treatments-xml/data/36/58/05/36580511724E57F9B82F260D188B29F9.xml

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<mods:title id="B8EBAFD909DDAA51A217C5120439257E">Taxonomic study on the subgenus Orientostichus, the Pterostichus pulcher species group (Coleoptera, Carabidae, Pterostichus)</mods:title>
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<mods:namePart id="AD936D36AA1EF5634E96ADABB52E7988">Yin, Wenqi</mods:namePart>
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<mods:namePart id="831396F87A58428343D33DB11321205C">Zhu, Pingzhou</mods:namePart>
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<mods:affiliation id="67F602DCB9359A983FBE333C9A883C06">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100093, China</mods:affiliation>
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<mods:namePart id="AA617BB90F4927CFD8018D4DDA7D2031">Shi, Hongliang</mods:namePart>
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<mods:affiliation id="7EEC5EED3EFBA20FB6895683CD907E70">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<treatment id="36580511724E57F9B82F260D188B29F9" LSID="urn:lsid:plazi:treatment:36580511724E57F9B82F260D188B29F9" httpUri="http://treatment.plazi.org/id/36580511724E57F9B82F260D188B29F9" lastPageNumber="163" pageId="0" pageNumber="163">
<subSubSection id="1A359E5B817D4B9F4E3B367DA545CAE4" pageId="0" pageNumber="163" type="nomenclature">
<paragraph id="E40989F470ADAD4D525B1BAD6A59C811" pageId="0" pageNumber="163">
<taxonomicName id="8A5F47F6BBD216A6C6E05EF1E3DD629B" ID-CoL="4Q46G" LSID="36580511-724E-57F9-B82F-260D188B29F9" authority="Sciaky &amp; Allegro, 2013" authorityName="Sciaky &amp; Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher" subGenus="Orientostichus">Pterostichus (Orientostichus) pulcher Sciaky &amp; Allegro, 2013</taxonomicName>
</paragraph>
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<subSubSection id="F1184D4B2D56AA7F3F03DDD9FE1300FF" pageId="0" pageNumber="163" type="description">
<paragraph id="9B732B5807DEFBE7539F1BC7400B1A2C" pageId="0" pageNumber="163">
<figureCitation id="A917864CE9C22D639573B17E253FBBBE" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">Figs 24</figureCitation>
<figureCitation id="AB8A4D978B7DF1EB954E0B16E9E99F1F" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. 25 - 28 endophallus bsp of Pterostichus pulcher species group, left lateral view 25 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 26 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 27 P. condylus sp. nov., paratype from Mianning county 28 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county 29, 30 Endophallus of Pterostichus (Orientostichus) spp., left lateral view and right lateral view 29 P. prattii Bates, 1890, a male from Sanming city, Fujian province 30 P. curtatus Fairmaire, a male from Heqing county, Yunnan province. Scale bars: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures25-30" httpUri="https://binary.pensoft.net/fig/893317" pageId="0" pageNumber="163">, 28</figureCitation>
<figureCitation id="E78D05E3433995C5866661559A336B86" captionStart="Figures 31–33" captionStartId="F6" captionText="Figures 31 - 33. Holotype of Pterostichus pulcher Sciaky &amp; Allegro, 2013 31 habitus and labels 32 pronotum 33 male genitalia, left lateral view and dorsal view of median lobe of aedeagus. Scale bars: 5 mm (31); 1 mm (32, 33)." figureDoi="10.3897/zookeys.1175.107636.figures31-33" httpUri="https://binary.pensoft.net/fig/893318" pageId="0" pageNumber="163">, 31-33</figureCitation>
<figureCitation id="D8903E733211CEFE5C049F342366E738" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">
, 39 Chinese vernacular name: 
<normalizedToken id="3D55EBA22F6150B944D098BAB31F1E08" originalValue="俊通缘步甲">俊通缘步甲</normalizedToken>
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</paragraph>
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<paragraph id="019BBF4CBB92C9613580D266A0A0DDB7" pageId="0" pageNumber="163">
<taxonomicName id="25353A5FAF3E175E5F7D67B88E8C2B6D" authorityName="Sciaky &amp; Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher">Pterostichus pulcher</taxonomicName>
Sciaky &amp; Allegro, 2013: 114. (holotype in IZAS; type locality: CHINA: Prov. Sichuan, Yizhi Yakou).
</paragraph>
</subSubSection>
<subSubSection id="618A8D3B4B8D417780C00CA6079E3072" pageId="0" pageNumber="163" type="materials_examined">
<paragraph id="0601BFA982CF9F534B12A4549BEDD43C" pageId="0" pageNumber="163">Material examined.</paragraph>
<paragraph id="73FBCB2B8FCC16EB42B2A901DB90ADE3" pageId="0" pageNumber="163">
<materialsCitation id="7CD67DB8785990D67F987A93E3F2654A" collectingDate="2007-07-19" collectorName="Cavazzuti P. F." country="China" elevation="2950" location="Yizhi Yakou" specimenCount="♂" stateProvince="Sichuan" typeStatus="Holotype">
<emphasis id="1400220AA8ABD3341F250259F0C6610E" bold="true" italics="true" pageId="0" pageNumber="163">
<typeStatus id="9603768F07F483224DCE9E7FD6D7356B">Holotype</typeStatus>
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: 
<specimenCount id="3BC2BB0DF10D61C1202C1AE70D8A0FF5">♂</specimenCount>
: &quot; 
<collectingCountry id="D569667B693F631170908E39091AFD84" name="China">China</collectingCountry>
, 
<collectingRegion id="4AE5A0364FE76A6B39ED8A21F56CE74D" country="China" name="Sichuan">Sichuan</collectingRegion>
merid. 
<normalizedToken id="95D78C4AD7D2544DC0E22FA9D1AB1858" originalValue="E’bian">E'bian</normalizedToken>
- 
<location id="15957C247871EE930041E4734C3BFC1D" LSID="urn:lsid:plazi:treatment:36580511724E57F9B82F260D188B29F9:15957C247871EE930041E4734C3BFC1D" country="China" name="Yizhi Yakou" stateProvince="Sichuan">Yizhi Yakou</location>
, Xiaoliang Shan - Si He. 
<quantity id="8FD809A8A42AD9E68AC900C2EDE8F036" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" unit="m" value="2950.0" valueMax="3100.0" valueMin="2800.0">
<elevation id="8BD055EAD6AB6C55A5A92452A3EC8D23" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" unit="m" value="2950.0" valueMax="3100.0" valueMin="2800.0">2800-3100 m</elevation>
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, 
<collectingDate id="DFB821A73F48D00910252B28E2EA2884" value="2007-07-19">2007.VII.19</collectingDate>
, 
<collectorName id="2FF657C343AEC2F16177B8ACD49A3A0A">Cavazzuti P.F.</collectorName>
leg.&quot;, &quot;2013 from Sciaky.&quot;, &quot; 
<typeStatus id="7DDC618441853F7FDCDC7315583DFE94">HOLOTYPE</typeStatus>
<specimenCount id="67D9EBECC8B3B0D874212656616D12DF">♂</specimenCount>
<taxonomicName id="BE5F0526202E4067F93322CD29864C50" authorityName="Sciaky &amp; Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher" subGenus="Orientostichus">Pterostichus (Orientostichus) pulcher</taxonomicName>
sp. nov. des. Sciaky &amp; Allegro, 2013&quot; [red label]
</materialsCitation>
. 
<materialsCitation id="0D0403B3E224D3CD0DF536BA6185EBFF" collectingDate="2012-06-15" collectingMethod="by pitfall trap" collectorName="Liu Ye" country="China" county="Yizi" elevation="2923" location="Institute of Zoology" municipality="Meigu county" specimenCount="8" specimenCount-male="8" stateProvince="Sichuan Prov." typeStatus="Paratypes">
<emphasis id="07D6545E26A08ED4B72A21BDF8DC7697" bold="true" italics="true" pageId="0" pageNumber="163">
<typeStatus id="D32A2CD6E5C2896696A342862D092BED">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="F78AAC3560835DEE42EAECFA88ED277F" type="male">8 ♂</specimenCount>
: &quot; 
<collectingCountry id="77904EF2871C323DE0472C1FF58FC6B5" name="China">China</collectingCountry>
, 
<collectingRegion id="08FB60557D349A1A42B5F74E66C115D7" country="China" name="Sichuan">Sichuan Prov.</collectingRegion>
, 
<collectingCounty id="5FC5F5045741E80B15BF31E247287895">Yizi</collectingCounty>
pass btw., 
<collectingMunicipality id="42D7354E27530FF06290386F2DC87DC7">Meigu county</collectingMunicipality>
and 
<normalizedToken id="868C30A7D663C81FA0CDF28050FDD6A7" originalValue="E’bian">E'bian</normalizedToken>
county, mixed forest; N28.67477, E103.05248&quot;, &quot; 
<quantity id="9DAFFF4172EAD2C65F4FA117954B1C86" metricMagnitude="3" metricUnit="m" metricValue="2.923" unit="m" value="2923.0">
<elevation id="C67FBDDE4BE52072EFADD9580B71E30A" metricMagnitude="3" metricUnit="m" metricValue="2.923" unit="m" value="2923.0">2923 m</elevation>
</quantity>
; 
<collectingDate id="12D6E66B7E965838E15620138A4539B7" value="2012-06-15">2012.VI.15</collectingDate>
; 
<collectingMethod id="D3DA4B8D17C52DFD4E77316467A144A5">by pitfall trap</collectingMethod>
; Shi Hongliang &amp; 
<collectorName id="3B9343A313182E8CBC7D4C72EBBEF29B">Liu Ye</collectorName>
leg. 
<location id="31425C1C4A02B7C78A94AC55C6B88CC2" LSID="urn:lsid:plazi:treatment:36580511724E57F9B82F260D188B29F9:31425C1C4A02B7C78A94AC55C6B88CC2" country="China" county="Yizi" municipality="Meigu county" name="Institute of Zoology" stateProvince="Sichuan Prov.">Institute of Zoology</location>
, CAS.&quot;, &quot; 
<typeStatus id="8C252C4AD850157A9E09BECA37D20FCB">PARATYPE</typeStatus>
of 
<taxonomicName id="C4235202F16D8AB8412890F095559BFE" authorityName="Sciaky &amp; Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher" subGenus="Orientostichus">Pterostichus (Orientostichus) pulcher</taxonomicName>
sp. nov. des. Sciaky &amp; Allegro, 2013&quot; [red label]
</materialsCitation>
. 
<emphasis id="FEB2810A831A6D8B5F5ED5A01D6EC59E" bold="true" italics="true" pageId="0" pageNumber="163">Non-type materials</emphasis>
: 
<materialsCitation id="5544696BC6E62E2E123AE6F91424594A" collectingDate="2019-07-22" country="China" elevation="2859" latitude="28.676586" location="Yi Autonomous county" longLatPrecision="1" longitude="103.05599" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Sichuan province">
<specimenCount id="9C5145F824F2DEFC0F459ADC1A7994CF" type="male">3 ♂</specimenCount>
and 
<specimenCount id="4938F1F00E94B2542843418AD1523955" type="female">6 ♀</specimenCount>
: &quot; 
<collectingRegion id="F1704DAE526D324E8791B56C5917D04A" country="China" name="Sichuan">Sichuan province</collectingRegion>
, Leshan city, 
<normalizedToken id="566AC5936F649B425A6B008ED6C41CF0" originalValue="e’bian">e'bian</normalizedToken>
<location id="264AC5A5CEF764CE8858B2D2FB336490" LSID="urn:lsid:plazi:treatment:36580511724E57F9B82F260D188B29F9:264AC5A5CEF764CE8858B2D2FB336490" country="China" latitude="28.676586" longLatPrecision="1" longitude="103.05599" name="Yi Autonomous county" stateProvince="Sichuan province">Yi Autonomous county</location>
, Heizhugou No. 615 forestry centre, 
<geoCoordinate id="55D70DFFCF482ACBA217F493E8FE04B3" degrees="103.055994" direction="east" orientation="longitude" precision="1" value="103.05599">103.055994E</geoCoordinate>
, 
<geoCoordinate id="2C3D9003B1F28B07CE03784401336631" degrees="28.676587" direction="north" orientation="latitude" precision="1" value="28.676586">28.676587N</geoCoordinate>
, 
<quantity id="EADD32AE1ABDFA5642198F36605CC1AF" metricMagnitude="3" metricUnit="m" metricValue="2.8589999999999995" unit="m" value="2859.0">
<elevation id="C108DAAC7E210B08C9BB3F8E59E46E7D" metricMagnitude="3" metricUnit="m" metricValue="2.8589999999999995" unit="m" value="2859.0">2859 m</elevation>
</quantity>
, Zhudong Liu, Zhiming Li and Tao Li lgt., 
<collectingDate id="A13F34F0D2F49BF837615804B9E9D8B9" value="2019-07-22">2019.VII.22</collectingDate>
&quot;
</materialsCitation>
; 
<materialsCitation id="B87CCA39446F98826AF41F4C3B3F850E" collectingDate="2020-05-28" country="China" elevation="2859" latitude="28.676586" location="Yi Autonomous county" longLatPrecision="1" longitude="103.05599" specimenCount="27" specimenCount-female="9" specimenCount-male="18" stateProvince="Sichuan province">
<specimenCount id="8A59F5468BEDB719DA20981B9785F543" type="male">18 ♂</specimenCount>
and 
<specimenCount id="2A33899FA35BBF441534317BAC0F3B96" type="female">9 ♀</specimenCount>
: &quot; 
<collectingRegion id="B8FB923A777C454411703DBD7208B204" country="China" name="Sichuan">Sichuan province</collectingRegion>
, Leshan city, 
<normalizedToken id="CA4FD037EFEB9DCBE281A339CBF03012" originalValue="e’bian">e'bian</normalizedToken>
<location id="FAF737ACE4F6714AB5222E985E9CDF35" LSID="urn:lsid:plazi:treatment:36580511724E57F9B82F260D188B29F9:FAF737ACE4F6714AB5222E985E9CDF35" country="China" latitude="28.676586" longLatPrecision="1" longitude="103.05599" name="Yi Autonomous county" stateProvince="Sichuan province">Yi Autonomous county</location>
, Heizhugou No.615 forestry centre, 
<geoCoordinate id="C97503BEAA3CC10A1EE2B6F56A369A3B" degrees="103.055994" direction="east" orientation="longitude" precision="1" value="103.05599">103.055994E</geoCoordinate>
, 
<geoCoordinate id="A94CE2FFE1D5AF48FFF9C2C2B5405122" degrees="28.676587" direction="north" orientation="latitude" precision="1" value="28.676586">28.676587N</geoCoordinate>
, 
<quantity id="BFB900F05852FD594DC58001C47E348B" metricMagnitude="3" metricUnit="m" metricValue="2.8589999999999995" unit="m" value="2859.0">
<elevation id="9F28FFE369B4422181A7B3297198CC52" metricMagnitude="3" metricUnit="m" metricValue="2.8589999999999995" unit="m" value="2859.0">2859 m</elevation>
</quantity>
, Zhudong Liu, Zhiming Li and Pingzhou Zhu lgt., 
<collectingDate id="F3DC753CF9B5C4D7093B4751B54A2550" value="2020-05-28">2020.V.28</collectingDate>
&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4AC1183D383B5A117AB662431B93E121" pageId="0" pageNumber="163" type="diagnosis">
<paragraph id="E12405A50CD2D6B350F8DAE9B6500771" pageId="0" pageNumber="163">Diagnosis.</paragraph>
<paragraph id="756A127C8474C305C364C614D54CF9CE" pageId="0" pageNumber="163">
Elytra with several large foveate discal pores on intervals 3, 5, and 7, forming strong catenulate sculpturing. Antennomere 3 without accessory setae. Pronotum subcordate, slightly narrowed to base (PBW/PW = 0.75-0.81); lateral margins evidently sinuate before posterior angles, posterior angles acute at apex, distinctly projected laterally; lateral margins with 2-4 mid-lateral setae; pronotal disc with shallow transverse wrinkles; basal foveae with inner and outer grooves well-defined (Fig. 
<figureCitation id="C8D27CF4882C895EA558938BEA68A722" captionStart="Figures 31–33" captionStartId="F6" captionText="Figures 31 - 33. Holotype of Pterostichus pulcher Sciaky &amp; Allegro, 2013 31 habitus and labels 32 pronotum 33 male genitalia, left lateral view and dorsal view of median lobe of aedeagus. Scale bars: 5 mm (31); 1 mm (32, 33)." figureDoi="10.3897/zookeys.1175.107636.figures31-33" httpUri="https://binary.pensoft.net/fig/893318" pageId="0" pageNumber="163">32</figureCitation>
), area between them strongly convex. Male sternite VII with single well-defined smooth tubercle (Fig. 
<figureCitation id="B1D3492D3D847F6363A8BB681ECA58E6" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">39</figureCitation>
). Apical lamella of male genitalia elongate (LL/LW = 1.75-1.95), lateral margins subparallel to apex, apex rounded (Fig. 
<figureCitation id="30ED8FA5A5C986AC7AD8A02BE7D059F5" captionStart="Figures 31–33" captionStartId="F6" captionText="Figures 31 - 33. Holotype of Pterostichus pulcher Sciaky &amp; Allegro, 2013 31 habitus and labels 32 pronotum 33 male genitalia, left lateral view and dorsal view of median lobe of aedeagus. Scale bars: 5 mm (31); 1 mm (32, 33)." figureDoi="10.3897/zookeys.1175.107636.figures31-33" httpUri="https://binary.pensoft.net/fig/893318" pageId="0" pageNumber="163">33</figureCitation>
); endophallus with U-shaped bsp, left branch evenly convex on dorsal margin (Figs 
<figureCitation id="5704FAC0CF711356F60E8C11918D7A21" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">24</figureCitation>
, 
<figureCitation id="9181454E02CC822D638D97B341B9015E" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. 25 - 28 endophallus bsp of Pterostichus pulcher species group, left lateral view 25 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 26 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 27 P. condylus sp. nov., paratype from Mianning county 28 P. pulcher Sciaky &amp; Allegro, paratype from Yizi Yakou, E'bian county to Meigu county 29, 30 Endophallus of Pterostichus (Orientostichus) spp., left lateral view and right lateral view 29 P. prattii Bates, 1890, a male from Sanming city, Fujian province 30 P. curtatus Fairmaire, a male from Heqing county, Yunnan province. Scale bars: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures25-30" httpUri="https://binary.pensoft.net/fig/893317" pageId="0" pageNumber="163">28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8B8E263380936C71BAC2462AB71F6D85" pageId="0" pageNumber="163" type="comparisons">
<paragraph id="971DB78CFDF4B35E67292943B2F3E61A" pageId="0" pageNumber="163">Comparison.</paragraph>
<paragraph id="7BB411FD072E51E20A9BEE7B8A99A98B" pageId="0" pageNumber="163">
<taxonomicName id="9F029F59FDA091B0DCD3618BAE00E680" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="9C0B86FED6E19F4890C2CBCCD9C3D7D2" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4B58FC208F413D358AA425910248FAD0" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
<emphasis id="F3D55428F92054530F14CF59CEB87A96" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
</taxonomicName>
sp. nov. for their interval 7 with several foveate discal pores. This character clearly distinguishes them from the other five species in the species group. The comparisons between these two species were provided under 
<taxonomicName id="B07FE7DB402182EAB23E93C15B9E4A3C" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
<emphasis id="B525C0539172159F3257E26490C6FFA0" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="A8F74F2B0C14818E4F6D4DBD80FE5D90" pageId="0" pageNumber="163" type="distribution">
<paragraph id="74D1571601C125BDA53730AA32CDE7AA" pageId="0" pageNumber="163">Distribution.</paragraph>
<paragraph id="C2A7B505BC964479D7D1B4BFBF370DF0" pageId="0" pageNumber="163">
This species has a narrow distribution range on the border of Meigu and 
<normalizedToken id="9372A203CD0D98D8C991601CB680E9A5" originalValue="E’bian">E'bian</normalizedToken>
counties, Liangshan Yi Autonomous Prefecture, but locally abundant (Fig. 
<figureCitation id="CA47086BACFF59C54D33BDAAFED7DAF4" captionStart="Figure 42" captionStartId="F9" captionText="Figure 42. Distribution of the taxa of Pterostichus pulcher species group in Sichuan province, China. Brown: P. pemphis sp. nov. Yellow: P. orbicollis sp. nov. Red: P. leo sp. nov. Purple: P. liyuani sp. nov. Blue: P. condylus sp. nov. Green: P. jialini sp. nov. Orange: P. pulcher Sciaky &amp; Allegro." figureDoi="10.3897/zookeys.1175.107636.figure42" httpUri="https://binary.pensoft.net/fig/893321" pageId="0" pageNumber="163">42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8550569191C1E057F6D8566E3065F327" pageId="0" pageNumber="163" type="species removed from subgenus orientostichus sciaky &amp; allegro">
<paragraph id="A7BFC5F6482E01D53BFF86602F1AF46F" pageId="0" pageNumber="163">
Species removed from subgenus 
<taxonomicName id="D98638FD229FE3AE6D5E15D2D246E34C" authorityName="Sciaky &amp; Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus)" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="subGenus" subGenus="Orientostichus">Pterostichus Orientostichus</taxonomicName>
Sciaky &amp; Allegro
</paragraph>
</subSubSection>
</treatment>
</document>