treatments-xml/data/19/75/B7/1975B758FFA98C10FF30FF67FAAE6371.xml

<document id="CC91B1709BBB367877E016978DB3875F" ID-DOI="10.11606/1807-0205/2020.60.42" ID-ISSN="1807-0205" ID-Zenodo-Dep="4614797" ID-ZooBank="99FB82B2-4D27-4387-B0A0-E3252A77F819" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616007701910" checkinUser="carolina" docAuthor="Cegolin, Bianca Melo, Bueno, Gabriel Marangão, Pereira, Gabriel Lopes, Santos, Daubian &amp; Santos, Charles Morphy Dias dos" docDate="2020" docId="1975B758FFA98C10FF30FF67FAAE6371" docLanguage="en" docName="PapAvZoo.60.e20206042.pdf" docOrigin="Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 60" docSource="http://dx.doi.org/10.11606/1807-0205/2020.60.42" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Chrysopilus kafkai Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="E54CCF20FFAD8C17FFD1FFBDFFEE6622" masterDocTitle="Description and molecular characterization of Chrysopilus kafkai sp. nov. (Diptera: Rhagionidae) from Serra da Bodoquena (Mato Grosso do Sul, Brazil)" masterLastPageNumber="10" masterPageNumber="1" pageNumber="5" updateTime="1698889136212" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<mods:title id="A4A860B1395972FA5FCABBAB94B955E0">Description and molecular characterization of Chrysopilus kafkai sp. nov. (Diptera: Rhagionidae) from Serra da Bodoquena (Mato Grosso do Sul, Brazil)</mods:title>
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<mods:namePart id="821C5562BFADE0EC97B91F9B3BB64369">Cegolin, Bianca Melo</mods:namePart>
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<mods:namePart id="58D0659B7858B985BCA1FF1C7DFE7D45">Bueno, Gabriel Marangão</mods:namePart>
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<mods:namePart id="D7A0F71BD582750049F34DF20DA2D334">Santos, Daubian</mods:namePart>
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<taxonomicName id="56DC7DCDFFA98C13FF30FF67FD6A66D1" authority="Cegolin &amp; Santos" authorityName="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos" authorityYear="2020" box="[225,644,218,243]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kafkai" status="sp. nov.">
<emphasis id="A3A8DA5CFFA98C13FF30FF67FE5866D1" bold="true" box="[225,438,218,243]" italics="true" pageId="4" pageNumber="5">Chrysopilus kafkai</emphasis>
Cegolin &amp; Santos
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<figureCitation id="09E71ACBFFA98C13FE60FF47FDFD6731" box="[433,531,250,275]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStartId-0="1.[94,153,990,1015]" captionStartId-1="1.[94,153,1741,1766]" captionStartId-2="2.[171,230,2042,2067]" captionStartId-3="3.[94,152,2042,2067]" captionStartId-4="4.[170,229,2106,2131]" captionStartId-5="5.[94,153,725,750]" captionStartId-6="5.[94,153,2074,2099]" captionTargetBox-0="[101,1417,204,960]" captionTargetBox-1="[96,718,1069,1709]" captionTargetBox-2="[170,1493,696,2012]" captionTargetBox-3="[94,1414,692,2012]" captionTargetBox-4="[170,1493,426,2076]" captionTargetBox-5="[94,718,204,695]" captionTargetBox-6="[126,1385,781,2041]" captionTargetId-0="figure-360@1.[94,1417,204,960]" captionTargetId-1="figure-315@1.[95,718,1068,1711]" captionTargetId-2="figure-210@2.[170,1493,689,2012]" captionTargetId-3="figure-133@3.[94,1417,689,2012]" captionTargetId-4="figure-95@4.[170,1493,413,2076]" captionTargetId-5="figure-228@5.[94,718,204,695]" captionTargetId-6="figure-240@5.[123,1389,778,2044]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="2" captionTargetPageId-3="3" captionTargetPageId-4="4" captionTargetPageId-5="5" captionTargetPageId-6="5" captionText-0="Figure 1. Chrysopilus kafkai, sp. nov., male holotype. (A) Habitus sp. nov., lateral view. (B) Head: frontal view. Abbreviations: ar, arista; cl,clypeus; ff, first flagellomere;ot, ocellar tubercle;pe, pedicel;pl, palpus;scp,scape. MATERIAL AND METHODS" captionText-1="Figure 2. Thorax of Chrysopilus kafkai sp. nov., male holotype: lateral view. Abbreviations: a spr, anterior spiracle; anepm, anepimeron; anepst, anepisternum;cx, coxa; kepst, katepisternum; ltg, laterotergite; mr, meron; mtepm, metepimeron; mtg, mediotergite; mtkepst,metakatepisternum; sct, scutum; sctl,scutellum,sbsctl, subscutellum. the Serra da Bodoquena (Mato Grosso do Sul, Brazil), a national park located in the Cerrado. These expeditions were organized by the SISBIOTA-Diptera project (Lamas et al., 2014). The new species described here is the first record of the family Rhagionidae in the Brazilian Midwest Brazilian region. The description includes both morphological and molecular data." captionText-2="Figure3.Wing ofChrysopiluskafkai sp.nov.,male holotype.(A) Illustration.(B)Photograph.Abbreviations:al, alula;a₁, firstanal cell;bm, basal medialcell;br, basal radial cell;C, costal vein;c,costal cell;CuA, cubital anterior vein;CuP, posterior branch of cubital vein;cua, anterior cubital cell;cup, posterior cubitalcell;d, distal cell; h, humeral;M, medial vein;m‑cu, crossvein m-cu;m‑m, crossvein m-m; R, radialvein;r‑m, crossvein r-m;Sc,subcostal vein." captionText-3="Figure 4.Terminalia ofChrysopilus kafkai sp.nov.,male holotype.(A) Genital capsule,dorsal view.(B)Ventralview.Abbreviations:gs, gonostylus;ce, cercus;t IX,tergite IX.(C) MEV of genital capsule,dorsal view.(D) Genital capsule,dorsal view.(E)Gonostylus,ventral view.Abbreviations:aed,aedeagus;crt, crest;ej ap, ejaculatory apodeme;gcapod,gonocoxalapodeme;gc,gonocoxite;gs,gonostylus;latejproc,lateralejaculatoryprocess;parap, parameralapodeme;par,paramere;spm sac,sperm sac." captionText-4="Figure 5. Habitus of Chrysopilus kafkai sp.nov.,female.(A) Dorsal view.(B) Lateral view.(C) Head,dorsolateral view." captionText-5="Figure 6. Female terminalia of Chrysopilus kafkai sp.nov., lateral view." captionText-6="Figure 7. (A) Female terminalia of Chrysopilus kafkai sp.nov.(B) Spermathecae.Abbreviations:ce, cercus;hyp val, hypogynial valve; pvl, posteroventral cercal lobe; s IX, sterniteIX; sensorg, sensorial organ." figureDoi-0="http://doi.org/10.5281/zenodo.4614801" figureDoi-1="http://doi.org/10.5281/zenodo.4614803" figureDoi-2="http://doi.org/10.5281/zenodo.4614805" figureDoi-3="http://doi.org/10.5281/zenodo.4614809" figureDoi-4="http://doi.org/10.5281/zenodo.4614813" figureDoi-5="http://doi.org/10.5281/zenodo.4614815" figureDoi-6="http://doi.org/10.5281/zenodo.4614819" httpUri-0="https://zenodo.org/record/4614801/files/figure.png" httpUri-1="https://zenodo.org/record/4614803/files/figure.png" httpUri-2="https://zenodo.org/record/4614805/files/figure.png" httpUri-3="https://zenodo.org/record/4614809/files/figure.png" httpUri-4="https://zenodo.org/record/4614813/files/figure.png" httpUri-5="https://zenodo.org/record/4614815/files/figure.png" httpUri-6="https://zenodo.org/record/4614819/files/figure.png" pageId="4" pageNumber="5">Figs. 1‑7</figureCitation>
</emphasis>
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<emphasis id="A3A8DA5CFFA98C13FF7BFE87FEFD6771" bold="true" box="[170,275,314,339]" pageId="4" pageNumber="5">Material:</emphasis>
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<typeStatus id="4E67B8ECFFA98C13FEF1FE87FE7E6771" box="[288,400,314,339]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
:
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, 
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, 
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Serra 
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, 
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, 06-22.
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<materialsCitation id="21B40C13FFA98C13FCB4FF67FA7D6711" collectingDate="2011-12-09" collectorName="Lamas &amp; Nihei" latitude="-20.704168" location="Serra da Bodoquena" longLatPrecision="21" longitude="-56.85" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul" typeStatus="paratype">
xii.2011, 
<collectorName id="3C296398FFA98C13FC14FF67FBE066D1" box="[965,1038,218,243]" pageId="4" pageNumber="5">Lamas</collectorName>
, 
<collectorName id="3C296398FFA98C13FBC6FF67FBBF66D1" box="[1047,1105,218,243]" pageId="4" pageNumber="5">Nihei</collectorName>
&amp; eq. col (MZUSP). 
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<typeStatus id="4E67B8ECFFA98C13FAF6FF67FA7166D1" box="[1319,1439,218,243]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
:
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, 
<collectingRegion id="5318C8ACFFA98C13FCB4FF47FBA06731" box="[869,1102,250,275]" country="Brazil" name="Mato Grosso do Sul" pageId="4" pageNumber="5">Mato Grosso do Sul</collectingRegion>
, 
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, 
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, 
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<collectingDate id="F526D966FFA98C13FC27FEA7FB966711" box="[1014,1144,282,307]" pageId="4" pageNumber="5" value="2011-12-09">09/XII/2011</collectingDate>
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<collectorName id="3C296398FFA98C13FB56FEA7FB3E6711" box="[1159,1232,282,307]" pageId="4" pageNumber="5">Lamas</collectorName>
, 
<collectorName id="3C296398FFA98C13FB0EFEA7FAF76711" box="[1247,1305,282,307]" pageId="4" pageNumber="5">Nihei</collectorName>
&amp; eq. col
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.; 
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<specimenCount id="87DACDC7FFA98C13FA76FEA7FA3E6713" box="[1447,1488,282,307]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
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, 
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<collectingDate id="F526D966FFA98C13FBC1FEE7FB606751" box="[1040,1166,346,371]" pageId="4" pageNumber="5" value="2011-12-22">22/XII/2011</collectingDate>
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- 
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<collectingDate id="F526D966FFA98C13FB46FEE7FAED6751" box="[1175,1283,346,371]" pageId="4" pageNumber="5" value="2012-01-06">06/I/2012</collectingDate>
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, 
<collectorName id="3C296398FFA98C13FAC8FEE7FA8C6751" box="[1305,1378,346,371]" pageId="4" pageNumber="5">Lamas</collectorName>
, 
<collectorName id="3C296398FFA98C13FAA9FEE7FA5C6751" box="[1400,1458,346,371]" pageId="4" pageNumber="5">Nihei</collectorName>
&amp; eq. col
</materialsCitation>
.; 
<materialsCitation id="21B40C13FFA88C12FCA8FF67FB646751" collectingDate="2011-12-06" collectingDateMax="2012-11-21" collectingDateMin="2011-12-06" collectorName="Lamas &amp; Nihei" latitude="-20.698195" location="Serra da Bodoquena" longLatPrecision="1" longitude="-56.882137" municipality="Serra da Bodoquena" pageId="5" pageNumber="6" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul">
<specimenCount id="87DACDC7FFA88C12FCA8FF67FC7166D3" box="[889,927,218,243]" pageId="5" pageNumber="6" type="female">1 ♀</specimenCount>
, 
<collectingRegion id="5318C8ACFFA88C12FC7CFF67FB7E66D1" box="[941,1168,218,243]" country="Brazil" name="Mato Grosso do Sul" pageId="5" pageNumber="6">Mato Grosso do Sul</collectingRegion>
, 
<collectingMunicipality id="71079C34FFA88C12FB4DFF67FA6866D1" box="[1180,1414,218,243]" pageId="5" pageNumber="6">Serra da Bodoquena</collectingMunicipality>
, 
<geoCoordinate id="F4E86089FFA88C12FCCBFF47FC486731" box="[794,934,249,275]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="49.9" value="-20.697193">20°41′49.9″S</geoCoordinate>
, 
<geoCoordinate id="F4E86089FFA88C12FC60FF47FBA86731" box="[945,1094,249,275]" degrees="56" direction="west" minutes="52" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="54.0" value="-56.881664">56°52′54.0″W</geoCoordinate>
, 
<date id="E562208EFFA88C12FB80FF47FB1E6731" box="[1105,1264,250,275]" pageId="5" pageNumber="6" value="2012-11-06" valueMax="2012-11-21" valueMin="2012-11-06">
<collectingDate id="F526D966FFA88C12FB80FF47FB1E6731" box="[1105,1264,250,275]" pageId="5" pageNumber="6" value="2012-11-06" valueMax="2012-11-21" valueMin="2012-11-06">06-21/XI/2012</collectingDate>
</date>
, 
<collectorName id="3C296398FFA88C12FB2AFF47FAAA6731" box="[1275,1348,250,275]" pageId="5" pageNumber="6">Lamas</collectorName>
, 
<collectorName id="3C296398FFA88C12FA9EFF47FA676731" box="[1359,1417,250,275]" pageId="5" pageNumber="6">Nihei</collectorName>
&amp; eq. col. 
<emphasis id="A3A8DA5CFFA88C12FC57FEA7FB9D6711" bold="true" box="[902,1139,282,307]" pageId="5" pageNumber="6">Additional material:</emphasis>
<specimenCount id="87DACDC7FFA88C12FBA8FEA7FB4E6713" box="[1145,1184,282,307]" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
, 
<collectingRegion id="5318C8ACFFA88C12FB7DFEA7FA686711" box="[1196,1414,282,307]" country="Brazil" name="Mato Grosso do Sul" pageId="5" pageNumber="6">Mato Grosso do Sul</collectingRegion>
, 
<location id="94035095FFA88C12FCCAFE87FBEC6771" LSID="urn:lsid:plazi:treatment:1975B758FFA98C10FF30FF67FAAE6371:94035095FFA88C12FCCAFE87FBEC6771" box="[795,1026,314,339]" latitude="-20.698195" longLatPrecision="1" longitude="-56.882137" municipality="Serra da Bodoquena" name="Serra da Bodoquena" pageId="5" pageNumber="6" stateProvince="Mato Grosso do Sul">Serra da Bodoquena</location>
, 
<geoCoordinate id="F4E86089FFA88C12FBDCFE87FB766771" box="[1037,1176,313,339]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="53.5" value="-20.698195">20°41′53.5″S</geoCoordinate>
, 
<geoCoordinate id="F4E86089FFA88C12FB72FE87FAD96771" box="[1187,1335,313,339]" degrees="56" direction="west" minutes="52" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="55.7" value="-56.882137">56°52′55.7″W</geoCoordinate>
, 
<date id="E562208EFFA88C12FA93FE87FC966751" pageId="5" pageNumber="6" value="2011-12-06" valueMax="2011-12-22" valueMin="2011-12-06">
<collectingDate id="F526D966FFA88C12FA93FE87FC966751" pageId="5" pageNumber="6" value="2011-12-06" valueMax="2011-12-22" valueMin="2011-12-06">06-22/ XII/2011</collectingDate>
</date>
, 
<collectorName id="3C296398FFA88C12FC54FEE7FC206751" box="[901,974,346,371]" pageId="5" pageNumber="6">Lamas</collectorName>
, 
<collectorName id="3C296398FFA88C12FC0AFEE7FBFB6751" box="[987,1045,346,371]" pageId="5" pageNumber="6">Nihei</collectorName>
&amp; eq. col
</materialsCitation>
.; 
<materialsCitation id="21B40C13FFA88C12FB4AFEE7FB666791" collectingDate="2011-12-09" collectorName="Lamas &amp; Nihei" latitude="-20.707779" location="Serra da Bodoquena" longLatPrecision="21" longitude="-55.851944" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="1" stateProvince="Mato Grosso do Sul">
<specimenCount id="87DACDC7FFA88C12FB4AFEE7FB2C6753" box="[1179,1218,346,371]" pageId="5" pageNumber="6" type="female">1 ♀</specimenCount>
, 
<collectingRegion id="5318C8ACFFA88C12FB03FEE7FCD067B1" country="Brazil" name="Mato Grosso do Sul" pageId="5" pageNumber="6">Mato Grosso do Sul</collectingRegion>
, 
<location id="94035095FFA88C12FC9CFEC7FBD367B1" LSID="urn:lsid:plazi:treatment:1975B758FFA98C10FF30FF67FAAE6371:94035095FFA88C12FC9CFEC7FBD367B1" box="[845,1085,378,403]" latitude="-20.707779" longLatPrecision="21" longitude="-55.851944" name="Serra da Bodoquena" pageId="5" pageNumber="6" stateProvince="Mato Grosso do Sul">Serra da Bodoquena</location>
, 
<geoCoordinate id="F4E86089FFA88C12FB9DFEC7FB2867B1" box="[1100,1222,377,403]" degrees="20" direction="south" minutes="42" orientation="latitude" pageId="5" pageNumber="6" precision="15" seconds="28" value="-20.707779">20°42′28″S</geoCoordinate>
, 
<geoCoordinate id="F4E86089FFA88C12FB04FEC7FAB867B1" box="[1237,1366,377,403]" degrees="55" direction="west" minutes="51" orientation="longitude" pageId="5" pageNumber="6" precision="15" seconds="07" value="-55.851944">55°51′07″W</geoCoordinate>
, 
<date id="E562208EFFA88C12FAB4FEC7FC966791" pageId="5" pageNumber="6" value="2011-12-09">
<collectingDate id="F526D966FFA88C12FAB4FEC7FC966791" pageId="5" pageNumber="6" value="2011-12-09">09/ XII/2011</collectingDate>
</date>
, 
<collectorName id="3C296398FFA88C12FC55FE27FC236791" box="[900,973,410,435]" pageId="5" pageNumber="6">Lamas</collectorName>
, 
<collectorName id="3C296398FFA88C12FC0BFE27FBFA6791" box="[986,1044,410,435]" pageId="5" pageNumber="6">Nihei</collectorName>
&amp; eq. col
</materialsCitation>
.; 
<materialsCitation id="21B40C13FFA88C12FB48FE27FB7267D1" collectingDate="2011-12-22" collectorName="Lamas &amp; Nihei" latitude="-20.697193" location="Serra da Bodoquena" longLatPrecision="1" longitude="-56.881664" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul">
<specimenCount id="87DACDC7FFA88C12FB48FE27FB2A6793" box="[1177,1220,410,435]" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
, 
<collectingRegion id="5318C8ACFFA88C12FB02FE27FCD067F1" country="Brazil" name="Mato Grosso do Sul" pageId="5" pageNumber="6">Mato Grosso do Sul</collectingRegion>
, 
<location id="94035095FFA88C12FC9EFE07FBAD67F1" LSID="urn:lsid:plazi:treatment:1975B758FFA98C10FF30FF67FAAE6371:94035095FFA88C12FC9EFE07FBAD67F1" box="[847,1091,442,467]" latitude="-20.697193" longLatPrecision="1" longitude="-56.881664" name="Serra da Bodoquena" pageId="5" pageNumber="6" stateProvince="Mato Grosso do Sul">Serra da Bodoquena</location>
, 
<geoCoordinate id="F4E86089FFA88C12FB84FE07FB0F67F1" box="[1109,1249,441,467]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="49.9" value="-20.697193">20°41′49.9″S</geoCoordinate>
, 
<geoCoordinate id="F4E86089FFA88C12FB23FE07FA6667F1" box="[1266,1416,441,467]" degrees="56" direction="west" minutes="52" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="54.0" value="-56.881664">56°52′54.0″W</geoCoordinate>
, 
<date id="E562208EFFA88C12FCCBFE67FC7267D1" box="[794,924,474,499]" pageId="5" pageNumber="6" value="2011-12-22">
<collectingDate id="F526D966FFA88C12FCCBFE67FC7267D1" box="[794,924,474,499]" pageId="5" pageNumber="6" value="2011-12-22">22/XII/2011</collectingDate>
</date>
, 
<collectorName id="3C296398FFA88C12FC74FE67FC0067D1" box="[933,1006,474,499]" pageId="5" pageNumber="6">Lamas</collectorName>
, 
<collectorName id="3C296398FFA88C12FC29FE67FBDC67D1" box="[1016,1074,474,499]" pageId="5" pageNumber="6">Nihei</collectorName>
&amp; eq. col
</materialsCitation>
.; 
<materialsCitation id="21B40C13FFA88C12FB7BFE67FB246411" collectingDate="2012-01-21" collectingDateMax="2012-02-05" collectingDateMin="2012-01-21" collectorName="Lamas &amp; Nihei" latitude="-20.697193" location="Serra da Bodoquena" longLatPrecision="1" longitude="-56.881664" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="1" stateProvince="Mato Grosso do Sul">
<specimenCount id="87DACDC7FFA88C12FB7BFE67FB2367D3" box="[1194,1229,474,499]" pageId="5" pageNumber="6" type="female">1 ♀</specimenCount>
, 
<collectingRegion id="5318C8ACFFA88C12FB08FE67FCD16431" country="Brazil" name="Mato Grosso do Sul" pageId="5" pageNumber="6">Mato Grosso do Sul</collectingRegion>
, 
<location id="94035095FFA88C12FC99FE47FBC56431" LSID="urn:lsid:plazi:treatment:1975B758FFA98C10FF30FF67FAAE6371:94035095FFA88C12FC99FE47FBC56431" box="[840,1067,506,531]" latitude="-20.697193" longLatPrecision="1" longitude="-56.881664" name="Serra da Bodoquena" pageId="5" pageNumber="6" stateProvince="Mato Grosso do Sul">Serra da Bodoquena</location>
, 
<geoCoordinate id="F4E86089FFA88C12FBE5FE47FB516431" box="[1076,1215,505,531]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="49.9" value="-20.697193">20°41′49.9″S</geoCoordinate>
, 
<geoCoordinate id="F4E86089FFA88C12FB19FE47FAB36431" box="[1224,1373,505,531]" degrees="56" direction="west" minutes="52" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="54.0" value="-56.881664">56°52′54.0″W</geoCoordinate>
, 
<date id="E562208EFFA88C12FAB4FE47FCB16411" pageId="5" pageNumber="6" value="2012-01-21">
<collectingDate id="F526D966FFA88C12FAB4FE47FCB16411" pageId="5" pageNumber="6" value="2012-01-21">21/ I/2012</collectingDate>
</date>
- 
<date id="E562208EFFA88C12FCB6FDA7FC346411" box="[871,986,538,563]" pageId="5" pageNumber="6" value="2012-02-05">
<collectingDate id="F526D966FFA88C12FCB6FDA7FC346411" box="[871,986,538,563]" pageId="5" pageNumber="6" value="2012-02-05">05/II/2012</collectingDate>
</date>
, 
<collectorName id="3C296398FFA88C12FC31FDA7FBC76411" box="[992,1065,538,563]" pageId="5" pageNumber="6">Lamas</collectorName>
, 
<collectorName id="3C296398FFA88C12FBE1FDA7FB846411" box="[1072,1130,538,563]" pageId="5" pageNumber="6">Nihei</collectorName>
&amp; eq.col
</materialsCitation>
.; 
<materialsCitation id="21B40C13FFA88C12FB05FDA7FB596451" collectingDate="2012-02-05" collectingDateMax="2012-02-20" collectingDateMin="2012-02-05" collectorName="Lamas &amp; Nihei" latitude="-20.697193" location="Serra da Bodoquena" longLatPrecision="1" longitude="-56.881664" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul">
<specimenCount id="87DACDC7FFA88C12FB05FDA7FB166413" box="[1236,1272,538,563]" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
, 
<collectingRegion id="5318C8ACFFA88C12FAD0FDA7FC8E6471" country="Brazil" name="Mato Grosso do Sul" pageId="5" pageNumber="6">Mato Grosso do Sul</collectingRegion>
, 
<location id="94035095FFA88C12FCBBFD87FBBE6471" LSID="urn:lsid:plazi:treatment:1975B758FFA98C10FF30FF67FAAE6371:94035095FFA88C12FCBBFD87FBBE6471" box="[874,1104,570,595]" latitude="-20.697193" longLatPrecision="1" longitude="-56.881664" name="Serra da Bodoquena" pageId="5" pageNumber="6" stateProvince="Mato Grosso do Sul">Serra da Bodoquena</location>
, 
<geoCoordinate id="F4E86089FFA88C12FB8BFD87FB076471" box="[1114,1257,569,595]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="49.9" value="-20.697193">20°41′49.9″S</geoCoordinate>
, 
<geoCoordinate id="F4E86089FFA88C12FB23FD87FA666471" box="[1266,1416,569,595]" degrees="56" direction="west" minutes="52" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="54.0" value="-56.881664">56°52′54.0″W</geoCoordinate>
, 
<date id="E562208EFFA88C12FCCBFDE7FC5E6451" box="[794,944,602,627]" pageId="5" pageNumber="6" value="2012-02-05" valueMax="2012-02-20" valueMin="2012-02-05">
<collectingDate id="F526D966FFA88C12FCCBFDE7FC5E6451" box="[794,944,602,627]" pageId="5" pageNumber="6" value="2012-02-05" valueMax="2012-02-20" valueMin="2012-02-05">05-20/II/2012</collectingDate>
</date>
, 
<collectorName id="3C296398FFA88C12FC6AFDE7FBEA6451" box="[955,1028,602,627]" pageId="5" pageNumber="6">Lamas</collectorName>
, 
<collectorName id="3C296398FFA88C12FBDEFDE7FBA76451" box="[1039,1097,602,627]" pageId="5" pageNumber="6">Nihei</collectorName>
&amp; eq. col
</materialsCitation>
.; 
<materialsCitation id="21B40C13FFA88C12FB17FDE7FAD16491" collectingDate="2012-09-21" collectingDateMax="2012-11-06" collectingDateMin="2012-09-21" collectorName="Lamas &amp; Nihei" latitude="-20.69886" location="Serra da Bodoquena" longLatPrecision="1" longitude="-56.880386" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul">
<specimenCount id="87DACDC7FFA88C12FB17FDE7FB016453" box="[1222,1263,602,627]" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
, 
<collectingRegion id="5318C8ACFFA88C12FB2DFDE7FC8C64B1" country="Brazil" name="Mato Grosso do Sul" pageId="5" pageNumber="6">Mato Grosso do Sul</collectingRegion>
, 
<location id="94035095FFA88C12FCBDFDC7FBBC64B1" LSID="urn:lsid:plazi:treatment:1975B758FFA98C10FF30FF67FAAE6371:94035095FFA88C12FCBDFDC7FBBC64B1" box="[876,1106,634,659]" latitude="-20.69886" longLatPrecision="1" longitude="-56.880386" name="Serra da Bodoquena" pageId="5" pageNumber="6" stateProvince="Mato Grosso do Sul">Serra da Bodoquena</location>
, 
<geoCoordinate id="F4E86089FFA88C12FB8DFDC7FB0764B1" box="[1116,1257,633,659]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="55.9" value="-20.69886">20°41′55.9″S</geoCoordinate>
, 
<geoCoordinate id="F4E86089FFA88C12FB23FDC7FA6664B1" box="[1266,1416,633,659]" degrees="56" direction="west" minutes="52" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="49.4" value="-56.880386">56°52′49.4″W</geoCoordinate>
, 
<date id="E562208EFFA88C12FCCBFD27FC7D6491" box="[794,915,666,691]" pageId="5" pageNumber="6" value="2012-09-21">
<collectingDate id="F526D966FFA88C12FCCBFD27FC7D6491" box="[794,915,666,691]" pageId="5" pageNumber="6" value="2012-09-21">21/IX/2012</collectingDate>
</date>
- 
<date id="E562208EFFA88C12FC4AFD27FBF96491" box="[923,1047,666,691]" pageId="5" pageNumber="6" value="2012-11-06">
<collectingDate id="F526D966FFA88C12FC4AFD27FBF96491" box="[923,1047,666,691]" pageId="5" pageNumber="6" value="2012-11-06">06/XI/2012</collectingDate>
</date>
, 
<collectorName id="3C296398FFA88C12FBF9FD27FB9F6491" box="[1064,1137,666,691]" pageId="5" pageNumber="6">Lamas</collectorName>
, 
<collectorName id="3C296398FFA88C12FB52FD27FB536491" box="[1155,1213,666,691]" pageId="5" pageNumber="6">Nihei</collectorName>
&amp; eq. col
</materialsCitation>
.; 
<materialsCitation id="21B40C13FFA88C12FA84FD27FABA64D1" collectingDate="2012-12-06" collectingDateMax="2012-12-21" collectingDateMin="2012-12-06" collectorName="Lamas &amp; Nihei" latitude="-20.697193" location="Serra da Bodoquena" longLatPrecision="1" longitude="-56.881664" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso do Sul">
<specimenCount id="87DACDC7FFA88C12FA84FD27FA6A6493" box="[1365,1412,666,691]" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
, 
<collectingRegion id="5318C8ACFFA88C12FCCBFD07FC1464F1" box="[794,1018,698,723]" country="Brazil" name="Mato Grosso do Sul" pageId="5" pageNumber="6">Mato Grosso do Sul</collectingRegion>
, 
<location id="94035095FFA88C12FBD7FD07FB0064F1" LSID="urn:lsid:plazi:treatment:1975B758FFA98C10FF30FF67FAAE6371:94035095FFA88C12FBD7FD07FB0064F1" box="[1030,1262,698,723]" latitude="-20.697193" longLatPrecision="1" longitude="-56.881664" name="Serra da Bodoquena" pageId="5" pageNumber="6" stateProvince="Mato Grosso do Sul">Serra da Bodoquena</location>
, 
<geoCoordinate id="F4E86089FFA88C12FB2BFD07FA6964F1" box="[1274,1415,697,723]" degrees="20" direction="south" minutes="41" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="49.9" value="-20.697193">20°41′49.9″S</geoCoordinate>
, 
<geoCoordinate id="F4E86089FFA88C12FCCBFD67FC4064D1" box="[794,942,729,755]" degrees="56" direction="west" minutes="52" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="54.0" value="-56.881664">56°52′54.0″W</geoCoordinate>
, 
<date id="E562208EFFA88C12FC67FD67FBB564D1" box="[950,1115,730,755]" pageId="5" pageNumber="6" value="2012-12-06" valueMax="2012-12-21" valueMin="2012-12-06">
<collectingDate id="F526D966FFA88C12FC67FD67FBB564D1" box="[950,1115,730,755]" pageId="5" pageNumber="6" value="2012-12-06" valueMax="2012-12-21" valueMin="2012-12-06">06-21/XII/2012</collectingDate>
</date>
, 
<collectorName id="3C296398FFA88C12FBB5FD67FB4364D1" box="[1124,1197,730,755]" pageId="5" pageNumber="6">Lamas</collectorName>
, 
<collectorName id="3C296398FFA88C12FB64FD67FB0164D1" box="[1205,1263,730,755]" pageId="5" pageNumber="6">Nihei</collectorName>
&amp; eq. col
</materialsCitation>
.
</paragraph>
<caption id="C5A356C6FFA98C13FF7BF787FBB16E71" ID-DOI="http://doi.org/10.5281/zenodo.4614813" ID-Zenodo-Dep="4614813" box="[170,1119,2106,2131]" httpUri="https://zenodo.org/record/4614813/files/figure.png" pageId="4" pageNumber="5" startId="4.[170,229,2106,2131]" targetBox="[170,1493,426,2076]" targetPageId="4">
<paragraph id="9163064EFFA98C13FF7BF787FBB16E71" blockId="4.[170,1119,2106,2131]" box="[170,1119,2106,2131]" pageId="4" pageNumber="5">
Figure 5. Habitus of 
<taxonomicName id="56DC7DCDFFA98C13FE86F787FE096E71" authority="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos, 2020" authorityName="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos" authorityYear="2020" box="[343,487,2106,2131]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kafkai" status="sp. nov.">
<emphasis id="A3A8DA5CFFA98C13FE86F787FE096E71" bold="true" box="[343,487,2106,2131]" italics="true" pageId="4" pageNumber="5">Chrysopilus kafkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89B6727FFA98C13FE3AF787FDC56E71" box="[491,555,2106,2131]" pageId="4" pageNumber="5" rank="species">sp.nov.</taxonomicNameLabel>
,female.(A) Dorsal view.(B) Lateral view.(C) Head,dorsolateral view.
</paragraph>
</caption>
<caption id="C5A356C6FFA88C12FF8FFD68FD7164CC" ID-DOI="http://doi.org/10.5281/zenodo.4614815" ID-Zenodo-Dep="4614815" box="[94,671,725,750]" httpUri="https://zenodo.org/record/4614815/files/figure.png" pageId="5" pageNumber="6" startId="5.[94,153,725,750]" targetBox="[94,718,204,695]" targetPageId="5">
<paragraph id="9163064EFFA88C12FF8FFD68FD7164CC" blockId="5.[94,671,725,750]" box="[94,671,725,750]" pageId="5" pageNumber="6">
Figure 6. Female terminalia of 
<taxonomicName id="56DC7DCDFFA88C12FEB1FD68FE1E64CC" authority="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos, 2020" authorityName="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos" authorityYear="2020" box="[352,496,725,750]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kafkai" status="sp. nov.">
<emphasis id="A3A8DA5CFFA88C12FEB1FD68FE1E64CC" bold="true" box="[352,496,725,750]" italics="true" pageId="5" pageNumber="6">Chrysopilus kafkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89B6727FFA88C12FE25FD68FDDA64CC" box="[500,564,725,750]" pageId="5" pageNumber="6" rank="species">sp.nov.</taxonomicNameLabel>
, lateral view.
</paragraph>
</caption>
<caption id="C5A356C6FFA88C12FF8FF7A7FE456E71" ID-DOI="http://doi.org/10.5281/zenodo.4614819" ID-Zenodo-Dep="4614819" httpUri="https://zenodo.org/record/4614819/files/figure.png" pageId="5" pageNumber="6" startId="5.[94,153,2074,2099]" targetBox="[126,1385,781,2041]" targetPageId="5">
<paragraph id="9163064EFFA88C12FF8FF7A7FE456E71" blockId="5.[94,1418,2074,2131]" pageId="5" pageNumber="6">
Figure 7. (A) Female terminalia of 
<taxonomicName id="56DC7DCDFFA88C12FEAFF7A7FDE06E11" authority="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos, 2020" authorityName="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos" authorityYear="2020" box="[382,526,2074,2099]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kafkai" status="sp. nov.">
<emphasis id="A3A8DA5CFFA88C12FEAFF7A7FDE06E11" bold="true" box="[382,526,2074,2099]" italics="true" pageId="5" pageNumber="6">Chrysopilus kafkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89B6727FFA88C12FDC3F7A7FDBE6E11" box="[530,592,2074,2099]" pageId="5" pageNumber="6" rank="species">sp.nov.</taxonomicNameLabel>
(B) Spermathecae.Abbreviations: 
<emphasis id="A3A8DA5CFFA88C12FCB8F7A7FC906E11" bold="true" box="[873,894,2074,2099]" italics="true" pageId="5" pageNumber="6">ce,</emphasis>
cercus; 
<emphasis id="A3A8DA5CFFA88C12FC6FF7A7FC156E11" bold="true" box="[958,1019,2074,2099]" italics="true" pageId="5" pageNumber="6">hyp val,</emphasis>
hypogynial valve; 
<emphasis id="A3A8DA5CFFA88C12FB42F7A7FB416E11" bold="true" box="[1171,1199,2074,2099]" italics="true" pageId="5" pageNumber="6">pvl,</emphasis>
posteroventral cercal lobe; 
<emphasis id="A3A8DA5CFFA88C12FF8EF787FF916E71" bold="true" box="[95,127,2106,2131]" italics="true" pageId="5" pageNumber="6">s IX,</emphasis>
sternite IX; 
<emphasis id="A3A8DA5CFFA88C12FF0EF787FECA6E71" bold="true" box="[223,292,2106,2131]" italics="true" pageId="5" pageNumber="6">sens org,</emphasis>
sensorial organ.
</paragraph>
</caption>
<paragraph id="9163064EFFAB8C11FF7BFF67FD656771" blockId="6.[170,795,218,339]" pageId="6" pageNumber="7">
<emphasis id="A3A8DA5CFFAB8C11FF7BFF67FEA766D1" bold="true" box="[170,329,218,243]" pageId="6" pageNumber="7">Preservation:</emphasis>
<typeStatus id="4E67B8ECFFAB8C11FE86FF67FE5066D1" box="[343,446,218,243]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
and 
<typeStatus id="4E67B8ECFFAB8C11FDD5FF67FD9C66D1" box="[516,626,218,243]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
dry and well preserved, with a few imperfections; additional material in 70% alcohol, with some missing parts, but complete enough to be identified as the new species.
</paragraph>
</subSubSection>
<subSubSection id="D9C655C5FFAB8C10FF7BFEC7FD486291" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="9163064EFFAB8C11FF7BFEC7FE566431" blockId="6.[169,794,378,531]" pageId="6" pageNumber="7">
<emphasis id="A3A8DA5CFFAB8C11FF7BFEC7FEC867B1" bold="true" box="[170,294,378,403]" pageId="6" pageNumber="7">Diagnosis:</emphasis>
Gonostylus crest dark brown,strongly curved distally with thick long setae (
<figureCitation id="09E71ACBFFAB8C11FDD6FE27FDA76791" box="[519,585,410,435]" captionStart="Figure 4" captionStartId="3.[94,152,2042,2067]" captionTargetBox="[94,1414,692,2012]" captionTargetId="figure-133@3.[94,1417,689,2012]" captionTargetPageId="3" captionText="Figure 4.Terminalia ofChrysopilus kafkai sp.nov.,male holotype.(A) Genital capsule,dorsal view.(B)Ventralview.Abbreviations:gs, gonostylus;ce, cercus;t IX,tergite IX.(C) MEV of genital capsule,dorsal view.(D) Genital capsule,dorsal view.(E)Gonostylus,ventral view.Abbreviations:aed,aedeagus;crt, crest;ej ap, ejaculatory apodeme;gcapod,gonocoxalapodeme;gc,gonocoxite;gs,gonostylus;latejproc,lateralejaculatoryprocess;parap, parameralapodeme;par,paramere;spm sac,sperm sac." figureDoi="http://doi.org/10.5281/zenodo.4614809" httpUri="https://zenodo.org/record/4614809/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
), yellowish brown anteriorly. Another diagnose characteristic is the wing non-hyaline with only the middle of r₃ and r₄ cells lighter than the borders (
<figureCitation id="09E71ACBFFAB8C11FEA1FE47FE436431" box="[368,429,506,531]" captionStart="Figure3" captionStartId="2.[171,230,2042,2067]" captionTargetBox="[170,1493,696,2012]" captionTargetId="figure-210@2.[170,1493,689,2012]" captionTargetPageId="2" captionText="Figure3.Wing ofChrysopiluskafkai sp.nov.,male holotype.(A) Illustration.(B)Photograph.Abbreviations:al, alula;a₁, firstanal cell;bm, basal medialcell;br, basal radial cell;C, costal vein;c,costal cell;CuA, cubital anterior vein;CuP, posterior branch of cubital vein;cua, anterior cubital cell;cup, posterior cubitalcell;d, distal cell; h, humeral;M, medial vein;m‑cu, crossvein m-cu;m‑m, crossvein m-m; R, radialvein;r‑m, crossvein r-m;Sc,subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.4614805" httpUri="https://zenodo.org/record/4614805/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="9163064EFFAB8C11FF7BFD87FDAB65D1" blockId="6.[170,795,570,1107]" pageId="6" pageNumber="7">
<emphasis id="A3A8DA5CFFAB8C11FF7BFD87FE956471" bold="true" box="[170,379,570,595]" pageId="6" pageNumber="7">
Male (
<figureCitation id="09E71ACBFFAB8C11FF2CFD87FE836471" box="[253,365,570,595]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure3" captionStart-3="Figure 4" captionStartId-0="1.[94,153,990,1015]" captionStartId-1="1.[94,153,1741,1766]" captionStartId-2="2.[171,230,2042,2067]" captionStartId-3="3.[94,152,2042,2067]" captionTargetBox-0="[101,1417,204,960]" captionTargetBox-1="[96,718,1069,1709]" captionTargetBox-2="[170,1493,696,2012]" captionTargetBox-3="[94,1414,692,2012]" captionTargetId-0="figure-360@1.[94,1417,204,960]" captionTargetId-1="figure-315@1.[95,718,1068,1711]" captionTargetId-2="figure-210@2.[170,1493,689,2012]" captionTargetId-3="figure-133@3.[94,1417,689,2012]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="2" captionTargetPageId-3="3" captionText-0="Figure 1. Chrysopilus kafkai, sp. nov., male holotype. (A) Habitus sp. nov., lateral view. (B) Head: frontal view. Abbreviations: ar, arista; cl,clypeus; ff, first flagellomere;ot, ocellar tubercle;pe, pedicel;pl, palpus;scp,scape. MATERIAL AND METHODS" captionText-1="Figure 2. Thorax of Chrysopilus kafkai sp. nov., male holotype: lateral view. Abbreviations: a spr, anterior spiracle; anepm, anepimeron; anepst, anepisternum;cx, coxa; kepst, katepisternum; ltg, laterotergite; mr, meron; mtepm, metepimeron; mtg, mediotergite; mtkepst,metakatepisternum; sct, scutum; sctl,scutellum,sbsctl, subscutellum. the Serra da Bodoquena (Mato Grosso do Sul, Brazil), a national park located in the Cerrado. These expeditions were organized by the SISBIOTA-Diptera project (Lamas et al., 2014). The new species described here is the first record of the family Rhagionidae in the Brazilian Midwest Brazilian region. The description includes both morphological and molecular data." captionText-2="Figure3.Wing ofChrysopiluskafkai sp.nov.,male holotype.(A) Illustration.(B)Photograph.Abbreviations:al, alula;a₁, firstanal cell;bm, basal medialcell;br, basal radial cell;C, costal vein;c,costal cell;CuA, cubital anterior vein;CuP, posterior branch of cubital vein;cua, anterior cubital cell;cup, posterior cubitalcell;d, distal cell; h, humeral;M, medial vein;m‑cu, crossvein m-cu;m‑m, crossvein m-m; R, radialvein;r‑m, crossvein r-m;Sc,subcostal vein." captionText-3="Figure 4.Terminalia ofChrysopilus kafkai sp.nov.,male holotype.(A) Genital capsule,dorsal view.(B)Ventralview.Abbreviations:gs, gonostylus;ce, cercus;t IX,tergite IX.(C) MEV of genital capsule,dorsal view.(D) Genital capsule,dorsal view.(E)Gonostylus,ventral view.Abbreviations:aed,aedeagus;crt, crest;ej ap, ejaculatory apodeme;gcapod,gonocoxalapodeme;gc,gonocoxite;gs,gonostylus;latejproc,lateralejaculatoryprocess;parap, parameralapodeme;par,paramere;spm sac,sperm sac." figureDoi-0="http://doi.org/10.5281/zenodo.4614801" figureDoi-1="http://doi.org/10.5281/zenodo.4614803" figureDoi-2="http://doi.org/10.5281/zenodo.4614805" figureDoi-3="http://doi.org/10.5281/zenodo.4614809" httpUri-0="https://zenodo.org/record/4614801/files/figure.png" httpUri-1="https://zenodo.org/record/4614803/files/figure.png" httpUri-2="https://zenodo.org/record/4614805/files/figure.png" httpUri-3="https://zenodo.org/record/4614809/files/figure.png" pageId="6" pageNumber="7">Figs. 1‑4</figureCitation>
):
</emphasis>
length: body 
<emphasis id="A3A8DA5CFFAB8C11FDEBFD87FDB76471" box="[570,601,570,595]" italics="true" pageId="6" pageNumber="7">ca.</emphasis>
<quantity id="5624ABABFFAB8C11FDBAFD87FD236471" box="[619,717,570,595]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="7" unit="mm" value="6.5">6.5 mm</quantity>
, wing 
<emphasis id="A3A8DA5CFFAB8C11FF7BFDE7FF276451" box="[170,201,602,627]" italics="true" pageId="6" pageNumber="7">ca.</emphasis>
<quantity id="5624ABABFFAB8C11FF00FDE7FEC66451" box="[209,296,602,627]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="6" pageNumber="7" unit="mm" value="7.3">7.3 mm</quantity>
. General color dark brown (
<figureCitation id="09E71ACBFFAB8C11FDB2FDE7FD436451" box="[611,685,602,627]" captionStart="Figure 1" captionStartId="1.[94,153,990,1015]" captionTargetBox="[101,1417,204,960]" captionTargetId="figure-360@1.[94,1417,204,960]" captionTargetPageId="1" captionText="Figure 1. Chrysopilus kafkai, sp. nov., male holotype. (A) Habitus sp. nov., lateral view. (B) Head: frontal view. Abbreviations: ar, arista; cl,clypeus; ff, first flagellomere;ot, ocellar tubercle;pe, pedicel;pl, palpus;scp,scape. MATERIAL AND METHODS" figureDoi="http://doi.org/10.5281/zenodo.4614801" httpUri="https://zenodo.org/record/4614801/files/figure.png" pageId="6" pageNumber="7">Fig. 1a</figureCitation>
). Head in shades of brown (
<figureCitation id="09E71ACBFFAB8C11FEA3FDC7FE5364B1" box="[370,445,634,659]" captionStart="Figure 1" captionStartId="1.[94,153,990,1015]" captionTargetBox="[101,1417,204,960]" captionTargetId="figure-360@1.[94,1417,204,960]" captionTargetPageId="1" captionText="Figure 1. Chrysopilus kafkai, sp. nov., male holotype. (A) Habitus sp. nov., lateral view. (B) Head: frontal view. Abbreviations: ar, arista; cl,clypeus; ff, first flagellomere;ot, ocellar tubercle;pe, pedicel;pl, palpus;scp,scape. MATERIAL AND METHODS" figureDoi="http://doi.org/10.5281/zenodo.4614801" httpUri="https://zenodo.org/record/4614801/files/figure.png" pageId="6" pageNumber="7">Fig. 1b</figureCitation>
). Eyes holoptic, slightly grayish brown with the borders in blackish brown, ommatidium facets round; ocellar tubercle dark brown, ocellus light brown. Gena brown, lighter than the eyes. Bare and strongly curved clypeus. Antenna uniformly light brown with short thin setae, 1.6 times smaller than head; scape flattened, pedicel two times longer than scape, first flagellomere tapering toward apex, clearly narrow- er than pedicel and approximately as wide as scape. Maxillary palpus light brown, distally angled with thick medium-sized setae. Proboscis yellowish brown. Occiput grayish brown, with long-sized setae.
</paragraph>
<paragraph id="9163064EFFAB8C11FF01FC47FA3B6431" blockId="6.[170,795,570,1107]" lastBlockId="6.[868,1494,218,883]" pageId="6" pageNumber="7">
Thorax dark brown (
<figureCitation id="09E71ACBFFAB8C11FE62FC47FE136231" box="[435,509,1018,1043]" captionStart="Figure 1" captionStartId="1.[94,153,990,1015]" captionTargetBox="[101,1417,204,960]" captionTargetId="figure-360@1.[94,1417,204,960]" captionTargetPageId="1" captionText="Figure 1. Chrysopilus kafkai, sp. nov., male holotype. (A) Habitus sp. nov., lateral view. (B) Head: frontal view. Abbreviations: ar, arista; cl,clypeus; ff, first flagellomere;ot, ocellar tubercle;pe, pedicel;pl, palpus;scp,scape. MATERIAL AND METHODS" figureDoi="http://doi.org/10.5281/zenodo.4614801" httpUri="https://zenodo.org/record/4614801/files/figure.png" pageId="6" pageNumber="7">Fig. 1a</figureCitation>
), wider than long. Meron almost two times wider than long, katepisternum almost as wide as high (
<figureCitation id="09E71ACBFFAB8C11FE8BFB87FE786271" box="[346,406,1082,1107]" captionStart="Figure 2" captionStartId="1.[94,153,1741,1766]" captionTargetBox="[96,718,1069,1709]" captionTargetId="figure-315@1.[95,718,1068,1711]" captionTargetPageId="1" captionText="Figure 2. Thorax of Chrysopilus kafkai sp. nov., male holotype: lateral view. Abbreviations: a spr, anterior spiracle; anepm, anepimeron; anepst, anepisternum;cx, coxa; kepst, katepisternum; ltg, laterotergite; mr, meron; mtepm, metepimeron; mtg, mediotergite; mtkepst,metakatepisternum; sct, scutum; sctl,scutellum,sbsctl, subscutellum. the Serra da Bodoquena (Mato Grosso do Sul, Brazil), a national park located in the Cerrado. These expeditions were organized by the SISBIOTA-Diptera project (Lamas et al., 2014). The new species described here is the first record of the family Rhagionidae in the Brazilian Midwest Brazilian region. The description includes both morphological and molecular data." figureDoi="http://doi.org/10.5281/zenodo.4614803" httpUri="https://zenodo.org/record/4614803/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
). Coxa dark brown,femur,tibia, and tarsus light yellow (
<figureCitation id="09E71ACBFFAB8C11FB95FF67FB7E66D1" box="[1092,1168,218,243]" captionStart="Figure 1" captionStartId="1.[94,153,990,1015]" captionTargetBox="[101,1417,204,960]" captionTargetId="figure-360@1.[94,1417,204,960]" captionTargetPageId="1" captionText="Figure 1. Chrysopilus kafkai, sp. nov., male holotype. (A) Habitus sp. nov., lateral view. (B) Head: frontal view. Abbreviations: ar, arista; cl,clypeus; ff, first flagellomere;ot, ocellar tubercle;pe, pedicel;pl, palpus;scp,scape. MATERIAL AND METHODS" figureDoi="http://doi.org/10.5281/zenodo.4614801" httpUri="https://zenodo.org/record/4614801/files/figure.png" pageId="6" pageNumber="7">Fig. 1a</figureCitation>
). Wing membrane brownish along most veins (
<figureCitation id="09E71ACBFFAB8C11FBFEFF47FB836731" box="[1071,1133,250,275]" captionStart="Figure3" captionStartId="2.[171,230,2042,2067]" captionTargetBox="[170,1493,696,2012]" captionTargetId="figure-210@2.[170,1493,689,2012]" captionTargetPageId="2" captionText="Figure3.Wing ofChrysopiluskafkai sp.nov.,male holotype.(A) Illustration.(B)Photograph.Abbreviations:al, alula;a₁, firstanal cell;bm, basal medialcell;br, basal radial cell;C, costal vein;c,costal cell;CuA, cubital anterior vein;CuP, posterior branch of cubital vein;cua, anterior cubital cell;cup, posterior cubitalcell;d, distal cell; h, humeral;M, medial vein;m‑cu, crossvein m-cu;m‑m, crossvein m-m; R, radialvein;r‑m, crossvein r-m;Sc,subcostal vein." figureDoi="http://doi.org/10.5281/zenodo.4614805" httpUri="https://zenodo.org/record/4614805/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
); pterostigma, approximately six times smaller than the wing length; R₁ and R₂₊₃ running parallel for the most part, only last third of R₂₊₃ converging towards R₁; R₄ barely curved; CuA and CuP joining far from wing margin, CuA+CuP as long as r-m crossvein; discal cell approximately 4.2x longer than wide, bm cell five times longer than wide, br cell six times longer than wide, cua cell six times longer than wide, c cell placed before pterostigma; alula broad. Halter light brown (
<figureCitation id="09E71ACBFFAB8C11FA50FE47FA266431" box="[1409,1480,506,531]" captionStart="Figure 1" captionStartId="1.[94,153,990,1015]" captionTargetBox="[101,1417,204,960]" captionTargetId="figure-360@1.[94,1417,204,960]" captionTargetPageId="1" captionText="Figure 1. Chrysopilus kafkai, sp. nov., male holotype. (A) Habitus sp. nov., lateral view. (B) Head: frontal view. Abbreviations: ar, arista; cl,clypeus; ff, first flagellomere;ot, ocellar tubercle;pe, pedicel;pl, palpus;scp,scape. MATERIAL AND METHODS" figureDoi="http://doi.org/10.5281/zenodo.4614801" httpUri="https://zenodo.org/record/4614801/files/figure.png" pageId="6" pageNumber="7">Fig. 1a</figureCitation>
).
</paragraph>
<paragraph id="9163064EFFAB8C11FC5AFDA7FAFE6551" blockId="6.[868,1494,218,883]" pageId="6" pageNumber="7">
Abdomen dark brown (
<figureCitation id="09E71ACBFFAB8C11FB79FDA7FB176411" box="[1192,1273,538,563]" captionStart="Figure 1" captionStartId="1.[94,153,990,1015]" captionTargetBox="[101,1417,204,960]" captionTargetId="figure-360@1.[94,1417,204,960]" captionTargetPageId="1" captionText="Figure 1. Chrysopilus kafkai, sp. nov., male holotype. (A) Habitus sp. nov., lateral view. (B) Head: frontal view. Abbreviations: ar, arista; cl,clypeus; ff, first flagellomere;ot, ocellar tubercle;pe, pedicel;pl, palpus;scp,scape. MATERIAL AND METHODS" figureDoi="http://doi.org/10.5281/zenodo.4614801" httpUri="https://zenodo.org/record/4614801/files/figure.png" pageId="6" pageNumber="7">Fig. 1a</figureCitation>
). Gonocoxite dark brown with long setae; gonostylus apex dark brown, protruding curved crest with long setae (
<figureCitation id="09E71ACBFFAB8C11FA9DFDE7FA7E6451" box="[1356,1424,602,627]" captionStart="Figure 4" captionStartId="3.[94,152,2042,2067]" captionTargetBox="[94,1414,692,2012]" captionTargetId="figure-133@3.[94,1417,689,2012]" captionTargetPageId="3" captionText="Figure 4.Terminalia ofChrysopilus kafkai sp.nov.,male holotype.(A) Genital capsule,dorsal view.(B)Ventralview.Abbreviations:gs, gonostylus;ce, cercus;t IX,tergite IX.(C) MEV of genital capsule,dorsal view.(D) Genital capsule,dorsal view.(E)Gonostylus,ventral view.Abbreviations:aed,aedeagus;crt, crest;ej ap, ejaculatory apodeme;gcapod,gonocoxalapodeme;gc,gonocoxite;gs,gonostylus;latejproc,lateralejaculatoryprocess;parap, parameralapodeme;par,paramere;spm sac,sperm sac." figureDoi="http://doi.org/10.5281/zenodo.4614809" httpUri="https://zenodo.org/record/4614809/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
), bottom yellowish brown; gonostylus with a distal concavity (
<figureCitation id="09E71ACBFFAB8C11FC5BFD27FC376491" box="[906,985,666,691]" captionStart="Figure 4" captionStartId="3.[94,152,2042,2067]" captionTargetBox="[94,1414,692,2012]" captionTargetId="figure-133@3.[94,1417,689,2012]" captionTargetPageId="3" captionText="Figure 4.Terminalia ofChrysopilus kafkai sp.nov.,male holotype.(A) Genital capsule,dorsal view.(B)Ventralview.Abbreviations:gs, gonostylus;ce, cercus;t IX,tergite IX.(C) MEV of genital capsule,dorsal view.(D) Genital capsule,dorsal view.(E)Gonostylus,ventral view.Abbreviations:aed,aedeagus;crt, crest;ej ap, ejaculatory apodeme;gcapod,gonocoxalapodeme;gc,gonocoxite;gs,gonostylus;latejproc,lateralejaculatoryprocess;parap, parameralapodeme;par,paramere;spm sac,sperm sac." figureDoi="http://doi.org/10.5281/zenodo.4614809" httpUri="https://zenodo.org/record/4614809/files/figure.png" pageId="6" pageNumber="7">Fig. 4e</figureCitation>
). Aedeagus long, half the gonocoxite length (
<figureCitation id="09E71ACBFFAB8C11FCBDFD07FC4464F1" box="[876,938,698,723]" captionStart="Figure 4" captionStartId="3.[94,152,2042,2067]" captionTargetBox="[94,1414,692,2012]" captionTargetId="figure-133@3.[94,1417,689,2012]" captionTargetPageId="3" captionText="Figure 4.Terminalia ofChrysopilus kafkai sp.nov.,male holotype.(A) Genital capsule,dorsal view.(B)Ventralview.Abbreviations:gs, gonostylus;ce, cercus;t IX,tergite IX.(C) MEV of genital capsule,dorsal view.(D) Genital capsule,dorsal view.(E)Gonostylus,ventral view.Abbreviations:aed,aedeagus;crt, crest;ej ap, ejaculatory apodeme;gcapod,gonocoxalapodeme;gc,gonocoxite;gs,gonostylus;latejproc,lateralejaculatoryprocess;parap, parameralapodeme;par,paramere;spm sac,sperm sac." figureDoi="http://doi.org/10.5281/zenodo.4614809" httpUri="https://zenodo.org/record/4614809/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
). Ejaculatory apodeme long, nearly as long as aedeagus. Gonocoxal apodeme approximately half of the aedeagus length; parameral apodeme short, do not extending beyond the gonocoxal apodeme; lateral ejaculatory process short. Cercus dark brown with short setae, distancing each other distally (
<figureCitation id="09E71ACBFFAB8C11FB68FCE7FAEB6551" box="[1209,1285,858,883]" captionStart="Figure 4" captionStartId="3.[94,152,2042,2067]" captionTargetBox="[94,1414,692,2012]" captionTargetId="figure-133@3.[94,1417,689,2012]" captionTargetPageId="3" captionText="Figure 4.Terminalia ofChrysopilus kafkai sp.nov.,male holotype.(A) Genital capsule,dorsal view.(B)Ventralview.Abbreviations:gs, gonostylus;ce, cercus;t IX,tergite IX.(C) MEV of genital capsule,dorsal view.(D) Genital capsule,dorsal view.(E)Gonostylus,ventral view.Abbreviations:aed,aedeagus;crt, crest;ej ap, ejaculatory apodeme;gcapod,gonocoxalapodeme;gc,gonocoxite;gs,gonostylus;latejproc,lateralejaculatoryprocess;parap, parameralapodeme;par,paramere;spm sac,sperm sac." figureDoi="http://doi.org/10.5281/zenodo.4614809" httpUri="https://zenodo.org/record/4614809/files/figure.png" pageId="6" pageNumber="7">Fig. 4b</figureCitation>
).
</paragraph>
<paragraph id="9163064EFFAB8C11FCB4FC27FA7665F1" blockId="6.[869,1494,922,1107]" pageId="6" pageNumber="7">
<emphasis id="A3A8DA5CFFAB8C11FCB4FC27FBAA6591" bold="true" box="[869,1092,922,947]" pageId="6" pageNumber="7">
Female (
<figureCitation id="09E71ACBFFAB8C11FC1DFC27FBD96591" box="[972,1079,922,947]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-2="Figure 7" captionStartId-0="4.[170,229,2106,2131]" captionStartId-1="5.[94,153,725,750]" captionStartId-2="5.[94,153,2074,2099]" captionTargetBox-0="[170,1493,426,2076]" captionTargetBox-1="[94,718,204,695]" captionTargetBox-2="[126,1385,781,2041]" captionTargetId-0="figure-95@4.[170,1493,413,2076]" captionTargetId-1="figure-228@5.[94,718,204,695]" captionTargetId-2="figure-240@5.[123,1389,778,2044]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="Figure 5. Habitus of Chrysopilus kafkai sp.nov.,female.(A) Dorsal view.(B) Lateral view.(C) Head,dorsolateral view." captionText-1="Figure 6. Female terminalia of Chrysopilus kafkai sp.nov., lateral view." captionText-2="Figure 7. (A) Female terminalia of Chrysopilus kafkai sp.nov.(B) Spermathecae.Abbreviations:ce, cercus;hyp val, hypogynial valve; pvl, posteroventral cercal lobe; s IX, sterniteIX; sensorg, sensorial organ." figureDoi-0="http://doi.org/10.5281/zenodo.4614813" figureDoi-1="http://doi.org/10.5281/zenodo.4614815" figureDoi-2="http://doi.org/10.5281/zenodo.4614819" httpUri-0="https://zenodo.org/record/4614813/files/figure.png" httpUri-1="https://zenodo.org/record/4614815/files/figure.png" httpUri-2="https://zenodo.org/record/4614819/files/figure.png" pageId="6" pageNumber="7">Figs. 5‑7</figureCitation>
):
</emphasis>
Same as males, except as follows: length: body 
<quantity id="5624ABABFFAB8C11FC28FC07FBA065F1" box="[1017,1102,954,979]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="7" unit="mm" value="7.8">7.8 mm</quantity>
, wing 
<quantity id="5624ABABFFAB8C11FB45FC07FB0765F1" box="[1172,1257,954,979]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="6" pageNumber="7" unit="mm" value="7.3">7.3 mm</quantity>
. Eyes dichoptic.
</paragraph>
<paragraph id="9163064EFFAB8C10FC5AFC67FD546271" blockId="6.[869,1494,922,1107]" lastBlockId="7.[94,718,1018,1107]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Cercus dark brown, two-segmented with slightly rounded apex in the distal segment (
<figureCitation id="09E71ACBFFAB8C11FACFFC47FA8F6231" box="[1310,1377,1018,1043]" captionStart="Figure 6" captionStartId="5.[94,153,725,750]" captionTargetBox="[94,718,204,695]" captionTargetId="figure-228@5.[94,718,204,695]" captionTargetPageId="5" captionText="Figure 6. Female terminalia of Chrysopilus kafkai sp.nov., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4614815" httpUri="https://zenodo.org/record/4614815/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
), posteroventral cercal lobe two times longer than wide (
<figureCitation id="09E71ACBFFAB8C11FAAEFBA7FA256211" box="[1407,1483,1050,1075]" captionStart="Figure 7" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[126,1385,781,2041]" captionTargetId="figure-240@5.[123,1389,778,2044]" captionTargetPageId="5" captionText="Figure 7. (A) Female terminalia of Chrysopilus kafkai sp.nov.(B) Spermathecae.Abbreviations:ce, cercus;hyp val, hypogynial valve; pvl, posteroventral cercal lobe; s IX, sterniteIX; sensorg, sensorial organ." figureDoi="http://doi.org/10.5281/zenodo.4614819" httpUri="https://zenodo.org/record/4614819/files/figure.png" pageId="6" pageNumber="7">Fig. 7a</figureCitation>
). Sternite VIII slightly rounded anteriorly; hypogynial valves with long and thick setae (
<figureCitation id="09E71ACBFFAA8C10FE36FC47FDD96231" box="[487,567,1018,1043]" captionStart="Figure 7" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[126,1385,781,2041]" captionTargetId="figure-240@5.[123,1389,778,2044]" captionTargetPageId="5" captionText="Figure 7. (A) Female terminalia of Chrysopilus kafkai sp.nov.(B) Spermathecae.Abbreviations:ce, cercus;hyp val, hypogynial valve; pvl, posteroventral cercal lobe; s IX, sterniteIX; sensorg, sensorial organ." figureDoi="http://doi.org/10.5281/zenodo.4614819" httpUri="https://zenodo.org/record/4614819/files/figure.png" pageId="7" pageNumber="8">Fig. 7a</figureCitation>
); genital fork with a slightly elongated anterior protrusion and distal short apodemes.Three spermathecae oblong (
<figureCitation id="09E71ACBFFAA8C10FDB3FB87FD416271" box="[610,687,1082,1107]" captionStart="Figure 7" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[126,1385,781,2041]" captionTargetId="figure-240@5.[123,1389,778,2044]" captionTargetPageId="5" captionText="Figure 7. (A) Female terminalia of Chrysopilus kafkai sp.nov.(B) Spermathecae.Abbreviations:ce, cercus;hyp val, hypogynial valve; pvl, posteroventral cercal lobe; s IX, sterniteIX; sensorg, sensorial organ." figureDoi="http://doi.org/10.5281/zenodo.4614819" httpUri="https://zenodo.org/record/4614819/files/figure.png" pageId="7" pageNumber="8">Fig. 7b</figureCitation>
).
</paragraph>
<caption id="C5A356C6FFAB8C11FF7BF787FD4F6E71" ID-DOI="http://doi.org/10.5281/zenodo.4614821" ID-Zenodo-Dep="4614821" box="[170,673,2106,2131]" httpUri="https://zenodo.org/record/4614821/files/figure.png" pageId="6" pageNumber="7" startId="6.[170,229,2106,2131]" targetBox="[170,1493,1131,2076]" targetPageId="6">
<paragraph id="9163064EFFAB8C11FF7BF787FD4F6E71" blockId="6.[170,673,2106,2131]" box="[170,673,2106,2131]" pageId="6" pageNumber="7">
Figure 8. Distribution map of Brazilian species of 
<taxonomicName id="56DC7DCDFFAB8C11FD91F787FD736E71" authorityName="Macquart" authorityYear="1826" box="[576,669,2106,2131]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Chrysopilus</taxonomicName>
.
</paragraph>
</caption>
<caption id="C5A356C6FFAA8C10FF8FFC05FC4965F3" ID-DOI="http://doi.org/10.5281/zenodo.4614825" ID-Zenodo-Dep="4614825" box="[94,935,952,977]" httpUri="https://zenodo.org/record/4614825/files/figure.png" pageId="7" pageNumber="8" startId="7.[94,153,952,977]" targetBox="[96,1411,206,918]" targetPageId="7">
<paragraph id="9163064EFFAA8C10FF8FFC05FC4965F3" blockId="7.[94,935,952,977]" box="[94,935,952,977]" pageId="7" pageNumber="8">
Figure 9. Comparison between gonostylus crests:(A) 
<taxonomicName id="56DC7DCDFFAA8C10FDC6FC05FD4965F3" authorityName="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos" authorityYear="2020" box="[535,679,952,977]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kafkai">
<emphasis id="A3A8DA5CFFAA8C10FDC6FC05FD4965F3" bold="true" box="[535,679,952,977]" italics="true" pageId="7" pageNumber="8">Chrysopilus kafkai</emphasis>
</taxonomicName>
and (B) 
<taxonomicName id="56DC7DCDFFAA8C10FD3AFC05FC4D65F3" authorityName="Santos &amp; Amorim" authorityYear="2007" box="[747,931,952,977]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="plaumanni">Chrysopilus plaumanni</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="9163064EFFAA8C10FF8FFBC7FD486291" blockId="7.[94,718,1146,1203]" pageId="7" pageNumber="8">
<emphasis id="A3A8DA5CFFAA8C10FF8FFBC7FE6A62B1" bold="true" box="[94,388,1146,1171]" pageId="7" pageNumber="8">Geographic distribution:</emphasis>
Serra da Bodoquena, state of 
<collectingRegion id="5318C8ACFFAA8C10FFB1FB27FED66291" box="[96,312,1178,1203]" country="Brazil" name="Mato Grosso do Sul" pageId="7" pageNumber="8">Mato Grosso do Sul</collectingRegion>
, Midwest Brazilian region (
<figureCitation id="09E71ACBFFAA8C10FD8FFB27FD756291" box="[606,667,1178,1203]" captionStart="Figure 8" captionStartId="6.[170,229,2106,2131]" captionTargetBox="[170,1493,1131,2076]" captionTargetId="figure-465@6.[170,1493,1131,2076]" captionTargetPageId="6" captionText="Figure 8. Distribution map of Brazilian species of Chrysopilus." figureDoi="http://doi.org/10.5281/zenodo.4614821" httpUri="https://zenodo.org/record/4614821/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D9C655C5FFAA8C10FF8FFB67FEF26371" pageId="7" pageNumber="8" type="etymology">
<paragraph id="9163064EFFAA8C10FF8FFB67FEF26371" blockId="7.[94,718,1242,1363]" pageId="7" pageNumber="8">
<emphasis id="A3A8DA5CFFAA8C10FF8FFB67FF0862D1" bold="true" box="[94,230,1242,1267]" pageId="7" pageNumber="8">Etymology:</emphasis>
Based on its general dark color, the species is named after the fiction author Franz Kafka, whose literature mostly deals with the darkness and absurdity of human behavior.
</paragraph>
</subSubSection>
<subSubSection id="D9C655C5FFAA8C10FCCBFC47FAAE6371" pageId="7" pageNumber="8" type="discussion">
<paragraph id="9163064EFFAA8C10FCCBFC47FB2E6291" blockId="7.[794,1418,1018,1363]" pageId="7" pageNumber="8">
<emphasis id="A3A8DA5CFFAA8C10FCCBFC47FC4E6231" bold="true" box="[794,928,1018,1043]" pageId="7" pageNumber="8">Comments:</emphasis>
<taxonomicName id="56DC7DCDFFAA8C10FC66FC47FB696231" authority="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos, 2020" authorityName="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos" authorityYear="2020" box="[951,1159,1018,1043]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kafkai" status="sp. nov.">
<emphasis id="A3A8DA5CFFAA8C10FC66FC47FB696231" box="[951,1159,1018,1043]" italics="true" pageId="7" pageNumber="8">Chrysopilus kafkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89B6727FFAA8C10FB4FFC47FAEC6231" box="[1182,1282,1018,1043]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
resembles 
<emphasis id="A3A8DA5CFFAA8C10FCCBFBA7FC636211" box="[794,909,1050,1075]" italics="true" pageId="7" pageNumber="8">Chysopilus</emphasis>
<taxonomicName id="56DC7DCDFFAA8C10FC44FBA7FB336211" authority="Santos &amp; Amorim" authorityName="Santos &amp; Amorim" authorityYear="2007" box="[917,1245,1050,1075]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="A3A8DA5CFFAA8C10FC44FBA7FBE56211" box="[917,1035,1050,1075]" italics="true" pageId="7" pageNumber="8">plaumanni</emphasis>
Santos &amp; Amorim
</taxonomicName>
in the general aspect of the male terminalia. However, 
<taxonomicName id="56DC7DCDFFAA8C10FB07FB87FADE6271" authority="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos, 2020" authorityName="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos" authorityYear="2020" box="[1238,1328,1082,1107]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kafkai" status="sp. nov.">
<emphasis id="A3A8DA5CFFAA8C10FB07FB87FADE6271" box="[1238,1328,1082,1107]" italics="true" pageId="7" pageNumber="8">C. kafkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89B6727FFAA8C10FAE7FB87FA646271" box="[1334,1418,1082,1107]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
has a distinctive crest occupying almost half of the gonostylus, while the crest in 
<taxonomicName id="56DC7DCDFFAA8C10FBFBFBC7FB5462B1" authorityName="Santos &amp; Amorim" authorityYear="2007" box="[1066,1210,1146,1171]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="A3A8DA5CFFAA8C10FBFBFBC7FB5462B1" box="[1066,1210,1146,1171]" italics="true" pageId="7" pageNumber="8">C. plaumanni</emphasis>
</taxonomicName>
is restricted to the distal region of the gonostylus (
<figureCitation id="09E71ACBFFAA8C10FBA6FB27FB5B6291" box="[1143,1205,1178,1203]" captionStart="Figure 9" captionStartId="7.[94,153,952,977]" captionTargetBox="[96,1411,206,918]" captionTargetId="figure-507@7.[94,1417,204,922]" captionTargetPageId="7" captionText="Figure 9. Comparison between gonostylus crests:(A) Chrysopilus kafkai and (B) Chrysopilus plaumanni." figureDoi="http://doi.org/10.5281/zenodo.4614825" httpUri="https://zenodo.org/record/4614825/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="9163064EFFAA8C10FCEEFB07FAAE6371" blockId="7.[794,1418,1018,1363]" pageId="7" pageNumber="8">
Our molecular characterization of 28S rRNA of 
<taxonomicName id="56DC7DCDFFAA8C10FCCBFB67FC3562D1" authority="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos, 2020" authorityName="Cegolin &amp; Bueno &amp; Pereira &amp; Santos &amp; Santos" authorityYear="2020" box="[794,987,1242,1267]" class="Insecta" family="Rhagionidae" genus="Chrysopilus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kafkai" status="sp. nov.">
<emphasis id="A3A8DA5CFFAA8C10FCCBFB67FC3562D1" box="[794,987,1242,1267]" italics="true" pageId="7" pageNumber="8">Chrysopilus kafkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B89B6727FFAA8C10FC31FB67FBDD62D1" box="[992,1075,1242,1267]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
expands the geographical representation of molecular data for the genus, now with sequences from species distributed in three biogeographical regions (Nearctic, Neotropical, and Australian).
</paragraph>
</subSubSection>
</treatment>
</document>