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<mods:title id="C4281F1D97975F6F4E53C4CACE2DB5FE">Three new fern names of hybrid genera and four new combinations in Athyriaceae, Osmundaceae, and Thelypteridaceae</mods:title>
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<mods:namePart id="E7B4812AA0152394AFECDFF414D5E26E">Wan, Xia</mods:namePart>
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<mods:affiliation id="9107406BECF8F8BE119A711355C7CC9B">Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China &amp; College of Life Sciences, Sichuan University, Chengdu 610065, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China &amp; Missouri Botanical Garden, St. Louis, Missouri 63110, USA &amp; xwan @ mobot. org; https: // orcid. org / 0000 - 0001 - 8312 - 6917</mods:affiliation>
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<mods:affiliation id="34EC1BCCE53E6EB2E35594ACFFB1F334">Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China &amp; Missouri Botanical Garden, St. Louis, Missouri 63110, USA &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
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<paragraph id="524F4D21FF9FFFDEFF6C9E8C93B1D2D6" blockId="2.[136,1451,233,400]" box="[136,1189,233,256]" pageId="2" pageNumber="106">
<emphasis id="60849133FF9FFFDEFF6C9E8C9780D2D6" bold="true" box="[136,148,233,255]" italics="true" pageId="2" pageNumber="106">×</emphasis>
<taxonomicName id="95F036A2FF9FFFDEFF709E8C9320D329" ID-CoL="BDJBG" authority="(W. C. Shieh &amp; J. L. Tsai 1987: 7) X. Wan &amp; Li Bing Zhang" authorityName="X. Wan &amp; Li Bing Zhang" authorityYear="2022" baseAuthorityName="W. C. Shieh &amp; J. L. Tsai" baseAuthorityPageNumber="7" baseAuthorityYear="1987" box="[148,1076,233,256]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyrocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="106" phylum="Tracheophyta" rank="species" species="erubesquirolicus" status="comb. nov.">
<emphasis id="60849133FF9FFFDEFF709E8C9512D2D6" bold="true" box="[148,518,233,255]" italics="true" pageId="2" pageNumber="106">Glaphyrocyclosorus erubesquirolicus</emphasis>
(W.C.Shieh &amp; J.L.Tsai 1987: 7) X.Wan &amp; Li Bing Zhang
</taxonomicName>
,
<emphasis id="60849133FF9FFFDEFBD99E8F93B1D2D6" box="[1085,1189,234,255]" italics="true" pageId="2" pageNumber="106">
<taxonomicNameLabel id="7BB72C48FF9FFFDEFBD99E8F93B1D2D6" box="[1085,1189,234,255]" pageId="2" pageNumber="106" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="524F4D21FF9FFFDEFF6C9F6B96A5D342" blockId="2.[136,1451,233,400]" pageId="2" pageNumber="106">
Basionym:
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<emphasis id="60849133FF9FFFDEFF189F6B967CD30A" box="[252,360,270,291]" italics="true" pageId="2" pageNumber="106">Thelypteris</emphasis>
×
<emphasis id="60849133FF9FFFDEFE659F6B950CD30A" box="[385,536,270,291]" italics="true" pageId="2" pageNumber="106">erubesquirolica</emphasis>
W.C.Shieh &amp; J.L.Tsai (1987: 7)
</taxonomicName>
<taxonomicName id="95F036A2FF9FFFDEFC6A9F6B92BFD30D" ID-CoL="8S5SP" authority="(W. C. Shieh &amp; J. L. Tsai)" box="[910,1451,270,292]" class="Polypodiopsida" family="Thelypteridaceae" genus="Cyclosorus" isHybrid="true" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="106" phylum="Tracheophyta" rank="species" species="erubesquirolicus">
<emphasis id="60849133FF9FFFDEFC6A9F6B94EED30A" box="[910,1018,270,291]" italics="true" pageId="2" pageNumber="106">Cyclosorus</emphasis>
×
<emphasis id="60849133FF9FFFDEFBF79F6B93A7D30A" box="[1043,1203,270,291]" italics="true" pageId="2" pageNumber="106">erubesquirolicus</emphasis>
(W.C.Shieh &amp; J.L.Tsai)
</taxonomicName>
J.
<bibRefCitation id="366130D0FF9FFFDEFF2D9F5796B9D36E" author="Mazumdar, J." box="[201,429,306,328]" pageId="2" pageNumber="106" pagination="398 - 400" refId="ref2914" refString="Mazumdar, J. (2013) New combinations for some hybrids in the fern family Thelypteridaceae. Annales Botanici Fennici 50: 398 - 400. https: // doi. org / 10.5735 / 085.050.0604" type="journal article" year="2013">Mazumdar (2013: 398)</bibRefCitation>
≡ “
<emphasis id="60849133FF9FFFDEFE369F5796CBD36E" box="[466,479,306,327]" italics="true" pageId="2" pageNumber="106">×</emphasis>
<taxonomicName id="95F036A2FF9FFFDEFE3A9F5792B7D361" ID-CoL="8S5SP" authority="(W. C. Shieh &amp; J. L. Tsai) Hong M. Liu, Schuettp. &amp; H. Schneid." baseAuthorityName="W. C. Shieh &amp; J. L. Tsai" box="[478,1443,306,328]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphypseudosorus" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="106" phylum="Tracheophyta" rank="species" species="erubesquirolicus">
<emphasis id="60849133FF9FFFDEFE3A9F579453D36E" box="[478,839,306,327]" italics="true" pageId="2" pageNumber="106">Glaphypseudosorus erubesquirolicus</emphasis>
(W.C.Shieh &amp; J.L.Tsai) Hong M.Liu, Schuettp. &amp; H.Schneid.
</taxonomicName>
” (2020: 1000),
<emphasis id="60849133FF9FFFDEFEA29F3396BFD342" box="[326,427,342,363]" italics="true" pageId="2" pageNumber="106">nom. inval</emphasis>
.
</paragraph>
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<materialsCitation id="E298477CFF9FFFDEFF6C9F1F9352D3A6" ID-GBIF-Occurrence="3987424306" accessionNumber="T10379" box="[136,1094,378,400]" collectingDate="1984-12-17" collectionCode="TCF" collectorName="J. - L. Tsai" country="Taiwan Island" location="Nantou Hsien" municipality="Wanghsiang" pageId="2" pageNumber="106" specimenCount="1" typeStatus="holotype">
<typeStatus id="8D4BF383FF9FFFDEFF6C9F1F97ADD3B9" box="[136,185,378,400]" pageId="2" pageNumber="106">Type</typeStatus>
:—
<collectingCountry id="2AE70DB1FF9FFFDEFF319F1F9675D3B9" box="[213,353,378,400]" name="Taiwan Island" pageId="2" pageNumber="106">Taiwan Island</collectingCountry>
:
<location id="572F1BFAFF9FFFDEFE8E9F1F96E5D3B9" LSID="urn:lsid:plazi:treatment:DA59FC37FF9FFFDEFF6C9E8C96D7D06D:572F1BFAFF9FFFDEFE8E9F1F96E5D3B9" box="[362,497,378,400]" country="Taiwan Island" municipality="Wanghsiang" name="Nantou Hsien" pageId="2" pageNumber="106">Nantou Hsien</location>
,
<collectingMunicipality id="B22BD75BFF9FFFDEFE1E9F1F9567D3B9" box="[506,627,378,400]" pageId="2" pageNumber="106">Wanghsiang</collectingMunicipality>
,
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<collectingDate id="360A9209FF9FFFDEFD989F1F9421D3A6" box="[636,821,378,400]" pageId="2" pageNumber="106" value="1984-12-17">17 December 1984</collectingDate>
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,
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<emphasis id="60849133FF9FFFDEFCDB9F1F9486D3A6" box="[831,914,378,399]" italics="true" pageId="2" pageNumber="106">J.-L.Tsai</emphasis>
</collectorName>
T10379a-d (
<collectionCode id="34E1D5E4FF9FFFDEFBEA9F1F932BD3A6" box="[1038,1087,378,399]" pageId="2" pageNumber="106">TCF</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="1AEA1EAAFF9FFFDEFF6C9FDA9207D009" pageId="2" pageNumber="106" type="discussion">
<paragraph id="524F4D21FF9FFFDEFF6C9FDA9207D009" blockId="2.[136,1451,446,580]" pageId="2" pageNumber="106">
Notes: This nothospecies is a hybrid between
<taxonomicName id="95F036A2FF9FFFDEFD6A9FDB961ED3D5" authority="(Wallich ex Hooker 1863: 236) Ching (1963: 320)" authorityName="Ching" authorityPageNumber="320" authorityYear="1963" baseAuthorityName="Hooker" baseAuthorityPageNumber="236" baseAuthorityYear="1863" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="106" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="60849133FF9FFFDEFD6A9FDB94ECD3F1" box="[654,1016,446,472]" italics="true" pageId="2" pageNumber="106">Glaphyropteridopsis erubescens</emphasis>
(Wallich ex
<bibRefCitation id="366130D0FF9FFFDEFB6D9FDB924CD3F1" author="Hooker, W. J." box="[1161,1368,446,472]" pageId="2" pageNumber="106" refId="ref2581" refString="Hooker, W. J. (1863) Species Filicum. vol. 4. W. Pamplin, London, 292 pp." type="book" year="1863">Hooker 1863: 236</bibRefCitation>
)
<bibRefCitation id="366130D0FF9FFFDEFA839FDB961ED3D5" author="Ching, R. - C." pageId="2" pageNumber="106" pagination="289 - 335" refId="ref2185" refString="Ching, R. - C. (1963) A reclassification of the family Thelypteridaceae from the Mainland of Asia. Acta Pytotaxonomica Snica 8: 289 - 335." type="journal article" year="1963">Ching (1963: 320)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="95F036A2FF9FFFDEFED89F879345D3D5" authority="(Christ 1907: 144 - 145) Ching (1963: 324)" authorityName="Ching" authorityPageNumber="324" authorityYear="1963" baseAuthorityName="Christ" baseAuthorityPageNumber="144 - 145" baseAuthorityYear="1907" box="[316,1105,482,509]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="106" phylum="Tracheophyta" rank="species" species="esquirolii">
<emphasis id="60849133FF9FFFDEFED89F879561D3D5" box="[316,629,482,508]" italics="true" pageId="2" pageNumber="106">Pseudocyclosorus esquirolii</emphasis>
(
<bibRefCitation id="366130D0FF9FFFDEFD669F879462D3D4" author="Christ, K. H." box="[642,886,482,509]" pageId="2" pageNumber="106" pagination="144 - 145" refId="ref2216" refString="Christ, K. H. (1907) Filices Chinenses. Bulletin de l'Academie Internationale de Geographie Botanique 17: 144 - 145." type="journal article" year="1907">Christ 1907: 144145</bibRefCitation>
)
<bibRefCitation id="366130D0FF9FFFDEFC679F879345D3D5" author="Ching, R. - C." box="[899,1105,482,508]" pageId="2" pageNumber="106" pagination="289 - 335" refId="ref2185" refString="Ching, R. - C. (1963) A reclassification of the family Thelypteridaceae from the Mainland of Asia. Acta Pytotaxonomica Snica 8: 289 - 335." type="journal article" year="1963">Ching (1963: 324)</bibRefCitation>
</taxonomicName>
. This hybrid is an allotetraploid with 72 bivalent chromosomes, which has intermediate morphology of the parents (
<bibRefCitation id="366130D0FF9FFFDEFBD49C639212D009" author="Shieh, W. C. &amp; Tsai, J. L." box="[1072,1286,518,544]" pageId="2" pageNumber="106" pagination="1 - 12" refId="ref3227" refString="Shieh, W. C. &amp; Tsai, J. L. (1987) Three natural hybrids of the Thelypteridaceae in Taiwan. Journal of Science and Engineering 24: 1 - 12." type="journal article" year="1987">Shieh &amp; Tsai 1987</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1AEA1EAAFF9FFFDEFF6C9C4F96D7D06D" box="[136,451,554,580]" pageId="2" pageNumber="106" type="distribution">
<paragraph id="524F4D21FF9FFFDEFF6C9C4F96D7D06D" blockId="2.[136,1451,446,580]" box="[136,451,554,580]" pageId="2" pageNumber="106">
Distribution:
<collectingCountry id="2AE70DB1FF9FFFDEFEFA9C4F96ABD06D" box="[286,447,554,580]" name="Taiwan Island" pageId="2" pageNumber="106">Taiwan Island</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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