165 lines
20 KiB
XML
165 lines
20 KiB
XML
<document id="58038D63AB1317F23C6C7F3BE89711EA" ID-DOI="10.3853/j.0067-1975.60.2008.1497" ID-ISSN="2201-4349" ID-Zenodo-Dep="4676531" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617989110877" checkinUser="carolina" docAuthor="Webby, Barry D. & Zhen, Yong Yi" docDate="2008" docId="03EF4E2E80639D08938EF4B7FD4BF9EC" docLanguage="en" docName="RecAustMus.60.3.215-236.pdf" docOrigin="Records of the Australian Museum 60 (3)" docSource="http://dx.doi.org/10.3853/j.0067-1975.60.2008.1497" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Columnostroma Bogoyavlenskaya 1972" docType="treatment" docVersion="3" lastPageNumber="233" masterDocId="FFD6365680729D1A9054F23AFFD3FFE2" masterDocTitle="Devonian Syringostromatid Stromatoporoids from the Broken River Region, North Queensland" masterLastPageNumber="236" masterPageNumber="215" pageNumber="232" updateTime="1698903950538" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="E76BB765D3A42147151F3BCDB288DB34">Devonian Syringostromatid Stromatoporoids from the Broken River Region, North Queensland</mods:title>
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<mods:namePart id="B53CD5B1549089009F168A4C4425D766">Webby, Barry D.</mods:namePart>
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<mods:roleTerm id="53137E5A23689A2070F412C32FFCD64C">Author</mods:roleTerm>
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<mods:namePart id="3BE19511BCE044AC35A9C36BE27B6A68">Zhen, Yong Yi</mods:namePart>
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<mods:title id="6F519353D5A148F19239AC235DF87D62">Records of the Australian Museum</mods:title>
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<mods:part id="31CD19798EC050895F8CDE55CE5949A3">
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<mods:date id="92CE595DE66B01F01C2D5EE5CC9F750E">2008</mods:date>
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<mods:identifier id="73873431E12C91371BA9DE3E72B663BE" type="DOI">10.3853/j.0067-1975.60.2008.1497</mods:identifier>
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<mods:identifier id="B0BF0FDF996F7605D83CA6AF7C57C2E1" type="ISSN">2201-4349</mods:identifier>
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<subSubSection id="C35CACB380639D0B938EF4B7FB1EF945" box="[986,1229,1677,1703]" pageId="17" pageNumber="232" type="nomenclature">
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<paragraph id="8BF9FF3880639D0B938EF4B7FB1EF945" blockId="17.[986,1229,1677,1703]" box="[986,1229,1677,1703]" pageId="17" pageNumber="232">
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<heading id="D0B1485480639D0B938EF4B7FB1EF945" box="[986,1229,1677,1703]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="232" reason="2">
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<emphasis id="B932232A80639D0B938EF4B7FB1EF945" bold="true" box="[986,1229,1677,1703]" pageId="17" pageNumber="232">
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<taxonomicName id="4C4684BB80639D0B938EF4B7FB72F945" authorityName="Bogoyavlenskaya" authorityYear="1972" box="[986,1185,1677,1703]" class="Hydrozoa" family="Hydractiniidae" genus="Columnostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="17" pageNumber="232" phylum="Cnidaria" rank="genus">
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<emphasis id="B932232A80639D0B938EF4B7FB72F945" bold="true" box="[986,1185,1677,1703]" italics="true" pageId="17" pageNumber="232">Columnostroma</emphasis>
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</taxonomicName>
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sp.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CACB380639D0B9440F4F1FB40F907" box="[1044,1171,1739,1765]" pageId="17" pageNumber="232" type="description">
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<paragraph id="8BF9FF3880639D0B9440F4F1FB40F907" blockId="17.[1044,1171,1739,1765]" box="[1044,1171,1739,1765]" pageId="17" pageNumber="232">
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<figureCitation id="137DE3BD80639D0B9440F4F1FB8AF907" box="[1044,1113,1739,1765]" captionStart="Fig" captionStartId="19.[121,156,1627,1648]" captionTargetBox="[121,1421,151,1615]" captionTargetId="figure-62@19.[121,1421,151,1620]" captionTargetPageId="19" captionText="Fig. 7. Columnostroma sp., A–C, specimen AM F.134920, from the Dosey Limestone, 11 m above the base of the SD260 section, NNW of Storm Hill; (A) longitudinal section (AM FT.15152), ×10; (B) longitudinal section (AM FT.15152), showing within much thickened pachysteles small areas where incompletely aligned longitudinal rows of microgalleries and dark melanospheric rows of dots occur, confirming the poorly preserved microreticulation, ×20; (C) tangential section (AM FT.15154), ×10. Parallelopora sp., D, E, both specimens come from the same locality at UQL.2974 of Mallett 1968, as reworked pebbles from Chinaman Creek Limestone along South Chinaman Creek; (D) tangential section of specimen UQF.47979, ×10; (E), longitudinal section of specimen UQF.47983, ×10. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4676553" httpUri="https://zenodo.org/record/4676553/files/figure.png" pageId="17" pageNumber="232">Fig. 7</figureCitation>
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A–C
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CACB380639D0B934EF538FA8DF82B" pageId="17" pageNumber="232" type="materials_examined">
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<paragraph id="8BF9FF3880639D0B934EF538FA8DF82B" blockId="17.[794,1422,1794,1993]" pageId="17" pageNumber="232">
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<emphasis id="B932232A80639D0B934EF538FCAEF8F8" bold="true" box="[794,893,1794,1818]" pageId="17" pageNumber="232">Material</emphasis>
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. Available material is fragmentary and recrystallized, comprising two specimens, SD260-11-3 (
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<collectionCode id="ED5767FD80639D0B950AF525FA5EF8D4" box="[1374,1421,1823,1846]" country="Australia" name="Australian Museum" pageId="17" pageNumber="232" type="Museum">AM</collectionCode>
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F.134919,
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<collectionCode id="ED5767FD80639D0B93DEF507FC66F8B6" box="[906,949,1853,1876]" country="Australia" name="Australian Museum" pageId="17" pageNumber="232" type="Museum">AM</collectionCode>
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FT.15150,
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<collectionCode id="ED5767FD80639D0B9465F507FB8FF8B6" box="[1073,1116,1853,1876]" country="Australia" name="Australian Museum" pageId="17" pageNumber="232" type="Museum">AM</collectionCode>
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FT.15151) and SD260-11-4 (
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<collectionCode id="ED5767FD80639D0B9376F560FC80F893" box="[802,851,1882,1905]" country="Australia" name="Australian Museum" pageId="17" pageNumber="232" type="Museum">AM</collectionCode>
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F.134920,
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<collectionCode id="ED5767FD80639D0B93B5F560FBDCF893" box="[993,1039,1882,1905]" country="Australia" name="Australian Museum" pageId="17" pageNumber="232" type="Museum">AM</collectionCode>
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FT.15152,
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<collectionCode id="ED5767FD80639D0B94F5F560FB1CF893" box="[1185,1231,1882,1905]" country="Australia" name="Australian Museum" pageId="17" pageNumber="232" type="Museum">AM</collectionCode>
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FT.15153,
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<collectionCode id="ED5767FD80639D0B9534F560FA5EF893" box="[1376,1421,1882,1905]" country="Australia" name="Australian Museum" pageId="17" pageNumber="232" type="Museum">AM</collectionCode>
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FT.15154), from
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<quantity id="4CBE52DD80639D0B939CF54DFC28F86C" box="[968,1019,1911,1934]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="232" unit="m" value="11.0">11 m</quantity>
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above the base of the Dosey Limestone in SD260 section of
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<bibRefCitation id="EFD782C980639D0B93BFF5AFFB7FF84E" author="Sloan, T & Talent, R & Mawson, A & Simpson, G & Brock, M & Englbretsen, J & Jell, A & Aung, C & Pfaffenritter, J" box="[1003,1196,1941,1964]" pageId="17" pageNumber="232" pagination="1 - 77" refId="ref19008" refString="Sloan, T. R., J. A. Talent, R. Mawson, A. J. Simpson, G. A. Brock, M. J. Englbretsen, J. S. Jell, A. K. Aung, C. Pfaffenritter, J. Trotter & I. W. Withnall, 1995. Conodont data from Silurian-Middle Devonian carbonate fans, debris flows, allochthonous blocks and adjacent platform margins. Broken River and Camel Creek areas, north Queensland, Australia. Courier Forschungsinstitut Senckenberg 182: 1 - 77, 10 text-figs, 14 Tables, 15 pls." type="book chapter" year="1995">
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Sloan
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<emphasis id="B932232A80639D0B9478F5AFFB8CF84E" box="[1068,1119,1941,1964]" italics="true" pageId="17" pageNumber="232">et al.</emphasis>
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(1995)
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</bibRefCitation>
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,
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<quantity id="4CBE52DD80639D0B94E1F5AFFB3AF84E" box="[1205,1257,1941,1964]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="232" unit="km" value="2.0">2 km</quantity>
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NNW of Storm Hill; the latter specimen is somewhat better preserved.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CACB380609D0890F2F2A0FD8EFC93" pageId="18" pageNumber="233" type="description">
|
||
<paragraph id="8BF9FF3880609D0890F2F2A0FD8EFC93" blockId="18.[166,794,154,881]" pageId="18" pageNumber="233">
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<emphasis id="B932232A80609D0890F2F2A0FEE2FF50" bold="true" box="[166,305,154,178]" pageId="18" pageNumber="233">Description.</emphasis>
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Skeleton laminar to weakly mamelonate and latilaminate; dominated by long, thickened and radially aligned pachysteles,
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<quantity id="4CBE52DD80609D0891DDF2EFFDE0FF0E" box="[393,563,213,236]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="18" pageNumber="233" unit="mm" value="0.275" valueMax="0.35" valueMin="0.2">0.2 to 0.35 mm</quantity>
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across, with spacing from
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<quantity id="4CBE52DD80609D0890B1F2C8FE9AFEEB" box="[229,329,242,265]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="18" pageNumber="233" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3 to 5 in</quantity>
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<quantity id="4CBE52DD80609D089107F2C8FE4BFEEB" box="[339,408,242,265]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="233" unit="mm" value="2.0">2 mm</quantity>
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; commonly they are close spaced, and often in contact, but rarely seen to merge or branch; because of alteration, pachysteles are preserved mainly as compact, dense, rod-like elements, but a few aligned dark melanospheric dots are preserved confirming an incomplete microreticulation of micropillars and microlaminae, from
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<quantity id="4CBE52DD80609D0890F2F398FE85FE5B" box="[166,342,418,441]" metricMagnitude="-5" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="18" pageNumber="233" unit="mm" value="0.025" valueMax="0.03" valueMin="0.02">0.02 to 0.03 mm</quantity>
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thick, defining subrounded microgalleries about
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<quantity id="4CBE52DD80609D0890A4F385FE60FE34" box="[240,435,447,470]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="18" pageNumber="233" unit="mm" value="0.045" valueMax="0.05" valueMin="0.04">0.04 to 0.05 mm</quantity>
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in diameter. Pachystromes are irregularly developed between pachysteles; they may be represented as thicker pachystromal extensions about
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<quantity id="4CBE52DD80609D089358F3C0FF1CFDCC" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="233" unit="mm" value="5.0">5 mm</quantity>
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in length and up to
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<quantity id="4CBE52DD80609D0891E4F02DFDD0FDCC" box="[432,515,535,558]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="233" unit="mm" value="0.2">0.2 mm</quantity>
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in thickness, but in other areas represented by short lengths of a poorly differentiated network that may include one or more microlaminae; they may be inconsistently aligned laterally, and much interrupted by intervening allotubes. Typically allotubes are elongated to rather irregular, and divided by slightly domal to obliquely aligned dissepiments, at intervals of
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<quantity id="4CBE52DD80609D089218F0FDFD1DFD3C" box="[588,718,711,734]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="18" pageNumber="233" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10 to 15 in</quantity>
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<quantity id="4CBE52DD80609D08928CF0FDFCC9FD3C" box="[728,794,711,734]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="233" unit="mm" value="2.0">2 mm</quantity>
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longitudinally; in tangential section allotubes are small and circular in outline, about
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<quantity id="4CBE52DD80609D0891E9F138FDC3FCFB" box="[445,528,770,793]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="233" unit="mm" value="0.2">0.2 mm</quantity>
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in diameter. Astrorhizal canals may be centred in broad mamelons, their branches extend tangentially through other parts of skeleton; some exhibit tabulae from
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<quantity id="4CBE52DD80609D0891D0F160FDCAFC93" box="[388,537,858,881]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="18" pageNumber="233" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2 to 0.4 mm</quantity>
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apart.
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C35CACB380609D0890F2F1B5FD4BF9EC" pageId="18" pageNumber="233" type="discussion">
|
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<paragraph id="8BF9FF3880609D0890F2F1B5FD4BF9EC" blockId="18.[166,794,911,1550]" pageId="18" pageNumber="233">
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<emphasis id="B932232A80609D0890F2F1B5FEDDFC45" bold="true" box="[166,270,911,935]" pageId="18" pageNumber="233">Remarks</emphasis>
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. The Broken River species should probably be assigned to the genus
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<taxonomicName id="4C4684BB80609D0891C9F197FD9BFC26" authorityName="Bogoyavlenskaya" authorityYear="1972" box="[413,584,941,964]" class="Hydrozoa" family="Hydractiniidae" genus="Columnostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="18" pageNumber="233" phylum="Cnidaria" rank="genus">
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<emphasis id="B932232A80609D0891C9F197FD9BFC26" box="[413,584,941,964]" italics="true" pageId="18" pageNumber="233">Columnostroma</emphasis>
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</taxonomicName>
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on the basis of its long, stout, rounded, close-spaced pachysteles, less important pachystromes and traces of a fine reticular microstructure, though the particular
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<typeStatus id="54FD419A80609D0891DCF63FFE65FBFE" box="[392,438,1029,1052]" pageId="18" pageNumber="233">type</typeStatus>
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||
of microreticulation,
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<emphasis id="B932232A80609D0892C8F63FFD04FBFE" box="[668,727,1029,1052]" italics="true" pageId="18" pageNumber="233">sensu</emphasis>
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||
<bibRefCitation id="EFD782C980609D089289F63FFF27FBDB" author="Stock, C" pageId="18" pageNumber="233" pagination="119 - 155" refId="ref19566" refString="Stock, C. W., 1989. Microreticulate microstructure in the Stromatoporoidea. Memoir of the Association of Australasian Palaeontologists 8: 119 - 155." type="journal article" year="1989">Stock (1989)</bibRefCitation>
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, cannot be ascertained. However, owing to the incompletenss of available material and poor preservation overall, the species must be left in open nomenclature. The
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<typeStatus id="54FD419A80609D089088F640FED8FB73" box="[220,267,1146,1169]" pageId="18" pageNumber="233">type</typeStatus>
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species of
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<emphasis id="B932232A80609D0891C6F640FD99FB73" box="[402,586,1146,1169]" italics="true" pageId="18" pageNumber="233">
|
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<taxonomicName id="4C4684BB80609D0891C6F640FD96FB73" authorityName="Bogoyavlenskaya" authorityYear="1972" box="[402,581,1146,1169]" class="Hydrozoa" family="Hydractiniidae" genus="Columnostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="18" pageNumber="233" phylum="Cnidaria" rank="genus">Columnostroma</taxonomicName>
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,
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</emphasis>
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<taxonomicName id="4C4684BB80609D089201F640FE99FB4C" authority="(Spencer, 1884)" baseAuthorityName="Spencer" baseAuthorityYear="1884" class="Hydrozoa" family="Hydractiniidae" genus="Coenostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthoathecata" pageId="18" pageNumber="233" phylum="Cnidaria" rank="species" species="ristigouchense">
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<emphasis id="B932232A80609D089201F640FCCAFB73" box="[597,793,1146,1169]" italics="true" pageId="18" pageNumber="233">C. ristigouchense</emphasis>
|
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(
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<bibRefCitation id="EFD782C980609D0890F9F6ADFE90FB4C" author="Spencer, J" box="[173,323,1175,1198]" pageId="18" pageNumber="233" pagination="593 - 610" refId="ref19119" refString="Spencer, J. W. W., 1884. Niagaran Fossils II: Stromatoporidae of the Upper Silurian System. Bulletin of the Museum of the University of Missouri 1: 43 - 52; also Transactions of the Academy of Science of St. Louis 4: 593 - 610." type="journal article" year="1884">Spencer, 1884</bibRefCitation>
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||
)
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||
</taxonomicName>
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from the Early Devonian of
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<collectingRegion id="498231DA80609D089238F6ADFCC5FB4C" box="[620,790,1175,1198]" country="Canada" name="New Brunswick" pageId="18" pageNumber="233">New Brunswick</collectingRegion>
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,
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<collectingCountry id="F351BFA880609D0890F2F68FFF2AFB2E" box="[166,249,1205,1228]" name="Canada" pageId="18" pageNumber="233">Canada</collectingCountry>
|
||
, shows some resemblances, having rather similar overall dimensions, but the pachysteles of the Broken River species are slightly thicker and more closely spaced, and the laminae within pachystromal segments are not demonstrably colliculate. Only one Australian species of
|
||
<emphasis id="B932232A80609D08923CF710FCC9FAA3" box="[616,794,1322,1345]" italics="true" pageId="18" pageNumber="233">
|
||
<taxonomicName id="4C4684BB80609D08923CF710FCC6FAA3" authorityName="Bogoyavlenskaya" authorityYear="1972" box="[616,789,1322,1345]" class="Hydrozoa" family="Hydractiniidae" genus="Columnostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="18" pageNumber="233" phylum="Cnidaria" rank="genus">Columnostroma</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="4C4684BB80609D0890F2F772FD7AFABC" authority="Webby, Stearn & Zhen, 1993" authorityName="Webby, Stearn & Zhen" authorityYear="1993" box="[166,681,1351,1375]" class="Hydrozoa" family="Hydractiniidae" genus="Columnostroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Syringostromatida" pageId="18" pageNumber="233" phylum="Cnidaria" rank="species" species="clathratum">
|
||
<emphasis id="B932232A80609D0890F2F772FE94FABD" box="[166,327,1352,1375]" italics="true" pageId="18" pageNumber="233">C. clathratum</emphasis>
|
||
Webby, Stearn & Zhen, 1993
|
||
</taxonomicName>
|
||
has been described previously from the Pragian Lilydale Limestone of Victoria, and this bears little resemblance to the Broken River species, being much finer in overall skeletal structure, and the pachystromes more regular, continuous, and finetextured, with microlaminae mainly in conformity with the horizontal dissepiments crossing the allotubes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |