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<document id="E6863A88E2684FB546F570629ECCB36D" ID="10.11646/zootaxa.4226.4.5" ID-CLB-Dataset="37246" ID-DOI="10.11646/zootaxa.4226.4.5" ID-GBIF-Dataset="241fa9c6-84dc-4be4-9318-3c120ac52671" ID-ISSN="1175-5326" ID-Zenodo-Dep="265135" ID-ZooBank="3EE63479-7793-42FB-BDC6-7602068C347A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1485851793066" checkinUser="plazi" docAuthor="Hamasaki, Katsuyuki, Tsuru, Takuma, Sanda, Tetsuya, Fujikawa, Shunsuke, Dan, Shigeki &amp; Kitada, Shuichi" docDate="2017" docId="3F1D0E356345701C68EB4623BEB6F9B9" docLanguage="en" docName="zootaxa.4226.4.5.pdf" docOrigin="Zootaxa 4226 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenobita cavipes" docType="treatment" docVersion="7" lastPageNumber="529" masterDocId="C324764D634D7010687C4051BB27FFCC" masterDocTitle="Ontogenetic change of body color patterns in laboratory-raised juveniles of six terrestrial hermit crab species" masterLastPageNumber="545" masterPageNumber="521" pageNumber="529" updateTime="1698436537460" updateUser="ExternalLinkService">
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<mods:title id="BAAD810DE8775425BD1602C6A9CE30CF">Ontogenetic change of body color patterns in laboratory-raised juveniles of six terrestrial hermit crab species</mods:title>
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<mods:namePart id="0C144F2613FECFE1D8CCFE7DCD5A0457">Hamasaki, Katsuyuki</mods:namePart>
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<mods:namePart id="67A1188109BF1E0975E413D3C732D9F9">Tsuru, Takuma</mods:namePart>
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<mods:namePart id="85F14989CDD0540830535685FDA19C1E">Sanda, Tetsuya</mods:namePart>
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<mods:namePart id="16B934391B11EE17FFC7919F15F4F24F">Fujikawa, Shunsuke</mods:namePart>
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<mods:namePart id="FD3738ABA1EDD3D989352EA41D61B567">Dan, Shigeki</mods:namePart>
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<mods:namePart id="ADA1FA9FD01EB1F1A16FF9B9CAE408D2">Kitada, Shuichi</mods:namePart>
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<treatment id="3F1D0E356345701C68EB4623BEB6F9B9" ID-DOI="http://doi.org/10.5281/zenodo.6004792" ID-GBIF-Taxon="127628083" ID-Zenodo-Dep="6004792" LSID="urn:lsid:plazi:treatment:3F1D0E356345701C68EB4623BEB6F9B9" httpUri="http://treatment.plazi.org/id/3F1D0E356345701C68EB4623BEB6F9B9" lastPageId="12" lastPageNumber="529" pageId="8" pageNumber="529">
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<taxonomicName id="70B4C4A06345701868EB4623BA4FF947" ID-CoL="f904166e-5d83-4c25-aaba-9eb45fda0d5c" box="[151,360,1650,1675]" class="Malacostraca" family="Coenobitidae" genus="Coenobita" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="cavipes">
<emphasis id="85C063316345701868EB4623BA4FF947" bold="true" box="[151,360,1650,1675]" italics="true" pageId="8" pageNumber="529">Coenobita cavipes</emphasis>
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The shield of the first stage crab (
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SL) has faint brownish pigments (
<figureCitation id="2F8FA3A6634570186C1C46C7BF84F963" box="[1120,1187,1686,1711]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[178,1416,194,1862]" captionTargetId="figure@13.[172,1416,193,1862]" captionTargetPageId="13" captionText="FIGURE 7. Coenobita cavipes. A (whole animal, dorsal view), B (animal showing shield, posterior carapace, and some appendages, right lateral view), C (animal showing ocular peduncles, ventral view), O 2 (sample no. in Table 1, 1.03 mm SL, 0 months old); D (whole animal, dorsal view), E (whole animal, left lateral view), O 5 (1.15 mm SL, 1 month old); F (animal showing shield, posterior carapace, and some appendages, dorsal view), O 7 (1.43 mm SL, 2 months old)." httpUri="https://zenodo.org/record/265142/files/figure.png" pageId="8" pageNumber="529">Fig. 7</figureCitation>
A, B); they develop as two longitudinal brown bands/patches on the middle portion and a dark brown band at the anterior lower portion of each side wall of the whitish/creamy shield in juveniles with an SL of&gt;
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after one month of age (
<figureCitation id="2F8FA3A6634570186D5F468FBE42F93A" box="[1315,1381,1758,1783]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[178,1416,194,1862]" captionTargetId="figure@13.[172,1416,193,1862]" captionTargetPageId="13" captionText="FIGURE 7. Coenobita cavipes. A (whole animal, dorsal view), B (animal showing shield, posterior carapace, and some appendages, right lateral view), C (animal showing ocular peduncles, ventral view), O 2 (sample no. in Table 1, 1.03 mm SL, 0 months old); D (whole animal, dorsal view), E (whole animal, left lateral view), O 5 (1.15 mm SL, 1 month old); F (animal showing shield, posterior carapace, and some appendages, dorsal view), O 7 (1.43 mm SL, 2 months old)." httpUri="https://zenodo.org/record/265142/files/figure.png" pageId="8" pageNumber="529">Fig. 7</figureCitation>
DF, 8AD, G). The ocular peduncle of the first stage crab has a brownish pigment area ventrally (
<figureCitation id="2F8FA3A6634570186C9A4753BE0AF8D7" box="[1254,1325,1794,1819]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[178,1416,194,1862]" captionTargetId="figure@13.[172,1416,193,1862]" captionTargetPageId="13" captionText="FIGURE 7. Coenobita cavipes. A (whole animal, dorsal view), B (animal showing shield, posterior carapace, and some appendages, right lateral view), C (animal showing ocular peduncles, ventral view), O 2 (sample no. in Table 1, 1.03 mm SL, 0 months old); D (whole animal, dorsal view), E (whole animal, left lateral view), O 5 (1.15 mm SL, 1 month old); F (animal showing shield, posterior carapace, and some appendages, dorsal view), O 7 (1.43 mm SL, 2 months old)." httpUri="https://zenodo.org/record/265142/files/figure.png" pageId="8" pageNumber="529">Fig. 7</figureCitation>
C), and it gradually develops to a dark brown color after one month of age (
<figureCitation id="2F8FA3A6634570186B0D4777B894F8F3" box="[881,947,1830,1855]" captionStart="FIGURE 8" captionStartId="14.[151,250,1798,1820]" captionTargetBox="[155,1436,196,1776]" captionTargetId="figure@14.[151,1436,193,1776]" captionTargetPageId="14" captionText="FIGURE 8. Coenobita cavipes. A (animal showing ocular acicles and shield, dorsal view), B (animal showing shield and posterior carapace, left lateral view), O 7 (sample no. in Table 1, 1.43 mm SL, 2 months old); C (animal showing shield, posterior carapace, and some appendages, dorsal view), D (whole animal, left dorsal view); E (animal showing ocular peduncles, ventral view), O 12 (1.42 mm SL, 3 months old); F (animal showing left appendages, dorsal view), G (whole animal, left lateral view), O 31 (2.49 mm SL, 10 months old)." httpUri="https://zenodo.org/record/265143/files/figure.png" pageId="8" pageNumber="529">Fig. 8</figureCitation>
E). The ocular acicles and the upper edge of the shield exhibit a brown color in juveniles with an SL of&gt;
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after two months of age (
<figureCitation id="2F8FA3A6634570186CA8471BBE30F8AE" box="[1236,1303,1866,1891]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[178,1416,194,1862]" captionTargetId="figure@13.[172,1416,193,1862]" captionTargetPageId="13" captionText="FIGURE 7. Coenobita cavipes. A (whole animal, dorsal view), B (animal showing shield, posterior carapace, and some appendages, right lateral view), C (animal showing ocular peduncles, ventral view), O 2 (sample no. in Table 1, 1.03 mm SL, 0 months old); D (whole animal, dorsal view), E (whole animal, left lateral view), O 5 (1.15 mm SL, 1 month old); F (animal showing shield, posterior carapace, and some appendages, dorsal view), O 7 (1.43 mm SL, 2 months old)." httpUri="https://zenodo.org/record/265142/files/figure.png" pageId="8" pageNumber="529">Fig. 7</figureCitation>
F, 8A). The second and third pereiopods of the first stage crabs are reddish brown, excluding the distal and proximal parts of each segment; they appear to have whitish rings at the boundaries between segments (
<figureCitation id="2F8FA3A6634570186C2747C3BFBAF867" box="[1115,1181,1938,1963]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[178,1416,194,1862]" captionTargetId="figure@13.[172,1416,193,1862]" captionTargetPageId="13" captionText="FIGURE 7. Coenobita cavipes. A (whole animal, dorsal view), B (animal showing shield, posterior carapace, and some appendages, right lateral view), C (animal showing ocular peduncles, ventral view), O 2 (sample no. in Table 1, 1.03 mm SL, 0 months old); D (whole animal, dorsal view), E (whole animal, left lateral view), O 5 (1.15 mm SL, 1 month old); F (animal showing shield, posterior carapace, and some appendages, dorsal view), O 7 (1.43 mm SL, 2 months old)." httpUri="https://zenodo.org/record/265142/files/figure.png" pageId="8" pageNumber="529">Fig. 7</figureCitation>
A, B). The palm of the left cheliped begins to develop brownish pigment areas on the upper and lower parts after one month of age (
<figureCitation id="2F8FA3A6634570186D1347E7BB82F83E" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[178,1416,194,1862]" captionTargetId="figure@13.[172,1416,193,1862]" captionTargetPageId="13" captionText="FIGURE 7. Coenobita cavipes. A (whole animal, dorsal view), B (animal showing shield, posterior carapace, and some appendages, right lateral view), C (animal showing ocular peduncles, ventral view), O 2 (sample no. in Table 1, 1.03 mm SL, 0 months old); D (whole animal, dorsal view), E (whole animal, left lateral view), O 5 (1.15 mm SL, 1 month old); F (animal showing shield, posterior carapace, and some appendages, dorsal view), O 7 (1.43 mm SL, 2 months old)." httpUri="https://zenodo.org/record/265142/files/figure.png" pageId="8" pageNumber="529">Fig. 7</figureCitation>
D, E); specifically, the pigment on the lower part develops as a distinct large brown patch, and the palm appears to have a longitudinal white stripe on the middle portion (
<figureCitation id="2F8FA3A66341701C6B7A4669B86FF99C" box="[774,840,1592,1617]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[178,1416,194,1862]" captionTargetId="figure@13.[172,1416,193,1862]" captionTargetPageId="13" captionText="FIGURE 7. Coenobita cavipes. A (whole animal, dorsal view), B (animal showing shield, posterior carapace, and some appendages, right lateral view), C (animal showing ocular peduncles, ventral view), O 2 (sample no. in Table 1, 1.03 mm SL, 0 months old); D (whole animal, dorsal view), E (whole animal, left lateral view), O 5 (1.15 mm SL, 1 month old); F (animal showing shield, posterior carapace, and some appendages, dorsal view), O 7 (1.43 mm SL, 2 months old)." httpUri="https://zenodo.org/record/265142/files/figure.png" pageId="12" pageNumber="533">Fig. 7</figureCitation>
F, 8C, D, F, G). The brownish bands also develop on the carpi and meri of the chelipeds, and the bands of the pereiopods become dark brown (
<figureCitation id="2F8FA3A66341701C6CFC460DBFE6F9B9" box="[1152,1217,1628,1653]" captionStart="FIGURE 7" captionStartId="13.[151,250,1910,1932]" captionTargetBox="[178,1416,194,1862]" captionTargetId="figure@13.[172,1416,193,1862]" captionTargetPageId="13" captionText="FIGURE 7. Coenobita cavipes. A (whole animal, dorsal view), B (animal showing shield, posterior carapace, and some appendages, right lateral view), C (animal showing ocular peduncles, ventral view), O 2 (sample no. in Table 1, 1.03 mm SL, 0 months old); D (whole animal, dorsal view), E (whole animal, left lateral view), O 5 (1.15 mm SL, 1 month old); F (animal showing shield, posterior carapace, and some appendages, dorsal view), O 7 (1.43 mm SL, 2 months old)." httpUri="https://zenodo.org/record/265142/files/figure.png" pageId="12" pageNumber="533">Fig. 7</figureCitation>
DF, 8C, D, F, G).
</paragraph>
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