]" gridrow="21" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CB99CFF3FC05" box="[116,231,909,923]" gridcol="0" gridrow="21" pageId="8" pageNumber="31">SL8 (t)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCB99CE7AFC05" box="[297,366,909,923]" gridcol="1" gridrow="21" pageId="8" pageNumber="31">22.8</td>
|
||
<td id="142C5DC3FFF588A0FE39CB99CD1AFC05" box="[431,526,909,923]" gridcol="2" gridrow="21" pageId="8" pageNumber="31">19.5–24.6</td>
|
||
<td id="142C5DC3FFF588A0FDC6CB99CD6FFC05" box="[592,635,909,923]" gridcol="3" gridrow="21" pageId="8" pageNumber="31">22.3</td>
|
||
<td id="142C5DC3FFF588A0FD2BCB99CDE6FC05" box="[701,754,909,923]" gridcol="4" gridrow="21" pageId="8" pageNumber="31">1.4, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CBB7CDE6FC2C" box="[116,754,931,946]" gridrow="22" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CBB7CFF3FC2C" box="[116,231,931,946]" gridcol="0" gridrow="22" pageId="8" pageNumber="31">LA8 (ac)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCBB7CE7AFC2C" box="[297,366,931,946]" gridcol="1" gridrow="22" pageId="8" pageNumber="31">7.3</td>
|
||
<td id="142C5DC3FFF588A0FE39CBB7CD1AFC2C" box="[431,526,931,946]" gridcol="2" gridrow="22" pageId="8" pageNumber="31">5.0–9.9</td>
|
||
<td id="142C5DC3FFF588A0FDC6CBB7CD6FFC2C" box="[592,635,931,946]" gridcol="3" gridrow="22" pageId="8" pageNumber="31">7.6</td>
|
||
<td id="142C5DC3FFF588A0FD2BCBB7CDE6FC2C" box="[701,754,931,946]" gridcol="4" gridrow="22" pageId="8" pageNumber="31">0.8, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CBAECDE6FC57" box="[116,754,954,969]" gridrow="23" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CBAECFF3FC57" box="[116,231,954,969]" gridcol="0" gridrow="23" pageId="8" pageNumber="31">LD10 (t)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCBAECE7AFC57" box="[297,366,954,969]" gridcol="1" gridrow="23" pageId="8" pageNumber="31">18.5</td>
|
||
<td id="142C5DC3FFF588A0FE39CBAECD1AFC57" box="[431,526,954,969]" gridcol="2" gridrow="23" pageId="8" pageNumber="31">16.5–35.9</td>
|
||
<td id="142C5DC3FFF588A0FDC6CBAECD6FFC57" box="[592,635,954,969]" gridcol="3" gridrow="23" pageId="8" pageNumber="31">21.6</td>
|
||
<td id="142C5DC3FFF588A0FD2BCBAECDE6FC57" box="[701,754,954,969]" gridcol="4" gridrow="23" pageId="8" pageNumber="31">5.3, 6</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CBC5CDE6FC7E" box="[116,754,977,992]" gridrow="24" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CBC5CFF3FC7E" box="[116,231,977,992]" gridcol="0" gridrow="24" pageId="8" pageNumber="31">LTS</th>
|
||
<td id="142C5DC3FFF588A0FEBFCBC5CE7AFC7E" box="[297,366,977,992]" gridcol="1" gridrow="24" pageId="8" pageNumber="31">154.1</td>
|
||
<td id="142C5DC3FFF588A0FE39CBC5CD1AFC7E" box="[431,526,977,992]" gridcol="2" gridrow="24" pageId="8" pageNumber="31">134.7–160.5</td>
|
||
<td id="142C5DC3FFF588A0FDC6CBC5CD6FFC7E" box="[592,635,977,992]" gridcol="3" gridrow="24" pageId="8" pageNumber="31">148.9</td>
|
||
<td id="142C5DC3FFF588A0FD2BCBC5CDE6FC7E" box="[701,754,977,992]" gridcol="4" gridrow="24" pageId="8" pageNumber="31">7.8, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CBFCCDE6FC68" box="[116,754,1000,1014]" gridrow="25" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CBFCCFF3FC68" box="[116,231,1000,1014]" gridcol="0" gridrow="25" pageId="8" pageNumber="31">LTS/TL (%)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCBFCCE7AFC68" box="[297,366,1000,1014]" gridcol="1" gridrow="25" pageId="8" pageNumber="31">49.6</td>
|
||
<td id="142C5DC3FFF588A0FE39CBFCCD1AFC68" box="[431,526,1000,1014]" gridcol="2" gridrow="25" pageId="8" pageNumber="31">43.4–62.4</td>
|
||
<td id="142C5DC3FFF588A0FDC6CBFCCD6FFC68" box="[592,635,1000,1014]" gridcol="3" gridrow="25" pageId="8" pageNumber="31">52.9</td>
|
||
<td id="142C5DC3FFF588A0FD2BCBFCCDE6FC68" box="[701,754,1000,1014]" gridcol="4" gridrow="25" pageId="8" pageNumber="31">5.4, 7</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CBEBCDE6FB93" box="[116,754,1023,1037]" gridrow="26" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CBEBCFF3FB93" box="[116,231,1023,1037]" gridcol="0" gridrow="26" pageId="8" pageNumber="31">LTAS</th>
|
||
<td id="142C5DC3FFF588A0FEBFCBEBCE7AFB93" box="[297,366,1023,1037]" gridcol="1" gridrow="26" pageId="8" pageNumber="31">/</td>
|
||
<td id="142C5DC3FFF588A0FE39CBEBCD1AFB93" box="[431,526,1023,1037]" gridcol="2" gridrow="26" pageId="8" pageNumber="31">/</td>
|
||
<td id="142C5DC3FFF588A0FDC6CBEBCD6FFB93" box="[592,635,1023,1037]" gridcol="3" gridrow="26" pageId="8" pageNumber="31">16.3</td>
|
||
<td id="142C5DC3FFF588A0FD2BCBEBCDE6FB93" box="[701,754,1023,1037]" gridcol="4" gridrow="26" pageId="8" pageNumber="31">0.0, 1</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CC02CDE6FBBA" box="[116,754,1046,1060]" gridrow="27" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CC02CFF3FBBA" box="[116,231,1046,1060]" gridcol="0" gridrow="27" pageId="8" pageNumber="31">LTAS/LTS (%)</th>
|
||
<td id="142C5DC3FFF588A0FEBFCC02CE7AFBBA" box="[297,366,1046,1060]" gridcol="1" gridrow="27" pageId="8" pageNumber="31">/</td>
|
||
<td id="142C5DC3FFF588A0FE39CC02CD1AFBBA" box="[431,526,1046,1060]" gridcol="2" gridrow="27" pageId="8" pageNumber="31">/</td>
|
||
<td id="142C5DC3FFF588A0FDC6CC02CD6FFBBA" box="[592,635,1046,1060]" gridcol="3" gridrow="27" pageId="8" pageNumber="31">10.1</td>
|
||
<td id="142C5DC3FFF588A0FD2BCC02CDE6FBBA" box="[701,754,1046,1060]" gridcol="4" gridrow="27" pageId="8" pageNumber="31">0.0, 1</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FF12CC63CE92FA3E" blockId="8.[100,770,1114,1468]" pageId="8" pageNumber="31">
|
||
Introvert with six rings of scalids (one ring of primary spinoscalids and five rings of regular-sized scalids) and 10 longitudinal sectors delimited by the arrangement of the primary spinoscalids. Ring 01 of introvert with 10 primary spinoscalids, larger than remaining ones, laterally compressed, composed of a medially fringed basal sheath and a distal, elongated, flexible, distally pointed end-piece. Remaining rings of introvert with scalids variable in size but always smaller than the primary spinoscalids, morphologically similar to the latter (
|
||
<figureCitation id="71F62A78FFF57757FDE8CD2DCDD0FAD2" box="[638,708,1337,1356]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="8" pageNumber="31">Fig. 6C</figureCitation>
|
||
). Scalids tend to collapse when mounted for LM, and specimens for SEM with fully everted heads were not available, so details on the scalid arrangement are not provided.
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FF12CDBDCBD8FE88" blockId="8.[100,770,1114,1468]" lastBlockId="8.[818,1488,148,1478]" pageId="8" pageNumber="31">
|
||
Neck: 16 trapezoidal placids, slightly wider at base, closely adjacent, with distinct joint between the neck and first trunk segment. Midventral placid widest (ca. 13–15 μm wide at base), remaining ones narrower (ca. 8–10 μm wide at base). A ring of six long, superficially haired trichoscalids associated with the placids, attached to small, rounded trichoscalid plates (
|
||
<figureCitation id="71F62A78FFF57757FC25C917CB05FE89" box="[947,1041,259,279]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FB8DC917CB31FE88" box="[1051,1061,259,278]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="8" pageNumber="31">6</figureCitation>
|
||
C-D; 7A-C; 9A-B).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FCC7C90BCB44FCE2" blockId="8.[818,1488,148,1478]" pageId="8" pageNumber="31">
|
||
Trunk: fusiform to softly rectangular, composed of 11 segments, heart-shaped in cross-section. Segments 1–2 closed, ring-like cuticular plates, remaining ones with one tergal and two sternal cuticular plates (
|
||
<figureCitation id="71F62A78FFF57757FCACC967CC8EFE18" box="[826,922,371,390]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FC33C967CCABFE18" box="[933,959,371,390]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="8" pageNumber="31">6A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FC5AC967CCF2FE18" box="[972,998,371,390]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="8" pageNumber="31">9A</figureCitation>
|
||
). Maximum sternal width at segment 8, trunk progressively tapering toward anterior and posterior ends. Cuticular hairs throughout segments 1–10, acicular, non-bracteate, distally pointed, emerging from rounded to slightly oval perforation sites. Cuticular hairs arranged as 8–10 (dorsal) and 4–6 (ventral) straight, irregular, transversal rows densely covering the whole surface of segment 1 dorsally and the most posterior half of the segment ventrally; as 6–8 transversal rows densely covering the whole surface of segment 2; as 6–16 transversal rows, becoming wavy at the lateral and ventral areas, densely covering the whole surface of segments 3–9 except in the vicinity of the droplet-shaped sensory spots and the large, oval areas (muscular scars) in laterodorsal and paraventral positions (also of segment
|
||
<quantity id="2E359B18FFF57757FAD1CAB2CA61FD27" box="[1351,1397,678,697]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="31" unit="in" value="10.0">10 in</quantity>
|
||
females); as 3–5 transversal rows densely covering the midlateral to ventrolateral region of segment
|
||
<quantity id="2E359B18FFF57757FC74CACACB07FD6E" box="[994,1043,733,753]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="31" unit="in" value="10.0">10 in</quantity>
|
||
males (
|
||
<figureCitation id="71F62A78FFF57757FBC9CAC9CBABFD6F" box="[1119,1215,733,753]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FB5FCAC9CBC7FD6E" box="[1225,1235,733,752]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="8" pageNumber="31">6</figureCitation>
|
||
E-G; 7A-G; 8A-B, G; 9B-L). Muscular scars in laterodorsal position on segments 3–9, with several spherical perforations (
|
||
<figureCitation id="71F62A78FFF57757FB80CB01CB6CFCB7" box="[1046,1144,789,809]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FB11CB01CB85FCB6" box="[1159,1169,789,808]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="8" pageNumber="31">7</figureCitation>
|
||
A-B, D; 8A; 9B, E, J). Posterior segment margins straight, with a strongly serrated primary pectinate fringe (
|
||
<figureCitation id="71F62A78FFF57757FCE0CB59CCC1FCFE" box="[886,981,845,864]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FC48CB59CCFCFCFE" box="[990,1000,845,864]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="8" pageNumber="31">6</figureCitation>
|
||
F-G; 7A-G; 8A-C, E, G; 9A-B, D-F, H-J, L). Secondary pectinate fringes not detected.
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FCC7CB91CB57FBE9" blockId="8.[818,1488,148,1478]" pageId="8" pageNumber="31">
|
||
Segment 1:
|
||
<typeStatus id="3676885FFFF57757FC5ECB91CCE5FC06" box="[968,1009,901,920]" pageId="8" pageNumber="31">type</typeStatus>
|
||
1 glandular cell outlets in middorsal and lateroventral positions, the former near the anterior segment margin, the latter located at the anterior half of the segment (
|
||
<figureCitation id="71F62A78FFF57757FB69CBA9CA4FFC4E" box="[1279,1371,957,976]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FAF2CBA9CA79FC4E" box="[1380,1389,957,976]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="8" pageNumber="31">7</figureCitation>
|
||
A-C);
|
||
<typeStatus id="3676885FFFF57757FA30CBA9CADBFC4E" box="[1446,1487,957,976]" pageId="8" pageNumber="31">type</typeStatus>
|
||
1 glandular cell outlets on this and following segments rounded, with a single pore. Rounded sensory spots (sensu
|
||
<bibRefCitation id="8D5C4B0CFFF57757FB4ACBE0CABEFB96" author="Lundbye, H. & Rho, H. S. & Sorensen, M. V." box="[1244,1450,1012,1032]" pageId="8" pageNumber="31" pagination="41 - 51" refId="ref17487" refString="Lundbye, H., Rho, H. S., Sorensen, M. V., 2011. Echinoderes rex n. sp. (Kinorhyncha: Cyclorhagida), the largest Echinoderes species found so far. Sci. Mar. 75, 41 - 51. https: // doi. org / 10.3989 / scimar. 2011.75 n 1041." type="journal article" year="2011">Lundbye et al., 2011</bibRefCitation>
|
||
) in subdorsal, laterodorsal and lateroventral positions (
|
||
<figureCitation id="71F62A78FFF57757FABACC04CA99FBBA" box="[1324,1421,1040,1060]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FA0DCC05CAB0FBBA" box="[1435,1444,1041,1060]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="8" pageNumber="31">7</figureCitation>
|
||
A-C; 9B-C). Rounded sensory spots as circular, depressed areas with 5–6 rings of micropapillae surrounding a single, central pore with a short, emerging cilium (
|
||
<figureCitation id="71F62A78FFF57757FC4CCC70CB2CFBE9" box="[986,1080,1124,1143]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="8" pageNumber="31">Fig. 9B–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FCC7CC94CC4BFA80" blockId="8.[818,1488,148,1478]" pageId="8" pageNumber="31">
|
||
Segment 2: droplet-shaped sensory spots (sensu
|
||
<bibRefCitation id="8D5C4B0CFFF57757FAAECC94CC76FB31" author="Lundbye, H. & Rho, H. S. & Sorensen, M. V." pageId="8" pageNumber="31" pagination="41 - 51" refId="ref17487" refString="Lundbye, H., Rho, H. S., Sorensen, M. V., 2011. Echinoderes rex n. sp. (Kinorhyncha: Cyclorhagida), the largest Echinoderes species found so far. Sci. Mar. 75, 41 - 51. https: // doi. org / 10.3989 / scimar. 2011.75 n 1041." type="journal article" year="2011">Lundbye et al., 2011</bibRefCitation>
|
||
) in subdorsal, laterodorsal and ventrolateral positions (
|
||
<figureCitation id="71F62A78FFF57757FAF8CC88CADFFB31" box="[1390,1483,1180,1199]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FCA4CCACCC2FFB55" box="[818,827,1208,1227]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="8" pageNumber="31">7</figureCitation>
|
||
A-C; 9B). Droplet-shaped sensory spots, on this and following segments, as oval areas with 7–8 rings of micropapillae with a single, anterior pore from which a long, thin cilium may emerge (
|
||
<figureCitation id="71F62A78FFF57757FACECCFBCA88FA9D" box="[1368,1436,1263,1283]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="8" pageNumber="31">Fig. 9B</figureCitation>
|
||
, E-H; J-K).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FCC7CD33CB2FFAC8" blockId="8.[818,1488,148,1478]" pageId="8" pageNumber="31">
|
||
Segment 3: droplet-shaped sensory spots in subdorsal and midlateral position (
|
||
<figureCitation id="71F62A78FFF57757FC1ACD57CCC4FAC9" box="[908,976,1347,1367]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FC4CCD57CCF7FAC8" box="[986,995,1347,1366]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="8" pageNumber="31">7</figureCitation>
|
||
A-C; 9B).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FCC7CD4BCADBFA58" blockId="8.[818,1488,148,1478]" pageId="8" pageNumber="31">
|
||
Segment 4: acicular spine in middorsal position (
|
||
<figureCitation id="71F62A78FFF57757FA88CD4BCA77FAED" box="[1310,1379,1375,1395]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FAFACD4BCA62FAEC" box="[1388,1398,1375,1394]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="8" pageNumber="31">7</figureCitation>
|
||
A-B; 9D); acicular spines on this and following segments are short (ca. 3–8 μm long), poorly sclerotized and weakly articulated (
|
||
<figureCitation id="71F62A78FFF57757FB6ECD83CA47FA34" box="[1272,1363,1431,1450]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FACDCD83CA71FA34" box="[1371,1381,1431,1450]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="8" pageNumber="31">7</figureCitation>
|
||
E-G; 9D, H- I).
|
||
<typeStatus id="3676885FFFF57757FCDACDA7CC6EFA58" box="[844,890,1459,1478]" pageId="8" pageNumber="31">Type</typeStatus>
|
||
2 glandular cell outlets in midlateral position (
|
||
<figureCitation id="71F62A78FFF57757FAA0CDA7CA6EFA58" box="[1334,1402,1459,1478]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="8" pageNumber="31">Fig. 5B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FA12CDA7CA89FA58" box="[1412,1437,1459,1478]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="8" pageNumber="31">7C</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF57757FA3FCDA7CAD5FA58" box="[1449,1473,1459,1478]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="8" pageNumber="31">9E</figureCitation>
|
||
);
|
||
</paragraph>
|
||
<caption id="BDB26675FFF57757FFF2CDE2CEC3F918" pageId="8" pageNumber="31" startId="8.[100,150,1526,1542]" targetType="table">
|
||
<paragraph id="E97236FDFFF57757FFF2CDE2CFB3F998" blockId="8.[100,1488,1526,1671]" box="[100,167,1526,1542]" pageId="8" pageNumber="31">
|
||
<emphasis id="DBB9EAEFFFF57757FFF2CDE2CFB3F998" bold="true" box="[100,167,1526,1542]" pageId="8" pageNumber="31">Table 5</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF57757FFF2CE04CEC3F918" blockId="8.[100,1488,1526,1671]" pageId="8" pageNumber="31">
|
||
Summary of nature and arrangement of spines, tubes, sensory spots, glandular cell outlets, nephridiopores and other cuticular structures in adults of
|
||
<taxonomicName id="2ECD4D7EFFF57757FAE6CE1BCFA8F9A4" authority="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli, 2022" authorityName="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli" authorityYear="2022" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="31" phylum="Kinorhyncha" rank="species" species="parthenope" status="sp. nov.">
|
||
<emphasis id="DBB9EAEFFFF57757FAE6CE1BCFA8F9A4" bold="true" italics="true" pageId="8" pageNumber="31">Echinoderes parthenope</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C08A5794FFF57757FF57CE3DCE10F9A7" box="[193,260,1577,1593]" pageId="8" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Abbreviations: ac, acicular spine; ce, cuticular extension; cp, cuticular process; dss, droplet-shaped sensory spot; gcoI/II, glandular cell outlet type 1/2; LA, lateral accessory; LD, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; LV, lateroventral; MD, middorsal; ML, midlateral; ne, nephridiopore; PD, paradorsal; ps, penile spines; rss, rounded sensory spot; SD, subdorsal; SL, sublateral; th, tuft of hairs; t, tube; VL, ventrolateral; VM, ventromedial; Ƌ/♀ indicate sexually dimorphic structures.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E97236FDFFF57757FFE2CE8ECBCEF82E" pageId="8" pageNumber="31">
|
||
<table id="9BCDC45DFFF588A0FFE2CE8ECAABF82E" box="[116,1471,1690,1968]" gridcols="11" gridrows="12" pageId="8" pageNumber="31">
|
||
<tr id="57FD34BFFFF588A0FFE2CE8ECAABF937" box="[116,1471,1690,1705]" gridrow="0" pageId="8" pageNumber="31">
|
||
<th id="142C5DC3FFF588A0FFE2CE8ECFA0F937" box="[116,180,1690,1705]" gridcol="0" gridrow="0" pageId="8" pageNumber="31">Segment</th>
|
||
<th id="142C5DC3FFF588A0FF6DCE8ECE47F937" box="[251,339,1690,1705]" gridcol="1" gridrow="0" pageId="8" pageNumber="31">MD</th>
|
||
<th id="142C5DC3FFF588A0FE0FCE8ECEA4F937" box="[409,432,1690,1705]" gridcol="2" gridrow="0" pageId="8" pageNumber="31">PD</th>
|
||
<th id="142C5DC3FFF588A0FE61CE8ECD4DF937" box="[503,601,1690,1705]" gridcol="3" gridrow="0" pageId="8" pageNumber="31">SD</th>
|
||
<th id="142C5DC3FFF588A0FD36CE8ECDD1F937" box="[672,709,1690,1705]" gridcol="4" gridrow="0" pageId="8" pageNumber="31">LD</th>
|
||
<th id="142C5DC3FFF588A0FC9ACE8ECC25F937" box="[780,817,1690,1705]" gridcol="5" gridrow="0" pageId="8" pageNumber="31">ML</th>
|
||
<th id="142C5DC3FFF588A0FCEECE8ECCACF937" box="[888,952,1690,1705]" gridcol="6" gridrow="0" pageId="8" pageNumber="31">SL</th>
|
||
<th id="142C5DC3FFF588A0FC68CE8ECB20F937" box="[1022,1076,1690,1705]" gridcol="7" gridrow="0" pageId="8" pageNumber="31">LA</th>
|
||
<th id="142C5DC3FFF588A0FBECCE8ECBCEF937" box="[1146,1242,1690,1705]" gridcol="8" gridrow="0" pageId="8" pageNumber="31">LV</th>
|
||
<th id="142C5DC3FFF588A0FAB7CE8ECA2CF937" box="[1313,1336,1690,1705]" gridcol="9" gridrow="0" pageId="8" pageNumber="31">VL</th>
|
||
<th id="142C5DC3FFF588A0FAE8CE8ECAABF937" box="[1406,1471,1690,1705]" gridcol="10" gridrow="0" pageId="8" pageNumber="31">VM</th>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CEA9CAABF952" box="[116,1471,1725,1740]" gridrow="1" pageId="8" pageNumber="31" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CEA9CFA0F952" box="[116,180,1725,1740]" gridcol="0" gridrow="1" pageId="8" pageNumber="31">1</th>
|
||
<td id="142C5DC3FFF588A0FF6DCEA9CE47F952" box="[251,339,1725,1740]" gridcol="1" gridrow="1" pageId="8" pageNumber="31">gcoI</td>
|
||
<td id="142C5DC3FFF588A0FE61CEA9CD4DF952" box="[503,601,1725,1740]" gridcol="3" gridrow="1" pageId="8" pageNumber="31">rss</td>
|
||
<td id="142C5DC3FFF588A0FD36CEA9CDD1F952" box="[672,709,1725,1740]" gridcol="4" gridrow="1" pageId="8" pageNumber="31">rss</td>
|
||
<td id="142C5DC3FFF588A0FBECCEA9CBCEF952" box="[1146,1242,1725,1740]" gridcol="8" gridrow="1" pageId="8" pageNumber="31">gcoI, rss</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CEC0CAABF97C" box="[116,1471,1748,1762]" gridrow="2" pageId="8" pageNumber="31" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="142C5DC3FFF588A0FFE2CEC0CFA0F97C" box="[116,180,1748,1762]" gridcol="0" gridrow="2" pageId="8" pageNumber="31">2</th>
|
||
<td id="142C5DC3FFF588A0FE61CEC0CD4DF97C" box="[503,601,1748,1762]" gridcol="3" gridrow="2" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FD36CEC0CDD1F97C" box="[672,709,1748,1762]" gridcol="4" gridrow="2" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FAB7CEC0CA2CF97C" box="[1313,1336,1748,1762]" gridcol="9" gridrow="2" pageId="8" pageNumber="31">dss</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CEFFCAABF967" box="[116,1471,1771,1785]" gridrow="3" pageId="8" pageNumber="31" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CEFFCFA0F967" box="[116,180,1771,1785]" gridcol="0" gridrow="3" pageId="8" pageNumber="31">3</th>
|
||
<td id="142C5DC3FFF588A0FE61CEFFCD4DF967" box="[503,601,1771,1785]" gridcol="3" gridrow="3" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FC9ACEFFCC25F967" box="[780,817,1771,1785]" gridcol="5" gridrow="3" pageId="8" pageNumber="31">dss</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CF16CAABF88E" box="[116,1471,1794,1808]" gridrow="4" pageId="8" pageNumber="31" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CF16CFA0F88E" box="[116,180,1794,1808]" gridcol="0" gridrow="4" pageId="8" pageNumber="31">4</th>
|
||
<td id="142C5DC3FFF588A0FF6DCF16CE47F88E" box="[251,339,1794,1808]" gridcol="1" gridrow="4" pageId="8" pageNumber="31">Ac</td>
|
||
<td id="142C5DC3FFF588A0FE61CF16CD4DF88E" box="[503,601,1794,1808]" gridcol="3" gridrow="4" pageId="8" pageNumber="31">dss (Ƌ)</td>
|
||
<td id="142C5DC3FFF588A0FC9ACF16CC25F88E" box="[780,817,1794,1808]" gridcol="5" gridrow="4" pageId="8" pageNumber="31">gcoII</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CF0DCAABF8B9" box="[116,1471,1817,1831]" gridrow="5" pageId="8" pageNumber="31" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CF0DCFA0F8B9" box="[116,180,1817,1831]" gridcol="0" gridrow="5" pageId="8" pageNumber="31">5</th>
|
||
<td id="142C5DC3FFF588A0FE61CF0DCD4DF8B9" box="[503,601,1817,1831]" gridcol="3" gridrow="5" pageId="8" pageNumber="31">gcoII</td>
|
||
<td id="142C5DC3FFF588A0FC9ACF0DCC25F8B9" box="[780,817,1817,1831]" gridcol="5" gridrow="5" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FBECCF0DCBCEF8B9" box="[1146,1242,1817,1831]" gridcol="8" gridrow="5" pageId="8" pageNumber="31">t</td>
|
||
<td id="142C5DC3FFF588A0FAE8CF0DCAABF8B9" box="[1406,1471,1817,1831]" gridcol="10" gridrow="5" pageId="8" pageNumber="31">gcoI, dss</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CF24CAABF8A0" box="[116,1471,1840,1854]" gridrow="6" pageId="8" pageNumber="31" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CF24CFA0F8A0" box="[116,180,1840,1854]" gridcol="0" gridrow="6" pageId="8" pageNumber="31">6</th>
|
||
<td id="142C5DC3FFF588A0FE61CF24CD4DF8A0" box="[503,601,1840,1854]" gridcol="3" gridrow="6" pageId="8" pageNumber="31">gcoII, dss</td>
|
||
<td id="142C5DC3FFF588A0FC9ACF24CC25F8A0" box="[780,817,1840,1854]" gridcol="5" gridrow="6" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FC68CF24CB20F8A0" box="[1022,1076,1840,1854]" gridcol="7" gridrow="6" pageId="8" pageNumber="31">gcoII</td>
|
||
<td id="142C5DC3FFF588A0FBECCF24CBCEF8A0" box="[1146,1242,1840,1854]" gridcol="8" gridrow="6" pageId="8" pageNumber="31">ac</td>
|
||
<td id="142C5DC3FFF588A0FAE8CF24CAABF8A0" box="[1406,1471,1840,1854]" gridcol="10" gridrow="6" pageId="8" pageNumber="31">gcoI, dss</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CF53CAABF8CB" box="[116,1471,1863,1877]" gridrow="7" pageId="8" pageNumber="31" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CF53CFA0F8CB" box="[116,180,1863,1877]" gridcol="0" gridrow="7" pageId="8" pageNumber="31">7</th>
|
||
<td id="142C5DC3FFF588A0FE61CF53CD4DF8CB" box="[503,601,1863,1877]" gridcol="3" gridrow="7" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FC9ACF53CC25F8CB" box="[780,817,1863,1877]" gridcol="5" gridrow="7" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FCEECF53CCACF8CB" box="[888,952,1863,1877]" gridcol="6" gridrow="7" pageId="8" pageNumber="31">gcoII, ac</td>
|
||
<td id="142C5DC3FFF588A0FAE8CF53CAABF8CB" box="[1406,1471,1863,1877]" gridcol="10" gridrow="7" pageId="8" pageNumber="31">gcoI, dss</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CF49CAABF8F2" box="[116,1471,1885,1900]" gridrow="8" pageId="8" pageNumber="31" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CF49CFA0F8F2" box="[116,180,1885,1900]" gridcol="0" gridrow="8" pageId="8" pageNumber="31">8</th>
|
||
<td id="142C5DC3FFF588A0FE61CF49CD4DF8F2" box="[503,601,1885,1900]" gridcol="3" gridrow="8" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FCEECF49CCACF8F2" box="[888,952,1885,1900]" gridcol="6" gridrow="8" pageId="8" pageNumber="31">t</td>
|
||
<td id="142C5DC3FFF588A0FC68CF49CB20F8F2" box="[1022,1076,1885,1900]" gridcol="7" gridrow="8" pageId="8" pageNumber="31">Ac</td>
|
||
<td id="142C5DC3FFF588A0FAE8CF49CAABF8F2" box="[1406,1471,1885,1900]" gridcol="10" gridrow="8" pageId="8" pageNumber="31">gcoI, dss</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CF60CAABF81D" box="[116,1471,1908,1923]" gridrow="9" pageId="8" pageNumber="31" rowspan-1="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="142C5DC3FFF588A0FFE2CF60CFA0F81D" box="[116,180,1908,1923]" gridcol="0" gridrow="9" pageId="8" pageNumber="31">9</th>
|
||
<td id="142C5DC3FFF588A0FE0FCF60CEA4F81D" box="[409,432,1908,1923]" gridcol="2" gridrow="9" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FE61CF60CD4DF81D" box="[503,601,1908,1923]" gridcol="3" gridrow="9" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FD36CF60CDD1F81D" box="[672,709,1908,1923]" gridcol="4" gridrow="9" pageId="8" pageNumber="31">gcoII</td>
|
||
<td id="142C5DC3FFF588A0FC9ACF60CC25F81D" box="[780,817,1908,1923]" gridcol="5" gridrow="9" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FCEECF60CCACF81D" box="[888,952,1908,1923]" gridcol="6" gridrow="9" pageId="8" pageNumber="31">ne</td>
|
||
<td id="142C5DC3FFF588A0FAB7CF60CA2CF81D" box="[1313,1336,1908,1923]" gridcol="9" gridrow="9" pageId="8" pageNumber="31">dss</td>
|
||
<td id="142C5DC3FFF588A0FAE8CF60CAABF81D" box="[1406,1471,1908,1923]" gridcol="10" gridrow="9" pageId="8" pageNumber="31">gcoI</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CF9FCAABF807" box="[116,1471,1931,1945]" gridrow="10" pageId="8" pageNumber="31" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CF9FCFA0F807" box="[116,180,1931,1945]" gridcol="0" gridrow="10" pageId="8" pageNumber="31">10</th>
|
||
<td id="142C5DC3FFF588A0FF6DCF9FCE47F807" box="[251,339,1931,1945]" gridcol="1" gridrow="10" pageId="8" pageNumber="31">gcoI, gcoI</td>
|
||
<td id="142C5DC3FFF588A0FE61CF9FCD4DF807" box="[503,601,1931,1945]" gridcol="3" gridrow="10" pageId="8" pageNumber="31">ce (Ƌ), th (♀)</td>
|
||
<td id="142C5DC3FFF588A0FD36CF9FCDD1F807" box="[672,709,1931,1945]" gridcol="4" gridrow="10" pageId="8" pageNumber="31">t (Ƌ)</td>
|
||
</tr>
|
||
<tr id="57FD34BFFFF588A0FFE2CFB6CAABF82E" box="[116,1471,1954,1968]" gridrow="11" pageId="8" pageNumber="31" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-9="1">
|
||
<th id="142C5DC3FFF588A0FFE2CFB6CFA0F82E" box="[116,180,1954,1968]" gridcol="0" gridrow="11" pageId="8" pageNumber="31">11</th>
|
||
<td id="142C5DC3FFF588A0FF6DCFB6CE47F82E" box="[251,339,1954,1968]" gridcol="1" gridrow="11" pageId="8" pageNumber="31">gcoI, cp (Ƌ)</td>
|
||
<td id="142C5DC3FFF588A0FC68CFB6CB20F82E" box="[1022,1076,1954,1968]" gridcol="7" gridrow="11" pageId="8" pageNumber="31">ltas (♀)</td>
|
||
<td id="142C5DC3FFF588A0FBECCFB6CBCEF82E" box="[1146,1242,1954,1968]" gridcol="8" gridrow="11" pageId="8" pageNumber="31">lts, ps x3 (Ƌ)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF47756FA81CF69CADBF810" blockId="9.[1303,1487,1917,1934]" box="[1303,1487,1917,1934]" pageId="9" pageNumber="32">
|
||
<emphasis id="DBB9EAEFFFF47756FA81CF69CADBF810" bold="true" box="[1303,1487,1917,1934]" italics="true" pageId="9" pageNumber="32">(caption on next page)</emphasis>
|
||
</paragraph>
|
||
<caption id="BDB26675FFF77755FFF2C880CA22FEEF" pageId="10" pageNumber="33" startId="10.[100,130,148,165]" targetType="figure">
|
||
<paragraph id="E97236FDFFF77755FFF2C880CA22FEEF" blockId="10.[100,1488,148,369]" pageId="10" pageNumber="33">
|
||
<emphasis id="DBB9EAEFFFF77755FFF2C880CF8AFF3A" bold="true" box="[100,158,148,165]" pageId="10" pageNumber="33">Fig. 5.</emphasis>
|
||
Line art drawing of
|
||
<taxonomicName id="2ECD4D7EFFF77755FEC5C880CD05FF3B" authority="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli, 2022" authorityName="Cepeda & González-Casarrubios & Sánchez & Spedicato & Michaud & Zeppilli" authorityYear="2022" box="[339,529,148,165]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="10" pageNumber="33" phylum="Kinorhyncha" rank="species" species="parthenope" status="sp. nov.">
|
||
<emphasis id="DBB9EAEFFFF77755FEC5C880CD05FF3B" bold="true" box="[339,529,148,165]" italics="true" pageId="10" pageNumber="33">Echinoderes parthenope</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C08A5794FFF77755FD81C880CD4FFF3A" box="[535,603,148,164]" pageId="10" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E97236FDFFF77755FFF2C9B2CFF7FD93" blockId="10.[100,770,422,1947]" pageId="10" pageNumber="33">
|
||
type 2 glandular cell outlets, on this and following segments, as minute, circular areas smaller than sensory spots bearing a single pore (
|
||
<figureCitation id="71F62A78FFF77755FD2CC9D6CDEAFE4B" box="[698,766,450,469]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">Fig. 9E</figureCitation>
|
||
, G-H, J). Droplet-shaped sensory spots in subdorsal position only in males (
|
||
<figureCitation id="71F62A78FFF77755FFFAC9EECFA5FD93" box="[108,177,506,525]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF2DC9EECFC1FD93" box="[187,213,506,525]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FF12CA02CE5FFD06" blockId="10.[100,770,422,1947]" pageId="10" pageNumber="33">
|
||
Segment 5: tubes in lateroventral position (
|
||
<figureCitation id="71F62A78FFF77755FD83CA02CD4EFDB7" box="[533,602,534,553]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FDF4CA02CD78FDB7" box="[610,620,534,553]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7</figureCitation>
|
||
E-F; 9E).
|
||
<typeStatus id="3676885FFFF77755FD55CA02CDE5FDB7" box="[707,753,534,553]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
2 glandular cell outlets in subdorsal position (
|
||
<figureCitation id="71F62A78FFF77755FD85CA25CD4EFDDB" box="[531,602,561,581]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FDF0CA26CD90FDDB" box="[614,644,562,581]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD06CA26CDB8FDDB" box="[656,684,562,581]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9E</figureCitation>
|
||
).
|
||
<typeStatus id="3676885FFFF77755FD29CA26CDF9FDDB" box="[703,749,562,581]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
1 glandular cell outlets in ventromedial position (
|
||
<figureCitation id="71F62A78FFF77755FDA5CA59CD6FFDFF" box="[563,635,589,609]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD11CA5ACD8AFDFF" box="[647,670,590,609]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7F</figureCitation>
|
||
). Droplet-shaped sensory spots in midlateral and ventromedial positions (
|
||
<figureCitation id="71F62A78FFF77755FFFACA91CFDEFD07" box="[108,202,645,665]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF42CA92CE1DFD07" box="[212,265,646,665]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7D, F</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE85CA91CE09FD06" box="[275,285,645,664]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9</figureCitation>
|
||
E-F).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FF12CAB5CE5AFCC2" blockId="10.[100,770,422,1947]" pageId="10" pageNumber="33">
|
||
Segment 6: acicular spines in lateroventral position (
|
||
<figureCitation id="71F62A78FFF77755FDEFCAB5CDABFD2A" box="[633,703,673,692]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD5FCAB5CDC7FD2A" box="[713,723,673,692]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7</figureCitation>
|
||
E-F).
|
||
<typeStatus id="3676885FFFF77755FFF2CAA9CF86FD4E" box="[100,146,701,720]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
2 glandular cell outlets in subdorsal and lateral accessory positions (
|
||
<figureCitation id="71F62A78FFF77755FFFACACDCFDEFD72" box="[108,202,729,748]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF42CACDCFCAFD72" box="[212,222,729,748]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7</figureCitation>
|
||
D-F; 9E, G).
|
||
<typeStatus id="3676885FFFF77755FEC0CACDCE90FD72" box="[342,388,729,748]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
1 glandular cell outlets in ventromedial position (
|
||
<figureCitation id="71F62A78FFF77755FF29CAE1CE12FC96" box="[191,262,757,776]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE86CAE1CE33FC96" box="[272,295,757,776]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7F</figureCitation>
|
||
). Droplet-shaped sensory spots in subdorsal, midlateral and ventromedial positions, the former located more lateral than the subdorsal
|
||
<typeStatus id="3676885FFFF77755FF71CB39CE04FCDE" box="[231,272,813,832]" pageId="10" pageNumber="33">type</typeStatus>
|
||
2 glandular cell outlets but still in subdorsal position (
|
||
<figureCitation id="71F62A78FFF77755FFFACB5CCFDEFCC2" box="[108,202,840,860]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF42CB5DCE1DFCC2" box="[212,265,841,860]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7D, F</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE85CB5DCE09FCC2" box="[275,285,841,860]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9</figureCitation>
|
||
E-G).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FF12CB70CDFEFC79" blockId="10.[100,770,422,1947]" pageId="10" pageNumber="33">
|
||
Segment 7: acicular spines in sublateral position (
|
||
<figureCitation id="71F62A78FFF77755FDF7CB70CDBCFCE6" box="[609,680,868,888]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD24CB71CDDFFCE6" box="[690,715,869,888]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7E</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD4ECB70CDE0FCE9" box="[728,756,868,887]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9H</figureCitation>
|
||
).
|
||
<typeStatus id="3676885FFFF77755FFF2CB95CF86FC0A" box="[100,146,897,916]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
2 glandular cell outlets in sublateral position, slightly lateral or above the acicular spines (
|
||
<figureCitation id="71F62A78FFF77755FECCCB88CE8BFC31" box="[346,415,924,943]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE31CB89CED4FC2E" box="[423,448,925,944]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7E</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE5DCB88CEF3FC31" box="[459,487,924,943]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9H</figureCitation>
|
||
).
|
||
<typeStatus id="3676885FFFF77755FE6CCB89CD3CFC2E" box="[506,552,925,944]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
1 glandular cell outlets in ventromedial position (
|
||
<figureCitation id="71F62A78FFF77755FEFECBACCEA0FC55" box="[360,436,952,971]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
). Droplet-shaped sensory spots in subdorsal, midlateral and ventromedial positions (
|
||
<figureCitation id="71F62A78FFF77755FDD6CBC0CD8AFC79" box="[576,670,980,999]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD3ECBC0CDCBFC79" box="[680,735,980,999]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9F, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FF12CBE4CDCDFBC9" blockId="10.[100,770,422,1947]" pageId="10" pageNumber="33">
|
||
Segment 8: acicular spines in lateral accessory position, and large tubes in sublateral position (
|
||
<figureCitation id="71F62A78FFF77755FEEECC18CED4FB81" box="[376,448,1036,1055]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE5DCC18CD15FB81" box="[459,513,1036,1055]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7E, G</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD9BCC18CD34FB81" box="[525,544,1036,1055]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9I</figureCitation>
|
||
).
|
||
<typeStatus id="3676885FFFF77755FDA0CC18CD70FB81" box="[566,612,1036,1055]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
1 glandular cell outlets in ventromedial position (
|
||
<figureCitation id="71F62A78FFF77755FE35CC33CEFFFBA5" box="[419,491,1063,1083]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE63CC3CCD04FBA5" box="[501,528,1064,1083]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7G</figureCitation>
|
||
). Droplet-shaped sensory spots in subdorsal and ventromedial positions (
|
||
<figureCitation id="71F62A78FFF77755FDB4CC57CD94FBC9" box="[546,640,1091,1111]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD1CCC50CDB0FBC9" box="[650,676,1092,1111]" captionStart="Fig" captionStartId="12.[100,130,1792,1809]" captionTargetBox="[141,1447,151,1762]" captionTargetId="figure-8@12.[139,1449,148,1764]" captionTargetPageId="12" captionText="Fig. 7. Light micrographs of male holotype MNHN-613 Ma (A, D, F-G), male paratype MNHN-641 Ma (C) and female paratype MNHN-637 Ma (B, E) of Echinoderes parthenope sp. nov. showing cuticular details of segments 1–8. A: Dorsal and lateral overview of neck and segments 1–4; B: Dorsal view of neck and segments 1–4; C: midlateral to ventromedial view on right side of cuticular plates of segments 1–4; D: middorsal to midlateral view on right side of tergal plates of segments 5–6; E: midlateral to lateroventral view on left side of tergal plates of segments 5–8; F: midlateral to ventromedial view on right side of cuticular plates of segments 5–6; G: ventral view of segment 8. Abbreviations: dpl, dorsal placid; gco2, type 2 glandular cell outlet; las, lateral accessory spine; lvs, lateroventral spine; lvt, lateroventral tube; m, male condition of sexually dimorphic character; mds, middorsal spine; mls, midlateral spine; mvpl, midventral placid; ppf, primary pectinate fringe; slt, sublateral tube; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375214" httpUri="https://zenodo.org/record/10375214/files/figure.png" pageId="10" pageNumber="33">7G</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD27CC50CDDFFBC9" box="[689,715,1092,1111]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FF12CC4BCE25FAA8" blockId="10.[100,770,422,1947]" pageId="10" pageNumber="33">
|
||
Segment 9:
|
||
<typeStatus id="3676885FFFF77755FF6DCC74CE30FBED" box="[251,292,1120,1139]" pageId="10" pageNumber="33">type</typeStatus>
|
||
2 glandular cell outlets in laterodorsal position (
|
||
<figureCitation id="71F62A78FFF77755FFFACC6FCFA4FB11" box="[108,176,1147,1167]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF2CCC6FCFC0FB10" box="[186,212,1147,1166]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF76CC6FCFE2FB10" box="[224,246,1147,1166]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9J</figureCitation>
|
||
).
|
||
<typeStatus id="3676885FFFF77755FE9CCC68CE2CFB11" box="[266,312,1148,1167]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
1 glandular cell outlets in ventromedial position (
|
||
<figureCitation id="71F62A78FFF77755FFFACC83CFA2FB34" box="[108,182,1175,1194]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF54CC83CFCFFB34" box="[194,219,1175,1194]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8B</figureCitation>
|
||
). Droplet-shaped sensory spots in paradorsal, subdorsal, midlateral and ventrolateral positions (
|
||
<figureCitation id="71F62A78FFF77755FE49CCA7CD2BFB58" box="[479,575,1203,1222]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FDDDCCA7CD40FB58" box="[587,596,1203,1222]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8</figureCitation>
|
||
A-B; 9J-K). Nephridiopores in sublateral position as triangular, enlarged sieve plates consisting of a slightly convex area with numerous pores followed by a small posterior, slightly concave region with a single, enlarged pore (
|
||
<figureCitation id="71F62A78FFF77755FFFACD37CFDEFAA8" box="[108,202,1315,1334]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF42CD37CFCAFAA8" box="[212,222,1315,1334]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="10" pageNumber="33">6</figureCitation>
|
||
F-G; 9J).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FF12CD2BCE01F98B" blockId="10.[100,770,422,1947]" pageId="10" pageNumber="33">
|
||
Segment 10: retractable into segment 9 (
|
||
<figureCitation id="71F62A78FFF77755FD9DCD2ACD7FFACC" box="[523,619,1342,1362]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FDE1CD2ACD85FACF" box="[631,657,1342,1361]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="10" pageNumber="33">6A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD08CD2BCDACFACC" box="[670,696,1343,1362]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9A</figureCitation>
|
||
). Large tubes in laterodorsal position only in males (
|
||
<figureCitation id="71F62A78FFF77755FD87CD4ECD42FAF0" box="[529,598,1370,1390]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FDF7CD4FCD97FAF0" box="[609,643,1371,1390]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8 E</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD18CD4FCDDCFAF0" box="[654,712,1371,1390]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9 J, L</figureCitation>
|
||
).
|
||
<specimenCount id="FFCBFD74FFF77755FD4FCD4FCF99FA14" count="2" pageId="10" pageNumber="33" type="generic" typeStatus="type">Two type</specimenCount>
|
||
1 glandular cell outlets in middorsal position, longitudinally aligned (
|
||
<figureCitation id="71F62A78FFF77755FF2FCD86CE0EFA38" box="[185,282,1426,1446]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5B, D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FEB0CD86CE2AFA3B" box="[294,318,1426,1445]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8E</figureCitation>
|
||
). Tergal plate of males with a pair of cuticular extensions in subdorsal position, rectangular, with straight, abruptly tapering, slightly curved tips protruding into the cuticular surface of the following segment; tergal plates of females with tufts of hairs instead (
|
||
<figureCitation id="71F62A78FFF77755FFFACE16CFF1F98B" box="[108,229,1538,1557]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B, D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FF79CE16CE1CF98B" box="[239,264,1538,1557]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FF12CE0ACE71F805" blockId="10.[100,770,422,1947]" pageId="10" pageNumber="33">
|
||
<materialsCitation id="59A53CA0FFF77755FF12CE0ACE76F805" collectionCode="LTS, LTAS" pageId="10" pageNumber="33" specimenCount="1">
|
||
Segment 11: retractable into segments 9–10 (
|
||
<figureCitation id="71F62A78FFF77755FDD7CE0ACDB7F9AF" box="[577,675,1566,1585]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD26CE0ACDDEF9AF" box="[688,714,1566,1585]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="10" pageNumber="33">6A</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FD4CCE0ACDE0F9AF" box="[730,756,1566,1585]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9A</figureCitation>
|
||
). Relatively long lateral terminal spines (
|
||
<collectionCode id="8FDCAE38FFF77755FE40CE2ECEE8F9D3" box="[470,508,1594,1613]" pageId="10" pageNumber="33">LTS</collectionCode>
|
||
:TL average ratio = 52.9%), apparently well sclerotized but still flexible, distally pointed, with a hollow central cavity (
|
||
<figureCitation id="71F62A78FFF77755FEAECE65CE83F91B" box="[312,407,1649,1669]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE37CE65CEADF91A" box="[417,441,1649,1668]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="10" pageNumber="33">6E</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE53CE65CEF4F91A" box="[453,480,1649,1668]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE7DCE65CD09F91B" box="[491,541,1649,1669]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9A, L</figureCitation>
|
||
). Males with three pairs of penile spines, dorsal and ventral pairs longer and slender, smooth and distally rounded, medial pair shorter and stouter, hairy and distally blunt (
|
||
<figureCitation id="71F62A78FFF77755FF34CED1CFEAF947" box="[162,254,1733,1753]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–B</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FE91CED1CE0BF946" box="[263,287,1733,1752]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8C</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FEBCCED1CE55F946" box="[298,321,1733,1752]" captionStart="Fig" captionStartId="15.[100,130,148,165]" captionText="Fig. 9. Scanning electron micrographs of male additional specimen of Echinoderes parthenope sp. nov. showing trunk overview and cuticular details of trunk segments. A: Lateral trunk overview; B: lateral view of segments 1–3; C: detail of laterodorsal, rounded sensory spot of segment 1; D: detail of middorsal spine of segment 4; E: lateral view of segments 4–6; F: ventral view of segments 5–7; G: detail of subdorsal droplet-shaped sensory spot and type 2 glandular cell outlet of segment 6; H: lateral view of segment 7; I: detail of sublateral tube and lateral accessory spine of segment 8; J: lateral view of segment 9 (inset shows the midlateral sieve plate); K: detail of midlateral, droplet-shaped sensory spot of segment 9; L: lateral view of segment 11. Abbreviations: gco2, type 2 glandular cell outlet; in, introvert; las, lateral accessory spine; ldt, laterodorsal tube; lts, lateral terminal spine; lvt, lateroventral tube; mds, middorsal spine; mls, midlateral spine; ms, muscular scar; ne, neck; pr, protuberance; ps, penile spines; S, segment (followed by numbers of corresponding segments); slt, sublateral tube; te, tergal extensions; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/droplet-shaped figures." pageId="10" pageNumber="33">9L</figureCitation>
|
||
); females with short (
|
||
<collectionCode id="8FDCAE38FFF77755FD9CCED1CD2BF946" box="[522,575,1733,1752]" pageId="10" pageNumber="33">LTAS</collectionCode>
|
||
:
|
||
<collectionCode id="8FDCAE38FFF77755FDD7CED1CD72F946" box="[577,614,1733,1752]" pageId="10" pageNumber="33">LTS</collectionCode>
|
||
value =10.1%), slender, lateral terminal accessory spines (
|
||
<figureCitation id="71F62A78FFF77755FE6ECEF5CD43F96A" box="[504,599,1761,1780]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5C–D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FDF7CEF5CD6DF96A" box="[609,633,1761,1780]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8F</figureCitation>
|
||
). Female gonopores near the anterolateral margins of sternal plates (
|
||
<figureCitation id="71F62A78FFF77755FDFDCEE9CDA7F88E" box="[619,691,1789,1808]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">Fig. 8G</figureCitation>
|
||
).
|
||
<typeStatus id="3676885FFFF77755FD54CEE9CDE4F88E" box="[706,752,1789,1808]" pageId="10" pageNumber="33">Type</typeStatus>
|
||
1 glandular cell outlet in middorsal position (
|
||
<figureCitation id="71F62A78FFF77755FE6BCF0DCD48F8B2" box="[509,604,1817,1836]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5B, D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FDF3CF0DCD6AF8B2" box="[613,638,1817,1836]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8E</figureCitation>
|
||
). Tergal plate of males with a middorsal, triangular, bulged process that extends slightly beyond the tergal extensions; blade-like tergal extensions, distally pointed (
|
||
<figureCitation id="71F62A78FFF77755FE92CF78CE77F81E" box="[260,355,1900,1920]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A–D</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FEFBCF79CE62F81E" box="[365,374,1901,1920]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8</figureCitation>
|
||
D-E; 9L). Sternal extensions short, distally rounded (
|
||
<figureCitation id="71F62A78FFF77755FF57CF9CCE35F805" box="[193,289,1928,1947]" captionStart="Fig" captionStartId="10.[100,130,148,165]" captionText="Fig. 5. Line art drawing of Echinoderes parthenope sp. nov. A: Ventral male overview; B: Dorsal male overview; C: ventral female overview of segments 10–11; D: dorsal female overview of segments 10–11. Abbreviations: dpl, dorsal placid; lagco2, lateral accessory type 2 glandular cell outlet; las, lateral accessory spine; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral type 1 glandular cell outlet; lvs, lateroventral spine; lvt, lateroventral tube; mdgco1, middorsal type 1 glandular cell outlet; mds, middorsal spine; mldss, midlateral droplet-shaped sensory spot; mlgco2, midlateral type 2 glandular cell outlet; mvpl, midventral placid; pc, pachycyclus; pddss, paradorsal droplet-shaped sensory spot; ppf, primary pectinate fringe; ps, penile spine (followed by number of corresponding pair); S, segment (followed by number of corresponding segment); sdce, subdorsal cuticular extension; sddss, subdorsal dropletshaped sensory spot; sdgco2, subdorsal type 2 glandular cell outlet; sdrss, subdorsal rounded sensory spot; sdth, subdorsal tuft of hairs; se, sternal extension; slgco2, sublateral type 2 glandular cell outlet; slne, sublateral nephridiopore; sls, sublateral spine; slt, sublateral tube; te, tergal extension; tsp, trichoscalid plate; vldss, ventrolateral droplet-shaped sensory spot; vmdss, ventromedial droplet-shaped sensory spot; vmgco1, ventromedial type 1 glandular cell outlet." pageId="10" pageNumber="33">Fig. 5A, C</figureCitation>
|
||
;
|
||
<figureCitation id="71F62A78FFF77755FEBDCF9CCE20F805" box="[299,308,1928,1947]" captionStart="Fig" captionStartId="13.[100,130,1639,1656]" captionTargetBox="[209,1379,150,1609]" captionTargetId="figure-125@13.[206,1381,148,1611]" captionTargetPageId="13" captionText="Fig. 8. Light micrographs of male holotype MNHN-613 Ma (A–D), male paratype MNHN-642 Ma (E) and female paratype MNHN-637 Ma (F–G) of Echinoderes parthenope sp. nov. showing cuticular details of segments 8–11. A: Dorsal view of segments 8–9; B: Ventral view of segments 9–10; C: penile spines; D: dorsal view of segment 11; E: dorsal view of segments 10–11; F: ventral view of segments 10–11; G: ventral view of segments 10–11. Abbreviations: ce, cuticular extension; f; female condition of sexually dimorphic character; gco2, type 2 glandular cell outlet; go, gonopore; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; m, male condition of sexually dimorphic character; ppf, primary pectinate fringe; pr, protuberance; ps, penile spine (followed by number of corresponding pair); se, sternal extension; te, tergal extension; glandular cell outlets are marked as continuous circles, and sensory spots as dashed circles/dropletshaped figures." figureDoi="http://doi.org/10.5281/zenodo.10375216" httpUri="https://zenodo.org/record/10375216/files/figure.png" pageId="10" pageNumber="33">8</figureCitation>
|
||
F-G)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FCA4C9B2CCD5FE27" blockId="10.[818,1487,422,525]" box="[818,961,422,441]" pageId="10" pageNumber="33">
|
||
<emphasis id="DBB9EAEFFFF77755FCA4C9B2CCD5FE27" bold="true" box="[818,961,422,441]" italics="true" pageId="10" pageNumber="33">3.2.5. Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E97236FDFFF77755FCC7C9D6CA7BFD93" blockId="10.[818,1487,422,525]" pageId="10" pageNumber="33">
|
||
Several specimens were found with ingested clusters of diatoms in the hindgut (segments 8–10) (
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<figureCitation id="71F62A78FFF77755FBD9C9CACB83FE6F" box="[1103,1175,478,497]" captionStart="Fig" captionStartId="11.[100,130,1802,1819]" captionTargetBox="[209,1379,150,1773]" captionTargetId="figure-8@11.[206,1381,148,1775]" captionTargetPageId="11" captionText="Fig. 6. Light micrographs of male holotype MNHN-613 Ma (E), male paratypes MNHN-649 Ma (B–C) and MNHN-651 Ma (G), and female paratype MNHN-637 Ma (A, D, F, H) of Echinoderes parthenope sp. nov. showing trunk overviews and details on the head, neck, nephridiopore and lateral terminal spines. A: Dorsal trunk overview; B: Mouth cone; C: Introvert; D: Ventral neck view: E: lateral terminal spines; F: detail of nephridiopore in midlateral view; G: overview of nephridiopore in midlateral view; H: ingested diatoms. Abbreviations: aa (sp), anterior area of nephridiopore (sieve plate); bfs, basal fringed sheath; bs, basal sheath; ep, distal endpiece; epo, enlarged pore; lts, lateral terminal spines; mlne, midlateral nephridiopore; mvpl, midventral placid; oos, outer oral style; pa, posterior area of nephridiopore; phc, pharyngeal crown; psc, primary spinoscalid; rsc, regular-sized scalid; tsp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.10375212" httpUri="https://zenodo.org/record/10375212/files/figure.png" pageId="10" pageNumber="33">Fig. 6H</figureCitation>
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). Interestingly, the diatoms seem to be unaltered despite they were swallowed by the animals.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |